149 lines
24 KiB
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149 lines
24 KiB
XML
<document id="D43B8EDBECF8A8070152DDB21FBC0BB1" ID-DOI="10.5281/zenodo.5352924" ID-ISSN="1942-1354" ID-Zenodo-Dep="5352924" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630209239027" checkinUser="felipe" docAuthor="Edmonds, W. D. & Zidek, J." docDate="2010" docId="3A7F7B2C7F708871FF34FD1325F0EEF2" docLanguage="en" docName="0129.pdf" docOrigin="Insecta Mundi 2010 (129)" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Coprophanaeus Olsoufieff 1924" docType="key" docVersion="4" lastPageNumber="9" masterDocId="C64603547F7C887DFFF4FFB02050E849" masterDocTitle="A taxonomic review of the neotropical genus Coprophanaeus Olsoufieff, 1924 (Coleoptera: Scarabaeidae, Scarabaeinae)" masterLastPageNumber="111" masterPageNumber="1" pageNumber="9" updateTime="1699186523840" updateUser="plazi" zenodo-license-document="CC-BY-NC-3.0">
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<mods:mods id="1E2A065B78A819329D1BB6842D3192E6" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="BB3637E4B1940013BB40F594705C914B">
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<mods:title id="E2E399E4D968D3DE37699BE948EED661">A taxonomic review of the neotropical genus Coprophanaeus Olsoufieff, 1924 (Coleoptera: Scarabaeidae, Scarabaeinae)</mods:title>
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<mods:name id="DD6B219FD9C415CC4E3914E7DEAB3655" type="personal">
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<mods:namePart id="219A2D30D1ADAF9B43473C735EF73A46">Edmonds, W. D.</mods:namePart>
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<mods:name id="CB49683C8ABBED6A4055CFAD5772E28A" type="personal">
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<mods:namePart id="8C529F5CD0FFDD455273ADF34D3F05F0">Zidek, J.</mods:namePart>
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<mods:affiliation id="1379F5A046D33367CDB2A699DC5EA01A">Coprophanaeus bonariensis (Gory)</mods:affiliation>
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<mods:title id="3A8D946324CBECFEAE27F352BA736DC3">Insecta Mundi</mods:title>
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<mods:part id="D7C10F9E86B802A10DAF8FA48992B7AE">
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<mods:date id="360F8D971E889BF28B16648CE159FA55">2010</mods:date>
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<mods:number id="703DE77A62FBFD1D41328A6503CB2459">2010-07-02</mods:number>
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<mods:number id="17A0FBE197085F1C0FA55D08D4B2ED8A">2010</mods:number>
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<mods:number id="D09E4BE96C1CAA6868D3C616D7081AE9">129</mods:number>
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<mods:identifier id="6818033BC47547DA4A06C4276F84AA4E" type="DOI">10.5281/zenodo.5352924</mods:identifier>
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<mods:identifier id="38C41131077E32071002C21334D1076D" type="ISSN">1942-1354</mods:identifier>
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<treatment id="3A7F7B2C7F708871FF34FD1325F0EEF2" ID-DOI="http://doi.org/10.5281/zenodo.5450866" ID-Zenodo-Dep="5450866" LSID="urn:lsid:plazi:treatment:3A7F7B2C7F708871FF34FD1325F0EEF2" httpUri="http://treatment.plazi.org/id/3A7F7B2C7F708871FF34FD1325F0EEF2" lastPageNumber="9" pageId="12" pageNumber="9">
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<subSubSection id="FACC99B17F708871FF34FD13227AEA92" pageId="12" pageNumber="9" type="nomenclature">
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<paragraph id="B269CA3A7F708871FF34FD132366EAF2" blockId="12.[192,822,674,731]" box="[192,822,674,699]" pageId="12" pageNumber="9">
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<heading id="E9217D567F708871FF34FD132366EAF2" bold="true" box="[192,822,674,699]" fontSize="10" level="2" pageId="12" pageNumber="9" reason="3">
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<emphasis id="80A216287F708871FF34FD132366EAF2" bold="true" box="[192,822,674,699]" pageId="12" pageNumber="9">
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Key to subgenera of
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<taxonomicName id="75D6B1B97F708871FE25FD122366EAF2" ID-CoL="3TRJ" authority="Olsoufieff" authorityName="Olsoufieff" authorityYear="1924" box="[465,822,674,699]" class="Insecta" family="Scarabaeidae" genus="Coprophanaeus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="9" phylum="Arthropoda" rank="genus">
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<emphasis id="80A216287F708871FE25FD1222F9EAF2" bold="true" box="[465,681,674,699]" italics="true" pageId="12" pageNumber="9">Coprophanaeus</emphasis>
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Olsoufieff
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="B269CA3A7F708871FF34FD73227AEA92" blockId="12.[192,822,674,731]" box="[192,554,707,731]" pageId="12" pageNumber="9">
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(Modified from
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<bibRefCitation id="D647B7CB7F708871FE86FD732272EA92" author="Edmonds, W. D." box="[370,546,707,731]" pageId="12" pageNumber="9" pagination="731 - 874" refId="ref56889" refString="Edmonds, W. D. 1972. Comparative skeletal morphology, systematic and evolution of the phanaeine dung beetles (Coleoptera: Scarabaeidae). The University of Kansas Science Bulletin 49: 731 - 874." type="journal article" year="1972">Edmonds 1972</bibRefCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="FACC99B17F708871FF34FCB325F0EEF2" pageId="12" pageNumber="9" type="key">
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<keyStep id="0922D29F7F708871FF34FCB325F0EC92" pageId="12" pageNumber="9">
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<paragraph id="B269CA3A7F708871FF34FCB325F0EC52" blockId="12.[192,1441,771,1243]" pageId="12" pageNumber="9">
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<keyLead id="0927690F7F708871FF34FCB325F0EC52" pageId="12" pageNumber="9">
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1. Posterior surface of each protibial tooth with dense, oblique clump of short setae (
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<figureCitation id="2AEDD6BF7F708871FB02FCB3256CEB52" box="[1270,1340,771,795]" captionStart="Figure 5-12" captionStartId="13.[195,279,1777,1799]" captionTargetBox="[196,1435,194,1764]" captionTargetId="figure-18@13.[196,1435,194,1764]" captionTargetPageId="13" captionText="Figure 5-12. Characters of Coprophanaeus. 5) Ventral view of front left tibia of C. ensifer (arrow marks elongate tuft of setae). 6) Same, C. dardanus. 7) Left anterolateral pronotal margin of C. spitzi (arrow indicates unbroken circumnotal carina. 8) Same, C. magnoi (arrow indicates broken circumnotal carina). 9) Base of left hind wing of C. cerberus. 10) Same, C. horus. 11) Lateral view left occipital lobe of C. ensifer (asterisk marks swelling). 12) Same, C. spitzi (asterisk marks flat area)." figureDoi="http://doi.org/10.5281/zenodo.5352932" httpUri="https://zenodo.org/record/5352932/files/figure.png" pageId="12" pageNumber="9">Fig. 5</figureCitation>
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, arrow). Striae conspicuous, always carinulate, with straight or sinuous margins (
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<figureCitation id="2AEDD6BF7F708871FB5CFC932571EB72" box="[1192,1313,803,827]" captionStart="Figure 13-27" captionStartId="15.[192,276,1691,1713]" captionTargetBox="[280,1357,196,1662]" captionTargetId="figure-18@15.[280,1357,195,1662]" captionTargetPageId="15" captionText="Figure 13-27. Characters of the Coprophanaeus subgenus Megaphanaeus. 13) Elytral surface of C. lancifer. 14) Same, C. ensifer. 15) Same, C. bonariensis. 16) Same, C. bellicosus. 17) Mesosternum and anterior portion of metasternum of C. lancifer (arrow indicates smooth area of metasternum). 18) Metasternum of C.lancifer. 19) Same, C. ensifer. 20) Dorsal view of posteromedian pronotal prominence of female C. ensifer (broad form). 21) Same (narrow form). 22) Partial lateral view of pronotum male C. ensifer (arrow indicates lateral spine). 23) Same, C. lancifer (arrow indicates vertical ridge). 24) Dorsal view of posteromedian pronotal prominence of male C. lancifer. 25) Same, female (broad form). 26) Same (narrow form). 27) Anterior view of cephalic process of female C. bellicosus." figureDoi="http://doi.org/10.5281/zenodo.5352934" httpUri="https://zenodo.org/record/5352934/files/figure.png" pageId="12" pageNumber="9">Fig. 13-16</figureCitation>
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). Protarsi present in female (except
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<taxonomicName id="75D6B1B97F708871FD98FCF4234AEB12" ID-CoL="Y6CB" baseAuthorityName="Gory" baseAuthorityYear="1844" box="[620,794,835,859]" class="Insecta" family="Scarabaeidae" genus="Coprophanaeus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="9" phylum="Arthropoda" rank="species" species="bonariensis">
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<emphasis id="80A216287F708871FD98FCF4234AEB12" box="[620,794,835,859]" italics="true" pageId="12" pageNumber="9">C. bonariensis</emphasis>
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</taxonomicName>
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). Posterior margin of hind wing lacking distinct basal notch (as in
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<figureCitation id="2AEDD6BF7F708871FE32FCD3224AEB32" box="[454,538,867,891]" captionStart="Figure 5-12" captionStartId="13.[195,279,1777,1799]" captionTargetBox="[196,1435,194,1764]" captionTargetId="figure-18@13.[196,1435,194,1764]" captionTargetPageId="13" captionText="Figure 5-12. Characters of Coprophanaeus. 5) Ventral view of front left tibia of C. ensifer (arrow marks elongate tuft of setae). 6) Same, C. dardanus. 7) Left anterolateral pronotal margin of C. spitzi (arrow indicates unbroken circumnotal carina. 8) Same, C. magnoi (arrow indicates broken circumnotal carina). 9) Base of left hind wing of C. cerberus. 10) Same, C. horus. 11) Lateral view left occipital lobe of C. ensifer (asterisk marks swelling). 12) Same, C. spitzi (asterisk marks flat area)." figureDoi="http://doi.org/10.5281/zenodo.5352932" httpUri="https://zenodo.org/record/5352932/files/figure.png" pageId="12" pageNumber="9">Fig. 10</figureCitation>
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). Sexual dimorphism not pronounced; male and female (except
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<taxonomicName id="75D6B1B97F708871FAF0FCD425C7EB32" baseAuthorityName="Olivier" baseAuthorityYear="1789" box="[1284,1431,867,891]" class="Insecta" family="Scarabaeidae" genus="Phanaeus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="9" phylum="Arthropoda" rank="species" species="bellicosus">
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<emphasis id="80A216287F708871FAF0FCD425C7EB32" box="[1284,1431,867,891]" italics="true" pageId="12" pageNumber="9">C. bellicosus</emphasis>
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</taxonomicName>
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) with long, thick, tapering head horn and massive, saddle-shaped pronotal prominence projecting forward above broad concavity. Occipital areas each with angulate prominence (
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<figureCitation id="2AEDD6BF7F708871FB22FC132578EBF2" box="[1238,1320,931,955]" captionStart="Figure 5-12" captionStartId="13.[195,279,1777,1799]" captionTargetBox="[196,1435,194,1764]" captionTargetId="figure-18@13.[196,1435,194,1764]" captionTargetPageId="13" captionText="Figure 5-12. Characters of Coprophanaeus. 5) Ventral view of front left tibia of C. ensifer (arrow marks elongate tuft of setae). 6) Same, C. dardanus. 7) Left anterolateral pronotal margin of C. spitzi (arrow indicates unbroken circumnotal carina. 8) Same, C. magnoi (arrow indicates broken circumnotal carina). 9) Base of left hind wing of C. cerberus. 10) Same, C. horus. 11) Lateral view left occipital lobe of C. ensifer (asterisk marks swelling). 12) Same, C. spitzi (asterisk marks flat area)." figureDoi="http://doi.org/10.5281/zenodo.5352932" httpUri="https://zenodo.org/record/5352932/files/figure.png" pageId="12" pageNumber="9">Fig. 11</figureCitation>
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, asterisk); paraocular areas (“genae”) carinate lateral to upper eye. Large to very large, length 30-50+ mm. Dorsum weakly shining metallic green or blue, or dull, dark violet. South America east of the Andes ...........................................................................................
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<taxonomicName id="75D6B1B97F708871FBB4FBB225F0EC52" authority="Olsoufieff" authorityName="Olsoufieff" box="[1088,1440,1026,1051]" class="Insecta" family="Scarabaeidae" genus="Megaphanaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="9" phylum="Arthropoda" rank="genus">
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<emphasis id="80A216287F708871FBB4FBB225F0EC52" bold="true" box="[1088,1440,1026,1051]" pageId="12" pageNumber="9">
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<emphasis id="80A216287F708871FBB4FBB22541EC52" bold="true" box="[1088,1297,1026,1051]" italics="true" pageId="12" pageNumber="9">Megaphanaeus</emphasis>
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Olsoufieff
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</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="B269CA3A7F708871FF34FB9425F0EC92" blockId="12.[192,1441,771,1243]" pageId="12" pageNumber="9">
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<keyLead id="0927690F7F708871FF34FB9425F0EC92" pageId="12" pageNumber="9">
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— Posterior surface of each protibial tooth with single oblique row of setae at base (
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<figureCitation id="2AEDD6BF7F708871FB0CFB93256FEC72" box="[1272,1343,1059,1083]" captionStart="Figure 5-12" captionStartId="13.[195,279,1777,1799]" captionTargetBox="[196,1435,194,1764]" captionTargetId="figure-18@13.[196,1435,194,1764]" captionTargetPageId="13" captionText="Figure 5-12. Characters of Coprophanaeus. 5) Ventral view of front left tibia of C. ensifer (arrow marks elongate tuft of setae). 6) Same, C. dardanus. 7) Left anterolateral pronotal margin of C. spitzi (arrow indicates unbroken circumnotal carina. 8) Same, C. magnoi (arrow indicates broken circumnotal carina). 9) Base of left hind wing of C. cerberus. 10) Same, C. horus. 11) Lateral view left occipital lobe of C. ensifer (asterisk marks swelling). 12) Same, C. spitzi (asterisk marks flat area)." figureDoi="http://doi.org/10.5281/zenodo.5352932" httpUri="https://zenodo.org/record/5352932/files/figure.png" pageId="12" pageNumber="9">Fig. 6</figureCitation>
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). Striae conspicuous or not; rarely carinulate. Protarsi absent in female. Base of hind wing notched or not. Sexual dimorphism pronounced; configuration of male and female head and pronotal shape usually very distinct. Occipital areas more-or-less flattened, lacking angular prominences (
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<figureCitation id="2AEDD6BF7F708871FA85FB332108ECF2" captionStart="Figure 5-12" captionStartId="13.[195,279,1777,1799]" captionTargetBox="[196,1435,194,1764]" captionTargetId="figure-18@13.[196,1435,194,1764]" captionTargetPageId="13" captionText="Figure 5-12. Characters of Coprophanaeus. 5) Ventral view of front left tibia of C. ensifer (arrow marks elongate tuft of setae). 6) Same, C. dardanus. 7) Left anterolateral pronotal margin of C. spitzi (arrow indicates unbroken circumnotal carina. 8) Same, C. magnoi (arrow indicates broken circumnotal carina). 9) Base of left hind wing of C. cerberus. 10) Same, C. horus. 11) Lateral view left occipital lobe of C. ensifer (asterisk marks swelling). 12) Same, C. spitzi (asterisk marks flat area)." figureDoi="http://doi.org/10.5281/zenodo.5352932" httpUri="https://zenodo.org/record/5352932/files/figure.png" pageId="12" pageNumber="9">Fig. 12</figureCitation>
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, asterisk); paraocular areas (genae) carinate or not lateral to upper eye. Small to mediumsized, length
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<quantity id="752E67DF7F708871FE23FB732204EC92" box="[471,596,1219,1243]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="3.5" metricValueMin="1.5" pageId="12" pageNumber="9" unit="mm" value="25.0" valueMax="35.0" valueMin="15.0">15-35 mm</quantity>
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. Color and distribution variable .........................................................
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<emphasis id="80A216287F708871FA65FB7325F0EC92" bold="true" box="[1425,1440,1219,1243]" pageId="12" pageNumber="9">2</emphasis>
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</keyLead>
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</paragraph>
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</keyStep>
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<keyStep id="0922D29F7F708871FF34FAB325F0EEF2" pageId="12" pageNumber="9">
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<paragraph id="B269CA3A7F708871FF34FAB325F0EDF2" blockId="12.[192,1440,1283,1723]" pageId="12" pageNumber="9">
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<keyLead id="0927690F7F708871FF34FAB325F0EDF2" pageId="12" pageNumber="9">
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2(1). Hind wing not notched basally (
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<figureCitation id="2AEDD6BF7F708871FD5CFAB32350ED52" box="[680,768,1283,1307]" captionStart="Figure 5-12" captionStartId="13.[195,279,1777,1799]" captionTargetBox="[196,1435,194,1764]" captionTargetId="figure-18@13.[196,1435,194,1764]" captionTargetPageId="13" captionText="Figure 5-12. Characters of Coprophanaeus. 5) Ventral view of front left tibia of C. ensifer (arrow marks elongate tuft of setae). 6) Same, C. dardanus. 7) Left anterolateral pronotal margin of C. spitzi (arrow indicates unbroken circumnotal carina. 8) Same, C. magnoi (arrow indicates broken circumnotal carina). 9) Base of left hind wing of C. cerberus. 10) Same, C. horus. 11) Lateral view left occipital lobe of C. ensifer (asterisk marks swelling). 12) Same, C. spitzi (asterisk marks flat area)." figureDoi="http://doi.org/10.5281/zenodo.5352932" httpUri="https://zenodo.org/record/5352932/files/figure.png" pageId="12" pageNumber="9">Fig. 10</figureCitation>
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). Width of lower portion of eye less than twice (usually about equal) that of adjacent oculogular space. Beading of anterior portion of circumnotal carina entire, not effaced behind eyes (as in
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<figureCitation id="2AEDD6BF7F708871FD0CFAF3236FED12" box="[760,831,1347,1371]" captionStart="Figure 5-12" captionStartId="13.[195,279,1777,1799]" captionTargetBox="[196,1435,194,1764]" captionTargetId="figure-18@13.[196,1435,194,1764]" captionTargetPageId="13" captionText="Figure 5-12. Characters of Coprophanaeus. 5) Ventral view of front left tibia of C. ensifer (arrow marks elongate tuft of setae). 6) Same, C. dardanus. 7) Left anterolateral pronotal margin of C. spitzi (arrow indicates unbroken circumnotal carina. 8) Same, C. magnoi (arrow indicates broken circumnotal carina). 9) Base of left hind wing of C. cerberus. 10) Same, C. horus. 11) Lateral view left occipital lobe of C. ensifer (asterisk marks swelling). 12) Same, C. spitzi (asterisk marks flat area)." figureDoi="http://doi.org/10.5281/zenodo.5352932" httpUri="https://zenodo.org/record/5352932/files/figure.png" pageId="12" pageNumber="9">Fig. 7</figureCitation>
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). Male with long, evenly tapering cylindrical head horn or shorter cylindrical horn with expanded base; female with trituberculate carina. Color often bright, metallic, occasionally dark. South America east of Andes and south of Amazonia to northern
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<collectingCountry id="CAC18AAA7F708871FE5DFA132274EDF2" box="[425,548,1443,1467]" name="Argentina" pageId="12" pageNumber="9">Argentina</collectingCountry>
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..................................................................
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<taxonomicName id="75D6B1B97F708871FBD0FA1225F0EDF2" authority="Olsoufieff" authorityName="Olsoufieff" box="[1060,1440,1442,1467]" class="Insecta" family="Scarabaeidae" genus="Metallophanaeus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="9" phylum="Arthropoda" rank="genus">
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<emphasis id="80A216287F708871FBD0FA1225F0EDF2" bold="true" box="[1060,1440,1442,1467]" pageId="12" pageNumber="9">
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<emphasis id="80A216287F708871FBD0FA122542EDF2" bold="true" box="[1060,1298,1442,1467]" italics="true" pageId="12" pageNumber="9">Metallophanaeus</emphasis>
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Olsoufieff
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</emphasis>
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</taxonomicName>
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</keyLead>
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</paragraph>
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<paragraph id="B269CA3A7F708871FF34FA7425F0EEF2" blockId="12.[192,1440,1283,1723]" pageId="12" pageNumber="9">
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<keyLead id="0927690F7F708871FF34FA7425F0EEF2" pageId="12" pageNumber="9">
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— Hind wing distinctly notched basally, vein V
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<subScript id="2E52C87F7F708871FCC9FA632316EDA8" attach="left" box="[829,838,1491,1505]" fontSize="6" pageId="12" pageNumber="9">2</subScript>
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usually hooked apically (
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<figureCitation id="2AEDD6BF7F708871FB8FFA732492ED92" box="[1147,1218,1475,1499]" captionStart="Figure 5-12" captionStartId="13.[195,279,1777,1799]" captionTargetBox="[196,1435,194,1764]" captionTargetId="figure-18@13.[196,1435,194,1764]" captionTargetPageId="13" captionText="Figure 5-12. Characters of Coprophanaeus. 5) Ventral view of front left tibia of C. ensifer (arrow marks elongate tuft of setae). 6) Same, C. dardanus. 7) Left anterolateral pronotal margin of C. spitzi (arrow indicates unbroken circumnotal carina. 8) Same, C. magnoi (arrow indicates broken circumnotal carina). 9) Base of left hind wing of C. cerberus. 10) Same, C. horus. 11) Lateral view left occipital lobe of C. ensifer (asterisk marks swelling). 12) Same, C. spitzi (asterisk marks flat area)." figureDoi="http://doi.org/10.5281/zenodo.5352932" httpUri="https://zenodo.org/record/5352932/files/figure.png" pageId="12" pageNumber="9">Fig. 9</figureCitation>
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). Lower portion of eye large, bulging, width more than twice that of adjacent oculogular space. Beading of anterior margin of circumnotal carina effaced behind eye (
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<figureCitation id="2AEDD6BF7F708871FC6BF9B323B5EE52" box="[927,997,1539,1563]" captionStart="Figure 5-12" captionStartId="13.[195,279,1777,1799]" captionTargetBox="[196,1435,194,1764]" captionTargetId="figure-18@13.[196,1435,194,1764]" captionTargetPageId="13" captionText="Figure 5-12. Characters of Coprophanaeus. 5) Ventral view of front left tibia of C. ensifer (arrow marks elongate tuft of setae). 6) Same, C. dardanus. 7) Left anterolateral pronotal margin of C. spitzi (arrow indicates unbroken circumnotal carina. 8) Same, C. magnoi (arrow indicates broken circumnotal carina). 9) Base of left hind wing of C. cerberus. 10) Same, C. horus. 11) Lateral view left occipital lobe of C. ensifer (asterisk marks swelling). 12) Same, C. spitzi (asterisk marks flat area)." figureDoi="http://doi.org/10.5281/zenodo.5352932" httpUri="https://zenodo.org/record/5352932/files/figure.png" pageId="12" pageNumber="9">Fig. 8</figureCitation>
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) (entire in the jasius group,
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<figureCitation id="2AEDD6BF7F708871FABDF9B325C2EE52" box="[1353,1426,1539,1563]" captionStart="Figure 5-12" captionStartId="13.[195,279,1777,1799]" captionTargetBox="[196,1435,194,1764]" captionTargetId="figure-18@13.[196,1435,194,1764]" captionTargetPageId="13" captionText="Figure 5-12. Characters of Coprophanaeus. 5) Ventral view of front left tibia of C. ensifer (arrow marks elongate tuft of setae). 6) Same, C. dardanus. 7) Left anterolateral pronotal margin of C. spitzi (arrow indicates unbroken circumnotal carina. 8) Same, C. magnoi (arrow indicates broken circumnotal carina). 9) Base of left hind wing of C. cerberus. 10) Same, C. horus. 11) Lateral view left occipital lobe of C. ensifer (asterisk marks swelling). 12) Same, C. spitzi (asterisk marks flat area)." figureDoi="http://doi.org/10.5281/zenodo.5352932" httpUri="https://zenodo.org/record/5352932/files/figure.png" pageId="12" pageNumber="9">Fig. 7</figureCitation>
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). Male head with horn or trituberculate carina; horn, if present, never a long, cylindrical, tapering protuberance. Female head with trituberculate carina. Color usually dark, somber; metallic reflections, if any, usually restricted to margins of pronotum; only seldom with metallic color conspicuous to unaided eye. Northern
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<collectingCountry id="CAC18AAA7F708871FCE4F93323DEEED2" box="[784,910,1667,1691]" name="Argentina" pageId="12" pageNumber="9">Argentina</collectingCountry>
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to northern
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<collectingCountry id="CAC18AAA7F708871FBDDF93324D0EED2" box="[1065,1152,1667,1691]" name="Mexico" pageId="12" pageNumber="9">Mexico</collectingCountry>
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(and extreme southern Texas) ...................................................................................................
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<emphasis id="80A216287F708871FB8EF91225F0EEF2" bold="true" box="[1146,1440,1698,1723]" italics="true" pageId="12" pageNumber="9">
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<taxonomicName id="75D6B1B97F708871FB8EF9122503EEF2" ID-CoL="3TRJ" authorityName="Olsoufieff" authorityYear="1924" box="[1146,1363,1698,1723]" class="Insecta" family="Scarabaeidae" genus="Coprophanaeus" kingdom="Animalia" order="Coleoptera" pageId="12" pageNumber="9" phylum="Arthropoda" rank="genus" sensu="stricto">Coprophanaeus</taxonomicName>
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<taxonomicNameLabel id="9B91AB537F708871FAACF91425F0EEF2" box="[1368,1440,1700,1723]" pageId="12" pageNumber="9" sensu="stricto">s. str.</taxonomicNameLabel>
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</emphasis>
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</keyLead>
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</paragraph>
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</keyStep>
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</subSubSection>
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</treatment>
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</document> |