173 lines
20 KiB
XML
173 lines
20 KiB
XML
<document id="D867D890A7CAF76353A9F6B9F40125AE" ID-DOI="10.5070/P9401557134" ID-ISSN="0031-0298" ID-Zenodo-Dep="10913374" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1712168371039" checkinUser="felipe" docAuthor="Powell, II, Charles L. & Dineen, Ashley A." docDate="2023" docId="295A87E1FFF2064884CCFDDE79AE0980" docLanguage="en" docName="PaleoBios.40.15.1-28.pdf" docOrigin="PaleoBios 40 (1904)" docSource="http://dx.doi.org/10.5070/p9401557134" docStyle="DocumentStyle:76F55B8832C4952BA7DD61B33E1F0E81.1:PaleoBios.2023-.journal_article" docStyleId="76F55B8832C4952BA7DD61B33E1F0E81" docStyleName="PaleoBios.2023-.journal_article" docStyleVersion="1" docTitle="Euspira AGASSIZ 1837, N. SP." docType="treatment" docVersion="2" lastPageNumber="3" masterDocId="D563FF99FFF1064C8724FFDA7C5E0E31" masterDocTitle="A new fossil Euspira? (Mollusca: Gastropoda: Naticidae) from the Gubik Formation on the North Slope of Arctic Alaska" masterLastPageNumber="28" masterPageNumber="1" pageNumber="2" updateTime="1712168691348" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-SA-4.0">
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<mods:title id="78B6A43D29E86F47F0592ED8EC43296A">A new fossil Euspira? (Mollusca: Gastropoda: Naticidae) from the Gubik Formation on the North Slope of Arctic Alaska</mods:title>
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<mods:namePart id="35479813D438A17CC3AA199583662A12">Powell, II, Charles L.</mods:namePart>
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<mods:namePart id="D0C3F3E305B47AE2E9D2CCF1573424B4">Dineen, Ashley A.</mods:namePart>
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<mods:title id="3951B6C1B1B490E6905F35235A816329">PaleoBios</mods:title>
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<mods:part id="44543E7890269A1300B3FE24520CC9F3">
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<mods:date id="419137605B35CA1DE5A1337E683ECFDB">2023</mods:date>
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<mods:number id="A5CDEB47418ACBF375CCACE8BD415E59">2023-11-05</mods:number>
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<mods:number id="3DBFB261291776E35729D54DD81E28B7">40</mods:number>
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<mods:number id="DB3371B030F7A363DB8B07C996A4874A">1904</mods:number>
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<mods:url id="CD69BF3256A71011C80A483237975D10">http://dx.doi.org/10.5070/p9401557134</mods:url>
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<mods:identifier id="BCB5708F7BC02771CBC387812AFE17C9" type="DOI">10.5070/P9401557134</mods:identifier>
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<treatment id="295A87E1FFF2064884CCFDDE79AE0980" LSID="urn:lsid:plazi:treatment:295A87E1FFF2064884CCFDDE79AE0980" httpUri="http://treatment.plazi.org/id/295A87E1FFF2064884CCFDDE79AE0980" lastPageId="4" lastPageNumber="3" pageId="3" pageNumber="2">
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<subSubSection id="E9E9657CFFF2064F84CCFDDE79630C70" pageId="3" pageNumber="2" type="nomenclature">
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<paragraph id="A14C36F7FFF2064F84CCFDDE79170C2F" blockId="3.[1000,1353,516,612]" box="[1000,1353,516,542]" pageId="3" pageNumber="2">
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<heading id="FA04819BFFF2064F84CCFDDE79170C2F" allCaps="true" box="[1000,1353,516,542]" centered="true" fontSize="10" level="2" pageId="3" pageNumber="2" reason="2">
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<emphasis id="9387EAE5FFF2064F84CCFDDE79170C2F" box="[1000,1353,516,542]" italics="true" pageId="3" pageNumber="2">
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<taxonomicName id="66F34D74FFF2064F84CCFDDE78130C2F" ID-CoL="7NZVH" authority="LOUIEMARINCOVICHI" authorityName="AGASSIZ" authorityYear="1837" box="[1000,1101,516,542]" class="Gastropoda" family="Naticidae" genus="Euspira" kingdom="Animalia" order="Littorinimorpha" pageId="3" pageNumber="2" phylum="Mollusca" rank="genus">EUSPIRA</taxonomicName>
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? LOUIEMARINCOVICHI
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="A14C36F7FFF2064F84D0FDFD79630C70" blockId="3.[1000,1353,516,612]" box="[1012,1341,551,577]" pageId="3" pageNumber="2">
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<heading id="FA04819BFFF2064F84D0FDFD79630C70" allCaps="true" box="[1012,1341,551,577]" centered="true" fontSize="10" level="2" pageId="3" pageNumber="2" reason="2">
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POWELL AND DINEEN,
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<taxonomicNameLabel id="88B4579EFFF2064F8224FDFD79630C70" box="[1280,1341,551,577]" pageId="3" pageNumber="2" rank="species">N. SP.</taxonomicNameLabel>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="E9E9657CFFF2064F836FFD9078B30CFA" pageId="3" pageNumber="2" type="description">
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<paragraph id="A14C36F7FFF2064F836FFD9078BB0C55" blockId="3.[1000,1353,516,612]" box="[1099,1253,586,612]" pageId="3" pageNumber="2">
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<figureCitation id="39C82A72FFF2064F836FFD9078BB0C55" box="[1099,1253,586,612]" captionStart="Figures 1–4" captionStartId="4.[110,201,1405,1429]" captionTargetBox="[113,1520,191,1390]" captionTargetId="figure-262@4.[112,1520,190,1390]" captionTargetPageId="4" captionText="Figures 1–4. Euspira? louiemarincovichi n. sp. UCMP holotype 123663, UCMP locality IP23235 (USGS M7346). Bar = 1 cm. 1. Aperical view, 2. Apical view, 3. Umbilical view, 4. Inclined side view. Photographs by Dave Strauss, UCMP." figureDoi="http://doi.org/10.5281/zenodo.10913376" httpUri="https://zenodo.org/record/10913376/files/figure.png" pageId="3" pageNumber="2">FIGURES 1–4</figureCitation>
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</paragraph>
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<paragraph id="A14C36F7FFF2064F8444FD5778B30CFA" blockId="3.[832,1521,653,1946]" pageId="3" pageNumber="2">
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<emphasis id="9387EAE5FFF2064F8444FD57785E0C96" bold="true" box="[864,1024,653,679]" pageId="3" pageNumber="2">Zoobank ID</emphasis>
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—
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<uri id="D5623AF5FFF2064F8303FD5778B30CFA" pageId="3" pageNumber="2">
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urn:lsid:zoobank.org:act:
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<uuid id="D5550C22FFF2064F827FFD5778B30CFA" pageId="3" pageNumber="2">2969452C- D99E-45B3-8851-D726958FB5C9</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="E9E9657CFFF2064F8444FD09781D0D66" pageId="3" pageNumber="2" type="diagnosis">
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<paragraph id="A14C36F7FFF2064F8444FD09781D0D66" blockId="3.[832,1521,653,1946]" pageId="3" pageNumber="2">
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<emphasis id="9387EAE5FFF2064F8444FD097FBD0CDC" bold="true" box="[864,995,723,749]" pageId="3" pageNumber="2">Diagnosis</emphasis>
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—The fine, closely spaced, narrow radial, and spiral sculpture forming a faint checkerboard pattern set this species apart from all northeastern Pacific and Arctic
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<taxonomicName id="66F34D74FFF2064F84E2FCE778610D66" authorityName="Guilding" authorityYear="1834" box="[966,1087,829,855]" class="Gastropoda" family="Naticidae" kingdom="Animalia" order="Littorinimorpha" pageId="3" pageNumber="2" phylum="Mollusca" rank="family">Naticidae</taxonomicName>
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.
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</paragraph>
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</subSubSection>
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<subSubSection id="E9E9657CFFF206488444FCBA7DA8088D" lastPageId="4" lastPageNumber="3" pageId="3" pageNumber="2" type="materials_examined">
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<paragraph id="A14C36F7FFF2064F8444FCBA7FEC0DAF" blockId="3.[832,1521,653,1946]" pageId="3" pageNumber="2">
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<materialsCitation id="119B3CAAFFF2064F8444FCBA7FEC0DAF" collectionCode="UCMP" pageId="3" pageNumber="2" specimenCode="UCMP 123663" specimenCount="1" typeStatus="holotype">
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<emphasis id="9387EAE5FFF2064F8444FCBA7FBB0D4B" bold="true" box="[864,997,864,890]" pageId="3" pageNumber="2">
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<typeStatus id="7E488855FFF2064F8444FCBA7FBB0D4B" box="[864,997,864,890]" pageId="3" pageNumber="2" type="holotype">Holotype</typeStatus>
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</emphasis>
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—
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<specimenCode id="F1559E8CFFF2064F8327FCBA78970D4B" box="[1027,1225,864,890]" collectionCode="UCMP" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="3" pageNumber="2" type="University or college">UCMP 123663</specimenCode>
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from
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<collectionCode id="C7E2AE32FFF2064F8207FCBA792D0D4B" box="[1315,1395,864,890]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="3" pageNumber="2" type="University or college">UCMP</collectionCode>
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locality- IP23235.
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</materialsCitation>
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</paragraph>
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<paragraph id="A14C36F7FFF2064F8444FC7D7FC90A36" blockId="3.[832,1521,653,1946]" pageId="3" pageNumber="2">
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<materialsCitation id="119B3CAAFFF2064F8444FC7D796D0DF1" box="[864,1331,934,961]" collectionCode="UCMP" pageId="3" pageNumber="2" specimenCode="UCMP 123666" specimenCount="1" typeStatus="paratype">
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<emphasis id="9387EAE5FFF2064F8444FC7D7FA80DF0" bold="true" box="[864,1014,935,961]" pageId="3" pageNumber="2">
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<typeStatus id="7E488855FFF2064F8444FC7D7FA80DF0" box="[864,1014,935,961]" pageId="3" pageNumber="2" type="paratype">Paratypes</typeStatus>
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</emphasis>
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—
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<specimenCode id="F1559E8CFFF2064F8332FC7D78BC0DF1" box="[1046,1250,934,961]" collectionCode="UCMP" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="3" pageNumber="2" type="University or college">UCMP 123666</specimenCode>
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from
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</materialsCitation>
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<collectionCode id="C7E2AE32FFF2064F8265FC7D79CD0DF0" box="[1345,1427,935,961]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="3" pageNumber="2" type="University or college">UCMP</collectionCode>
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locality IP23232 and
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<materialsCitation id="119B3CAAFFF2064F8337FC1079500DD5" box="[1043,1294,969,996]" collectionCode="UCMP" pageId="3" pageNumber="2" specimenCode="UCMP 123677" specimenCount="1" typeStatus="paratype">
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<specimenCode id="F1559E8CFFF2064F8337FC1078940DD2" box="[1043,1226,969,996]" collectionCode="UCMP" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="3" pageNumber="2" type="University or college">UCMP 123677</specimenCode>
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from
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</materialsCitation>
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<collectionCode id="C7E2AE32FFF2064F8232FC10793C0DD5" box="[1302,1378,970,996]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="3" pageNumber="2" type="University or college">UCMP</collectionCode>
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locality IP 23248.
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</paragraph>
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<paragraph id="A14C36F7FFF2064F8444FBCA789308B0" blockId="3.[832,1521,653,1946]" pageId="3" pageNumber="2">
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<emphasis id="9387EAE5FFF2064F8444FBCA78320A1B" bold="true" box="[864,1132,1040,1066]" pageId="3" pageNumber="2">Referred Collections</emphasis>
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—The number of specimens and partial specimens in each collection are documented as follows: complete or nearly complete specimens followed in square brackets by partial specimens or larger identifi- able fragments – i.e., 5[4] = 5 complete or nearly complete specimens (4 significantly broken or fragmented speci- mens).
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<collectionCode id="C7E2AE32FFF2064F84BFFB397FB90ACC" box="[923,999,1251,1277]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="3" pageNumber="2" type="University or college">UCMP</collectionCode>
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localities IP22958 (3[3]), IP23232 (5[1]), IP23233 (0[1]), IP23235 (3[0]), IP23238 (3[0]), IP23239 (3[0]), IP23248 (1[2]), IP23264 (4[0]), IP23266 (1[0]), IP23272 (2[0]), IP23279 (1[0]), IP23280 (7[0]), IP23316 (1[0]), IP23511 (3[3]), IP23555 (1[0]), IP23559 (12[3]), IP23561 (1[1]), IP23574 (7[0]), IP23576 (4[0]), IP23577 (8[0]), IP23588 (1[2]), IP23589 (6[2]), IP23590 (16[5]), IP23593 (9[1]), IP23596 (4[1]), IP23597 (1[0]), IP23602 (1[0]), IP23607 (1[0]), IP23721 (2[0]), IP23722 (5[0]), IP23724 (6[1]), IP23812 (1[0]), IP23856 (4[3]), IP23857 (3[0]), IP23868 (1[2]), IP23870 (6[1]), IP23879 (1[0]), IP23884 (5[0]), IP24104 (7[0]).
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</paragraph>
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<paragraph id="A14C36F7FFF2064F8444F95079950965" blockId="3.[832,1521,653,1946]" pageId="3" pageNumber="2">
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Additional collections observed:
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<collectionCode id="C7E2AE32FFF2064F8201F95079340895" box="[1317,1386,1674,1700]" country="USA" httpUri="http://grbio.org/cool/3n47-ewde" name="U.S. Geological Survey" pageId="3" pageNumber="2">USGS</collectionCode>
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localities M8151 and M8997. These collections could not be locat- ed at
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<collectionCode id="C7E2AE32FFF2064F84A7F90A7F9008DB" box="[899,974,1744,1770]" country="USA" httpUri="http://grbio.org/cool/2fxn-eays" name="University of California Museum of Paleontology" pageId="3" pageNumber="2" type="University or college">UCMP</collectionCode>
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, but were identified for internal
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<collectionCode id="C7E2AE32FFF2064F827AF90A79FE08DB" box="[1374,1440,1744,1770]" country="USA" httpUri="http://grbio.org/cool/3n47-ewde" name="U.S. Geological Survey" pageId="3" pageNumber="2">USGS</collectionCode>
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E&R’s (Evaluation and Report documents). The information is added here for completeness as either Louie Marincovich or the senior author (
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<collectionCode id="C7E2AE32FFF2064F836EF8E078DA0965" box="[1098,1156,1850,1876]" pageId="3" pageNumber="2">CPII</collectionCode>
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) identified the specimens.
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</paragraph>
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<paragraph id="A14C36F7FFF2064F8444F8877FAA09AB" blockId="3.[832,1521,653,1946]" pageId="3" pageNumber="2">Locality information for all collections can be found in Appendix 1.</paragraph>
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<caption id="F58C667FFFF50648874AFAA779480B84" ID-DOI="http://doi.org/10.5281/zenodo.10913376" ID-Zenodo-Dep="10913376" httpUri="https://zenodo.org/record/10913376/files/figure.png" pageId="4" pageNumber="3" startId="4.[110,201,1405,1429]" targetBox="[113,1520,191,1390]" targetPageId="4" targetType="figure">
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<paragraph id="A14C36F7FFF50648874AFAA779480B84" blockId="4.[110,1517,1405,1461]" pageId="4" pageNumber="3">
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<emphasis id="9387EAE5FFF50648874AFAA77CA20BA4" bold="true" box="[110,252,1405,1429]" pageId="4" pageNumber="3">Figures 1–4</emphasis>
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.
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<emphasis id="9387EAE5FFF50648862CFAA47E6B0BA4" box="[264,565,1405,1429]" italics="true" pageId="4" pageNumber="3">
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<taxonomicName id="66F34D74FFF50648862CFAA47D000BA4" authorityName="AGASSIZ" authorityYear="1837" box="[264,350,1406,1429]" class="Gastropoda" family="Naticidae" genus="Euspira" kingdom="Animalia" order="Littorinimorpha" pageId="4" pageNumber="3" phylum="Mollusca" rank="genus">Euspira</taxonomicName>
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? louiemarincovichi
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</emphasis>
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n. sp. UCMP holotype 123663, UCMP locality IP23235 (USGS M7346). Bar = 1 cm. 1. Aperical view, 2. Apical view, 3. Umbilical view, 4. Inclined side view. Photographs by Dave Strauss, UCMP.
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</paragraph>
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</caption>
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<paragraph id="A14C36F7FFF5064887B4FA177DA8088D" blockId="4.[112,800,1485,1970]" pageId="4" pageNumber="3">
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<emphasis id="9387EAE5FFF5064887B4FA177D760BD6" bold="true" box="[144,296,1485,1511]" pageId="4" pageNumber="3">Occurrence</emphasis>
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— Known only from the Gubik Formation along the northwestern part of the Arctic Coastal Plain (North Slope) of
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<collectingRegion id="6337F815FFF50648861BF9CE7DCC081F" box="[319,402,1556,1582]" country="United States of America" name="Alaska" pageId="4" pageNumber="3">Alaska</collectingRegion>
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from Skull Cliff south of Barrow in the west to along the Colville River and Kogru River/ Teshekpuk Lake areas in the east, North Slope Borough,
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<collectingRegion id="6337F815FFF506488754F9A77C9D08A6" box="[112,195,1661,1687]" country="United States of America" name="Alaska" pageId="4" pageNumber="3">Alaska</collectingRegion>
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(
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<figureCitation id="39C82A72FFF5064887F7F9A77D6008A6" box="[211,318,1661,1687]" captionStart="Figure 6" captionStartId="7.[112,191,1229,1253]" captionTargetBox="[112,1519,192,1215]" captionTargetId="figure-292@7.[112,1520,192,1215]" captionTargetPageId="7" captionText="Figure 6. Map showing generalized occurrences of Euspira? louiemarincovichi along the northern coastal plain of Alaska. These occurrences are concentrated in three areas: adjacent to the Colville River (southern eastern occurrences), near and along the Kogru River/Teshekpuk Lake (northern eastern occurrences), and along Skull Cliff (occurrences southwest of Barrow to the west). Not all collections are represented in this figure, just enough to determine the general geographic areas where this new species occurs. Some addition geographic names are mentioned to help interpret the geographic ranges of some gastropods." figureDoi="http://doi.org/10.5281/zenodo.10913380" httpUri="https://zenodo.org/record/10913380/files/figure.png" pageId="4" pageNumber="3">Figure 6</figureCitation>
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and Appendix 1). Early Pleistocene in age (see discussion for details).
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</paragraph>
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</subSubSection>
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<subSubSection id="E9E9657CFFF5064887B4F91E7D49095A" pageId="4" pageNumber="3" type="etymology">
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<paragraph id="A14C36F7FFF5064887B4F91E7D49095A" blockId="4.[112,800,1485,1970]" pageId="4" pageNumber="3">
|
||
<emphasis id="9387EAE5FFF5064887B4F91E7D4008EF" bold="true" box="[144,286,1732,1758]" pageId="4" pageNumber="3">Etymology</emphasis>
|
||
— This new species is named for Louie Marincovich, Jr. who first introduced this species to the senior author and for his work on unraveling many mysteries of Paleogene–Neogene Arctic geology and paleontology.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="E9E9657CFFF5064887B4F8AE79AE0980" pageId="4" pageNumber="3" type="description">
|
||
<paragraph id="A14C36F7FFF5064887B4F8AE79AE0980" blockId="4.[112,800,1485,1970]" lastBlockId="4.[832,1520,1485,1970]" pageId="4" pageNumber="3">
|
||
<emphasis id="9387EAE5FFF5064887B4F8AE7D0609BF" bold="true" box="[144,344,1908,1934]" pageId="4" pageNumber="3">Description —</emphasis>
|
||
This new species has a naticiform shell about as high as wide and with a moderate thick, substantial shell. The spire is very short and consists of about four whorls and is less than a tenth the height of the entire shell. The protoconch is worn on all specimens and not available for description.With a generally smooth shell profile the whorls show a slight indentation or suture and a very fine, channel at the top of its whorls. The exterior of the shell shows very fine, closely spaced, irregular, spiral threads crossed by very fine, closely spaced, mostly continuous, radial threads forming a microscopic crosshatch pattern covering the exterior of the shell (
|
||
<figureCitation id="39C82A72FFF5064884A8F8F77FA40979" box="[908,1018,1837,1864]" captionStart="Figure 5" captionStartId="5.[110,189,674,698]" captionTargetBox="[112,800,191,661]" captionTargetId="figure-823@5.[112,800,191,661]" captionTargetPageId="5" captionText="Figure 5. Close–up of spiral and radial sculpture on the body whorl of the holotype specimen. Field of view approximately 0.5 cm. Blue arrow indicates radial sculpture, red arrow indicates spiral sculpture. angle of the aperture is acute and points towards the apex. A deep, partially open umbilicus is present, but is obscured in part by the umbilical callus making it tear– drop shaped." figureDoi="http://doi.org/10.5281/zenodo.10913378" httpUri="https://zenodo.org/record/10913378/files/figure.png" pageId="4" pageNumber="3">Figure 5</figureCitation>
|
||
). The aperture is large, near–tear–drop shape and is about three–quarters of the shell's height with a parental callus that mirrors the inner margin of the aperture a few millimeters wide. The posterior apertural
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |