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<document id="8248E2D1A0339B3647EB571380B302B7" ID-CLB-Dataset="302193" ID-DOI="10.11646/phytotaxa.659.3.1" ID-GBIF-Dataset="52524fe3-dfb8-4afc-829a-31089bb4e306" ID-ISSN="1179-3163" ID-Zenodo-Dep="13217894" IM.bibliography_approvedBy="valdenar" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="valdenar" IM.taxonomicNames_approvedBy="valdenar" IM.treatments_approvedBy="valdenar" checkinTime="1722613780751" checkinUser="felipe" docAuthor="Zhang, Liang, Lu, Ngan Thi, Zhou, Xin-Mao, Zhou, Zhuo, Thepkayson, Khamphanh, Luong, Thien Tam &amp; Zhang, Li-Bing" docDate="2024" docId="4B175324C942A501D1BAFF5CFEDDAD5D" docLanguage="en" docName="phytotaxa.659.3.1.pdf" docOrigin="Phytotaxa 659 (3)" docSource="http://dx.doi.org/10.11646/phytotaxa.659.3.1" docStyle="DocumentStyle:96748F8F1B6C902996E134952A3A36B9.13:Phytotaxa.2014-.journal_article" docStyleId="96748F8F1B6C902996E134952A3A36B9" docStyleName="Phytotaxa.2014-.journal_article" docStyleVersion="13" docTitle="Leptochilus vietnamensis Liang Zhang, N. T. Lu &amp; Li Bing Zhang, sp. nov." docType="treatment" docVersion="5" lastPageNumber="231" masterDocId="B72E2B5CC953A513D132FFC2FFE5AB4D" masterDocTitle="Exploring the diversity of the Java fern genus Leptochilus (Polypodiaceae) in the Indo-Burma Biodiversity Hotspot" masterLastPageNumber="235" masterPageNumber="213" pageNumber="230" updateTime="1725399130512" updateUser="ExternalLinkService" zenodo-license-figures="UNSPECIFIED" zenodo-license-treatments="UNSPECIFIED">
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<mods:title id="C06297E6C86DA415733F6391319A9E60">Exploring the diversity of the Java fern genus Leptochilus (Polypodiaceae) in the Indo-Burma Biodiversity Hotspot</mods:title>
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<mods:namePart id="F692603B59B90755CD186130172B0BC8">Zhang, Liang</mods:namePart>
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<mods:affiliation id="8CD7CBCFC58E3022074EE816B9057658">Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming, Yunnan 650201, China</mods:affiliation>
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<mods:namePart id="837BBB7446B6F3AA4F6BF4E8E5C8C38B">Lu, Ngan Thi</mods:namePart>
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<mods:affiliation id="E0EE96D2E18D26BB242E77ABCF042566">Department of Biology, Vietnam National Museum of Nature, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Hanoi, Vietnam; Graduate University of Science and Technology, Vietnam Academy of Science and Technology, 18 Hoang Quoc Viet, Hanoi, Vietnam</mods:affiliation>
<mods:nameIdentifier id="6687FCA23F633DE0B3A24780E574CD5C" type="email">vnmngan@gmail.com</mods:nameIdentifier>
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<mods:namePart id="17FF09BE4377C5710A902A182B943030">Zhou, Xin-Mao</mods:namePart>
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<mods:namePart id="67D05FF847D1688F5367F6ECBA4A32FF">Zhou, Zhuo</mods:namePart>
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<mods:affiliation id="D58C0F45AC21B4EDCB14BB62EEF5FE02">Key Laboratory for Plant Diversity and Biogeography of East Asia, Kunming Institute of Botany, Chinese Academy of Sciences, Kunming, Yunnan 650201, China</mods:affiliation>
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<mods:namePart id="9B9BE716A06E8A79E1D00BF7510DB8A7">Thepkayson, Khamphanh</mods:namePart>
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<mods:namePart id="80BCFB59DD4C82AA75684A541183B2E3">Luong, Thien Tam</mods:namePart>
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<mods:affiliation id="2F642DF7F08A96878D18746D5D668130">Department of Ecology - Evolutionary Biology, Vietnam National University Ho Chi Minh City (VNUHCM) - University of Science, 227 Nguyen Van Cu Street, District 5, Ho Chi Minh City, Vietnam</mods:affiliation>
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<mods:namePart id="9E10F54FF5CD04A527D68E577794F1FC">Zhang, Li-Bing</mods:namePart>
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<mods:affiliation id="76809B2EFE3449152ECF238E7E32B27A">Missouri Botanical Garden, 4344 Shaw Blvd, St. Louis, Missouri, 63110 U. S. A.; Chengdu Institute of Biology, Chinese Academy of Sciences, Chengdu 610041, Sichuan, China</mods:affiliation>
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<mods:date id="70BB59673B912999FA772DA8DEAC6913">2024</mods:date>
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10. 
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<emphasis id="F1CA3E20C942A502D180FF5CFE30ABF5" bold="true" box="[178,469,158,185]" italics="true" pageId="17" pageNumber="230">Leptochilus vietnamensis</emphasis>
Liang Zhang, N.T. Lu &amp; Li Bing Zhang
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, 
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(
<figureCitation id="5B85FEB7C942A502D53EFF5CFB85ABF5" box="[1036,1120,158,184]" captionStart="FIGURE 12" captionStartId="17.[136,229,1989,2011]" captionTargetBox="[195,1392,365,1952]" captionTargetId="figure-85@17.[183,1404,353,1964]" captionTargetPageId="17" captionText="FIGURE 12. Leptochilus vietnamensis Liang Zhang, N.T. Lu &amp; Li Bing Zhang,—A. Habit.—B. Abaxial laminae.—C. Abaxial view of portion of sterile lamina.—D. Rhizomes and petioles.—E. Rhizome scales. Photo credit: Liang Zhang." figureDoi="http://doi.org/10.5281/zenodo.13217918" httpUri="https://zenodo.org/record/13217918/files/figure.png" pageId="17" pageNumber="230">Fig. 12</figureCitation>
).
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<emphasis id="F1CA3E20C942A502D1BAFF24FF26AA4D" bold="true" box="[136,195,230,256]" pageId="17" pageNumber="230">Type</emphasis>
: 
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. 
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: Lac Son District, Ngoc Son Ngo Luong Nature Reserve, elev. 
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, 
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, on a tree trunk in limestone mountains, 
<date id="B700C4F2C942A502D30BFEC8FCF4AA69" box="[569,785,266,292]" pageId="17" pageNumber="230" value="2013-11-20">20 November 2013</date>
, 
<emphasis id="F1CA3E20C942A502D228FEC9FADEAA69" box="[794,1339,266,293]" italics="true" pageId="17" pageNumber="230">Li Bing Zhang, Liang Zhang &amp; Ngan Thi Lu 6334</emphasis>
(
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VNMN, 
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!).
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<emphasis id="F1CA3E20C942A502D1BAF807FEE1AC97" bold="true" box="[136,260,1989,2011]" pageId="17" pageNumber="230">FIGURE 12</emphasis>
. 
<taxonomicName id="04BE99B1C942A502D023F807FC6BAC96" authority="Liang Zhang, N. T. Lu &amp; Li Bing Zhang" authorityName="Liang Zhang, N. T. Lu &amp; Li Bing Zhang" box="[273,910,1989,2011]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="17" pageNumber="230" phylum="Tracheophyta" rank="species" species="vietnamensis">
<emphasis id="F1CA3E20C942A502D023F807FDE6AC97" box="[273,515,1989,2010]" italics="true" pageId="17" pageNumber="230">Leptochilus vietnamensis</emphasis>
Liang Zhang, N.T. Lu &amp; Li Bing Zhang
</taxonomicName>
,—A. Habit.—B. Abaxial laminae.—C. Abaxial view of portion of sterile lamina.—D. Rhizomes and petioles.—E. Rhizome scales. Photo credit: Liang Zhang.
</paragraph>
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<subSubSection id="8BA4B1B9C941A501D18FFF5CFD7EA9FD" pageId="18" pageNumber="231" type="diagnosis">
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<emphasis id="F1CA3E20C941A501D18FFF5CFED5ABF5" bold="true" box="[189,304,158,184]" pageId="18" pageNumber="231">Diagnosis</emphasis>
: 
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<emphasis id="F1CA3E20C941A501D073FF5CFDB8ABF5" box="[321,605,158,184]" italics="true" pageId="18" pageNumber="231">Leptochilus vietnamensis</emphasis>
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differs from other species in the genus by the combination of having long petioles in both fertile and sterile fronds, sterile laminae that can reach up to 
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in length, and fertile laminae that are less than half the length of the sterile laminae.
</paragraph>
<paragraph id="C301E232C941A501D18FFEC8FD7EA9FD" blockId="18.[136,1452,158,1552]" pageId="18" pageNumber="231">
<emphasis id="F1CA3E20C941A501D18FFEC8FEEDAA69" bold="true" box="[189,264,266,292]" pageId="18" pageNumber="231">Plants</emphasis>
perennial, evergreen, epiphytic. Rhizome long creeping, robust, ca. 2.5–5.0 mm in diam., dorsiventrally flattened, scaled, with a few short phyllopodia and roots.Apex of rhizome and basal petiole densely scaly, scales subulatelanceolate, 1.8–2.9 × 
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, brown, basifixed, concolorous, clathrate, margins entire or sparely denticulate, apex acuminate. 
<emphasis id="F1CA3E20C941A501D03EFEB4FE84AADD" bold="true" box="[268,353,374,400]" pageId="18" pageNumber="231">Fronds</emphasis>
dimorphic, fertile fronds longer than sterile fronds. 
<emphasis id="F1CA3E20C941A501D2F0FEB4FB82AADD" bold="true" box="[962,1127,374,400]" pageId="18" pageNumber="231">Sterile fronds</emphasis>
(15–) 
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tall, petiole shorter than lamina; 
<emphasis id="F1CA3E20C941A501D05DFE58FE2BAAF9" bold="true" box="[367,462,410,436]" pageId="18" pageNumber="231">laminae</emphasis>
ribbon-lanceolate, margins entire, slightly repand, base cuneate, apex acuminate, 
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long, (2.5–) 
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wide at middle, chartaceous, glabrous; 
<emphasis id="F1CA3E20C941A501D25AFE7CFC27AA95" bold="true" box="[872,962,446,472]" pageId="18" pageNumber="231">petioles</emphasis>
stramineous, grooved adaxially, glabrous, 
<quantity id="04464FD7C941A501D1BAFE20FEF4AAB0" box="[136,273,482,509]" metricMagnitude="-3" metricUnit="m" metricValue="1.85" metricValueMax="2.2" metricValueMin="1.5" pageId="18" pageNumber="231" unit="mm" value="1.85" valueMax="2.2" valueMin="1.5">1.5–2.2 mm</quantity>
in diam., (3–) 
<quantity id="04464FD7C941A501D084FE20FDE9AAB0" box="[438,524,482,509]" metricMagnitude="-2" metricUnit="m" metricValue="7.0" metricValueMax="9.0" metricValueMin="5.0" pageId="18" pageNumber="231" unit="cm" value="7.0" valueMax="9.0" valueMin="5.0">5–9 cm</quantity>
long. 
<emphasis id="F1CA3E20C941A501D36AFE20FD18AAB1" bold="true" box="[600,765,482,508]" pageId="18" pageNumber="231">Fertile fronds</emphasis>
<quantity id="04464FD7C941A501D235FE20FC9DAAB0" box="[775,888,482,509]" metricMagnitude="-1" metricUnit="m" metricValue="2.55" metricValueMax="3.0" metricValueMin="2.1" pageId="18" pageNumber="231" unit="cm" value="25.5" valueMax="30.0" valueMin="21.0">21–30 cm</quantity>
tall, petiole usually longer or the same length as lamina; 
<emphasis id="F1CA3E20C941A501D1D1FDC4FEA7A96D" bold="true" box="[227,322,518,544]" pageId="18" pageNumber="231">laminae</emphasis>
elliptic-lanceolate to ovate-lanceolate, margins entire, base cuneate, apex long acuminate, (8–) 
<quantity id="04464FD7C941A501D456FDC4FF4FA908" metricMagnitude="-1" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="18" pageNumber="231" unit="cm" value="12.5" valueMax="15.0" valueMin="10.0">10–15 cm</quantity>
long, 
<quantity id="04464FD7C941A501D1C6FDE8FEB8A908" box="[244,349,554,581]" metricMagnitude="-2" metricUnit="m" metricValue="4.25" metricValueMax="5.5" metricValueMin="3.0" pageId="18" pageNumber="231" unit="cm" value="4.25" valueMax="5.5" valueMin="3.0">3–5.5 cm</quantity>
wide at middle, chartaceous, glabrous; 
<emphasis id="F1CA3E20C941A501D211FDE8FC98A909" bold="true" box="[803,893,554,580]" pageId="18" pageNumber="231">petioles</emphasis>
<quantity id="04464FD7C941A501D2B7FDE8FBEBA908" box="[901,1038,554,581]" metricMagnitude="-3" metricUnit="m" metricValue="1.75" metricValueMax="2.1" metricValueMin="1.4" pageId="18" pageNumber="231" unit="mm" value="1.75" valueMax="2.1" valueMin="1.4">1.4–2.1 mm</quantity>
in diam., (8–) 
<quantity id="04464FD7C941A501D582FDE8FAC4A908" box="[1200,1313,554,581]" metricMagnitude="-1" metricUnit="m" metricValue="1.35" metricValueMax="1.5" metricValueMin="1.2" pageId="18" pageNumber="231" unit="cm" value="13.5" valueMax="15.0" valueMin="12.0">12–15 cm</quantity>
long. 
<emphasis id="F1CA3E20C941A501D459FDE8FA4EA909" bold="true" box="[1387,1451,554,580]" pageId="18" pageNumber="231">Veins</emphasis>
anastomosing, lateral vein distinct, veinlets visible. 
<emphasis id="F1CA3E20C941A501D3EAFD8CFCEFA925" bold="true" box="[728,778,590,616]" pageId="18" pageNumber="231">Sori</emphasis>
linear, reach lamina margins, 25–42 (–50) on each side of midrib, 
<quantity id="04464FD7C941A501D1D3FDB0FEBAA9C0" box="[225,351,626,653]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="3.2" metricValueMin="1.8" pageId="18" pageNumber="231" unit="cm" value="2.5" valueMax="3.2" valueMin="1.8">1.8–3.2 cm</quantity>
long, 
<quantity id="04464FD7C941A501D095FDB0FDCBA9C0" box="[423,558,626,653]" metricMagnitude="-3" metricUnit="m" metricValue="2.0" metricValueMax="2.2" metricValueMin="1.8" pageId="18" pageNumber="231" unit="mm" value="2.0" valueMax="2.2" valueMin="1.8">1.8–2.2 mm</quantity>
wide when mature, at angles of 44–62 degree with midribs, centers ca. 
<quantity id="04464FD7C941A501D465FDB0FF51A9FC" metricMagnitude="-3" metricUnit="m" metricValue="1.8" metricValueMax="2.1" metricValueMin="1.5" pageId="18" pageNumber="231" unit="mm" value="1.8" valueMax="2.1" valueMin="1.5">1.5–2.1 mm</quantity>
apart from one another. Paraphyses absent.
</paragraph>
</subSubSection>
<subSubSection id="8BA4B1B9C941A501D18FFD78FD4DAF80" pageId="18" pageNumber="231" type="materials_examined">
<paragraph id="C301E232C941A501D18FFD78FD4DAF80" blockId="18.[136,1452,158,1552]" pageId="18" pageNumber="231">
<emphasis id="F1CA3E20C941A501D18FFD78FDD3A999" bold="true" box="[189,566,698,724]" pageId="18" pageNumber="231">Additional specimens examined</emphasis>
: 
<materialsCitation id="73D6E86FC941A501D376FD78FB90A850" ID-GBIF-Occurrence="4928466308" collectingDate="1998-04-06" collectingDateMax="2003-11-29" collectingDateMin="1998-04-06" collectorName="Daniel K. Harder, P. K &amp; Loc &amp; T. T. Tai &amp; Su-Gong Wu, L. K &amp; Phan, J. Y &amp; Xiang, X &amp; Gong, V. T &amp; Nguyen &amp; K. S. Nguyen" country="Vietnam" county="Tra Tinh District" elevation="749" latitude="20.54" location="Trang Hen" longLatPrecision="761" longitude="105.18" municipality="Quoc Toan Municipality" pageId="18" pageNumber="231" specimenCount="1" stateProvince="Cao Bang">
<collectingCountry id="BBA9A2A2C941A501D376FD78FD2CA999" box="[580,713,698,724]" name="Vietnam" pageId="18" pageNumber="231">VIETNAM</collectingCountry>
. 
<collectingRegion id="017A2CD0C941A501D3E1FD78FCA3A998" box="[723,838,698,725]" country="Vietnam" name="Cao Bang" pageId="18" pageNumber="231">Cao Bang</collectingRegion>
: 
<collectingCounty id="2A609ABEC941A501D262FD79FBE8A999" box="[848,1037,698,725]" pageId="18" pageNumber="231">Tra Tinh District</collectingCounty>
, 
<collectingMunicipality id="23657848C941A501D52AFD78FACFA999" box="[1048,1322,698,725]" pageId="18" pageNumber="231">Quoc Toan Municipality</collectingMunicipality>
, 
<location id="C661B4E9C941A501D406FD79FA49A998" LSID="urn:lsid:plazi:treatment:4B175324C942A501D1BAFF5CFEDDAD5D:C661B4E9C941A501D406FD79FA49A998" box="[1332,1452,699,725]" country="Vietnam" county="Tra Tinh District" latitude="20.54" longLatPrecision="761" longitude="105.18" municipality="Quoc Toan Municipality" name="Trang Hen" pageId="18" pageNumber="231" stateProvince="Cao Bang">Trang Hen</location>
(
<location id="C661B4E9C941A501D1A2FD1CFED3A9B5" LSID="urn:lsid:plazi:treatment:4B175324C942A501D1BAFF5CFEDDAD5D:C661B4E9C941A501D1A2FD1CFED3A9B5" box="[144,310,734,760]" country="Vietnam" county="Tra Tinh District" latitude="20.54" longLatPrecision="761" longitude="105.18" municipality="Quoc Toan Municipality" name="Lakes District" pageId="18" pageNumber="231" stateProvince="Cao Bang">Lakes District</location>
), elev. 666–831 m, 
<geoCoordinate id="A68A84F5C941A501D325FD1CFD93A9B4" box="[535,630,734,761]" degrees="22.75" direction="north" orientation="latitude" pageId="18" pageNumber="231" precision="555" value="22.75">22.75°N</geoCoordinate>
/ 
<geoCoordinate id="A68A84F5C941A501D34DFD1CFD02A9B4" box="[639,743,734,761]" degrees="106.28" direction="east" orientation="longitude" pageId="18" pageNumber="231" precision="555" value="106.28">106.28°E</geoCoordinate>
, on tree branches in disturbed forests in mountains with solid limestone rocks, 
<date id="B700C4F2C941A501D076FCC0FE34A851" box="[324,465,770,796]" pageId="18" pageNumber="231" value="1998-04-06">
<collectingDate id="A7443D1AC941A501D076FCC0FE34A851" box="[324,465,770,796]" pageId="18" pageNumber="231" value="1998-04-06">6 April 1998</collectingDate>
</date>
, 
<emphasis id="F1CA3E20C941A501D0E9FCC0FC27A850" box="[475,962,770,797]" italics="true" pageId="18" pageNumber="231">
<collectorName id="6E4B87E4C941A501D0E9FCC0FD34A851" box="[475,721,770,796]" pageId="18" pageNumber="231">Daniel K. Harder, P.K</collectorName>
. 
<collectorName id="6E4B87E4C941A501D3E8FCC1FCE1A851" box="[730,772,771,796]" pageId="18" pageNumber="231">Loc</collectorName>
&amp; 
<collectorName id="6E4B87E4C941A501D217FCC1FC92A851" box="[805,887,771,796]" pageId="18" pageNumber="231">T.T. Tai</collectorName>
4254 (
</emphasis>
MO-3200946!
<emphasis id="F1CA3E20C941A501D556FCC1FB88A850" box="[1124,1133,771,797]" italics="true" pageId="18" pageNumber="231">)</emphasis>
.
</materialsCitation>
<materialsCitation id="73D6E86FC941A501D548FCC1FE7CA8C4" ID-GBIF-Occurrence="4928466306" collectingDate="1998-04-06" collectingDateMax="2003-11-29" collectingDateMin="1998-04-06" collectorName="Daniel K. Harder, P. K &amp; Loc &amp; T. T. Tai &amp; Su-Gong Wu, L. K &amp; Phan, J. Y &amp; Xiang, X &amp; Gong, V. T &amp; Nguyen &amp; K. S. Nguyen" country="Vietnam" county="Tra Tinh District" elevation="749" latitude="20.54" location="Trang Hen" longLatPrecision="761" longitude="105.18" municipality="Quoc Toan Municipality" pageId="18" pageNumber="231" specimenCount="1" stateProvince="Cao Bang">
<collectingRegion id="017A2CD0C941A501D548FCC1FB0CA851" box="[1146,1257,770,797]" country="Vietnam" name="Hoa Binh" pageId="18" pageNumber="231">Hoa Binh</collectingRegion>
: 
<location id="C661B4E9C941A501D5C0FCC1FA42A851" LSID="urn:lsid:plazi:treatment:4B175324C942A501D1BAFF5CFEDDAD5D:C661B4E9C941A501D5C0FCC1FA42A851" box="[1266,1447,770,797]" country="Vietnam" county="Tra Tinh District" latitude="20.54" longLatPrecision="761" longitude="105.18" municipality="Quoc Toan Municipality" name="Tan Lac District" pageId="18" pageNumber="231" stateProvince="Cao Bang">Tan Lac District</location>
, 
<location id="C661B4E9C941A501D1BAFCE5FE77A80D" LSID="urn:lsid:plazi:treatment:4B175324C942A501D1BAFF5CFEDDAD5D:C661B4E9C941A501D1BAFCE5FE77A80D" box="[136,402,806,833]" country="Vietnam" county="Tra Tinh District" latitude="20.54" longLatPrecision="761" longitude="105.18" municipality="Quoc Toan Municipality" name="Lung Van Municipality" pageId="18" pageNumber="231" stateProvince="Cao Bang">Lung Van Municipality</location>
, 
<location id="C661B4E9C941A501D0ADFCE5FDD0A80D" LSID="urn:lsid:plazi:treatment:4B175324C942A501D1BAFF5CFEDDAD5D:C661B4E9C941A501D0ADFCE5FDD0A80D" box="[415,565,806,833]" country="Vietnam" county="Tra Tinh District" latitude="20.54" longLatPrecision="761" longitude="105.18" municipality="Quoc Toan Municipality" name="Huop village" pageId="18" pageNumber="231" stateProvince="Cao Bang">Huop village</location>
, elev. 1095–1100 m, 
<geoCoordinate id="A68A84F5C941A501D219FCE4FC6EA80C" box="[811,907,806,833]" degrees="20.54" direction="north" orientation="latitude" pageId="18" pageNumber="231" precision="555" value="20.54">20.54°N</geoCoordinate>
/ 
<geoCoordinate id="A68A84F5C941A501D2A6FCE4FC1EA80C" box="[916,1019,806,833]" degrees="105.18" direction="east" orientation="longitude" pageId="18" pageNumber="231" precision="555" value="105.18">105.18°E</geoCoordinate>
, epiphytic, in heavily logged primary evergreen broad-leaved forest, 
<date id="B700C4F2C941A501D0EDFC88FD94A829" box="[479,625,842,869]" pageId="18" pageNumber="231" value="2003-11-29">
<collectingDate id="A7443D1AC941A501D0EDFC88FD94A829" box="[479,625,842,869]" pageId="18" pageNumber="231" value="2003-11-29">29 Nov 2003</collectingDate>
</date>
, 
<emphasis id="F1CA3E20C941A501D349FC89FF01A8C5" italics="true" pageId="18" pageNumber="231">
<collectorName id="6E4B87E4C941A501D349FC89FCA4A829" box="[635,833,843,869]" pageId="18" pageNumber="231">Su-Gong Wu, L.K</collectorName>
. 
<collectorName id="6E4B87E4C941A501D278FC88FC50A829" box="[842,949,842,868]" pageId="18" pageNumber="231">Phan, J.Y</collectorName>
. 
<collectorName id="6E4B87E4C941A501D28CFC89FBC6A829" box="[958,1059,843,868]" pageId="18" pageNumber="231">Xiang, X</collectorName>
. 
<collectorName id="6E4B87E4C941A501D519FC89FB79A829" box="[1067,1180,843,869]" pageId="18" pageNumber="231">Gong, V.T</collectorName>
. 
<collectorName id="6E4B87E4C941A501D596FC89FB1FA829" box="[1188,1274,843,868]" pageId="18" pageNumber="231">Nguyen</collectorName>
&amp; 
<collectorName id="6E4B87E4C941A501D429FC89FA42A829" box="[1307,1447,843,869]" pageId="18" pageNumber="231">K.S. Nguyen</collectorName>
, WP-303
</emphasis>
(MO-5980968)
</materialsCitation>
; 
<materialsCitation id="73D6E86FC941A501D091FCADFC78A8E1" ID-GBIF-Occurrence="4928466310" collectingDate="2013-11-22" collectionCode="VNMN, CDBI" collectorName="Li Bing Zhang &amp; Liang Zhang &amp; Ngan Thi Lu" country="Vietnam" elevation="840" latitude="20.53" location="Tam Son Community" longLatPrecision="761" longitude="105.18" municipality="Tan Lac District" pageId="18" pageNumber="231" specimenCount="1">
<collectingMunicipality id="23657848C941A501D091FCADFDBEA8C5" box="[419,603,878,905]" pageId="18" pageNumber="231">Tan Lac District</collectingMunicipality>
, 
<location id="C661B4E9C941A501D357FCADFCB2A8C5" LSID="urn:lsid:plazi:treatment:4B175324C942A501D1BAFF5CFEDDAD5D:C661B4E9C941A501D357FCADFCB2A8C5" box="[613,855,878,905]" country="Vietnam" latitude="20.53" longLatPrecision="761" longitude="105.18" municipality="Tan Lac District" name="Tam Son Community" pageId="18" pageNumber="231">Tam Son Community</location>
, elev. 840 m, 
<geoCoordinate id="A68A84F5C941A501D2C2FCACFBABA8C4" box="[1008,1102,878,905]" degrees="20.53" direction="north" orientation="latitude" pageId="18" pageNumber="231" precision="555" value="20.53">20.53°N</geoCoordinate>
/ 
<geoCoordinate id="A68A84F5C941A501D565FCACFB5AA8C4" box="[1111,1215,878,905]" degrees="105.18" direction="east" orientation="longitude" pageId="18" pageNumber="231" precision="555" value="105.18">105.18°E</geoCoordinate>
, 
<date id="B700C4F2C941A501D5FFFCACFA42A8C5" box="[1229,1447,878,904]" pageId="18" pageNumber="231" value="2013-11-22">
<collectingDate id="A7443D1AC941A501D5FFFCACFA42A8C5" box="[1229,1447,878,904]" pageId="18" pageNumber="231" value="2013-11-22">22 November 2013</collectingDate>
</date>
, 
<emphasis id="F1CA3E20C941A501D1BAFC51FD29A8E1" box="[136,716,914,941]" italics="true" pageId="18" pageNumber="231">
<collectorName id="6E4B87E4C941A501D1BAFC51FED4A8E1" box="[136,305,914,940]" pageId="18" pageNumber="231">Li Bing Zhang</collectorName>
, 
<collectorName id="6E4B87E4C941A501D00CFC51FE34A8E1" box="[318,465,914,940]" pageId="18" pageNumber="231">Liang Zhang</collectorName>
&amp; 
<collectorName id="6E4B87E4C941A501D0CBFC51FD6EA8E1" box="[505,651,914,940]" pageId="18" pageNumber="231">Ngan Thi Lu</collectorName>
6378
</emphasis>
(
<collectionCode id="A5AF7AF7C941A501D3EFFC50FCD2A8E1" box="[733,823,914,940]" country="Viet Nam" name="Vietnam National Museum of Nature" pageId="18" pageNumber="231">VNMN</collectionCode>
, 
<collectionCode id="A5AF7AF7C941A501D276FC50FC74A8E1" box="[836,913,914,940]" country="China" lsid="urn:lsid:biocol.org:col:12796" name="Chengdu Institute of Biology" pageId="18" pageNumber="231" type="Herbarium">CDBI</collectionCode>
!)
</materialsCitation>
; 
<materialsCitation id="73D6E86FC941A501D299FC50FCD9A8B8" ID-GBIF-Occurrence="4928466313" collectingDate="2002-12-03" collectorName="Daniel &amp; Harder &amp; P. K. Loc &amp; B. Hall &amp; N. Q. Hung" country="Vietnam" elevation="1368" latitude="20.74" location="Pa Co Municipality" longLatPrecision="760" longitude="104.89" municipality="Mai Chau District" pageId="18" pageNumber="231" specimenCount="1">
<collectingMunicipality id="23657848C941A501D299FC50FB99A8E1" box="[939,1148,914,940]" pageId="18" pageNumber="231">Mai Chau District</collectingMunicipality>
, 
<location id="C661B4E9C941A501D5BBFC51FA8FA8E1" LSID="urn:lsid:plazi:treatment:4B175324C942A501D1BAFF5CFEDDAD5D:C661B4E9C941A501D5BBFC51FA8FA8E1" box="[1161,1386,914,941]" country="Vietnam" latitude="20.74" longLatPrecision="760" longitude="104.89" municipality="Mai Chau District" name="Pa Co Municipality" pageId="18" pageNumber="231">Pa Co Municipality</location>
, elev. 1368 m, 
<geoCoordinate id="A68A84F5C941A501D1C2FC74FEAAA89D" box="[240,335,950,976]" degrees="20.74" direction="north" orientation="latitude" pageId="18" pageNumber="231" precision="555" value="20.74">20.74°N</geoCoordinate>
/ 
<geoCoordinate id="A68A84F5C941A501D06AFC74FE5AA89C" box="[344,447,950,977]" degrees="104.89" direction="east" orientation="longitude" pageId="18" pageNumber="231" precision="555" value="104.89">104.89°E</geoCoordinate>
, epiphytic, in wet, semi-deciduous and evergreen forest, 
<date id="B700C4F2C941A501D560FC74FACAA89D" box="[1106,1327,950,976]" pageId="18" pageNumber="231" value="2002-12-03">
<collectingDate id="A7443D1AC941A501D560FC74FACAA89D" box="[1106,1327,950,976]" pageId="18" pageNumber="231" value="2002-12-03">03 December 2002</collectingDate>
</date>
, 
<emphasis id="F1CA3E20C941A501D40EFC74FD98A8B9" italics="true" pageId="18" pageNumber="231">
<collectorName id="6E4B87E4C941A501D40EFC74FA6DA89D" box="[1340,1416,950,976]" pageId="18" pageNumber="231">Daniel</collectorName>
K. 
<collectorName id="6E4B87E4C941A501D1BAFC18FF3CA8B9" box="[136,217,986,1012]" pageId="18" pageNumber="231">Harder</collectorName>
, 
<collectorName id="6E4B87E4C941A501D1D6FC19FEA3A8B9" box="[228,326,986,1012]" pageId="18" pageNumber="231">P.K. Lôc</collectorName>
, 
<collectorName id="6E4B87E4C941A501D063FC19FE47A8B9" box="[337,418,986,1012]" pageId="18" pageNumber="231">B. Hall</collectorName>
&amp; 
<collectorName id="6E4B87E4C941A501D0F7FC19FDDBA8B9" box="[453,574,987,1013]" pageId="18" pageNumber="231">N.Q. Hung</collectorName>
7701
</emphasis>
(MO-3321598!)
</materialsCitation>
; 
<materialsCitation id="73D6E86FC941A501D275FC18FDC9AF71" ID-GBIF-Occurrence="4928466303" collectingDate="2002-12-05" collectorName="Daniel &amp; Harder &amp; B. Hall &amp; N. Q. Hung &amp; P. H. Hoang" country="Vietnam" county="Mai Chau District" elevation="1198" latitude="20.72" location="Thung Ang Village" longLatPrecision="760" longitude="104.89" municipality="Hang Kia Municipality" pageId="18" pageNumber="231" specimenCount="1">
<collectingCounty id="2A609ABEC941A501D275FC18FBF6A8B9" box="[839,1043,986,1012]" pageId="18" pageNumber="231">Mai Chau District</collectingCounty>
, 
<collectingMunicipality id="23657848C941A501D52CFC19FAC6A8B9" box="[1054,1315,986,1013]" pageId="18" pageNumber="231">Hang Kia Municipality</collectingMunicipality>
, 
<location id="C661B4E9C941A501D41FFC18FF39AF55" LSID="urn:lsid:plazi:treatment:4B175324C942A501D1BAFF5CFEDDAD5D:C661B4E9C941A501D41FFC18FF39AF55" country="Vietnam" county="Mai Chau District" latitude="20.72" longLatPrecision="760" longitude="104.89" municipality="Hang Kia Municipality" name="Thung Ang Village" pageId="18" pageNumber="231">Thung Ang Village</location>
, elev. 1095–1300 m, 
<geoCoordinate id="A68A84F5C941A501D0F9FC3CFDCFAF55" box="[459,554,1022,1048]" degrees="20.72" direction="north" orientation="latitude" pageId="18" pageNumber="231" precision="555" value="20.72">20.72°N</geoCoordinate>
/ 
<geoCoordinate id="A68A84F5C941A501D301FC3CFD7FAF54" box="[563,666,1022,1049]" degrees="104.89" direction="east" orientation="longitude" pageId="18" pageNumber="231" precision="555" value="104.89">104.89°E</geoCoordinate>
, epiphytic, 
<date id="B700C4F2C941A501D22CFC3CFC12AF55" box="[798,1015,1022,1049]" pageId="18" pageNumber="231" value="2002-12-05">
<collectingDate id="A7443D1AC941A501D22CFC3CFC12AF55" box="[798,1015,1022,1049]" pageId="18" pageNumber="231" value="2002-12-05">05 December 2002</collectingDate>
</date>
, 
<emphasis id="F1CA3E20C941A501D530FC3CFE89AF71" italics="true" pageId="18" pageNumber="231">
<collectorName id="6E4B87E4C941A501D530FC3CFBABAF55" box="[1026,1102,1022,1048]" pageId="18" pageNumber="231">Daniel</collectorName>
K. 
<collectorName id="6E4B87E4C941A501D544FC3CFB22AF55" box="[1142,1223,1022,1048]" pageId="18" pageNumber="231">Harder</collectorName>
, 
<collectorName id="6E4B87E4C941A501D5E0FC3DFAC3AF55" box="[1234,1318,1022,1048]" pageId="18" pageNumber="231">B. Hall</collectorName>
, 
<collectorName id="6E4B87E4C941A501D403FC3DFA4EAF55" box="[1329,1451,1023,1049]" pageId="18" pageNumber="231">N.Q. Hung</collectorName>
&amp; 
<collectorName id="6E4B87E4C941A501D194FBE1FECEAF71" box="[166,299,1059,1084]" pageId="18" pageNumber="231">P.H. Hoang</collectorName>
7782
</emphasis>
(MO-6082983!)
</materialsCitation>
; 
<materialsCitation id="73D6E86FC941A501D30BFBE0FE18AFC8" ID-GBIF-Occurrence="4928466304" collectingDate="1994-03-08" collectingDateMax="2002-12-09" collectingDateMin="1994-03-08" collectorName="Daniel K. Harder, T. H &amp; Nguyen &amp; L. V. Averyanov &amp; D. K. Harder &amp; N. T. Hiep &amp; H. H. Schmidt &amp; N. Q. Binh" country="Vietnam" county="Mai Chau District" elevation="1147" latitude="22.33" location="Na Pai Village" longLatPrecision="756" longitude="105.42" municipality="Pa Co Municipality" pageId="18" pageNumber="231" specimenCount="1" stateProvince="Tuyen Quang">
<collectingCounty id="2A609ABEC941A501D30BFBE0FCEFAF71" box="[569,778,1058,1084]" pageId="18" pageNumber="231">Mai Chau District</collectingCounty>
, 
<collectingMunicipality id="23657848C941A501D224FBE1FC13AF71" box="[790,1014,1058,1085]" pageId="18" pageNumber="231">Pa Co Municipality</collectingMunicipality>
, near 
<location id="C661B4E9C941A501D509FBE1FB00AF71" LSID="urn:lsid:plazi:treatment:4B175324C942A501D1BAFF5CFEDDAD5D:C661B4E9C941A501D509FBE1FB00AF71" box="[1083,1253,1058,1085]" country="Vietnam" county="Mai Chau District" latitude="22.33" longLatPrecision="756" longitude="105.42" municipality="Pa Co Municipality" name="Na Pai Village" pageId="18" pageNumber="231" stateProvince="Tuyen Quang">Na Pai Village</location>
, elev. 1094–1200 m, 
<geoCoordinate id="A68A84F5C941A501D19EFB84FEEEAF2D" box="[172,267,1094,1120]" degrees="20.74" direction="north" orientation="latitude" pageId="18" pageNumber="231" precision="555" value="20.74">20.74°N</geoCoordinate>
/ 
<geoCoordinate id="A68A84F5C941A501D026FB84FE9EAF2C" box="[276,379,1094,1121]" degrees="104.94" direction="east" orientation="longitude" pageId="18" pageNumber="231" precision="555" value="104.94">104.94°E</geoCoordinate>
, lithophytic and epiphytic on rocks, 
<date id="B700C4F2C941A501D224FB84FC0BAF2D" box="[790,1006,1094,1120]" pageId="18" pageNumber="231" value="2002-12-09">
<collectingDate id="A7443D1AC941A501D224FB84FC0BAF2D" box="[790,1006,1094,1120]" pageId="18" pageNumber="231" value="2002-12-09">09 December 2002</collectingDate>
</date>
, 
<emphasis id="F1CA3E20C941A501D2CBFB84FED9AFC9" italics="true" pageId="18" pageNumber="231">
<collectorName id="6E4B87E4C941A501D2CBFB84FB13AF2D" box="[1017,1270,1094,1120]" pageId="18" pageNumber="231">Daniel K. Harder, T.H</collectorName>
. 
<collectorName id="6E4B87E4C941A501D432FB84FAB3AF2D" box="[1280,1366,1094,1120]" pageId="18" pageNumber="231">Nguyên</collectorName>
&amp; 
<collectorName id="6E4B87E4C941A501D44BFB85FF1BAFC8" pageId="18" pageNumber="231">L. V. Averyanov</collectorName>
8020
</emphasis>
(MO-3321599!).
</materialsCitation>
<materialsCitation id="73D6E86FC941A501D333FBA9FD41AF80" ID-GBIF-Occurrence="4928466320" collectingDate="1994-03-08" collectingDateMax="2002-12-09" collectingDateMin="1994-03-08" collectorName="Daniel K. Harder, T. H &amp; Nguyen &amp; L. V. Averyanov &amp; D. K. Harder &amp; N. T. Hiep &amp; H. H. Schmidt &amp; N. Q. Binh" country="Vietnam" county="Mai Chau District" elevation="1147" latitude="22.33" location="Na Pai Village" longLatPrecision="756" longitude="105.42" municipality="Pa Co Municipality" pageId="18" pageNumber="231" specimenCount="1" stateProvince="Tuyen Quang">
<collectingRegion id="017A2CD0C941A501D333FBA9FD79AFC9" box="[513,668,1130,1157]" country="Vietnam" name="Tuyen Quang" pageId="18" pageNumber="231">Tuyen Quang</collectingRegion>
: 
<location id="C661B4E9C941A501D397FBA9FC81AFC9" LSID="urn:lsid:plazi:treatment:4B175324C942A501D1BAFF5CFEDDAD5D:C661B4E9C941A501D397FBA9FC81AFC9" box="[677,868,1130,1157]" country="Vietnam" county="Mai Chau District" latitude="22.33" longLatPrecision="756" longitude="105.42" municipality="Pa Co Municipality" name="Na Hang District" pageId="18" pageNumber="231" stateProvince="Tuyen Quang">Na Hang District</location>
, 
<location id="C661B4E9C941A501D25FFBA9FBA4AFC8" LSID="urn:lsid:plazi:treatment:4B175324C942A501D1BAFF5CFEDDAD5D:C661B4E9C941A501D25FFBA9FBA4AFC8" box="[877,1089,1130,1157]" country="Vietnam" county="Mai Chau District" latitude="22.33" longLatPrecision="756" longitude="105.42" municipality="Pa Co Municipality" name="Nam Chang Forest" pageId="18" pageNumber="231" stateProvince="Tuyen Quang">Nam Chang Forest</location>
, elev. 210 m, 
<geoCoordinate id="A68A84F5C941A501D5E7FBA8FAD6AFC8" box="[1237,1331,1130,1157]" degrees="22.33" direction="north" orientation="latitude" pageId="18" pageNumber="231" precision="555" value="22.33">22.33°N</geoCoordinate>
/ 
<geoCoordinate id="A68A84F5C941A501D40EFBA8FA41AFC8" box="[1340,1444,1130,1157]" degrees="105.42" direction="east" orientation="longitude" pageId="18" pageNumber="231" precision="555" value="105.42">105.42°E</geoCoordinate>
, on tree branches, 1 m from ground, in primary forests on sandy quartz soils, 
<date id="B700C4F2C941A501D2C2FB4CFB77AFE5" box="[1008,1170,1166,1192]" pageId="18" pageNumber="231" value="1994-03-08">
<collectingDate id="A7443D1AC941A501D2C2FB4CFB77AFE5" box="[1008,1170,1166,1192]" pageId="18" pageNumber="231" value="1994-03-08">8 March 1994</collectingDate>
</date>
, 
<emphasis id="F1CA3E20C941A501D5ACFB4DFE16AF81" italics="true" pageId="18" pageNumber="231">
<collectorName id="6E4B87E4C941A501D5ACFB4DFACEAFE5" box="[1182,1323,1166,1192]" pageId="18" pageNumber="231">D.K. Harder</collectorName>
, 
<collectorName id="6E4B87E4C941A501D40AFB4DFA42AFE5" box="[1336,1447,1166,1192]" pageId="18" pageNumber="231">N.T. Hiêp</collectorName>
, 
<collectorName id="6E4B87E4C941A501D1BAFB71FEFAAF81" box="[136,287,1202,1228]" pageId="18" pageNumber="231">H.H. Schmidt</collectorName>
&amp; 
<collectorName id="6E4B87E4C941A501D071FB71FE51AF81" box="[323,436,1202,1229]" pageId="18" pageNumber="231">N.Q. Biñh</collectorName>
2412
</emphasis>
(MO-320056!)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="8BA4B1B9C941A501D18FFB14FB83AE59" pageId="18" pageNumber="231" type="distribution">
<paragraph id="C301E232C941A501D18FFB14FB83AE59" blockId="18.[136,1452,158,1552]" pageId="18" pageNumber="231">
<emphasis id="F1CA3E20C941A501D18FFB14FE09AFBD" bold="true" box="[189,492,1238,1264]" pageId="18" pageNumber="231">Distribution and habitat:</emphasis>
<taxonomicName id="04BE99B1C941A501D0C7FB14FCF4AFBD" box="[501,785,1238,1264]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="18" pageNumber="231" phylum="Tracheophyta" rank="species" species="vietnamensis">
<emphasis id="F1CA3E20C941A501D0C7FB14FCF4AFBD" box="[501,785,1238,1264]" italics="true" pageId="18" pageNumber="231">Leptochilus vietnamensis</emphasis>
</taxonomicName>
was discovered in four localities within three provinces in northern 
<collectingCountry id="BBA9A2A2C941A501D1DFFB38FEB5AE59" box="[237,336,1274,1300]" name="Vietnam" pageId="18" pageNumber="231">Vietnam</collectingCountry>
. The species is observed growing on tree branches, trunks or on rocks.
</paragraph>
</subSubSection>
<subSubSection id="8BA4B1B9C941A501D18FFADCFEDDAD5D" pageId="18" pageNumber="231" type="etymology">
<paragraph id="C301E232C941A501D18FFADCFF0AAE11" blockId="18.[136,1452,158,1552]" pageId="18" pageNumber="231">
<emphasis id="F1CA3E20C941A501D18FFADCFEDBAE75" bold="true" box="[189,318,1310,1336]" pageId="18" pageNumber="231">Etymology</emphasis>
: The specific epithet is derived from the name of the country where the new species was discovered, 
<collectingCountry id="BBA9A2A2C941A501D1BAFA80FF0EAE11" box="[136,235,1346,1372]" name="Vietnam" pageId="18" pageNumber="231">Vietnam</collectingCountry>
.
</paragraph>
<paragraph id="C301E232C941A501D18FFAA4FEDDAD5D" blockId="18.[136,1452,158,1552]" pageId="18" pageNumber="231">
<emphasis id="F1CA3E20C941A501D18FFAA4FF1AAECD" bold="true" box="[189,255,1382,1408]" pageId="18" pageNumber="231">Note:</emphasis>
<taxonomicName id="04BE99B1C941A501D03AFAA4FDC0AECD" box="[264,549,1382,1408]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="18" pageNumber="231" phylum="Tracheophyta" rank="species" species="vietnamensis">
<emphasis id="F1CA3E20C941A501D03AFAA4FDC0AECD" box="[264,549,1382,1408]" italics="true" pageId="18" pageNumber="231">Leptochilus vietnamensis</emphasis>
</taxonomicName>
shares similarities with both 
<taxonomicName id="04BE99B1C941A501D24BFAA5FC25AECD" authorityName="Liang Zhang &amp; Li Bing Zhang" baseAuthorityName="C. Christensen" baseAuthorityYear="1933" box="[889,960,1383,1408]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="18" pageNumber="231" phylum="Tracheophyta" rank="species" species="wui">
<emphasis id="F1CA3E20C941A501D24BFAA5FC25AECD" box="[889,960,1383,1408]" italics="true" pageId="18" pageNumber="231">L. wui</emphasis>
</taxonomicName>
and 
<taxonomicName id="04BE99B1C941A501D2CEFAA5FB8FAECD" authorityName="Liang Zhang &amp; Li Bing Zhang" authorityYear="2018" box="[1020,1130,1382,1408]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="18" pageNumber="231" phylum="Tracheophyta" rank="species" species="chingii">
<emphasis id="F1CA3E20C941A501D2CEFAA5FB8FAECD" box="[1020,1130,1382,1408]" italics="true" pageId="18" pageNumber="231">L. chingii</emphasis>
</taxonomicName>
, as the latter two both have slightly dimorphic fronds and linear sori. However, 
<taxonomicName id="04BE99B1C941A501D3D4FA49FC7CAEE9" box="[742,921,1419,1444]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="18" pageNumber="231" phylum="Tracheophyta" rank="species" species="vietnamensis">
<emphasis id="F1CA3E20C941A501D3D4FA49FC7CAEE9" box="[742,921,1419,1444]" italics="true" pageId="18" pageNumber="231">L. vietnamensis</emphasis>
</taxonomicName>
can be distinguished from them by its fertile fronds, which are shorter than its sterile fronds, and the long, ribbon-like lamina of its sterile fronds. The phylogenetic analysis resolved 
<taxonomicName id="04BE99B1C941A501D066FA11FDE3AEA1" box="[340,518,1491,1516]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="18" pageNumber="231" phylum="Tracheophyta" rank="species" species="vietnamensis">
<emphasis id="F1CA3E20C941A501D066FA11FDE3AEA1" box="[340,518,1491,1516]" italics="true" pageId="18" pageNumber="231">L. vietnamensis</emphasis>
</taxonomicName>
as sister to 
<taxonomicName id="04BE99B1C941A501D3A6FA11FC6EAEA1" box="[660,907,1490,1516]" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="18" pageNumber="231" phylum="Tracheophyta" rank="species" species="luangprabangensis">
<emphasis id="F1CA3E20C941A501D3A6FA11FC6EAEA1" box="[660,907,1490,1516]" italics="true" pageId="18" pageNumber="231">L. luangprabangensis</emphasis>
</taxonomicName>
, and these two species together are sister to 
<taxonomicName id="04BE99B1C941A501D4A7FA11FF33AD5D" authorityName="Liang Zhang &amp; Li Bing Zhang" authorityYear="2018" class="Polypodiopsida" family="Polypodiaceae" genus="Leptochilus" kingdom="Plantae" order="Polypodiales" pageId="18" pageNumber="231" phylum="Tracheophyta" rank="species" species="chingii">
<emphasis id="F1CA3E20C941A501D4A7FA11FF33AD5D" italics="true" pageId="18" pageNumber="231">L. chingii</emphasis>
</taxonomicName>
(
<figureCitation id="5B85FEB7C941A501D1D7FA34FEC2AD5D" box="[229,295,1526,1552]" captionStart="FIGURE 1" captionStartId="3.[136,229,1956,1978]" captionTargetBox="[153,1430,353,1922]" captionTargetId="figure-76@3.[151,1435,347,1928]" captionTargetPageId="3" captionText="FIGURE 1. Phylogeny of Leptochilus based on six plastid markers (modified from Zhang et al. 2024). Orange color indicates the new species described here." figureDoi="http://doi.org/10.5281/zenodo.13217896" httpUri="https://zenodo.org/record/13217896/files/figure.png" pageId="18" pageNumber="231">Fig. 1</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>