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<mods:title id="35E2316C971C1625AB8494478AC0F171">On seven undescribed leaf insect species revealed within the recent &quot; Tree of Leaves &quot; (Phasmatodea, Phylliidae)</mods:title>
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<paragraph id="A3EA9AB9DE5FF02AF1A27628DD664A81" pageId="0" pageNumber="145">Type species.</paragraph>
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(Audinet-Serville, 1838).
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Historically, many 
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<emphasis id="5352E77750E2246E28B49E4103D7E3E7" italics="true" pageId="0" pageNumber="145">Pulchriphyllium</emphasis>
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populations have been the subject of repeated taxonomic adjustments, oftentimes without being properly assessed or clearly defined. Originally, many of the populations were treated as distinct species, but eventually, as phylliids became better known and more common in museums, authors started to assume phylliids were species-poor with wide geographic ranges (
<bibRefCitation id="F4F637E21AD6D0E755B54BC9436A7277" author="Brock, PD" journalOrPublisher="Bhd., Kuala Lumpur" pageId="0" pageNumber="145" refId="B11" refString="Brock, PD, 1999. Stick and Leaf Insects of Peninsular Malaysia and Singapore. Malaysian Nature Society, Academe Art &amp; Printing Sdn. Bhd., Kuala Lumpur" title="Stick and Leaf Insects of Peninsular Malaysia and Singapore. Malaysian Nature Society, Academe Art &amp; Printing Sdn." year="1999">Brock 1999</bibRefCitation>
; 
<bibRefCitation id="97AF846DB586165C119C747A01E6599B" author="Groesser, D" journalOrPublisher="Edition Chimaira, Frankfurt am Main" pageId="0" pageNumber="145" refId="B33" refString="Groesser, D, 2008. Wandelnde Blaetter. Ein Katalog aller bisher beschriebenen Phylliinae- Arten und deren Eier mit drei Neubeschreibungen (2nd Ed.). Edition Chimaira, Frankfurt am Main" title="Wandelnde Blaetter. Ein Katalog aller bisher beschriebenen Phylliinae- Arten und deren Eier mit drei Neubeschreibungen (2 nd Ed.)." year="2008">
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2008
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; 
<bibRefCitation id="623ECF9966EC2C710857F606F35C93DA" author="Brock, PD" journalOrPublisher="NAP Editions, Verrieres le Buisson, France" pageId="0" pageNumber="145" publicationUrl="https://www.catalogueoflife.org/col" refId="B13" refString="Brock, PD, Buescher, TH, Baker, E, 2022. Phasmida SF: Phasmida Species File Version 5.0/5.0. In: Roskov Y et al. (Eds) Species 2000 and ITIS Catalogue of Life. Species 2000: Naturalis, Leiden. https://www.catalogueoflife.org/col" title="Phasmida SF: Phasmida Species File Version 5.0 / 5.0. In: Roskov Y et al. (Eds) Species 2000 and ITIS Catalogue of Life. Species 2000: Naturalis, Leiden." url="https://www.catalogueoflife.org/col" year="2022">Brock et al. 2022</bibRefCitation>
). Several taxa have been treated as forms, subspecies, or synonyms by past authors (
<bibRefCitation id="E569E29E97BBEF9D410C23E795E85A10" author="Giglio-Tos, E" journalOrPublisher="Bollettino dei Musei di Zoologia ed Anatomia comparata della Royal Universita di Torino" pageId="0" pageNumber="145" pagination="1 - 57" refId="B29" refString="Giglio-Tos, E, 1910. Fasmidi esotici del R. Museo zoologico di Torino e del Museo civico di Storia naturale di Genova. Bollettino dei Musei di Zoologia ed Anatomia comparata della Royal Universita di Torino 25 (625): 1 - 57" title="Fasmidi esotici del R. Museo zoologico di Torino e del Museo civico di Storia naturale di Genova." volume="25" year="1910">Giglio-Tos 1910</bibRefCitation>
; 
<bibRefCitation id="B2C99FDA2A043707470D2896B93BDFBC" author="Brock, PD" journalOrPublisher="Malayan Nature Journal" pageId="0" pageNumber="145" pagination="83 - 102" refId="B10" refString="Brock, PD, 1995. Catalogue of Stick and Leaf-Insects (Insecta: Phasmida) associated with Peninsular Malaysia and Singapore. Malayan Nature Journal 49: 83 - 102" title="Catalogue of Stick and Leaf-Insects (Insecta: Phasmida) associated with Peninsular Malaysia and Singapore." volume="49" year="1995">Brock 1995</bibRefCitation>
; 
<bibRefCitation id="33E770458EE185EE6BA4AA9C9097699C" DOI="https://doi.org/10.11646/zootaxa.2322.1.1" author="Hennemann, FH" journalOrPublisher="Zootaxa" pageId="0" pageNumber="145" pagination="1 - 83" refId="B37" refString="Hennemann, FH, Conle, OV, Gottardo, M, Bresseel, J, 2009. On certain species of the genus Phyllium Illiger, 1798, with proposals for an intrageneric systematization and the descriptions of five new species from the Philippines and Palawan (Phasmida: Phylliidae: Phylliinae: Phylliini). Zootaxa 2322: 1 - 83, DOI: https://doi.org/10.11646/zootaxa.2322.1.1" title="On certain species of the genus Phyllium Illiger, 1798, with proposals for an intrageneric systematization and the descriptions of five new species from the Philippines and Palawan (Phasmida: Phylliidae: Phylliinae: Phylliini)." url="https://doi.org/10.11646/zootaxa.2322.1.1" volume="2322" year="2009">Hennemann et al. 2009</bibRefCitation>
), and it was only recently that authors have begun to split certain lineages into distinct species again (
<bibRefCitation id="68A33C7403AD96973EA9BC87C1FFD9D9" author="Seow-Choen, F" journalOrPublisher="Natural History Publications (Borneo), Kota Kinabalu, Sabah" pageId="0" pageNumber="145" refId="B58" refString="Seow-Choen, F, 2017. A Taxonomic Guide to the Stick Insects of Borneo (Vol. II). Natural History Publications (Borneo), Kota Kinabalu, Sabah" title="A Taxonomic Guide to the Stick Insects of Borneo (Vol. II)." year="2017">Seow-Choen 2017</bibRefCitation>
). The 
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<emphasis id="95A25D99286A06EFE718A869C6010B17" italics="true" pageId="0" pageNumber="145">Pulchriphyllium</emphasis>
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are only found Northwest of 
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line of faunal balance (
<bibRefCitation id="148EF12829FC7769E4BB42EF888FFAFB" author="Huxley, TH" journalOrPublisher="Molecular Biology and Evolution" pageId="0" pageNumber="145" refId="B40" refString="Huxley, TH, 1868. On the Classification and Distribution of the Alectoromorphae and Heteromorphae. Proceedings of the Zoological Society, London, 294-319." title="On the Classification and Distribution of the Alectoromorphae and Heteromorphae. Proceedings of the Zoological Society, London, 294 - 319." year="1868">Huxley 1868</bibRefCitation>
) and range as far north as Bangladesh/northeastern India, and as far west as the Seychelles islands (Fig. 
<figureCitation id="ADEBA28B68662ABD11C4A66BED7EAEFC" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Distribution map for the herein discussed Pulchriphyllium species with known records plotted. For data on the distribution points used for this map, see Suppl. material 1 A distribution of the &quot; Pulchriphyllium bioculatum - like species &quot; B distribution of the Pulchriphyllium species from mainland Asia with a boxy abdominal shape (inset outline in A). Abbreviations: KH, Cambodia; LA, Laos; TH, Thailand; VN, Vietnam." figureDoi="10.3897/zookeys.1173.104413.figure15" httpUri="https://binary.pensoft.net/fig/887717" pageId="0" pageNumber="145">15A</figureCitation>
). The area with the highest 
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<emphasis id="9420FE3AAD7F60A1C7AB7C4861499988" italics="true" pageId="0" pageNumber="145">Pulchriphyllium</emphasis>
</taxonomicName>
diversity appears to be the biogeographical region of Sundaland, an area which was recovered as the likely geographic ancestral range for this clade (
<bibRefCitation id="D6E3F642C2350535CDC14B7E3CD44BB8" DOI="https://doi.org/10.1038/s42003-021-02436-z" author="Bank, S" journalOrPublisher="Libraire Encyclopedique de Roret, Paris" pageId="0" pageNumber="145" refId="B3" refString="Bank, S, Cumming, RT, Yunchang, L, Henze, K, Le Tirant, S, Bradler, S, 2021. A tree of leaves: Phylogeny and historical biogeography of the leaf insects (Phasmatodea: Phylliidae). Communications Biology 4(932): e932. https://doi.org/10.1038/s42003-021-02436-z" title="A tree of leaves: Phylogeny and historical biogeography of the leaf insects (Phasmatodea: Phylliidae). Communications Biology 4 (932): e 932." url="https://doi.org/10.1038/s42003-021-02436-z" year="2021">Bank et al. 2021</bibRefCitation>
).
</paragraph>
<caption id="5C0B933CBADBD9CF3B38A43AF0CF0086" doi="10.3897/zookeys.1173.104413.figure15" httpUri="https://binary.pensoft.net/fig/887717" pageId="0" pageNumber="145" start="Figure 15" startId="F15">
<paragraph id="A7290044E5994C173E85AFB0886DC505" pageId="0" pageNumber="145">
<emphasis id="03BB8E5E4C46BB839BECDAD8774E0536" bold="true" pageId="0" pageNumber="145">Figure 15.</emphasis>
Distribution map for the herein discussed 
<taxonomicName id="10C794A46800FBE96EF7BEE5C4720587" authorityName="Griffini" authorityYear="1898" class="Insecta" family="Phylliidae" genus="Pulchriphyllium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pulchriphyllium" order="Phasmida" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="E36B9340065776C747EC31DF2ECA8709" italics="true" pageId="0" pageNumber="145">Pulchriphyllium</emphasis>
</taxonomicName>
species with known records plotted. For data on the distribution points used for this map, see Suppl. material 1 
<emphasis id="05AE0FE048A8631CE400179E8089369A" bold="true" pageId="0" pageNumber="145">A</emphasis>
distribution of the &quot; 
<taxonomicName id="717E8182083C07046AC0D14BB1DF5D3C" baseAuthorityName="G.R.Gray" baseAuthorityYear="1832" class="Insecta" family="Phylliidae" genus="Pulchriphyllium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pulchriphyllium bioculatum" order="Phasmida" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="bioculatum">
<emphasis id="B78E37564541019DF8D4E633C8161703" italics="true" pageId="0" pageNumber="145">Pulchriphyllium bioculatum</emphasis>
</taxonomicName>
-like species&quot; 
<emphasis id="EC6F2A4F2E56D6065D437649ABF6E3A3" bold="true" pageId="0" pageNumber="145">B</emphasis>
distribution of the 
<taxonomicName id="BE4E4B868DE615BC8C3FC5DB2028D097" authorityName="Griffini" authorityYear="1898" class="Insecta" family="Phylliidae" genus="Pulchriphyllium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pulchriphyllium" order="Phasmida" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="40659A8A10494DC3EF06EB89C533BB49" italics="true" pageId="0" pageNumber="145">Pulchriphyllium</emphasis>
</taxonomicName>
species from mainland Asia with a boxy abdominal shape (inset outline in 
<emphasis id="B12F3D8B73E3FFEDBBCD88E385CA5961" bold="true" pageId="0" pageNumber="145">A</emphasis>
). Abbreviations: KH, Cambodia; LA, Laos; TH, Thailand; VN, Vietnam.
</paragraph>
</caption>
<paragraph id="1F5A52D569D2A39D91227D8F8DBD4E98" pageId="0" pageNumber="145">
Within the 
<taxonomicName id="3341721AD24489026685222E079CB50A" authorityName="Griffini" authorityYear="1898" class="Insecta" family="Phylliidae" genus="Pulchriphyllium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pulchriphyllium" order="Phasmida" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="406DBB75246658E52FF28B1DF6F0BD7A" italics="true" pageId="0" pageNumber="145">Pulchriphyllium</emphasis>
</taxonomicName>
there appear to be several recurring, leaf-shape mimicry patterns; broad leaf (like in 
<taxonomicName id="C4FA2C2C55C7284DC48D835D8CAB2E49" baseAuthorityName="Hausleithner" baseAuthorityYear="1984" class="Insecta" family="Phylliidae" genus="Pulchriphyllium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pulchriphyllium giganteum" order="Phasmida" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="giganteum">
<emphasis id="8C25E7C1EC6E39DE186619936D75A9AD" italics="true" pageId="0" pageNumber="145">Pulchriphyllium giganteum</emphasis>
</taxonomicName>
), boxy leaf (like in 
<taxonomicName id="29657A7E28D15F41CD9F11AC33DFD07D" baseAuthorityName="Audinet-Serville" baseAuthorityYear="1838" class="Insecta" family="Phylliidae" genus="Pulchriphyllium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pulchriphyllium pulchrifolium" order="Phasmida" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="pulchrifolium">
<emphasis id="6EA4A4DA50D54B5084ECD41745DA3778" italics="true" pageId="0" pageNumber="145">Pulchriphyllium pulchrifolium</emphasis>
</taxonomicName>
), and tapered leaf (like in 
<taxonomicName id="ACB3CC68E4802F9BAA41F4122E32B0E8" baseAuthorityName="G.R.Gray" baseAuthorityYear="1832" class="Insecta" family="Phylliidae" genus="Pulchriphyllium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pulchriphyllium bioculatum" order="Phasmida" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="bioculatum">
<emphasis id="03F5C0BC491C88F6435CCCE999C78124" italics="true" pageId="0" pageNumber="145">Pulchriphyllium bioculatum</emphasis>
</taxonomicName>
Gray, 1832). Of particular difficulty to differentiate are the numerous species where females have a tapered abdominal shape and males have rounded/ovoid abdomen, herein referred to as &quot; 
<taxonomicName id="D2E2A13C7A7F13AD0523F3CD047D1AC7" baseAuthorityName="G.R.Gray" baseAuthorityYear="1832" class="Insecta" family="Phylliidae" genus="Pulchriphyllium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pulchriphyllium bioculatum" order="Phasmida" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="bioculatum">
<emphasis id="A6A152F23DF1C7B70DB86125A451C4EC" italics="true" pageId="0" pageNumber="145">Pulchriphyllium bioculatum</emphasis>
</taxonomicName>
-like species&quot;. These &quot; 
<taxonomicName id="455A9ABBB69FA138118EDE94F3EE67D6" baseAuthorityName="G.R.Gray" baseAuthorityYear="1832" class="Insecta" family="Phylliidae" genus="Pulchriphyllium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pulchriphyllium bioculatum" order="Phasmida" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="bioculatum">
<emphasis id="F3C6E2D630D26F13F9248637E4A15DED" italics="true" pageId="0" pageNumber="145">Pulchriphyllium bioculatum</emphasis>
</taxonomicName>
-like species&quot; are a subset of the &quot; 
<taxonomicName id="30517F3DB354BD5C157848D9F01C415F" baseAuthorityName="G.R.Gray" baseAuthorityYear="1832" class="Insecta" family="Phylliidae" genus="Pulchriphyllium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pulchriphyllium bioculatum" order="Phasmida" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="bioculatum">
<emphasis id="CD3E0F7C434928D730B4CDE09ECA7CF5" italics="true" pageId="0" pageNumber="145">Pulchriphyllium bioculatum</emphasis>
</taxonomicName>
species group&quot;, the only species group within the 
<taxonomicName id="9D3BF4D0132DB18AC085990CBA22D24E" authorityName="Griffini" authorityYear="1898" class="Insecta" family="Phylliidae" genus="Pulchriphyllium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pulchriphyllium" order="Phasmida" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="B8CDD089632651CADD329332D1A7B258" italics="true" pageId="0" pageNumber="145">Pulchriphyllium</emphasis>
</taxonomicName>
proposed by 
<bibRefCitation id="3DBED4E9C6B2CC4051FD9858341E5528" DOI="https://doi.org/10.11646/zootaxa.2322.1.1" author="Hennemann, FH" journalOrPublisher="Zootaxa" pageId="0" pageNumber="145" pagination="1 - 83" refId="B37" refString="Hennemann, FH, Conle, OV, Gottardo, M, Bresseel, J, 2009. On certain species of the genus Phyllium Illiger, 1798, with proposals for an intrageneric systematization and the descriptions of five new species from the Philippines and Palawan (Phasmida: Phylliidae: Phylliinae: Phylliini). Zootaxa 2322: 1 - 83, DOI: https://doi.org/10.11646/zootaxa.2322.1.1" title="On certain species of the genus Phyllium Illiger, 1798, with proposals for an intrageneric systematization and the descriptions of five new species from the Philippines and Palawan (Phasmida: Phylliidae: Phylliinae: Phylliini)." url="https://doi.org/10.11646/zootaxa.2322.1.1" volume="2322" year="2009">Hennemann et al. (2009)</bibRefCitation>
which has not been taxonomically adjusted (the other three species groups proposed within that work have either been transferred to a different genus (such as the &quot; 
<emphasis id="1C48D01D9BB6AB64B9E9D09721B3C8C2" italics="true" pageId="0" pageNumber="145">frondosum</emphasis>
species group&quot; which were transferred to the 
<taxonomicName id="D24F23633921A1A0DDE5163DA63C98D2" authorityName="Redtenbacher" authorityYear="1906" class="Insecta" family="Phylliidae" genus="Nanophyllium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Nanophyllium" order="Phasmatodea" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="4A2C755A7F50A177D016DBD88CBD6A43" italics="true" pageId="0" pageNumber="145">Nanophyllium</emphasis>
</taxonomicName>
by 
<bibRefCitation id="D6C15DB18C02CE7BD66F1ED138E47106" DOI="https://doi.org/10.3897/zookeys.969.56214" author="Cumming, RT" journalOrPublisher="ZooKeys" pageId="0" pageNumber="145" pagination="43 - 84" refId="B24" refString="Cumming, RT, Le Tirant, S, Teemsma, SN, Hennemann, FH, Willemse, L, Buescher, TH, 2020d. Lost lovers linked at long last: Elusive female Nanophyllium mystery solved after a century of being placed in a different genus (Phasmatodea, Phylliidae). ZooKeys 969: 43 - 84, DOI: https://doi.org/10.3897/zookeys.969.56214" title="Lost lovers linked at long last: Elusive female Nanophyllium mystery solved after a century of being placed in a different genus (Phasmatodea, Phylliidae)." url="https://doi.org/10.3897/zookeys.969.56214" volume="969" year="2020 d">Cumming et al. 2020d</bibRefCitation>
), or have since been described as their own genera (like the &quot; 
<emphasis id="7E4BFBBCEAF3E20C8054E16EDD6111A3" italics="true" pageId="0" pageNumber="145">brevipenne</emphasis>
species group&quot; which is now 
<taxonomicName id="3A6A6BF1FD7128CA57CAF984848C37DE" authorityName="Cumming &amp; Le Tirant" authorityYear="2022" class="Insecta" family="Phylliidae" genus="Acentetaphyllium" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Acentetaphyllium" order="Phasmatodea" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="2DF52C6196E7C2D252F485589AD2118A" italics="true" pageId="0" pageNumber="145">Acentetaphyllium</emphasis>
</taxonomicName>
Cumming &amp; Le Tirant, 2022 and the &quot; 
<emphasis id="5B36E9F51BD4B15A4A3523BFB2568EAA" italics="true" pageId="0" pageNumber="145">schultzei</emphasis>
species group&quot; which is now 
<taxonomicName id="EA0207E1079035A4FB24774636DC9176" authorityName="Cumming &amp; Le Tirant" authorityYear="2022" class="Insecta" family="Phylliidae" genus="Rakaphyllium" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Rakaphyllium" order="Phasmatodea" pageId="0" pageNumber="145" phylum="Arthropoda" rank="genus">
<emphasis id="1CEE9EF4F81CB2F6786B568C91DF0F9A" italics="true" pageId="0" pageNumber="145">Rakaphyllium</emphasis>
</taxonomicName>
Cumming &amp; Le Tirant, 2022).
</paragraph>
<paragraph id="2B6EA840F7B6CF8D6E8BF5C10DD05B7F" pageId="0" pageNumber="145">
The &quot; 
<taxonomicName id="20ABA82A7D28A311934F25D2C1FAB3B1" baseAuthorityName="G.R.Gray" baseAuthorityYear="1832" class="Insecta" family="Phylliidae" genus="Pulchriphyllium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pulchriphyllium bioculatum" order="Phasmida" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="bioculatum">
<emphasis id="6CCC9DA17C26E8B166C2DA305D352981" italics="true" pageId="0" pageNumber="145">Pulchriphyllium bioculatum</emphasis>
</taxonomicName>
-like species&quot; are difficult to morphologically differentiate, have wide-ranging geographic distributions (Fig. 
<figureCitation id="10CBAF410F4EAB57AB5E565C94FABB2B" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Distribution map for the herein discussed Pulchriphyllium species with known records plotted. For data on the distribution points used for this map, see Suppl. material 1 A distribution of the &quot; Pulchriphyllium bioculatum - like species &quot; B distribution of the Pulchriphyllium species from mainland Asia with a boxy abdominal shape (inset outline in A). Abbreviations: KH, Cambodia; LA, Laos; TH, Thailand; VN, Vietnam." figureDoi="10.3897/zookeys.1173.104413.figure15" httpUri="https://binary.pensoft.net/fig/887717" pageId="0" pageNumber="145">15A</figureCitation>
), and coupled with the fact that the holotype of 
<taxonomicName id="2C6146401A2A07CDACF9D1C1BFC136BF" baseAuthorityName="G.R.Gray" baseAuthorityYear="1832" class="Insecta" family="Phylliidae" genus="Pulchriphyllium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pulchriphyllium bioculatum" order="Phasmida" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="bioculatum">
<emphasis id="09B18876A3357257F90660F0D7C13627" italics="true" pageId="0" pageNumber="145">Pulchriphyllium bioculatum</emphasis>
</taxonomicName>
lacks a precise collection locality, has made this group of similar-looking species difficult to clearly define. Based upon the type specimen morphology, the population considered true &quot; 
<taxonomicName id="E6929510AEC8DBE69BF6F6283B7065BD" baseAuthorityName="G.R.Gray" baseAuthorityYear="1832" class="Insecta" family="Phylliidae" genus="Pulchriphyllium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pulchriphyllium bioculatum" order="Phasmida" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="bioculatum">
<emphasis id="B82B89E346498594F1569AF3D596D1E4" italics="true" pageId="0" pageNumber="145">Pulchriphyllium bioculatum</emphasis>
</taxonomicName>
&quot; is the species centered around Sundaland and mainland Asia (Fig. 
<figureCitation id="9AE6C9A56987BC2BB62A7B7C290992FD" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Distribution map for the herein discussed Pulchriphyllium species with known records plotted. For data on the distribution points used for this map, see Suppl. material 1 A distribution of the &quot; Pulchriphyllium bioculatum - like species &quot; B distribution of the Pulchriphyllium species from mainland Asia with a boxy abdominal shape (inset outline in A). Abbreviations: KH, Cambodia; LA, Laos; TH, Thailand; VN, Vietnam." figureDoi="10.3897/zookeys.1173.104413.figure15" httpUri="https://binary.pensoft.net/fig/887717" pageId="0" pageNumber="145">15A</figureCitation>
; 
<bibRefCitation id="6B4C4E8CE635CE6B9CAEB3A27AC6A0DB" DOI="https://doi.org/10.11646/zootaxa.2322.1.1" author="Hennemann, FH" journalOrPublisher="Zootaxa" pageId="0" pageNumber="145" pagination="1 - 83" refId="B37" refString="Hennemann, FH, Conle, OV, Gottardo, M, Bresseel, J, 2009. On certain species of the genus Phyllium Illiger, 1798, with proposals for an intrageneric systematization and the descriptions of five new species from the Philippines and Palawan (Phasmida: Phylliidae: Phylliinae: Phylliini). Zootaxa 2322: 1 - 83, DOI: https://doi.org/10.11646/zootaxa.2322.1.1" title="On certain species of the genus Phyllium Illiger, 1798, with proposals for an intrageneric systematization and the descriptions of five new species from the Philippines and Palawan (Phasmida: Phylliidae: Phylliinae: Phylliini)." url="https://doi.org/10.11646/zootaxa.2322.1.1" volume="2322" year="2009">Hennemann et al. 2009</bibRefCitation>
). This leaves the numerous populations once considered subspecies or synonyms of 
<taxonomicName id="0B4AC1182DF8627C0BD61DDB87DD0CA2" baseAuthorityName="G.R.Gray" baseAuthorityYear="1832" class="Insecta" family="Phylliidae" genus="Pulchriphyllium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pulchriphyllium bioculatum" order="Phasmida" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="bioculatum">
<emphasis id="818944E1FFBB72EC99CC4C10AC25B8B8" italics="true" pageId="0" pageNumber="145">Pulchriphyllium bioculatum</emphasis>
</taxonomicName>
(but now recognized as independent species; Fig. 
<figureCitation id="FDC69AF02EF0B7AA8C384EAFD9D5BCA8" captionStart="Figure 15" captionStartId="F15" captionText="Figure 15. Distribution map for the herein discussed Pulchriphyllium species with known records plotted. For data on the distribution points used for this map, see Suppl. material 1 A distribution of the &quot; Pulchriphyllium bioculatum - like species &quot; B distribution of the Pulchriphyllium species from mainland Asia with a boxy abdominal shape (inset outline in A). Abbreviations: KH, Cambodia; LA, Laos; TH, Thailand; VN, Vietnam." figureDoi="10.3897/zookeys.1173.104413.figure15" httpUri="https://binary.pensoft.net/fig/887717" pageId="0" pageNumber="145">15A</figureCitation>
) in need of clarification. Thanks to extensive review of historic type specimen morphology, modern specimen morphology, geographic distribution, and molecular analyses, the &quot; 
<taxonomicName id="655C7C2B0AA71510E398D0A1066C5530" baseAuthorityName="G.R.Gray" baseAuthorityYear="1832" class="Insecta" family="Phylliidae" genus="Pulchriphyllium" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pulchriphyllium bioculatum" order="Phasmida" pageId="0" pageNumber="145" phylum="Arthropoda" rank="species" species="bioculatum">
<emphasis id="CB3CECB840B900A941402FCA20EC88C7" italics="true" pageId="0" pageNumber="145">Pulchriphyllium bioculatum</emphasis>
</taxonomicName>
-like species&quot; are now beginning to be clarified.
</paragraph>
</subSubSection>
</treatment>
</document>