treatments-xml/data/74/45/30/744530540ED659E788658B840BFB9B58.xml

<document ID-DOI="http://dx.doi.org/10.3897/jhr.84.68702" ID-GBIF-Dataset="d29ff8c2-6b65-4a92-9c54-3471dd7f0b71" ID-Pensoft-Pub="1314-2607-84-301" ID-Pensoft-UUID="5BA39CCDC71356889C3BF1CCF60C44E1" ID-Zenodo-Dep="5349670" ID-ZooBank="E6FDE915C31542E2ABA8B80CE3B922A2" ModsDocID="1314-2607-84-301" checkinTime="1629847041995" checkinUser="pensoft" docAuthor="Samartsev, Konstantin &amp; van Achterberg, Cornelis" docDate="2021" docId="744530540ED659E788658B840BFB9B58" docLanguage="en" docName="JourHymenoptRes 84: 301-325" docOrigin="Journal of Hymenoptera Research 84" docPubDate="2021-08-24" docSource="http://dx.doi.org/10.3897/jhr.84.68702" docTitle="Sculptolobus Yang, van Achterberg &amp; Chen 2008" docType="treatment" docVersion="4" id="5BA39CCDC71356889C3BF1CCF60C44E1" lastPageNumber="301" masterDocId="5BA39CCDC71356889C3BF1CCF60C44E1" masterDocTitle="Afrotropical species of the genus Sculptolobus Yang, van Achterberg &amp; Chen (Hymenoptera, Braconidae, Braconinae)" masterLastPageNumber="325" masterPageNumber="301" pageNumber="301" updateTime="1678756216309" updateUser="pensoft">
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<mods:title>Afrotropical species of the genus Sculptolobus Yang, van Achterberg &amp; Chen (Hymenoptera, Braconidae, Braconinae)</mods:title>
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<mods:namePart>Samartsev, Konstantin</mods:namePart>
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<mods:affiliation>Zoological Institute, Russian Academy of Sciences, St Petersburg 199034, Russia</mods:affiliation>
<mods:nameIdentifier type="email">ksamartsev@gmail.com</mods:nameIdentifier>
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<mods:namePart>van Achterberg, Cornelis</mods:namePart>
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<mods:affiliation>Naturalis Biodiversity Center, 2300 RA Leiden, The Netherlands</mods:affiliation>
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<treatment ID-GBIF-Taxon="184941240" LSID="urn:lsid:plazi:treatment:744530540ED659E788658B840BFB9B58" httpUri="http://treatment.plazi.org/id/744530540ED659E788658B840BFB9B58" lastPageNumber="301" pageId="0" pageNumber="301">
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Genus 
<taxonomicName ID-CoL="63T8L" ID-ENA="2572388" LSID="74453054-0ED6-59E7-8865-8B840BFB9B58" authority="Yang, van Achterberg &amp; Chen, 2008" authorityName="Yang, van Achterberg &amp; Chen" authorityYear="2008" class="Insecta" family="Braconidae" genus="Sculptolobus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sculptolobus" order="Hymenoptera" pageId="0" pageNumber="301" phylum="Arthropoda" rank="genus">Sculptolobus Yang, van Achterberg &amp; Chen, 2008</taxonomicName>
</paragraph>
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<subSubSection pageId="0" pageNumber="301" type="reference_group">
<paragraph pageId="0" pageNumber="301">
<taxonomicName authorityName="Yang, van Achterberg &amp; Chen" authorityYear="2008" class="Insecta" family="Braconidae" genus="Sculptolobus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sculptolobus" order="Hymenoptera" pageId="0" pageNumber="301" phylum="Arthropoda" rank="genus">Sculptolobus</taxonomicName>
van Achterberg, 2006: 
<bibRefCitation author="Chen, J" journalOrPublisher="Fujian Science and Technology Publishing House, Fujian" pageId="0" pageNumber="301" refId="B3" refString="Chen, J, Yang, J, 2006. Systematic studies on Braconinae of China. (Hymenoptera: Braconidae). Fujian Science and Technology Publishing House, Fujian" title="Systematic studies on Braconinae of China. (Hymenoptera: Braconidae)." year="2006">Chen and Yang 2006</bibRefCitation>
: 143 (unavailable: 
<bibRefCitation author="Yu, DS" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="301" refId="B19" refString="Yu, DS, van Achterberg, C, Horstmann, K, 2016. Taxapad 2016, Ichneumonoidea 2015. Database on flash-drive. Nepean (Ontario), Canada." title="Taxapad 2016, Ichneumonoidea 2015. Database on flash-drive. Nepean (Ontario), Canada." year="2016">Yu et al. 2016</bibRefCitation>
).
</paragraph>
<paragraph pageId="0" pageNumber="301">
<taxonomicName authorityName="Yang, van Achterberg &amp; Chen" authorityYear="2008" class="Insecta" family="Braconidae" genus="Sculptolobus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sculptolobus" order="Hymenoptera" pageId="0" pageNumber="301" phylum="Arthropoda" rank="genus">Sculptolobus</taxonomicName>
Yang, van Achterberg &amp; Chen, 2008: 95 (type species: 
<taxonomicName authorityName="Yang, van Achterberg &amp; Chen" authorityYear="2008" class="Insecta" family="Braconidae" genus="Sculptolobus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sculptolobus sulcifer" order="Hymenoptera" pageId="0" pageNumber="301" phylum="Arthropoda" rank="species" species="sulcifer">Sculptolobus sulcifer</taxonomicName>
Yang, van Achterberg &amp; Chen, 2008); 
<bibRefCitation DOI="https://doi.org/10.3161/00034541ANZ2017.67.4.013" author="Samartsev, KG" journalOrPublisher="Annales Zoologici" pageId="0" pageNumber="301" pagination="781 - 794" refId="B14" refString="Samartsev, KG, Belokobylskij, SA, Villemant, C, 2017. Reclassification of some Bracon species (Hymenoptera: Braconidae: Braconinae) described by Ch. Granger and G. Szepligeti from Madagascar. Annales Zoologici 67 (4): 781 - 794, DOI: https://doi.org/10.3161/00034541ANZ2017.67.4.013" title="Reclassification of some Bracon species (Hymenoptera: Braconidae: Braconinae) described by Ch. Granger and G. Szepligeti from Madagascar." url="https://doi.org/10.3161/00034541ANZ2017.67.4.013" volume="67" year="2017">Samartsev et al. 2017</bibRefCitation>
: 786.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="301" type="remarks">
<paragraph pageId="0" pageNumber="301">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="301">
By a curious incident, the genus 
<taxonomicName authorityName="Yang, van Achterberg &amp; Chen" authorityYear="2008" class="Insecta" family="Braconidae" genus="Sculptolobus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sculptolobus" order="Hymenoptera" pageId="0" pageNumber="301" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="301">Sculptolobus</emphasis>
</taxonomicName>
was established two years after its first species had been described. The species 
<taxonomicName lsidName="S. bannaensis" pageId="0" pageNumber="301" rank="species" species="bannaensis">
<emphasis italics="true" pageId="0" pageNumber="301">S. bannaensis</emphasis>
</taxonomicName>
Yang &amp; Chen, 
<taxonomicName lsidName="S. tobiasi" pageId="0" pageNumber="301" rank="species" species="tobiasi">
<emphasis italics="true" pageId="0" pageNumber="301">S. tobiasi</emphasis>
</taxonomicName>
Yang &amp; Chen, and 
<taxonomicName lsidName="S. tongmuensis" pageId="0" pageNumber="301" rank="species" species="tongmuensis">
<emphasis italics="true" pageId="0" pageNumber="301">S. tongmuensis</emphasis>
</taxonomicName>
Chen &amp; Yang have been described in the monograph on Chinese 
<taxonomicName lsidName="" pageId="0" pageNumber="301" rank="subFamily" subFamily="Braconinae">Braconinae</taxonomicName>
(
<bibRefCitation author="Chen, J" journalOrPublisher="Fujian Science and Technology Publishing House, Fujian" pageId="0" pageNumber="301" refId="B3" refString="Chen, J, Yang, J, 2006. Systematic studies on Braconinae of China. (Hymenoptera: Braconidae). Fujian Science and Technology Publishing House, Fujian" title="Systematic studies on Braconinae of China. (Hymenoptera: Braconidae)." year="2006">Chen and Yang 2006</bibRefCitation>
), but the genus itself was described there without an explicit indication of its novelty and with reference to an article by C. van Achterberg, which has not been published (
<bibRefCitation author="Yu, DS" journalOrPublisher="Zoological Journal of the Linnean Society" pageId="0" pageNumber="301" refId="B19" refString="Yu, DS, van Achterberg, C, Horstmann, K, 2016. Taxapad 2016, Ichneumonoidea 2015. Database on flash-drive. Nepean (Ontario), Canada." title="Taxapad 2016, Ichneumonoidea 2015. Database on flash-drive. Nepean (Ontario), Canada." year="2016">Yu et al. 2016</bibRefCitation>
). Thus, 
<taxonomicName authorityName="Yang, van Achterberg &amp; Chen" authorityYear="2008" class="Insecta" family="Braconidae" genus="Sculptolobus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sculptolobus" order="Hymenoptera" pageId="0" pageNumber="301" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="301">Sculptolobus</emphasis>
</taxonomicName>
Chen &amp; Yang, 2006 is an unavailable name according to the article 16.1 of the International Code of Zoological Nomenclature (
<bibRefCitation author="Granger, C" journalOrPublisher="Memoires de l'Institut Scientifique de Madagascar (A)" pageId="0" pageNumber="301" refId="B7" refString="ICZN (1999) International Code of Zoological Nomenclature. Fourth Edition.The International Trust for Zoological Nomenclature, London, 306 pp." title="International Code of Zoological Nomenclature. Fourth Edition." year="1949">ICZN 1999</bibRefCitation>
). The valid description of the genus has been published later by Yang et al. (2008).
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="301" type="diagnosis">
<paragraph pageId="0" pageNumber="301">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="301">
In the key to the Old World genera of 
<taxonomicName lsidName="" pageId="0" pageNumber="301" rank="subFamily" subFamily="Braconinae">Braconinae</taxonomicName>
(
<bibRefCitation DOI="https://doi.org/10.1080/00222938700770031" author="Quicke, DLJ" journalOrPublisher="Journal of Natural History" pageId="0" pageNumber="301" pagination="43 - 157" refId="B10" refString="Quicke, DLJ, 1987. The Old World genera of braconine wasps (Hymenoptera: Braconidae). Journal of Natural History 21: 43 - 157, DOI: https://doi.org/10.1080/00222938700770031" title="The Old World genera of braconine wasps (Hymenoptera: Braconidae)." url="https://doi.org/10.1080/00222938700770031" volume="21" year="1987">Quicke 1987</bibRefCitation>
), 
<taxonomicName authorityName="Yang, van Achterberg &amp; Chen" authorityYear="2008" class="Insecta" family="Braconidae" genus="Sculptolobus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sculptolobus" order="Hymenoptera" pageId="0" pageNumber="301" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="301">Sculptolobus</emphasis>
</taxonomicName>
runs to the couplets 56-74. Further diagnostics of the genus using this key is not possible, because the characters which are variable in species of 
<taxonomicName authorityName="Yang, van Achterberg &amp; Chen" authorityYear="2008" class="Insecta" family="Braconidae" genus="Sculptolobus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sculptolobus" order="Hymenoptera" pageId="0" pageNumber="301" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="301">Sculptolobus</emphasis>
</taxonomicName>
are used as diagnostic there (e.g., the relative length of the fore wing veins 2-SR+M and 3-SR, degree of development of the medio-longitudinal carina on propodeum and anterolateral areas on T3-T5). The set of 19 genera included in the couplets 56-74 currently is not complete as nine later described Old World genera (excluding 
<taxonomicName authorityName="Yang, van Achterberg &amp; Chen" authorityYear="2008" class="Insecta" family="Braconidae" genus="Sculptolobus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sculptolobus" order="Hymenoptera" pageId="0" pageNumber="301" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="301">Sculptolobus</emphasis>
</taxonomicName>
) are suitable for this part of the key. These genera are: 
<taxonomicName authorityName="van Achterberg" authorityYear="1989" class="Insecta" family="Braconidae" genus="Acrocerilia" kingdom="Animalia" lsidName="Acrocerilia" order="Hymenoptera" pageId="0" pageNumber="301" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="301">Acrocerilia</emphasis>
</taxonomicName>
van Achterberg, 1989, 
<taxonomicName authorityName="Quicke" authorityYear="1989" class="Insecta" family="Braconidae" genus="Ancilibracon" kingdom="Animalia" lsidName="Ancilibracon" order="Hymenoptera" pageId="0" pageNumber="301" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="301">Ancilibracon</emphasis>
</taxonomicName>
Quicke, 1989, 
<taxonomicName authorityName="Quicke" authorityYear="1988" class="Insecta" family="Braconidae" genus="Crinibracon" kingdom="Animalia" lsidName="Crinibracon" order="Hymenoptera" pageId="0" pageNumber="301" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="301">Crinibracon</emphasis>
</taxonomicName>
Quicke, 1988, 
<taxonomicName authorityName="Quicke" authorityYear="1989" class="Insecta" family="Braconidae" genus="Esengoides" kingdom="Animalia" lsidName="Esengoides" order="Hymenoptera" pageId="0" pageNumber="301" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="301">Esengoides</emphasis>
</taxonomicName>
Quicke, 1989, 
<taxonomicName authorityName="Quicke" authorityYear="1989" class="Insecta" family="Braconidae" genus="Gelasinibracon" kingdom="Animalia" lsidName="Gelasinibracon" order="Hymenoptera" pageId="0" pageNumber="301" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="301">Gelasinibracon</emphasis>
</taxonomicName>
Quicke, 1989, 
<taxonomicName authorityName="Quicke &amp; Walker" authorityYear="1989" class="Insecta" family="Braconidae" genus="Scutibracon" kingdom="Animalia" lsidName="Scutibracon" order="Hymenoptera" pageId="0" pageNumber="301" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="301">Scutibracon</emphasis>
</taxonomicName>
Quicke &amp; Walker, 1989, 
<taxonomicName authorityName="Quicke" authorityYear="1988" class="Insecta" family="Braconidae" genus="Simplicibracon" kingdom="Animalia" lsidName="Simplicibracon" order="Hymenoptera" pageId="0" pageNumber="301" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="301">Simplicibracon</emphasis>
</taxonomicName>
Quicke, 1988, 
<taxonomicName class="Magnoliopsida" family="Dipterocarpaceae" genus="Stephanobracon" higherTaxonomySource="GBIF" kingdom="Plantae" lsidName="Stephanobracon" order="Malvales" pageId="0" pageNumber="301" phylum="Tracheophyta" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="301">Stephanobracon</emphasis>
</taxonomicName>
Ranjith &amp; Quicke, 2016, and 
<taxonomicName authorityName="Papp" authorityYear="1996" class="Insecta" family="Braconidae" genus="Uncobracon" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Uncobracon" order="Hymenoptera" pageId="0" pageNumber="301" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="301">Uncobracon</emphasis>
</taxonomicName>
Papp, 1996. Preparation of a new version of the key to the genera related to 
<taxonomicName authorityName="Yang, van Achterberg &amp; Chen" authorityYear="2008" class="Insecta" family="Braconidae" genus="Sculptolobus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sculptolobus" order="Hymenoptera" pageId="0" pageNumber="301" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="301">Sculptolobus</emphasis>
</taxonomicName>
requires a larger-scale revision of many genera and is far beyond the scope of the current article. The diagnostic characters of the genus 
<taxonomicName authorityName="Yang, van Achterberg &amp; Chen" authorityYear="2008" class="Insecta" family="Braconidae" genus="Sculptolobus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Sculptolobus" order="Hymenoptera" pageId="0" pageNumber="301" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="301">Sculptolobus</emphasis>
</taxonomicName>
are as follows:
</paragraph>
<paragraph pageId="0" pageNumber="301">
Scape with dorsal side longer than ventral side (correctly figured but mistakenly reversed in the original description). Face with weakly differentiated triangular area above clypeus and more or less developed medio-longitudinal elevation (Figs 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Sculptolobus bipustulatus (Szepligeti, 1913), comb. nov. (A-I holotype, female, HNHM J-Q, S-U female, RMNH R female ZISP V male, RMNH) A, J habitus, lateral view B, R T 1, dorsal view C tarsal claw D, L head, lateral view E, O mesosoma, lateral view F head, dorsal view G, Q propodeum, dorsal view H, T mesosoma, dorsal view I, U metasoma, dorsal view K fore wing M head, ventrolateral view N head, anterior view P apex of ovipositor S hind tarsus, anterior view V habitus, dorsal view. ala anterolateral area dc dorsal carina dlc dorsolateral carina slc sublateral carina. Scale bars: 1 mm (A, J, K, V); 0.5 mm (B, E, H, I, O, T, U); 0.25 mm (C, D, F, G, L-N, P-S)." figureDoi="10.3897/jhr.84.68702.figure2" httpUri="https://binary.pensoft.net/fig/579974" pageId="0" pageNumber="301">2M</figureCitation>
, 
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Sculptolobus lembaensis harteni ssp. nov. (A-E, G-L holotype, female, RMNH F paratype, female, RMNH M, N paratype, male, RMNH) A, M habitus, lateral view B head, dorsal view C head, lateral view D head, anterior view E head, ventrolateral view F fore wing G propodeum, dorsolateral view H T 1, dorsal view I mesosoma, lateral view J metasoma, dorsolateral view K mesosoma, dorsal view L apex of ovipositor N propodeum and T 1 - T 3, dorsal view. Scale bars: 1 mm (A, F, M); 0.5 mm (I-K, N); 0.25 mm (B-E, G, H, L-N)." figureDoi="10.3897/jhr.84.68702.figure5" httpUri="https://binary.pensoft.net/fig/579977" pageId="0" pageNumber="301">5E</figureCitation>
, 
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Sculptolobus subellipticus (Granger, 1949), comb. nov. (lectotype, female, MNHN) A habitus, lateral view B head, lateral view C head, anterior view D head, ventrolateral view E head, dorsal view F mesosoma, lateral view G fore wing H propodeum, dorsal view I mesosoma, dorsal view J metasoma, dorsal view K fore tibia M middle and hind legs N T 1, dorsal view. Scale bars: 1 mm (A, G); 0.5 mm (F, M, I, J); 0.25 mm (B-E, H, K, L, N)." figureDoi="10.3897/jhr.84.68702.figure8" httpUri="https://binary.pensoft.net/fig/579980" pageId="0" pageNumber="301">8D</figureCitation>
), sometimes very prominent and incorporating clypeus (Fig. 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Sculptolobus biareolatus (Granger, 1949) (holotype, female, MNHN) A habitus, lateral view B head, lateral view C head, dorsal view D apex of ovipositor E head, ventrolateral view F head, anterior view G fore wing H metanotum, dorsal view I mesosoma, dorsal view J metasoma, dorsal view K mesosoma, lateral view L T 1 - T 2, dorsal view. Scale bars: 1 mm (A, G); 0.5 mm (I-L); 0.25 mm (B-F, H)." figureDoi="10.3897/jhr.84.68702.figure1" httpUri="https://binary.pensoft.net/fig/579973" pageId="0" pageNumber="301">1B</figureCitation>
). Clypeus flattened, with ventral rim (almost) not protruding and with no or weak dorsal carina. Malar suture weakly (Figs 
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Sculptolobus somnialis (Szepligeti, 1913), comb. nov. (A-C, E, G, H female, RMNH D, I male, RMNH F female, ZISP) and S. suavis (Szepligeti, 1918), comb. nov. (J-R lectotype, female, MNB) A, L head, anterior view B head, ventrolateral view C head, lateral view D, G metasoma, dorsolateral view E, N head and mesosoma, lateral view F, H, R metasoma, dorsal view I, J habitus, lateral view K head, dorsal view M labels O T 1, dorsal view P fore wing Q propodeum, dorsal view. Scale bars: 1 mm (I, J); 0.5 mm (D-H, N, P, R); 0.25 mm (A-C, K, L, O, Q)." figureDoi="10.3897/jhr.84.68702.figure7" httpUri="https://binary.pensoft.net/fig/579979" pageId="0" pageNumber="301">7B</figureCitation>
, 
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Sculptolobus subellipticus (Granger, 1949), comb. nov. (lectotype, female, MNHN) A habitus, lateral view B head, lateral view C head, anterior view D head, ventrolateral view E head, dorsal view F mesosoma, lateral view G fore wing H propodeum, dorsal view I mesosoma, dorsal view J metasoma, dorsal view K fore tibia M middle and hind legs N T 1, dorsal view. Scale bars: 1 mm (A, G); 0.5 mm (F, M, I, J); 0.25 mm (B-E, H, K, L, N)." figureDoi="10.3897/jhr.84.68702.figure8" httpUri="https://binary.pensoft.net/fig/579980" pageId="0" pageNumber="301">8B, D</figureCitation>
) or deeply (Figs 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Sculptolobus bipustulatus (Szepligeti, 1913), comb. nov. (A-I holotype, female, HNHM J-Q, S-U female, RMNH R female ZISP V male, RMNH) A, J habitus, lateral view B, R T 1, dorsal view C tarsal claw D, L head, lateral view E, O mesosoma, lateral view F head, dorsal view G, Q propodeum, dorsal view H, T mesosoma, dorsal view I, U metasoma, dorsal view K fore wing M head, ventrolateral view N head, anterior view P apex of ovipositor S hind tarsus, anterior view V habitus, dorsal view. ala anterolateral area dc dorsal carina dlc dorsolateral carina slc sublateral carina. Scale bars: 1 mm (A, J, K, V); 0.5 mm (B, E, H, I, O, T, U); 0.25 mm (C, D, F, G, L-N, P-S)." figureDoi="10.3897/jhr.84.68702.figure2" httpUri="https://binary.pensoft.net/fig/579974" pageId="0" pageNumber="301">2M</figureCitation>
, 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Sculptolobus leleji sp. nov. (A-F, I-O holotype, female, RMNH G, H paratype, female, RMNH) A, G habitus, lateral view B head, lateral view C head, dorsal view D head, ventrolateral view E head, anterior view F apex of ovipositor H fore wing I mesosoma, lateral view J metasoma, dorsolateral view K metasoma, dorsal view L mesosoma, dorsal view M T 1, dorsolateral view N metanotum, dorsal view O hind tarsus. ala anterolateral area dli dorsolateral impression s spiracle. Scale bars: 1 mm (A, G, H); 0.5 mm (I-L); 0.25 mm (B-F, M-O)." figureDoi="10.3897/jhr.84.68702.figure3" httpUri="https://binary.pensoft.net/fig/579975" pageId="0" pageNumber="301">3D</figureCitation>
) impressed; malar space not impressed. Notauli deep and often sculptured anteriorly, shallow posteriorly (Figs 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Sculptolobus bipustulatus (Szepligeti, 1913), comb. nov. (A-I holotype, female, HNHM J-Q, S-U female, RMNH R female ZISP V male, RMNH) A, J habitus, lateral view B, R T 1, dorsal view C tarsal claw D, L head, lateral view E, O mesosoma, lateral view F head, dorsal view G, Q propodeum, dorsal view H, T mesosoma, dorsal view I, U metasoma, dorsal view K fore wing M head, ventrolateral view N head, anterior view P apex of ovipositor S hind tarsus, anterior view V habitus, dorsal view. ala anterolateral area dc dorsal carina dlc dorsolateral carina slc sublateral carina. Scale bars: 1 mm (A, J, K, V); 0.5 mm (B, E, H, I, O, T, U); 0.25 mm (C, D, F, G, L-N, P-S)." figureDoi="10.3897/jhr.84.68702.figure2" httpUri="https://binary.pensoft.net/fig/579974" pageId="0" pageNumber="301">2T</figureCitation>
, 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Sculptolobus leleji sp. nov. (A-F, I-O holotype, female, RMNH G, H paratype, female, RMNH) A, G habitus, lateral view B head, lateral view C head, dorsal view D head, ventrolateral view E head, anterior view F apex of ovipositor H fore wing I mesosoma, lateral view J metasoma, dorsolateral view K metasoma, dorsal view L mesosoma, dorsal view M T 1, dorsolateral view N metanotum, dorsal view O hind tarsus. ala anterolateral area dli dorsolateral impression s spiracle. Scale bars: 1 mm (A, G, H); 0.5 mm (I-L); 0.25 mm (B-F, M-O)." figureDoi="10.3897/jhr.84.68702.figure3" httpUri="https://binary.pensoft.net/fig/579975" pageId="0" pageNumber="301">3L</figureCitation>
). Precoxal sulcus shallowly and widely impressed (Figs 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Sculptolobus leleji sp. nov. (A-F, I-O holotype, female, RMNH G, H paratype, female, RMNH) A, G habitus, lateral view B head, lateral view C head, dorsal view D head, ventrolateral view E head, anterior view F apex of ovipositor H fore wing I mesosoma, lateral view J metasoma, dorsolateral view K metasoma, dorsal view L mesosoma, dorsal view M T 1, dorsolateral view N metanotum, dorsal view O hind tarsus. ala anterolateral area dli dorsolateral impression s spiracle. Scale bars: 1 mm (A, G, H); 0.5 mm (I-L); 0.25 mm (B-F, M-O)." figureDoi="10.3897/jhr.84.68702.figure3" httpUri="https://binary.pensoft.net/fig/579975" pageId="0" pageNumber="301">3I</figureCitation>
, 
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Sculptolobus lembaensis (Cameron, 1912), comb. nov. (lectotype, female, RMCA) A habitus, lateral view B labels C apex of ovipositor D head, dorsal view E head, lateral view F mesosoma, dorsal view G head, anterior view H head, ventrolateral view I metasoma, dorsolateral view J fore wing K mesosoma, lateral view L propodeum, dorsolateral view. Scale bars: 1 mm (A, J); 0.5 mm (E, K, I); 0.25 mm (C, D, F-H, L)." figureDoi="10.3897/jhr.84.68702.figure4" httpUri="https://binary.pensoft.net/fig/579976" pageId="0" pageNumber="301">4K</figureCitation>
). Fore wing vein 1-SR+M straight, vein 1-SR very short. Angle between veins C+SC+R and 1-SR about 70-85°. Base of hind wing membrane evenly setose (Fig. 
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Sculptolobus biareolatus (Granger, 1949) (holotype, female, MNHN) A habitus, lateral view B head, lateral view C head, dorsal view D apex of ovipositor E head, ventrolateral view F head, anterior view G fore wing H metanotum, dorsal view I mesosoma, dorsal view J metasoma, dorsal view K mesosoma, lateral view L T 1 - T 2, dorsal view. Scale bars: 1 mm (A, G); 0.5 mm (I-L); 0.25 mm (B-F, H)." figureDoi="10.3897/jhr.84.68702.figure1" httpUri="https://binary.pensoft.net/fig/579973" pageId="0" pageNumber="301">1G</figureCitation>
). Fore tibia with thick setae only near its apex (Fig. 
<figureCitation captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Sculptolobus subellipticus (Granger, 1949), comb. nov. (lectotype, female, MNHN) A habitus, lateral view B head, lateral view C head, anterior view D head, ventrolateral view E head, dorsal view F mesosoma, lateral view G fore wing H propodeum, dorsal view I mesosoma, dorsal view J metasoma, dorsal view K fore tibia M middle and hind legs N T 1, dorsal view. Scale bars: 1 mm (A, G); 0.5 mm (F, M, I, J); 0.25 mm (B-E, H, K, L, N)." figureDoi="10.3897/jhr.84.68702.figure8" httpUri="https://binary.pensoft.net/fig/579980" pageId="0" pageNumber="301">8K</figureCitation>
). Hind tibia without subapical row of thick setae (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Sculptolobus leleji sp. nov. (A-F, I-O holotype, female, RMNH G, H paratype, female, RMNH) A, G habitus, lateral view B head, lateral view C head, dorsal view D head, ventrolateral view E head, anterior view F apex of ovipositor H fore wing I mesosoma, lateral view J metasoma, dorsolateral view K metasoma, dorsal view L mesosoma, dorsal view M T 1, dorsolateral view N metanotum, dorsal view O hind tarsus. ala anterolateral area dli dorsolateral impression s spiracle. Scale bars: 1 mm (A, G, H); 0.5 mm (I-L); 0.25 mm (B-F, M-O)." figureDoi="10.3897/jhr.84.68702.figure3" httpUri="https://binary.pensoft.net/fig/579975" pageId="0" pageNumber="301">3O</figureCitation>
). Tarsal claws with protruding basal lobes (Figs 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Sculptolobus bipustulatus (Szepligeti, 1913), comb. nov. (A-I holotype, female, HNHM J-Q, S-U female, RMNH R female ZISP V male, RMNH) A, J habitus, lateral view B, R T 1, dorsal view C tarsal claw D, L head, lateral view E, O mesosoma, lateral view F head, dorsal view G, Q propodeum, dorsal view H, T mesosoma, dorsal view I, U metasoma, dorsal view K fore wing M head, ventrolateral view N head, anterior view P apex of ovipositor S hind tarsus, anterior view V habitus, dorsal view. ala anterolateral area dc dorsal carina dlc dorsolateral carina slc sublateral carina. Scale bars: 1 mm (A, J, K, V); 0.5 mm (B, E, H, I, O, T, U); 0.25 mm (C, D, F, G, L-N, P-S)." figureDoi="10.3897/jhr.84.68702.figure2" httpUri="https://binary.pensoft.net/fig/579974" pageId="0" pageNumber="301">2S</figureCitation>
, 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Sculptolobus leleji sp. nov. (A-F, I-O holotype, female, RMNH G, H paratype, female, RMNH) A, G habitus, lateral view B head, lateral view C head, dorsal view D head, ventrolateral view E head, anterior view F apex of ovipositor H fore wing I mesosoma, lateral view J metasoma, dorsolateral view K metasoma, dorsal view L mesosoma, dorsal view M T 1, dorsolateral view N metanotum, dorsal view O hind tarsus. ala anterolateral area dli dorsolateral impression s spiracle. Scale bars: 1 mm (A, G, H); 0.5 mm (I-L); 0.25 mm (B-F, M-O)." figureDoi="10.3897/jhr.84.68702.figure3" httpUri="https://binary.pensoft.net/fig/579975" pageId="0" pageNumber="301">3O</figureCitation>
). Metasoma with six coarsely sclerotised tergites. Median area of T2 parallel-sided, incomplete, and laterally delineated by deep furrows. Anterolateral areas of T2 (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Sculptolobus bipustulatus (Szepligeti, 1913), comb. nov. (A-I holotype, female, HNHM J-Q, S-U female, RMNH R female ZISP V male, RMNH) A, J habitus, lateral view B, R T 1, dorsal view C tarsal claw D, L head, lateral view E, O mesosoma, lateral view F head, dorsal view G, Q propodeum, dorsal view H, T mesosoma, dorsal view I, U metasoma, dorsal view K fore wing M head, ventrolateral view N head, anterior view P apex of ovipositor S hind tarsus, anterior view V habitus, dorsal view. ala anterolateral area dc dorsal carina dlc dorsolateral carina slc sublateral carina. Scale bars: 1 mm (A, J, K, V); 0.5 mm (B, E, H, I, O, T, U); 0.25 mm (C, D, F, G, L-N, P-S)." figureDoi="10.3897/jhr.84.68702.figure2" httpUri="https://binary.pensoft.net/fig/579974" pageId="0" pageNumber="301">2U</figureCitation>
: ala) round, convex, separated by crenulate margin and/or furrows, with distal margins bordered by posteriorly converging sublateral carinae (Fig. 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Sculptolobus bipustulatus (Szepligeti, 1913), comb. nov. (A-I holotype, female, HNHM J-Q, S-U female, RMNH R female ZISP V male, RMNH) A, J habitus, lateral view B, R T 1, dorsal view C tarsal claw D, L head, lateral view E, O mesosoma, lateral view F head, dorsal view G, Q propodeum, dorsal view H, T mesosoma, dorsal view I, U metasoma, dorsal view K fore wing M head, ventrolateral view N head, anterior view P apex of ovipositor S hind tarsus, anterior view V habitus, dorsal view. ala anterolateral area dc dorsal carina dlc dorsolateral carina slc sublateral carina. Scale bars: 1 mm (A, J, K, V); 0.5 mm (B, E, H, I, O, T, U); 0.25 mm (C, D, F, G, L-N, P-S)." figureDoi="10.3897/jhr.84.68702.figure2" httpUri="https://binary.pensoft.net/fig/579974" pageId="0" pageNumber="301">2U</figureCitation>
: slc). Spiracle of T2 located on dorsum of tergite, often far from its lateral margin (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Sculptolobus leleji sp. nov. (A-F, I-O holotype, female, RMNH G, H paratype, female, RMNH) A, G habitus, lateral view B head, lateral view C head, dorsal view D head, ventrolateral view E head, anterior view F apex of ovipositor H fore wing I mesosoma, lateral view J metasoma, dorsolateral view K metasoma, dorsal view L mesosoma, dorsal view M T 1, dorsolateral view N metanotum, dorsal view O hind tarsus. ala anterolateral area dli dorsolateral impression s spiracle. Scale bars: 1 mm (A, G, H); 0.5 mm (I-L); 0.25 mm (B-F, M-O)." figureDoi="10.3897/jhr.84.68702.figure3" httpUri="https://binary.pensoft.net/fig/579975" pageId="0" pageNumber="301">3I, J</figureCitation>
: s). Median area of T3 absent. T3-T5 with small anterolateral areas around spiracles separated by crenulate furrows (Fig. 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Sculptolobus leleji sp. nov. (A-F, I-O holotype, female, RMNH G, H paratype, female, RMNH) A, G habitus, lateral view B head, lateral view C head, dorsal view D head, ventrolateral view E head, anterior view F apex of ovipositor H fore wing I mesosoma, lateral view J metasoma, dorsolateral view K metasoma, dorsal view L mesosoma, dorsal view M T 1, dorsolateral view N metanotum, dorsal view O hind tarsus. ala anterolateral area dli dorsolateral impression s spiracle. Scale bars: 1 mm (A, G, H); 0.5 mm (I-L); 0.25 mm (B-F, M-O)." figureDoi="10.3897/jhr.84.68702.figure3" httpUri="https://binary.pensoft.net/fig/579975" pageId="0" pageNumber="301">3J</figureCitation>
: ala). Apical margins of T3-T6 thick, laterally straight (only margin of T3 acutely protruding). Ovipositor sheath 0.5-2.0 
<normalizedToken originalValue="×">x</normalizedToken>
as long as hind tibia; apex of ovipositor with developed dorsal nodus and ventral serration (Figs 
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Sculptolobus bipustulatus (Szepligeti, 1913), comb. nov. (A-I holotype, female, HNHM J-Q, S-U female, RMNH R female ZISP V male, RMNH) A, J habitus, lateral view B, R T 1, dorsal view C tarsal claw D, L head, lateral view E, O mesosoma, lateral view F head, dorsal view G, Q propodeum, dorsal view H, T mesosoma, dorsal view I, U metasoma, dorsal view K fore wing M head, ventrolateral view N head, anterior view P apex of ovipositor S hind tarsus, anterior view V habitus, dorsal view. ala anterolateral area dc dorsal carina dlc dorsolateral carina slc sublateral carina. Scale bars: 1 mm (A, J, K, V); 0.5 mm (B, E, H, I, O, T, U); 0.25 mm (C, D, F, G, L-N, P-S)." figureDoi="10.3897/jhr.84.68702.figure2" httpUri="https://binary.pensoft.net/fig/579974" pageId="0" pageNumber="301">2P</figureCitation>
, 
<figureCitation captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Sculptolobus leleji sp. nov. (A-F, I-O holotype, female, RMNH G, H paratype, female, RMNH) A, G habitus, lateral view B head, lateral view C head, dorsal view D head, ventrolateral view E head, anterior view F apex of ovipositor H fore wing I mesosoma, lateral view J metasoma, dorsolateral view K metasoma, dorsal view L mesosoma, dorsal view M T 1, dorsolateral view N metanotum, dorsal view O hind tarsus. ala anterolateral area dli dorsolateral impression s spiracle. Scale bars: 1 mm (A, G, H); 0.5 mm (I-L); 0.25 mm (B-F, M-O)." figureDoi="10.3897/jhr.84.68702.figure3" httpUri="https://binary.pensoft.net/fig/579975" pageId="0" pageNumber="301">3F</figureCitation>
, 
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Sculptolobus lembaensis harteni ssp. nov. (A-E, G-L holotype, female, RMNH F paratype, female, RMNH M, N paratype, male, RMNH) A, M habitus, lateral view B head, dorsal view C head, lateral view D head, anterior view E head, ventrolateral view F fore wing G propodeum, dorsolateral view H T 1, dorsal view I mesosoma, lateral view J metasoma, dorsolateral view K mesosoma, dorsal view L apex of ovipositor N propodeum and T 1 - T 3, dorsal view. Scale bars: 1 mm (A, F, M); 0.5 mm (I-K, N); 0.25 mm (B-E, G, H, L-N)." figureDoi="10.3897/jhr.84.68702.figure5" httpUri="https://binary.pensoft.net/fig/579977" pageId="0" pageNumber="301">5L</figureCitation>
). Head, mesosoma, coxae and most of metasoma granulate or coriaceous.
</paragraph>
</subSubSection>
</treatment>
</document>