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<document id="001326A6F7A61FE7921A7A995760DE5D" ID-CLB-Dataset="302193" ID-DOI="10.11646/phytotaxa.659.3.1" ID-GBIF-Dataset="52524fe3-dfb8-4afc-829a-31089bb4e306" ID-ISSN="1179-3163" ID-Zenodo-Dep="13217894" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="valdenar" IM.taxonomicNames_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" checkinTime="1722613780751" checkinUser="felipe" docAuthor="Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, Zhou, Zhuo, Thepkayson, Khamphanh, Luong, Thien Tam &amp; Zhang, Li-Bing" docDate="2024" docId="4B175324C95AA518D1BAF8E5FEE3A80D" docLanguage="en" docName="phytotaxa.659.3.1.pdf" docOrigin="Phytotaxa 659 (3)" docSource="http://dx.doi.org/10.11646/phytotaxa.659.3.1" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Leptochilus locii Liang Zhang, N. T. Lu &amp; Li Bing Zhang, sp. nov." docType="treatment" docVersion="5" lastPageNumber="224" masterDocId="B72E2B5CC953A513D132FFC2FFE5AB4D" masterDocTitle="Exploring the diversity of the Java fern genus Leptochilus (Polypodiaceae) in the Indo-Burma Biodiversity Hotspot" masterLastPageNumber="235" masterPageNumber="213" pageNumber="222" updateTime="1725399130512" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="8F6E48E5E9324DF8900AB7373FE6FDE8">Exploring the diversity of the Java fern genus Leptochilus (Polypodiaceae) in the Indo-Burma Biodiversity Hotspot</mods:title>
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<mods:namePart id="29BA4DFF4B0E6DE2B18F0747963A0F10">Zhang, Liang</mods:namePart>
<mods:nameIdentifier id="7E453BBCD9C38C2709CA16FFDCD477C0" type="ORCID">0000-0003-3784-3135</mods:nameIdentifier>
<mods:affiliation id="615129A5219FE9FA8DE5C15500E85CB3">Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming, Yunnan 650201, China</mods:affiliation>
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<mods:namePart id="F3CB5055ECAB49D6DCABF0FA6682E604">Lu, Ngan Thi</mods:namePart>
<mods:nameIdentifier id="7C91E232B40EB2E0BAD712370606F84E" type="ORCID">0000-0001-9823-2018</mods:nameIdentifier>
<mods:affiliation id="C936997C4B99B81D3B0F65BC22536D7D">Department of Biology, Vietnam National Museum of Nature, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Hanoi, Vietnam; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Hanoi, Vietnam</mods:affiliation>
<mods:nameIdentifier id="971F52404CD399432D5ECB60180F8EC3" type="email">vnmngan@gmail.com</mods:nameIdentifier>
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<mods:namePart id="B4BC4D591A2454E023B8D9F2505DC2DB">Zhou, Xin-Mao</mods:namePart>
<mods:nameIdentifier id="2979804E57DB7C8401121DDA05995E67" type="ORCID">0000-0003-3555-7784</mods:nameIdentifier>
<mods:affiliation id="87EBF7898F1FEDE18663BE0066C081DB">School of Ecology and Environmental Science, Yunnan University, Kunming, Yunnan 650091, China</mods:affiliation>
<mods:nameIdentifier id="3122F99DE9BC4DABE4745D6189716495" type="email">xinmao.zhou@ynu.edu.cn</mods:nameIdentifier>
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<mods:namePart id="010DFE724E16E89F2444FF086D22FECA">Zhou, Zhuo</mods:namePart>
<mods:nameIdentifier id="F6FEE3A2C2BEB2C154C7513C5BE106E1" type="ORCID">0000-0001-9682-5717</mods:nameIdentifier>
<mods:affiliation id="6749BD534623C17EE7F3CC8C4EF20FBF">Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming, Yunnan 650201, China</mods:affiliation>
<mods:nameIdentifier id="956A246C01B9B2DB8801B7CE6C7EB08F" type="email">zhouzhuo@mail.kib.ac.cn</mods:nameIdentifier>
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<mods:namePart id="50390A7E3222F206E1ECD8E0B5E6F2BD">Thepkayson, Khamphanh</mods:namePart>
<mods:nameIdentifier id="046D090E937D3A551150105488B90A21" type="ORCID">0009-0001-1757-9907</mods:nameIdentifier>
<mods:affiliation id="7746039B96591B2563CFA1ED43F6CE36">Institute of Traditional Medicine, Vientiane, Lao PDR</mods:affiliation>
<mods:nameIdentifier id="3F5CFB8AA3915DC751B5E2F755DE928F" type="email">khamphanh@hotmail.com</mods:nameIdentifier>
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<mods:namePart id="CD1342F7FFB65F71568D6A10E9F93D8A">Luong, Thien Tam</mods:namePart>
<mods:nameIdentifier id="B6AD7FC9B6D96821D818E19210012043" type="ORCID">0000-0001-8354-6321</mods:nameIdentifier>
<mods:affiliation id="E405480BC771D3AA04944C65B104FE8D">Department of Ecology - Evolutionary Biology, Vietnam National University Ho Chi Minh City (VNUHCM) - University of Science, 227 Nguyen Van Cu Street, District 5, Ho Chi Minh City, Vietnam</mods:affiliation>
<mods:nameIdentifier id="06AB7340C5F44604AA536395E60F203F" type="email">lttam1511@gmail.com</mods:nameIdentifier>
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<mods:namePart id="EAC5E7CE15398365FFF125221B609F9B">Zhang, Li-Bing</mods:namePart>
<mods:nameIdentifier id="8C90B3E913DCBB2F3CABCDF2306205A8" type="ORCID">0000-0002-4905-040X</mods:nameIdentifier>
<mods:affiliation id="497E985CD84FC7BE18CC756B542CD3FF">Missouri Botanical Garden, 4344 Shaw Blvd, St. Louis, Missouri, 63110 U. S. A.; Chengdu Institute of Biology, Chinese Academy of Sciences, Chengdu 610041, Sichuan, China</mods:affiliation>
<mods:nameIdentifier id="E62D39B9D1184ACE9B0DADC039630CEB" type="email">Libing.Zhang@mobot.org</mods:nameIdentifier>
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<mods:title id="B16E354DAF29665EDEE9CD496CA804B4">Phytotaxa</mods:title>
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<mods:date id="0F52E7B66660D776367329332CD50816">2024</mods:date>
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<mods:number id="C3B5563E28972C8CED4F65182DF76328">2024-07-11</mods:number>
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<mods:number id="E076C0A860CE1D629250C57A7A367196">659</mods:number>
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<mods:number id="42099A14E4609AE7FED3123CEB674545">3</mods:number>
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<mods:start id="FCAA8E4C5DE4D237F27AB5A8596499DF">213</mods:start>
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<mods:url id="6D01D712A621BB6029165B7FF2685110">http://dx.doi.org/10.11646/phytotaxa.659.3.1</mods:url>
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<mods:classification id="7B3D490572729C2616B65868C2F273A6">journal article</mods:classification>
<mods:identifier id="D1AD0DE8FF088C5278D2FB957C083A52" type="CLB-Dataset">302193</mods:identifier>
<mods:identifier id="67E49582C6741A1B09F5F4A940ACD750" type="DOI">10.11646/phytotaxa.659.3.1</mods:identifier>
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<treatment id="4B175324C95AA518D1BAF8E5FEE3A80D" ID-DOI="http://doi.org/10.5281/zenodo.13652242" ID-GBIF-Taxon="240245552" ID-Zenodo-Dep="13652242" LSID="urn:lsid:plazi:treatment:4B175324C95AA518D1BAF8E5FEE3A80D" httpUri="http://treatment.plazi.org/id/4B175324C95AA518D1BAF8E5FEE3A80D" lastPageId="11" lastPageNumber="224" pageId="9" pageNumber="222">
<subSubSection id="8BA4B1B9C95AA51AD1BAF8E5FC08AC0C" box="[136,1005,1830,1858]" pageId="9" pageNumber="222" type="nomenclature">
<paragraph id="C301E232C95AA51AD1BAF8E5FC08AC0C" blockId="9.[136,1005,1830,1858]" box="[136,1005,1830,1858]" pageId="9" pageNumber="222">
<heading id="9849555EC95AA51AD1BAF8E5FC08AC0C" box="[136,1005,1830,1858]" fontSize="11" level="2" pageId="9" pageNumber="222" reason="4">
5.
<taxonomicName id="04BE99B1C95AA51AD196F8E4FCCEAC0C" authority="Liang Zhang, N.T. Lu &amp; Li Bing Zhang" authorityName="Liang Zhang, N. T. Lu &amp; Li Bing Zhang" box="[164,811,1830,1857]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="9" pageNumber="222" phylum="Tracheophyta" rank="species" species="locii" status="sp. nov.">
<emphasis id="F1CA3E20C95AA51AD196F8E4FE81AC0C" bold="true" box="[164,356,1830,1857]" italics="true" pageId="9" pageNumber="222">Leptochilus locii</emphasis>
Liang Zhang, N.T. Lu &amp; Li Bing Zhang
</taxonomicName>
,
<emphasis id="F1CA3E20C95AA51AD205F8EAFC69AC0C" box="[823,908,1832,1857]" italics="true" pageId="9" pageNumber="222">
<taxonomicNameLabel id="EAF9835BC95AA51AD205F8EAFC69AC0C" box="[823,908,1832,1857]" pageId="9" pageNumber="222" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
(
<figureCitation id="5B85FEB7C95AA51AD2A9F8E5FC3AAC0C" box="[923,991,1831,1857]" captionStart="FIGURE 7" captionStartId="10.[136,229,1804,1826]" captionTargetBox="[163,1424,202,1767]" captionTargetId="figure-25@10.[151,1436,190,1780]" captionTargetPageId="10" captionText="FIGURE 7. Leptochilus locii Liang Zhang, N.T. Lu &amp; Li Bing Zhang,—A. Habitat.—B. Adaxial view of sterile frond.—C. Habit.—D. Abaxial view of sterile frond.—E. Abaxial view of fertile lamina.—F. Adaxial view of fertile lamina. Photo credit: Liang Zhang." figureDoi="http://doi.org/10.5281/zenodo.13217908" httpUri="https://zenodo.org/record/13217908/files/figure.png" pageId="9" pageNumber="222">Fig. 7</figureCitation>
).
</heading>
</paragraph>
</subSubSection>
<subSubSection id="8BA4B1B9C95AA51AD1BAF8ADFED8AC9C" pageId="9" pageNumber="222" type="materials_examined">
<paragraph id="C301E232C95AA51AD1BAF8ADFED8AC9C" blockId="9.[136,1452,1903,2073]" pageId="9" pageNumber="222">
<emphasis id="F1CA3E20C95AA51AD1BAF8ADFF26ACC4" bold="true" box="[136,195,1903,1929]" pageId="9" pageNumber="222">Type</emphasis>
:
<collectingCountry id="BBA9A2A2C95AA51AD1E6F8ADFEBCACC4" box="[212,345,1903,1929]" name="Vietnam" pageId="9" pageNumber="222">VIETNAM</collectingCountry>
.
<collectingRegion id="017A2CD0C95AA51AD054F8ADFE10ACC4" box="[358,501,1903,1929]" country="Vietnam" name="Quang Binh" pageId="9" pageNumber="222">Quang Binh</collectingRegion>
: Phong Nha Ke Bang National Park, Tan Traih, elev.
<quantity id="04464FD7C95AA51AD554F8ADFB57ACC4" box="[1126,1202,1903,1929]" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="9" pageNumber="222" unit="m" value="700.0">700 m</quantity>
,
<geoCoordinate id="A68A84F5C95AA51AD5F2F8ADFAFBACC4" box="[1216,1310,1903,1929]" degrees="17.39" direction="north" orientation="latitude" pageId="9" pageNumber="222" precision="555" value="17.39">17.39°N</geoCoordinate>
/106.20°, on rocks in limestone mountains,
<date id="B700C4F2C95AA51AD0E9F851FD43ACE0" box="[475,678,1939,1965]" pageId="9" pageNumber="222" value="2014-11-06">6 November 2014</date>
,
<emphasis id="F1CA3E20C95AA51AD382F856FB3DACE0" box="[688,1240,1939,1965]" italics="true" pageId="9" pageNumber="222">Li Bing Zhang, Liang Zhang &amp; Ngan Thi Lu 7352</emphasis>
(
<typeStatus id="1C055C90C95AA51AD5D7F851FAACACE0" box="[1253,1353,1939,1965]" pageId="9" pageNumber="222" type="holotype">holotype</typeStatus>
VNMN,
<typeStatus id="1C055C90C95AA51AD1BAF875FF3CAC9C" box="[136,217,1975,2001]" pageId="9" pageNumber="222" type="isotype">isotype</typeStatus>
<collectionCode id="A5AF7AF7C95AA51AD1D2F875FEC8AC9C" box="[224,301,1975,2001]" country="China" lsid="urn:lsid:biocol.org:col:12796" name="Chengdu Institute of Biology" pageId="9" pageNumber="222" type="Herbarium">CDBI</collectionCode>
!).
</paragraph>
</subSubSection>
<subSubSection id="8BA4B1B9C95AA518D18FF819FD7EAADD" lastPageId="11" lastPageNumber="224" pageId="9" pageNumber="222" type="diagnosis">
<paragraph id="C301E232C95AA51AD18FF819FB28A354" blockId="9.[136,1452,1903,2073]" pageId="9" pageNumber="222">
<emphasis id="F1CA3E20C95AA51AD18FF819FED5ACB8" bold="true" box="[189,304,2011,2037]" pageId="9" pageNumber="222">Diagnosis</emphasis>
:
<taxonomicName id="04BE99B1C95AA51AD071F819FDE7ACB8" box="[323,514,2011,2037]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="9" pageNumber="222" phylum="Tracheophyta" rank="species" species="locii">
<emphasis id="F1CA3E20C95AA51AD071F819FDE7ACB8" box="[323,514,2011,2037]" italics="true" pageId="9" pageNumber="222">Leptochilus locii</emphasis>
</taxonomicName>
is characterized by its dimorphic fronds, fertile fronds approximately four times longer or more than sterile fronds, fertile fronds long-petiolated, and laminae thickly chartaceous.
</paragraph>
<caption id="97C1B2BAC959A519D1BAF8CEFABEAC0A" ID-DOI="http://doi.org/10.5281/zenodo.13217908" ID-Zenodo-Dep="13217908" httpUri="https://zenodo.org/record/13217908/files/figure.png" pageId="10" pageNumber="223" startId="10.[136,229,1804,1826]" targetBox="[163,1424,202,1767]" targetPageId="10" targetType="figure">
<paragraph id="C301E232C959A519D1BAF8CEFABEAC0A" blockId="10.[136,1452,1804,1863]" pageId="10" pageNumber="223">
<emphasis id="F1CA3E20C959A519D1BAF8CEFF1DAC6F" bold="true" box="[136,248,1804,1826]" pageId="10" pageNumber="223">FIGURE 7</emphasis>
.
<taxonomicName id="04BE99B1C959A519D037F8CFFCD6AC6E" authority="Liang Zhang, N. T. Lu &amp; Li Bing Zhang" authorityName="Liang Zhang, N. T. Lu &amp; Li Bing Zhang" box="[261,819,1805,1827]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="10" pageNumber="223" phylum="Tracheophyta" rank="species" species="locii">
<emphasis id="F1CA3E20C959A519D037F8CFFE42AC6F" box="[261,423,1805,1826]" italics="true" pageId="10" pageNumber="223">Leptochilus locii</emphasis>
Liang Zhang, N.T. Lu &amp; Li Bing Zhang
</taxonomicName>
,—A. Habitat.—B. Adaxial view of sterile frond.—C. Habit.—D. Abaxial view of sterile frond.—E. Abaxial view of fertile lamina.—F. Adaxial view of fertile lamina. Photo credit: Liang Zhang.
</paragraph>
</caption>
<paragraph id="C301E232C959A518D18FF8B7FD7EAADD" blockId="10.[136,1452,1909,2080]" lastBlockId="11.[136,1452,158,832]" lastPageId="11" lastPageNumber="224" pageId="10" pageNumber="223">
<emphasis id="F1CA3E20C959A519D18FF8B7FEEDACC2" bold="true" box="[189,264,1909,1935]" pageId="10" pageNumber="223">Plants</emphasis>
perennial, evergreen, lithophytic.
<emphasis id="F1CA3E20C959A519D34DF8B7FD0CACC2" bold="true" box="[639,745,1909,1935]" pageId="10" pageNumber="223">Rhizome</emphasis>
long creeping, ca.
<quantity id="04464FD7C959A519D286F8B7FBDCACDD" box="[948,1081,1909,1936]" metricMagnitude="-3" metricUnit="m" metricValue="2.35" metricValueMax="2.9" metricValueMin="1.8" pageId="10" pageNumber="223" unit="mm" value="2.35" valueMax="2.9" valueMin="1.8">1.8–2.9 mm</quantity>
in diam., dorsiventrally flattened, scaled, with a few short phyllopodia and roots. Apex of rhizome and basal petiole densely scaly, scales lanceolate, base enlarged, (0.7–)1.2–3.3 ×
<quantity id="04464FD7C959A519D080F87FFDD8AC95" box="[434,573,1981,2008]" metricMagnitude="-4" metricUnit="m" metricValue="9.0" metricValueMax="14.0" metricValueMin="4.0" pageId="10" pageNumber="223" unit="mm" value="0.8999999999999999" valueMax="1.4" valueMin="0.4">0.4–1.4 mm</quantity>
, brown, basifixed, concolorous, margin entire, apex long acuminate.
<emphasis id="F1CA3E20C959A519D464F87FFA4EAC9A" bold="true" box="[1366,1451,1981,2007]" pageId="10" pageNumber="223">Fronds</emphasis>
dimorphic, fertile fronds much longer than sterile fronds.
<emphasis id="F1CA3E20C959A519D22CF823FC27ACB6" bold="true" box="[798,962,2017,2043]" pageId="10" pageNumber="223">Sterile fronds</emphasis>
<quantity id="04464FD7C959A519D2F9F823FBCAACB1" box="[971,1071,2017,2044]" metricMagnitude="-1" metricUnit="m" metricValue="1.05" metricValueMax="1.2" metricValueMin="0.9" pageId="10" pageNumber="223" unit="cm" value="10.5" valueMax="12.0" valueMin="9.0">9–12 cm</quantity>
tall, laminae slightly longer than or the same length as petioles; laminae triangular-ovate, base truncate, apex obtuse-acuminate,
<quantity id="04464FD7C959A519D589F7C7FAEAA36D" box="[1211,1295,2053,2080]" metricMagnitude="-2" metricUnit="m" metricValue="5.5" metricValueMax="7.0" metricValueMin="4.0" pageId="10" pageNumber="223" unit="cm" value="5.5" valueMax="7.0" valueMin="4.0">4–7 cm</quantity>
long,
<quantity id="04464FD7C959A518D465F7C7FF4FABF4" lastPageId="11" lastPageNumber="224" metricMagnitude="-2" metricUnit="m" metricValue="3.2" metricValueMax="3.5" metricValueMin="2.9" pageId="10" pageNumber="223" unit="cm" value="3.2" valueMax="3.5" valueMin="2.9">2.9–3.5 cm</quantity>
wide near base, thickly chartaceous, glabrous;
<emphasis id="F1CA3E20C958A518D3F7FF5CFCFAABF5" bold="true" box="[709,799,158,184]" pageId="11" pageNumber="224">petioles</emphasis>
stramineous, grooved adaxially, glabrous,
<quantity id="04464FD7C958A518D437FF5CFA6BABF4" box="[1285,1422,158,185]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" metricValueMax="9.0" metricValueMin="7.0" pageId="11" pageNumber="224" unit="mm" value="0.8" valueMax="0.9" valueMin="0.7">0.7–0.9 mm</quantity>
in diam.,
<quantity id="04464FD7C958A518D1E7FF00FECDAB90" box="[213,296,194,221]" metricMagnitude="-2" metricUnit="m" metricValue="6.0" metricValueMax="7.0" metricValueMin="5.0" pageId="11" pageNumber="224" unit="cm" value="6.0" valueMax="7.0" valueMin="5.0">5–7 cm</quantity>
long.
<emphasis id="F1CA3E20C958A518D05DFF00FDF6AB91" bold="true" box="[367,531,194,220]" pageId="11" pageNumber="224">Fertile fronds</emphasis>
<quantity id="04464FD7C958A518D328FF00FD6CAB90" box="[538,649,194,221]" metricMagnitude="-1" metricUnit="m" metricValue="2.2" metricValueMax="2.5" metricValueMin="1.9" pageId="11" pageNumber="224" unit="cm" value="22.0" valueMax="25.0" valueMin="19.0">19–25 cm</quantity>
tall, petioles much longer than laminae;
<emphasis id="F1CA3E20C958A518D561FF00FB57AB91" bold="true" box="[1107,1202,194,220]" pageId="11" pageNumber="224">laminae</emphasis>
ovate, margins entire, base cuneate, apex long acuminate, (1.6–)
<quantity id="04464FD7C958A518D36BFF24FD3CAA4C" box="[601,729,230,257]" metricMagnitude="-2" metricUnit="m" metricValue="4.2" metricValueMax="4.5" metricValueMin="3.9" pageId="11" pageNumber="224" unit="cm" value="4.2" valueMax="4.5" valueMin="3.9">3.9–4.5 cm</quantity>
long, (0.8–)
<quantity id="04464FD7C958A518D251FF24FC05AA4C" box="[867,992,230,257]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" metricValueMax="1.7" metricValueMin="1.5" pageId="11" pageNumber="224" unit="cm" value="1.6" valueMax="1.7" valueMin="1.5">1.5–1.7 cm</quantity>
near base, thickly chartaceous, glabrous;
<emphasis id="F1CA3E20C958A518D1BAFEC8FF07AA69" bold="true" box="[136,226,266,292]" pageId="11" pageNumber="224">petioles</emphasis>
stramineous, grooved adaxially, glabrous,
<quantity id="04464FD7C958A518D3F7FEC8FCA9AA68" box="[709,844,266,293]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="11" pageNumber="224" unit="mm" value="1.0" valueMax="1.1" valueMin="0.9">0.9–1.1 mm</quantity>
in diam.,
<quantity id="04464FD7C958A518D28CFEC8FBC8AA68" box="[958,1069,266,293]" metricMagnitude="-1" metricUnit="m" metricValue="1.9" metricValueMax="2.0" metricValueMin="1.8" pageId="11" pageNumber="224" unit="cm" value="19.0" valueMax="20.0" valueMin="18.0">18–20 cm</quantity>
long.
<emphasis id="F1CA3E20C958A518D547FEC8FB50AA69" bold="true" box="[1141,1205,266,292]" pageId="11" pageNumber="224">Veins</emphasis>
anastomosing, lateral vein distinct, veinlets visible.
<emphasis id="F1CA3E20C958A518D0E6FEECFDE3AA05" bold="true" box="[468,518,302,328]" pageId="11" pageNumber="224">Sori</emphasis>
linear, ca.
<quantity id="04464FD7C958A518D34DFEECFCE0AA04" box="[639,773,302,329]" metricMagnitude="-4" metricUnit="m" metricValue="8.0" metricValueMax="11.0" metricValueMin="5.0" pageId="11" pageNumber="224" unit="mm" value="0.8" valueMax="1.1" valueMin="0.5">0.5–1.1 mm</quantity>
from margins and
<quantity id="04464FD7C958A518D2E5FEECFBACAA04" box="[983,1097,302,329]" metricMagnitude="-4" metricUnit="m" metricValue="1.0" metricValueMax="2.0" metricValueMin="0.0" pageId="11" pageNumber="224" unit="mm" value="0.1" valueMax="0.2" valueMin="0.0">0–0.2 mm</quantity>
from midrib, 9–12 on each side of midrib,
<quantity id="04464FD7C958A518D033FE90FE9AAA20" box="[257,383,338,365]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" metricValueMax="11.0" metricValueMin="8.0" pageId="11" pageNumber="224" unit="cm" value="0.95" valueMax="1.1" valueMin="0.8">0.8–1.1 cm</quantity>
long,
<quantity id="04464FD7C958A518D0FAFE90FDB5AA20" box="[456,592,338,365]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" metricValueMax="1.1" metricValueMin="0.9" pageId="11" pageNumber="224" unit="mm" value="1.0" valueMax="1.1" valueMin="0.9">0.9–1.1 mm</quantity>
wide when mature, at angles of 42–50 degree with midribs, centers ca.
<quantity id="04464FD7C958A518D4B3FE90FF51AADC" metricMagnitude="-3" metricUnit="m" metricValue="2.5" metricValueMax="3.0" metricValueMin="2.0" pageId="11" pageNumber="224" unit="mm" value="2.5" valueMax="3.0" valueMin="2.0">2–3 mm</quantity>
apart from one another. Paraphyses absent.
</paragraph>
</subSubSection>
<subSubSection id="8BA4B1B9C958A518D18FFE58FB58AAB1" pageId="11" pageNumber="224" type="materials_examined">
<paragraph id="C301E232C958A518D18FFE58FB58AAB1" blockId="11.[136,1452,158,832]" pageId="11" pageNumber="224">
<emphasis id="F1CA3E20C958A518D18FFE58FDDBAAF9" bold="true" box="[189,574,410,436]" pageId="11" pageNumber="224">Additional specimens examined:</emphasis>
<collectingCountry id="BBA9A2A2C958A518D376FE58FD2CAAF9" box="[580,713,410,436]" name="Vietnam" pageId="11" pageNumber="224">VIETNAM</collectingCountry>
.
<collectingRegion id="017A2CD0C958A518D3E0FE58FCBBAAF9" box="[722,862,410,436]" country="Vietnam" name="Quang Binh" pageId="11" pageNumber="224">Quang Binh</collectingRegion>
: Bo Trach District, Hung Trach, Phong Nha-Ke Bang National Park,
<quantity id="04464FD7C958A518D002FE7CFE54AA94" box="[304,433,446,473]" metricMagnitude="2" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="11" pageNumber="224" unit="m" value="175.0" valueMax="200.0" valueMin="150.0">150–200 m</quantity>
,
<geoCoordinate id="A68A84F5C958A518D08FFE7CFDF9AA94" box="[445,540,446,473]" degrees="17.50" direction="north" orientation="latitude" pageId="11" pageNumber="224" precision="555" value="17.5">17.50°N</geoCoordinate>
/106.28°, on limestone rock in slightly logged primary forest,
<date id="B700C4F2C958A518D5E2FE7CFA42AA95" box="[1232,1447,446,472]" pageId="11" pageNumber="224" value="2004-12-12">12 December 2004</date>
,
<emphasis id="F1CA3E20C958A518D1BAFE21FBF7AAB1" box="[136,1042,482,509]" italics="true" pageId="11" pageNumber="224">S.-K. Wu, L.K. Phan, X. Gong, J.-Y. Xiang, V.T. Nguyen &amp; K.S. Nguyen WP-1077</emphasis>
(KUN!, MO!).
</paragraph>
</subSubSection>
<subSubSection id="8BA4B1B9C958A518D18FFDC4FCDFA909" pageId="11" pageNumber="224" type="distribution">
<paragraph id="C301E232C958A518D18FFDC4FCDFA909" blockId="11.[136,1452,158,832]" pageId="11" pageNumber="224">
<emphasis id="F1CA3E20C958A518D18FFDC4FE0BA96D" bold="true" box="[189,494,518,544]" pageId="11" pageNumber="224">Distribution and habitat:</emphasis>
Three populations of
<taxonomicName id="04BE99B1C958A518D3DFFDC4FC4EA96D" box="[749,939,518,544]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="11" pageNumber="224" phylum="Tracheophyta" rank="species" species="locii">
<emphasis id="F1CA3E20C958A518D3DFFDC4FC4EA96D" box="[749,939,518,544]" italics="true" pageId="11" pageNumber="224">Leptochilus locii</emphasis>
</taxonomicName>
were discovered in central
<collectingCountry id="BBA9A2A2C958A518D5DBFDC4FAAEA96D" box="[1257,1355,518,544]" name="Vietnam" pageId="11" pageNumber="224">Vietnam</collectingCountry>
, and the species was exclusively observed growing on limestone rock.
</paragraph>
</subSubSection>
<subSubSection id="8BA4B1B9C958A518D18FFD8CFEE3A80D" pageId="11" pageNumber="224" type="etymology">
<paragraph id="C301E232C958A518D18FFD8CFD3FA9C1" blockId="11.[136,1452,158,832]" pageId="11" pageNumber="224">
<emphasis id="F1CA3E20C958A518D18FFD8CFEDBA925" bold="true" box="[189,318,590,616]" pageId="11" pageNumber="224">Etymology</emphasis>
: Named in honor of Professor Phan Ke Loc of
<collectingRegion id="017A2CD0C958A518D259FD8CFC4AA925" box="[875,943,590,616]" country="Vietnam" name="Ha Noi" pageId="11" pageNumber="224">Hanoi</collectingRegion>
University of Science, in recognition of his significant contributions to the fern flora of
<collectingCountry id="BBA9A2A2C958A518D346FDB0FD32A9C1" box="[628,727,626,652]" name="Vietnam" pageId="11" pageNumber="224">Vietnam</collectingCountry>
.
</paragraph>
<paragraph id="C301E232C958A518D18FFD54FEE3A80D" blockId="11.[136,1452,158,832]" pageId="11" pageNumber="224">
<emphasis id="F1CA3E20C958A518D18FFD54FF1AA9FD" bold="true" box="[189,255,662,688]" pageId="11" pageNumber="224">Note:</emphasis>
<taxonomicName id="04BE99B1C958A518D037FD54FE5AA9FD" box="[261,447,662,688]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="11" pageNumber="224" phylum="Tracheophyta" rank="species" species="locii">
<emphasis id="F1CA3E20C958A518D037FD54FE5AA9FD" box="[261,447,662,688]" italics="true" pageId="11" pageNumber="224">Leptochilus locii</emphasis>
</taxonomicName>
is most similar to
<taxonomicName id="04BE99B1C958A518D3B8FD55FD15A9FD" authorityName="Copeland" authorityYear="1914" box="[650,752,663,688]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="11" pageNumber="224" phylum="Tracheophyta" rank="species" species="ovatus">
<emphasis id="F1CA3E20C958A518D3B8FD55FD15A9FD" box="[650,752,663,688]" italics="true" pageId="11" pageNumber="224">L. ovatus</emphasis>
</taxonomicName>
by sharing features such as dimorphic fronds, sterile fronds ca. 4–6 times larger than fertile fronds, and petioles of fertile fronds ca. 8–10 times longer than fertile laminae. However, the new species differs from the latter in having triangular-ovate sterile lamina, and lamina base slightly cordate to truncate. In contrast,
<taxonomicName id="04BE99B1C958A518D049FCC1FE06A851" authorityName="Copeland" authorityYear="1914" box="[379,483,771,796]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="11" pageNumber="224" phylum="Tracheophyta" rank="species" species="ovatus">
<emphasis id="F1CA3E20C958A518D049FCC1FE06A851" box="[379,483,771,796]" italics="true" pageId="11" pageNumber="224">L. ovatus</emphasis>
</taxonomicName>
has elliptic-ovate to broadly elliptic sterile lamina, and sterile lamina base gradually decrescent.
</paragraph>
</subSubSection>
</treatment>
</document>
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