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<document id="60C9431A75D63D7E1C897F8E505614AA" ID-DOI="10.1206/0003-0090.442.1.1" ID-ISSN="0003-0090" ID-Zenodo-Dep="5411297" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630352812419" checkinUser="felipe" docAuthor="Prendini, Lorenzo &amp; Loria, Stephanie F." docDate="2020" docId="5E6CB374FF0A6DF2FD7865DEFB58DB6F" docLanguage="en" docName="B442 highres.pdf" docOrigin="Bulletin of the American Museum of Natural History 2020 (442)" docSource="https://bioone.org/journals/bulletin-of-the-american-museum-of-natural-history/volume-442/issue-1/0003-0090.442.1.1/Systematic-Revision-of-the-Asian-Forest-Scorpions-Heterometrinae-Simon-1879/10.1206/0003-0090.442.1.1.full" docStyle="DocumentStyle:C5E2DA72A22EF33813C92A197453A310.5:BulAmeMusNatHis.2011-.journal_article.0cover" docStyleId="C5E2DA72A22EF33813C92A197453A310" docStyleName="BulAmeMusNatHis.2011-.journal_article.0cover" docStyleVersion="5" docTitle="Heterometrus longimanus LP" docType="treatment" docVersion="4" lastPageNumber="252" masterDocId="A255CB0CFFF16D09FFAF6038FFF6DD40" masterDocTitle="Systematic Revision Of The Asian Forest Scorpions (Heterometrinae Simon, 1879), Revised Suprageneric Classification Of Scorpionidae Latreille, 1802, And Revalidation Of Rugodentidae Bastawade Et Al., 2005" masterLastPageNumber="480" masterPageNumber="1" pageNumber="252" updateTime="1699242878868" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="79F78FBCA680E2BDED2530360A95D2BD">Systematic Revision Of The Asian Forest Scorpions (Heterometrinae Simon, 1879), Revised Suprageneric Classification Of Scorpionidae Latreille, 1802, And Revalidation Of Rugodentidae Bastawade Et Al., 2005</mods:title>
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<mods:namePart id="C9839724AC02E0D70BE6EBCFD999A1F1">Prendini, Lorenzo</mods:namePart>
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<treatment id="5E6CB374FF0A6DF2FD7865DEFB58DB6F" ID-DOI="http://doi.org/10.5281/zenodo.5478615" ID-Zenodo-Dep="5478615" LSID="urn:lsid:plazi:treatment:5E6CB374FF0A6DF2FD7865DEFB58DB6F" httpUri="http://treatment.plazi.org/id/5E6CB374FF0A6DF2FD7865DEFB58DB6F" lastPageNumber="252" pageId="251" pageNumber="252">
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<paragraph id="D67A0262FF0A6DF2FD7865DEFB64D8BD" blockId="251.[727,1170,1509,1533]" box="[727,1170,1509,1533]" pageId="251" pageNumber="252">
<heading id="8D32B50EFF0A6DF2FD7865DEFB64D8BD" box="[727,1170,1509,1533]" centered="true" fontSize="9" level="2" pageId="251" pageNumber="252" reason="2">
<taxonomicName id="11C579E1FF0A6DF2FD7865DEFB64D8BD" ID-CoL="6LSX9" authority="(Herbst, 1800)" authorityName="LP" baseAuthorityName="Herbst" baseAuthorityYear="1800" box="[727,1170,1509,1533]" class="Arachnida" family="Scorpionidae" genus="Heterometrus" kingdom="Animalia" order="Scorpiones" pageId="251" pageNumber="252" phylum="Arthropoda" rank="species" species="longimanus">
<emphasis id="E4B1DE70FF0A6DF2FD7865DEFC1DD8BD" box="[727,1003,1509,1533]" italics="true" pageId="251" pageNumber="252">Heterometrus longimanus</emphasis>
(
<bibRefCitation id="B2547F93FF0A6DF2FC5465DDFB7DD8BD" author="Herbst, J. F. W." box="[1019,1163,1509,1533]" pageId="251" pageNumber="252" pagination="1 - 86" refId="ref239146" refString="Herbst, J. F. W. 1800. Naturgeschichte der Skorpionen. In Natursystem der Ungeflugelten Insekten 4: 1 - 86. Berlin: Gottlieb August Lange." type="journal article" year="1800">Herbst, 1800</bibRefCitation>
)
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<subSubSection id="9EDF51E9FF0A6DF2FD16662FFB58DB6F" box="[697,1198,1559,1583]" pageId="251" pageNumber="252" type="description">
<paragraph id="D67A0262FF0A6DF2FD16662FFB58DB6F" blockId="251.[697,1198,1559,1616]" box="[697,1198,1559,1583]" pageId="251" pageNumber="252">
<heading id="8D32B50EFF0A6DF2FD16662FFB58DB6F" box="[697,1198,1559,1583]" centered="true" fontSize="9" level="2" pageId="251" pageNumber="252" reason="2">
<figureCitation id="4EFE1EE7FF0A6DF2FD16662FFCDBDB6F" box="[697,813,1559,1583]" captionStart="FIGURE 10" captionStartId="25.[92,179,1416,1440]" captionTargetBox="[82,1612,415,1388]" captionTargetId="graphics-9@25.[82,1465,419,1375]" captionTargetPageId="25" captionText="FIGURE 10. Phylogeny of Asian forest scorpions (Scorpionidae: Heterometrinae Simon, 1879, stat. nov.), obtained by simultaneous phylogenetic analysis of 186 morphological characters and 4188 aligned DNA nucleotides from three mitochondrial and two nuclear gene loci (Loria and Prendini, in press). Maximum likelihood tree with unambiguous morphological synapomorphies optimized. Black circles indicate uniquely derived apomorphic states, white circles parallel derivations of apomorphic states. Numbers above circles indicate characters, numbers below indicate states (appendix 1)." figureDoi="http://doi.org/10.5281/zenodo.5411321" httpUri="https://zenodo.org/record/5411321/files/figure.png" pageId="251" pageNumber="252">Figures 10</figureCitation>
,
<figureCitation id="4EFE1EE7FF0A6DF2FC96662FFC71DB6E" box="[825,903,1559,1583]" captionStart="FIGURE 22" captionStartId="66.[108,191,1627,1648]" captionTargetBox="[107,1212,224,1599]" captionTargetId="figure-13@66.[107,1212,224,1600]" captionTargetPageId="66" captionText="FIGURE 22. Heterometrus Ehrenberg, 1828, carapace, dorsal aspect. A, B. Heterometrus glaucus (Thorell, 1876), comb. nov. et stat. rev.,A. ♂ (ZMH 3966), B. ♀ (AMNH), Nias Island, Indonesia. C, D. Heterometrus longimanus (Herbst, 1800), C. ♂, D. ♀ (AMNH), Kuching, Malaysia. E, F. Heterometrus thorellii (Pocock, 1892), E. ♂ (AMNH), Mandalay, Myanmar, F. ♀ (AMNH), Pho Kyar Elephant Camp, Yedashe, Myanmar. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5411353" httpUri="https://zenodo.org/record/5411353/files/figure.png" pageId="251" pageNumber="252">22C, D</figureCitation>
,
<figureCitation id="4EFE1EE7FF0A6DF2FC3C662FFC14DB6E" box="[915,994,1559,1583]" captionStart="FIGURE 36" captionStartId="94.[108,194,1624,1645]" captionTargetBox="[107,1211,224,1597]" captionTargetId="figure-12@94.[107,1211,224,1597]" captionTargetPageId="94" captionText="FIGURE 36. Heterometrus Ehrenberg, 1828, sternum and pectines, ventral aspect. A, B. Heterometrus glaucus (Thorell, 1876), comb. nov. et stat. rev., A. ♂ (ZMH), Padang, West Sumatra, Indonesia. B. ♀ (AMNH), Nias Island, Indonesia. C, D. Heterometrus longimanus (Herbst, 1800), C. ♂, D. ♀ (AMNH), Kuching, Malaysia. E, F. Heterometrus thorellii (Pocock, 1892), E. ♂ (AMNH), Mandalay, Myanmar, F. ♀ (AMNH), Pho Kyar Elephant Camp, Yedashe, Myanmar. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5411397" httpUri="https://zenodo.org/record/5411397/files/figure.png" pageId="251" pageNumber="252">36C, D</figureCitation>
,
<figureCitation id="4EFE1EE7FF0A6DF2FC42662FFBCADB6F" box="[1005,1084,1559,1583]" captionStart="FIGURE 52" captionStartId="121.[108,194,1167,1188]" captionTargetBox="[113,1204,225,1138]" captionTargetId="figure-249@121.[108,1212,224,1141]" captionTargetPageId="121" captionText="FIGURE 52. Heterometrus Ehrenberg, 1828, legs I–IV, tibiae, basitarsi and telotarsi, ventral aspect. A–D. Heterometrus glaucus (Thorell, 1876), comb. nov. et stat. rev., ♀ (AMNH), Nias Island, Indonesia. E–H. Heterometrus longimanus (Herbst, 1800), ♂ (AMNH), Kuching, Malaysia. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5411454" httpUri="https://zenodo.org/record/5411454/files/figure.png" pageId="251" pageNumber="252">52E–H</figureCitation>
,
<figureCitation id="4EFE1EE7FF0A6DF2FBE8662FFB85DB6F" box="[1095,1139,1559,1583]" captionStart="FIGURE 70" captionStartId="150.[108,194,1466,1487]" captionTargetBox="[108,1212,224,1438]" captionTargetId="figure-12@150.[108,1212,224,1438]" captionTargetPageId="150" captionText="FIGURE 70. A–D. Heterometrus Ehrenberg, 1828, and E. Javanimetrus Couzijn, 1981, stat. nov. et stat. rev., metasoma and telson, dorsal aspect. A. Heterometrus laoticus Couzijn, 1981, ♂ (AMNH), Phahom Camp, Laos. B. Heterometrus longimanus (Herbst, 1800), ♂ (AMNH), Kuching, Malaysia. C. Heterometrus silenus (Simon, 1884), comb. nov. et stat. rev., ♂ (AMNH), Ta Kou Mountain Nature Reserve, Vietnam. D. Heterometrus thorellii (Pocock, 1892), ♀ (AMNH), Pho Kyar Elephant Camp, Yedashe, Myanmar. E. Javanimetrus cyaneus (Simon, 1872), comb. nov., ♂ (AMNH), Ujung Kulon, Indonesia. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5411504" httpUri="https://zenodo.org/record/5411504/files/figure.png" pageId="251" pageNumber="252">70B</figureCitation>
,
<figureCitation id="4EFE1EE7FF0A6DF2FBD0662FFB5CDB6F" box="[1151,1194,1559,1583]" captionStart="FIGURE 71" captionStartId="151.[107,193,1470,1491]" captionTargetBox="[108,1212,224,1443]" captionTargetId="figure-9@151.[108,1212,224,1444]" captionTargetPageId="151" captionText="FIGURE 71. A–D. Heterometrus Ehrenberg, 1828, and E. Javanimetrus Couzijn, 1981, stat. nov. et stat. rev., metasoma and telson, lateral aspect. A. Heterometrus laoticus Couzijn, 1981, ♂ (AMNH), Phahom Camp, Laos. B. Heterometrus longimanus (Herbst, 1800), ♂ (AMNH), Kuching, Malaysia. C. Heterometrus silenus (Simon, 1884), comb. nov. et stat. rev., ♂ (AMNH), Ta Kou Mountain Nature Reserve, Vietnam. D. Heterometrus thorellii (Pocock, 1892), ♀ (AMNH), Pho Kyar Elephant Camp, Yedashe, Myanmar. E. Javanimetrus cyaneus (Simon, 1872), comb. nov., ♂ (AMNH), Ujung Kulon, Indonesia. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5411508" httpUri="https://zenodo.org/record/5411508/files/figure.png" pageId="251" pageNumber="252">71B</figureCitation>
,
</heading>
</paragraph>
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