treatments-xml/data/5E/4B/64/5E4B64CEA2305FE7BE0DE7E8AB65D1B9.xml

<document ID-DOI="http://dx.doi.org/10.3897/zookeys.463.8499" ID-PMC="PMC4294300" ID-Pensoft-Pub="1313-2970-463-21" ID-Pensoft-UUID="3E5CFFC6FFC91628FFF6E355951DFFF8" ID-PubMed="25589861" ID-Zenodo-Dep="578654" ID-ZooBank="71C1AE92B0CE4D26B1E5076E480180AD" ModsDocID="1313-2970-463-21" checkinTime="1451244962410" checkinUser="pensoft" docAuthor="Solodovnikov, Igor A., Zamotajlov, Alexandr S., Fominykh, Dmitriy D. &amp; Titarenko, Andrey Y." docDate="2014" docId="5E4B64CEA2305FE7BE0DE7E8AB65D1B9" docLanguage="en" docName="ZooKeys 463: 21-56" docOrigin="ZooKeys 463" docPubDate="2014-12-12" docSource="http://dx.doi.org/10.3897/zookeys.463.8499" docTitle="Carabus (Archiplectes) besleticus subsp. dsychvensis Gottwald 1985" docType="treatment" docVersion="3" id="3E5CFFC6FFC91628FFF6E355951DFFF8" lastPageNumber="31" masterDocId="3E5CFFC6FFC91628FFF6E355951DFFF8" masterDocTitle="Contribution to the knowledge of the Carabus (Archiplectes) satyrus Kurnakov, 1962, species complex in Abkhazia (Coleoptera, Carabidae, Carabini)" masterLastPageNumber="56" masterPageNumber="21" pageNumber="31" updateTime="1668159733340" updateUser="ExternalLinkService">
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<mods:title>Contribution to the knowledge of the Carabus (Archiplectes) satyrus Kurnakov, 1962, species complex in Abkhazia (Coleoptera, Carabidae, Carabini)</mods:title>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Solodovnikov, Igor A.</mods:namePart>
<mods:affiliation>Vitebsk State P. M. Masherov University, Moskovskyi prospect, 33, Vitebsk 210038, Belarus</mods:affiliation>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Zamotajlov, Alexandr S.</mods:namePart>
<mods:affiliation>Kuban State Agrarian University, Kalinin St. 13, Krasnodar 350044, Russia</mods:affiliation>
<mods:nameIdentifier type="email">a_zamotajlov@mail.ru</mods:nameIdentifier>
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Fominykh, Dmitriy D.</mods:namePart>
<mods:affiliation>Russian Entomological Society, Kuban Branch, Kalinin St. 13, Krasnodar 350044, Russia</mods:affiliation>
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<mods:name type="personal">
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<mods:roleTerm>Author</mods:roleTerm>
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<mods:namePart>Titarenko, Andrey Y.</mods:namePart>
<mods:affiliation>Publicly Traded Company &quot; Morpho Absoloni &quot;, Bolotnikovskaya St. 5 / 3, Moscow 117556, Russia</mods:affiliation>
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<mods:date>2014</mods:date>
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<mods:number>2014-12-12</mods:number>
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<mods:number>463</mods:number>
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<mods:url>http://dx.doi.org/10.3897/zookeys.463.8499</mods:url>
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<treatment ID-GBIF-Taxon="182234818" LSID="urn:lsid:plazi:treatment:5E4B64CEA2305FE7BE0DE7E8AB65D1B9" httpUri="http://treatment.plazi.org/id/5E4B64CEA2305FE7BE0DE7E8AB65D1B9" lastPageId="11" lastPageNumber="31" pageId="10" pageNumber="31">
<subSubSection pageId="10" pageNumber="31" type="nomenclature">
<paragraph pageId="10" pageNumber="31">
<taxonomicName LSID="5E4B64CE-A230-5FE7-BE0D-E7E8AB65D1B9" authority="Gottwald, 1985" authorityName="Gottwald" authorityYear="1985" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus subsp. dsychvensis" order="Coleoptera" pageId="10" pageNumber="31" phylum="Arthropoda" rank="subSpecies" species="besleticus" subGenus="Archiplectes" subSpecies="dsychvensis">Carabus (Archiplectes) besleticus dsychvensis Gottwald, 1985</taxonomicName>
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution of morphometric characters in males of Carabus (Archiplectes) besleticus and Carabus (Archiplectes) satyrus subspecies constructed using discriminant analysis based on 8 parameters." pageId="10" pageNumber="31">Figs 1</figureCitation>
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Distribution of morphometric characters in females of Carabus (Archiplectes) besleticus and Carabus (Archiplectes) satyrus subspecies constructed using discriminant analysis based on 8 parameters." pageId="10" pageNumber="31">, 2</figureCitation>
<figureCitation captionStart="Figure 7" captionStartId="F7" captionText="Figure 7. Distribution map of Carabus (Archiplectes) satyrus, Carabus (Archiplectes) besleticus, and Carabus (Archiplectes) pseudopshuensis in Abkhazia." pageId="10" pageNumber="31">, 7</figureCitation>
<figureCitation captionStart="Figures 28–33" captionStartId="F11" captionText="Figures 28 - 33. Fully inflated endophallus preparation and apical lamella of aedeagus of Carabus (Archiplectes) besleticus dsychvensis and Carabus (Archiplectes) besleticus resheviensis 28 - 30 Carabus (Archiplectes) besleticus dsychvensis 31 - 33 Carabus (Archiplectes) besleticus resheviensis." pageId="10" pageNumber="31">, 28-30</figureCitation>
<figureCitation captionStart="Figures 74–77" captionStartId="F18" captionText="Figures 74 - 77. Dorsal habitus, Carabus (Archiplectes) besleticus dsychvensis, Abkhazia, Bzybian Mt. Range, SW slopes of Mt. Dzykhva." pageId="10" pageNumber="31">, 74-77</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="31" type="reference_group">
<paragraph pageId="10" pageNumber="31">
<taxonomicName class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) juenthneri subsp. dsychvensis" order="Coleoptera" pageId="10" pageNumber="31" phylum="Arthropoda" rank="subSpecies" species="juenthneri" subGenus="Archiplectes" subSpecies="dsychvensis">Carabus (Archiplectes) juenthneri dsychvensis</taxonomicName>
Gottwald, 1985: 304 (&quot;Mt. Dsykhwa-Nordhang&quot;);
<bibRefCitation author="Retezar, I" journalOrPublisher="Mesterprint Kft., Budapest" pageId="23" pageNumber="44" refId="B8" refString="Retezar, I, 2008. The Carabus of Abkhazia, Caucasus (Coleoptera, Carabidae): Iconography, genital morphology, systematics and faunistics. Mesterprint Kft., Budapest" title="The Carabus of Abkhazia, Caucasus (Coleoptera, Carabidae): Iconography, genital morphology, systematics and faunistics." year="2008">
<normalizedToken originalValue="Retezár">Retezar</normalizedToken>
2008
</bibRefCitation>
: 39.
</paragraph>
<paragraph pageId="10" pageNumber="31">
<taxonomicName class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) reitteri subsp. dsychvensis" order="Coleoptera" pageId="10" pageNumber="31" phylum="Arthropoda" rank="subSpecies" species="reitteri" subGenus="Archiplectes" subSpecies="dsychvensis">Carabus (Archiplectes) reitteri dsychvensis</taxonomicName>
:
<bibRefCitation author="Bousquet, Y" journalOrPublisher="Catalogue of Palaearctic Coleoptera" pageId="23" pageNumber="44" pagination="118 - 201" refId="B2" refString="Bousquet, Y, Brezina, B, Davies, A, Farkac, J, Smetana, A, 2003. Tribe Carabini Latreille, 1802. Catalogue of Palaearctic Coleoptera Vol. 1: 118 - 201" title="Tribe Carabini Latreille, 1802." volume="Vol. 1" year="2003">Bousquet et al. 2003</bibRefCitation>
: 132.
</paragraph>
<paragraph pageId="10" pageNumber="31">
<taxonomicName class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Tribax) juenthneri subsp. dsychvensis" order="Coleoptera" pageId="10" pageNumber="31" phylum="Arthropoda" rank="subSpecies" species="juenthneri" subGenus="Tribax" subSpecies="dsychvensis">Carabus (Tribax) juenthneri dsychvensis</taxonomicName>
:
<bibRefCitation author="Deuve, T" journalOrPublisher="Pensoft Publishers, Soﬁa-Moscow" pageId="23" pageNumber="44" refId="B3" refString="Deuve, T, 2004. Illustrated Catalogue of the Genus Carabus of the World (Coleoptera: Carabidae). Pensoft Publishers, Soﬁa-Moscow" title="Illustrated Catalogue of the Genus Carabus of the World (Coleoptera: Carabidae)." year="2004">Deuve 2004</bibRefCitation>
: 273.
</paragraph>
<paragraph pageId="10" pageNumber="31">
<taxonomicName class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) satyrus subsp. dsychvensis" order="Coleoptera" pageId="10" pageNumber="31" phylum="Arthropoda" rank="subSpecies" species="satyrus" subGenus="Archiplectes" subSpecies="dsychvensis">Carabus (Archiplectes) satyrus dsychvensis</taxonomicName>
:
<bibRefCitation author="Retezar, I" journalOrPublisher="Folia Entomologica Hungarica" pageId="23" pageNumber="44" pagination="1 - 6" refId="B9" refString="Retezar, I, 2013. Taxonomic and faunistic studies on the subgenus Carabus (Archiplectes) Gottwald, 1982 in Abkhazia (Coleoptera: Carabidae). Folia Entomologica Hungarica 74: 1 - 6" title="Taxonomic and faunistic studies on the subgenus Carabus (Archiplectes) Gottwald, 1982 in Abkhazia (Coleoptera: Carabidae)." volume="74" year="2013">
<normalizedToken originalValue="Retezár">Retezar</normalizedToken>
2013
</bibRefCitation>
: 4.
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="31" type="comparative material examined">
<paragraph pageId="10" pageNumber="31">Comparative material examined.</paragraph>
<paragraph pageId="10" pageNumber="31">
278 specimens were examined (37 specimens measured, 12 male genitalia preparations studied): 151 males, 127 females, Abkhazia, Bzybian Mt. Range, SW slopes of Mt. Dzykhva, 2000-2300 m, alpine zone,
<geoCoordinate degrees="43" direction="north" minutes="13" orientation="latitude" precision="925" value="43.216667">N43°13'</geoCoordinate>
,
<geoCoordinate degrees="041" direction="east" minutes="08" orientation="longitude" precision="925" value="41.133335">E041°08'</geoCoordinate>
, 29.VI-03.VII.2013, leg. D. Fominykh (cFDD, cTAYu, cSAA, cSIA, cKVM).
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="31" type="description">
<paragraph pageId="10" pageNumber="31">Description.</paragraph>
<paragraph pageId="10" pageNumber="31">Small form, males 23.3-26.5 (25.2) mm and females 23.3-26.5 (25.2) mm long. Underside black, dorsum normally with bright metallic lustre, green, bronze, less often dark violet or dark blue. Red femoral forms are also rather frequent. Body slender.</paragraph>
<paragraph pageId="10" pageNumber="31">
Head not inflated. Pronotum variable in shape, subcordate, transverse. PW/PL = 1.20-1.58 (1.43) in males and 1.30-1.62 (1.46) in females, PW/PB = 1.07-1.37 (1.22) in males and 1.03-1.30 (1.17) in females, hind angles protruding backwards, pointed apically. Median groove distinct in females, and inconspicuous and smoothed in males, disk with fine rugosity, gradually strengthening towards median groove. Elytra oblong-ovate, more elongate and somewhat convex in males, with inconspicuous to missing middle depression in the females. EL/EW = 1.56-1.85 (1.68) in males and 1.50-1.79 (1.66) in females. Elytral sculpture smooth in males and precise, forming distinct series of short links in females. The main morphometric measurements are presented in Table
<tableCitation captionStart="Table 2" captionStartId="T2" captionText="Table 2. Morphometric characteristic of males (n = 16) and females (n = 31) of Carabus (Archiplectes) satyrus and males (n = 83) and females (n = 91) of Carabus (Archiplectes) besleticus subspecies." httpUri="http://table.plazi.org/id/E3AA304075ED326595338A0A005BF64B" pageId="10" pageNumber="31" tableUuid="E3AA304075ED326595338A0A005BF64B">2</tableCitation>
.
</paragraph>
<paragraph pageId="10" pageNumber="31">
Apical lamella of aedeagus of transitional between
<taxonomicName authorityName="Kurnakov" authorityYear="1972" baseAuthorityName="besleticus Kurnakov" baseAuthorityYear="1972" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus" order="Coleoptera" pageId="10" pageNumber="31" phylum="Arthropoda" rank="species" species="besleticus" subGenus="Archiplectes">Carabus (Archiplectes) besleticus</taxonomicName>
and
<taxonomicName authorityName="Zamotajlov" authorityYear="1991" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) pseudopshuensis" order="Coleoptera" pageId="10" pageNumber="31" phylum="Arthropoda" rank="species" species="pseudopshuensis" subGenus="Archiplectes">Carabus (Archiplectes) pseudopshuensis</taxonomicName>
shape, possessing a sharp hollow on the right between apex and tubercle. Also sharply differs in this respect from
<taxonomicName baseAuthorityName="Reitter" baseAuthorityYear="1899" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) juenthneri" order="Coleoptera" pageId="10" pageNumber="31" phylum="Arthropoda" rank="species" species="juenthneri" subGenus="Archiplectes">Carabus (Archiplectes) juenthneri</taxonomicName>
(previously considered as its subspecies). The endophallus differs from
<taxonomicName authorityName="Kurnakov" authorityYear="1962" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) satyrus" order="Coleoptera" pageId="10" pageNumber="31" phylum="Arthropoda" rank="species" species="satyrus" subGenus="Archiplectes">Carabus (Archiplectes) satyrus</taxonomicName>
in the shape of the dorsal appendix, which is more elongate and possesses a more extended form, characteristic for populations dwelling in an interfluve of the r. Kealasur and Bzyb.
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="31" type="differential diagnosis">
<paragraph pageId="10" pageNumber="31">Differential diagnosis.</paragraph>
<paragraph pageId="10" pageNumber="31">
Habitually resembles
<taxonomicName authorityName="Gottwald" authorityYear="1985" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) juenthneri subsp. adsypschi" order="Coleoptera" pageId="10" pageNumber="31" phylum="Arthropoda" rank="subSpecies" species="juenthneri" subGenus="Archiplectes" subSpecies="adsypschi">Carabus (Archiplectes) juenthneri adsypschi</taxonomicName>
Gottwald, 1985, from which differs in more elongate and more ovate elytra, hind angles of pronotum more strongly protruding, elytral sculpture more coarse, and larger average body size. Also easily distinguishable by male genitalia. The apical lamella of this form combines some features of both
<taxonomicName authorityName="Kurnakov" authorityYear="1972" baseAuthorityName="besleticus Kurnakov" baseAuthorityYear="1972" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus" order="Coleoptera" pageId="10" pageNumber="31" phylum="Arthropoda" rank="species" species="besleticus" subGenus="Archiplectes">Carabus (Archiplectes) besleticus</taxonomicName>
and
<taxonomicName authorityName="Zamotajlov" authorityYear="1991" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) pseudopshuensis" order="Coleoptera" pageId="10" pageNumber="31" phylum="Arthropoda" rank="species" species="pseudopshuensis" subGenus="Archiplectes">Carabus (Archiplectes) pseudopshuensis</taxonomicName>
. The shape of the saccellus and aggonoporius seems, however, to be closer to
<taxonomicName authorityName="Kurnakov" authorityYear="1972" baseAuthorityName="besleticus Kurnakov" baseAuthorityYear="1972" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Archiplectes) besleticus" order="Coleoptera" pageId="10" pageNumber="31" phylum="Arthropoda" rank="species" species="besleticus" subGenus="Archiplectes">Carabus (Archiplectes) besleticus</taxonomicName>
. The study of the further high-mountain populations of
<taxonomicName authorityName="Gottwald" authorityYear="1982" class="Insecta" family="Carabidae" genus="Archiplectes" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="Archiplectes" order="Coleoptera" pageId="10" pageNumber="31" phylum="Arthropoda" rank="genus">
<emphasis italics="true" pageId="10" pageNumber="31">Archiplectes</emphasis>
</taxonomicName>
from adjacent woodless alpine massifs is required for correct interpretation of its taxonomic status within the species-complex in question. Morphometric characters are illustrated in Figures
<figureCitation captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution of morphometric characters in males of Carabus (Archiplectes) besleticus and Carabus (Archiplectes) satyrus subspecies constructed using discriminant analysis based on 8 parameters." pageId="10" pageNumber="31">1</figureCitation>
-
<figureCitation captionStart="Figure 2" captionStartId="F2" captionText="Figure 2. Distribution of morphometric characters in females of Carabus (Archiplectes) besleticus and Carabus (Archiplectes) satyrus subspecies constructed using discriminant analysis based on 8 parameters." pageId="10" pageNumber="31">2</figureCitation>
.
</paragraph>
</subSubSection>
<subSubSection pageId="10" pageNumber="31" type="distribution">
<paragraph pageId="10" pageNumber="31">Distribution.</paragraph>
<paragraph pageId="10" pageNumber="31">Populates the alpine belt of Mt. Dzykhva at 2000-2300 m altitude.</paragraph>
</subSubSection>
<subSubSection lastPageId="11" lastPageNumber="32" pageId="10" pageNumber="31" type="habitat">
<paragraph pageId="10" pageNumber="31">Habitat.</paragraph>
<paragraph lastPageId="11" lastPageNumber="32" pageId="10" pageNumber="31">
Prefers alpine herb meadows. Activity of imago proceeds from May to August, beetles being active at melting of snow cover.
<taxonomicName authorityName="Gottwald" authorityYear="1983" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Procechenochilus) adangensis" order="Coleoptera" pageId="11" pageNumber="32" phylum="Arthropoda" rank="species" species="adangensis" subGenus="Procechenochilus">
<pageBreakToken pageId="11" pageNumber="32" start="start">Carabus</pageBreakToken>
(Procechenochilus) adangensis
</taxonomicName>
,
<taxonomicName authority="circassicus" class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Tribax) circassicus subsp. circassicus" order="Coleoptera" pageId="11" pageNumber="32" phylum="Arthropoda" rank="subSpecies" species="circassicus" subGenus="Tribax" subSpecies="circassicus">Carabus (Tribax) circassicus circassicus</taxonomicName>
(natio
<taxonomicName class="Insecta" family="Carabidae" form="tshchaltensis" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="" order="Coleoptera" pageId="11" pageNumber="32" phylum="Arthropoda" rank="form">
<emphasis italics="true" pageId="11" pageNumber="32">tshchaltensis</emphasis>
</taxonomicName>
<normalizedToken originalValue="Novotný">Novotny</normalizedToken>
&amp;
<normalizedToken originalValue="Voříšek">Vorisek</normalizedToken>
, 1988),
<taxonomicName class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Tribax) constantinovi subsp. otcharensis" order="Coleoptera" pageId="11" pageNumber="32" phylum="Arthropoda" rank="subSpecies" species="constantinovi" subGenus="Tribax" subSpecies="otcharensis">Carabus (Tribax) constantinovi otcharensis</taxonomicName>
,
<taxonomicName class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Tribax) agnatus subsp. pseudoagnatus" order="Coleoptera" pageId="11" pageNumber="32" phylum="Arthropoda" rank="subSpecies" species="agnatus" subGenus="Tribax" subSpecies="pseudoagnatus">Carabus (Tribax) agnatus pseudoagnatus</taxonomicName>
<normalizedToken originalValue="Novotný">Novotny</normalizedToken>
&amp;
<normalizedToken originalValue="Voříšek">Vorisek</normalizedToken>
, 1988, and
<taxonomicName class="Insecta" family="Carabidae" genus="Carabus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Carabus (Pachycarabus) imitator subsp. katharinae" order="Coleoptera" pageId="11" pageNumber="32" phylum="Arthropoda" rank="subSpecies" species="imitator" subGenus="Pachycarabus" subSpecies="katharinae">Carabus (Pachycarabus) imitator katharinae</taxonomicName>
Reitter, 1896 occur together with this subspecies.
</paragraph>
</subSubSection>
</treatment>
</document>