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<document id="A67F64F6D40C69C4430C349AC9C60568" ID-DOI="10.1206/0003-0090.442.1.1" ID-ISSN="0003-0090" ID-Zenodo-Dep="5411297" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630352812419" checkinUser="felipe" docAuthor="Prendini, Lorenzo &amp; Loria, Stephanie F." docDate="2020" docId="5E6CB374FE7E6C86FF11641CFD98D92E" docLanguage="en" docName="B442 highres.pdf" docOrigin="Bulletin of the American Museum of Natural History 2020 (442)" docSource="https://bioone.org/journals/bulletin-of-the-american-museum-of-natural-history/volume-442/issue-1/0003-0090.442.1.1/Systematic-Revision-of-the-Asian-Forest-Scorpions-Heterometrinae-Simon-1879/10.1206/0003-0090.442.1.1.full" docStyle="DocumentStyle:C5E2DA72A22EF33813C92A197453A310.5:BulAmeMusNatHis.2011-.journal_article.0cover" docStyleId="C5E2DA72A22EF33813C92A197453A310" docStyleName="BulAmeMusNatHis.2011-.journal_article.0cover" docStyleVersion="5" docTitle="Srilankametrus couzijni Prendini &amp; Loria 2020, sp. nov." docType="treatment" docVersion="4" lastPageNumber="400" masterDocId="A255CB0CFFF16D09FFAF6038FFF6DD40" masterDocTitle="Systematic Revision Of The Asian Forest Scorpions (Heterometrinae Simon, 1879), Revised Suprageneric Classification Of Scorpionidae Latreille, 1802, And Revalidation Of Rugodentidae Bastawade Et Al., 2005" masterLastPageNumber="480" masterPageNumber="1" pageNumber="400" updateTime="1699242878868" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="AE7C288E78D45B9E6FEF9E00E80C08FF">Systematic Revision Of The Asian Forest Scorpions (Heterometrinae Simon, 1879), Revised Suprageneric Classification Of Scorpionidae Latreille, 1802, And Revalidation Of Rugodentidae Bastawade Et Al., 2005</mods:title>
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<mods:namePart id="7916DC7441F7E184D494D4BC8533B1EE">Prendini, Lorenzo</mods:namePart>
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<mods:namePart id="35FE97E5449F7AB1004B3A5B464F8DC4">Loria, Stephanie F.</mods:namePart>
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<treatment id="5E6CB374FE7E6C86FF11641CFD98D92E" ID-DOI="http://doi.org/10.5281/zenodo.5478625" ID-Zenodo-Dep="5478625" LSID="urn:lsid:plazi:treatment:5E6CB374FE7E6C86FF11641CFD98D92E" httpUri="http://treatment.plazi.org/id/5E6CB374FE7E6C86FF11641CFD98D92E" lastPageNumber="400" pageId="399" pageNumber="400">
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<heading id="8D32B50EFE7E6C86FF11641CFDDDD97C" box="[190,555,1060,1085]" centered="true" fontSize="9" level="2" pageId="399" pageNumber="400" reason="2">
<emphasis id="E4B1DE70FE7E6C86FF11641CFE27D97D" bold="true" box="[190,465,1060,1085]" pageId="399" pageNumber="400">
<taxonomicName id="11C579E1FE7E6C86FF11641CFE3CD97D" ID-CoL="8LYZG" authority="Prendini &amp; Loria, 2020" authorityName="Prendini &amp; Loria" authorityYear="2020" box="[190,458,1060,1085]" class="Arachnida" family="Scorpionidae" genus="Srilankametrus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Scorpiones" pageId="399" pageNumber="400" phylum="Arthropoda" rank="species" species="couzijni" status="sp. nov.">
<emphasis id="E4B1DE70FE7E6C86FF11641CFE3CD97D" bold="true" box="[190,458,1060,1085]" italics="true" pageId="399" pageNumber="400">Srilankametrus couzijni</emphasis>
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,
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<taxonomicNameLabel id="FF82630BFE7E6C86FE77641DFDDDD97C" box="[472,555,1061,1084]" pageId="399" pageNumber="400" rank="species">sp. nov.</taxonomicNameLabel>
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<subSubSection id="9EDF51E9FE7E6C86FFD6646EFD98D92E" box="[121,622,1110,1135]" pageId="399" pageNumber="400" type="description">
<paragraph id="D67A0262FE7E6C86FFD6646EFD98D92E" blockId="399.[121,622,1110,1168]" box="[121,622,1110,1135]" pageId="399" pageNumber="400">
<heading id="8D32B50EFE7E6C86FFD6646EFD98D92E" box="[121,622,1110,1135]" centered="true" fontSize="9" level="2" pageId="399" pageNumber="400" reason="2">
<figureCitation id="4EFE1EE7FE7E6C86FFD6646EFF1BD92F" box="[121,237,1110,1135]" captionStart="FIGURE 10" captionStartId="25.[92,179,1416,1440]" captionTargetBox="[82,1612,415,1388]" captionTargetId="graphics-9@25.[82,1465,419,1375]" captionTargetPageId="25" captionText="FIGURE 10. Phylogeny of Asian forest scorpions (Scorpionidae: Heterometrinae Simon, 1879, stat. nov.), obtained by simultaneous phylogenetic analysis of 186 morphological characters and 4188 aligned DNA nucleotides from three mitochondrial and two nuclear gene loci (Loria and Prendini, in press). Maximum likelihood tree with unambiguous morphological synapomorphies optimized. Black circles indicate uniquely derived apomorphic states, white circles parallel derivations of apomorphic states. Numbers above circles indicate characters, numbers below indicate states (appendix 1)." figureDoi="http://doi.org/10.5281/zenodo.5411321" httpUri="https://zenodo.org/record/5411321/files/figure.png" pageId="399" pageNumber="400">Figures 10</figureCitation>
,
<figureCitation id="4EFE1EE7FE7E6C86FF56646FFEB1D92E" box="[249,327,1111,1135]" captionStart="FIGURE 27" captionStartId="75.[108,194,1628,1649]" captionTargetBox="[106,1210,224,1599]" captionTargetId="figure-10@75.[106,1210,224,1599]" captionTargetPageId="75" captionText="FIGURE 27. Srilankametrus Couzijn, 1981, stat. nov. et stat. rev., carapace, dorsal aspect. A, B. Srilankametrus caesar (C.L. Koch, 1841), comb. nov. et stat. rev., A. ♂, B. ♀ (MNHN RS 0079). C, D. Srilankametrus couzijni, sp. nov., C. holotype ♂ (BMNH 1896.7.30.121), D. paratype ♂ (SMF 5332/1). E, F. Srilankametrus gravimanus (Pocock, 1894), comb. nov., E. ♂, F. ♀ (AMNH), Kokmotte, Sri Lanka. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5411367" httpUri="https://zenodo.org/record/5411367/files/figure.png" pageId="399" pageNumber="400">27C, D</figureCitation>
,
<figureCitation id="4EFE1EE7FE7E6C86FEFC646FFE54D92E" box="[339,418,1111,1135]" captionStart="FIGURE 41" captionStartId="103.[108,194,1624,1645]" captionTargetBox="[108,1212,224,1597]" captionTargetId="figure-9@103.[108,1212,224,1597]" captionTargetPageId="103" captionText="FIGURE 41. Srilankametrus Couzijn, 1981, stat. nov. et stat. rev., sternum and pectines, ventral aspect. A, B. Srilankametrus caesar (C.L. Koch, 1841), comb. nov. et stat. rev., A. ♂, B. ♀ (MNHN RS 0079). C, D. Srilankametrus couzijni, sp. nov., C. holotype ♂ (BMNH 1896.7.30.121), D. paratype ♂ (SMF 5332/1). E, F. Srilankametrus gravimanus (Pocock, 1894), comb. nov., E. ♂, F.♀ (AMNH), Kokmotte, Sri Lanka. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5411416" httpUri="https://zenodo.org/record/5411416/files/figure.png" pageId="399" pageNumber="400">41C, D</figureCitation>
,
<figureCitation id="4EFE1EE7FE7E6C86FE02646EFE0AD92E" box="[429,508,1110,1134]" captionStart="FIGURE 56" captionStartId="129.[109,195,1628,1649]" captionTargetBox="[114,1202,228,1599]" captionTargetId="figure-10@129.[107,1211,224,1599]" captionTargetPageId="129" captionText="FIGURE 56. Srilankametrus Couzijn, 1981, stat. nov. et stat. rev., legs I–IV, tibiae, basitarsi and telotarsi, ventral aspect. A–D. Srilankametrus caesar (C.L. Koch, 1841), comb. nov. et stat. rev., ♂ (MNHN RS 0079). E–H. Srilankametrus couzijni, sp. nov., holotype ♂ (BMNH 1896.7.30.121). I–L. Srilankametrus gravimanus (Pocock, 1894), comb. nov., ♂ (AMNH), Kokmotte, Sri Lanka. Scale bars = 2 mm." figureDoi="http://doi.org/10.5281/zenodo.5411464" httpUri="https://zenodo.org/record/5411464/files/figure.png" pageId="399" pageNumber="400">56E–H</figureCitation>
,
<figureCitation id="4EFE1EE7FE7E6C86FDA8646EFDC2D92E" box="[519,564,1110,1134]" captionStart="FIGURE 76" captionStartId="158.[108,194,1533,1554]" captionTargetBox="[109,1213,224,1505]" captionTargetId="figure-110@158.[109,1213,224,1507]" captionTargetPageId="158" captionText="FIGURE 76. Srilankametrus Couzijn, 1978, stat. nov. et stat. rev., metasoma and telson, dorsal aspect. A. Srilankametrus caesar (C.L. Koch, 1841), comb. nov. et stat. rev., ♂ (MNHN RS 0079). B. Srilankametrus couzijni, sp. nov., holotype ♂ (BMNH 1896.7.30.121). C. Srilankametrus gravimanus (Pocock, 1894), comb. nov., ♂ (AMNH), Kokmotte, Sri Lanka. D. Srilankametrus indus (DeGeer, 1778), comb. nov., ♂ (AMNH), Kandy, Sri Lanka. E. Srilankametrus pococki, sp. nov., holotype ♀ (AMNH), Kodigala, Sri Lanka. F. Srilankametrus serratus (Pocock, 1900), comb. nov., ♂ (ZMH), Ceylon [Sri Lanka]. Scale bars = 5 mm" figureDoi="http://doi.org/10.5281/zenodo.5411524" httpUri="https://zenodo.org/record/5411524/files/figure.png" pageId="399" pageNumber="400">76B</figureCitation>
,
<figureCitation id="4EFE1EE7FE7E6C86FD90646FFD9DD92E" box="[575,619,1111,1134]" captionStart="FIGURE 77" captionStartId="159.[109,195,1534,1555]" captionTargetBox="[108,1212,224,1505]" captionTargetId="figure-9@159.[108,1212,224,1508]" captionTargetPageId="159" captionText="FIGURE 77. Srilankametrus Couzijn, 1978, stat. nov. et stat. rev., metasoma and telson, lateral aspect. A. Srilankametrus caesar (C.L. Koch, 1841), comb. nov. et stat. rev., ♂ (MNHN RS 0079). B. Srilankametrus couzijni, sp. nov., holotype ♂ (BMNH 1896.7.30.121). C. Srilankametrus gravimanus (Pocock, 1894), comb. nov., ♂ (AMNH), Kokmotte, Sri Lanka. D. Srilankametrus indus (DeGeer, 1778), comb. nov., ♂ (AMNH), Kandy, Sri Lanka. E. Srilankametrus pococki, sp. nov., holotype ♀ (AMNH), Kodigala, Sri Lanka. F. Srilankametrus serratus (Pocock, 1900), comb. nov., ♂ (ZMH), Ceylon [Sri Lanka]. Scale bars = 5 mm." figureDoi="http://doi.org/10.5281/zenodo.5411526" httpUri="https://zenodo.org/record/5411526/files/figure.png" pageId="399" pageNumber="400">77B</figureCitation>
,
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