327 lines
33 KiB
XML
327 lines
33 KiB
XML
<document ID-DOI="10.1590/S0073-47212003000400008" ID-GBIF-Dataset="32bd22ce-7d3f-4eac-8203-e9f61a19c7f0" ID-ISSN="1678-4766" ID-Zenodo-Dep="3719467" approvedBy="tatiana" checkinTime="1584724827204" checkinUser="tatiana" docAuthor="Barcellos, Aline & Grazia, Jocélia" docDate="2003" docId="217887A7991F80224BF85C0BFE906036" docLanguage="en" docName="Iheringia.93.4.413-446.pdf.imf" docOrigin="Iheringia, Sér. Zool. 93 (4)" docStyle="DocumentStyle{}" docTitle="Brachystethus vexillum Breddin 1903" docType="treatment" docVersion="8" lastPageNumber="432" masterDocId="DD41FFDF990D80314A705E42FF826415" masterDocTitle="Revision Of Brachystethus (Heteroptera, Pentatomidae, Edessinae)" masterLastPageNumber="446" masterPageNumber="413" pageNumber="431" updateTime="1668426789504" updateUser="valdenar" zenodo-license-document="CC-BY-4.0">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Revision Of Brachystethus (Heteroptera, Pentatomidae, Edessinae)</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Barcellos, Aline</mods:namePart>
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<mods:affiliation>Museu de Ciências Naturais, Fundação Zoobotânica do Rio Grande do Sul, Caixa Postal 1188, 90001 - 970, Porto Alegre, RS, Brazil.</mods:affiliation>
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<mods:nameIdentifier type="email">alinebar@ig.com.br</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Grazia, Jocélia</mods:namePart>
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<mods:affiliation>Departamento de Zoologia, Universidade Federal do Rio Grande do Sul, Av. Bento Gonçalves, 9500, Prédio 43435.2, 91501 - 970, Porto Alegre, RS, Brazil.</mods:affiliation>
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<mods:nameIdentifier type="email">jocelia@ufrgs.br</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>Iheringia, Sér. Zool.</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2003</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2003-12-30</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>93</mods:number>
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</mods:detail>
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<mods:detail type="issue">
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<mods:number>4</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>413</mods:start>
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<mods:end>446</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">10.1590/S0073-47212003000400008</mods:identifier>
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<mods:identifier type="GBIF-Dataset">32bd22ce-7d3f-4eac-8203-e9f61a19c7f0</mods:identifier>
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<mods:identifier type="ISSN">1678-4766</mods:identifier>
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<mods:identifier type="Zenodo-Dep">3719467</mods:identifier>
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</mods:mods>
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<treatment ID-DOI="http://doi.org/10.5281/zenodo.3719837" ID-GBIF-Taxon="162653796" ID-Zenodo-Dep="3719837" LSID="urn:lsid:plazi:treatment:217887A7991F80224BF85C0BFE906036" httpUri="http://treatment.plazi.org/id/217887A7991F80224BF85C0BFE906036" lastPageId="19" lastPageNumber="432" pageId="18" pageNumber="431">
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<subSubSection box="[392,816,585,609]" pageId="18" pageNumber="431" type="nomenclature">
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<paragraph blockId="18.[392,816,585,633]" box="[392,816,585,609]" pageId="18" pageNumber="431">
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<heading bold="true" box="[392,816,585,609]" centered="true" fontSize="10" level="1" pageId="18" pageNumber="431" reason="0">
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<taxonomicName authority="Breddin, 1903" authorityName="Breddin" authorityYear="1903" box="[392,816,585,609]" class="Insecta" family="Pentatomidae" genus="Brachystethus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="431" phylum="Arthropoda" rank="species" species="vexillum">
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<emphasis bold="true" box="[392,816,585,609]" pageId="18" pageNumber="431">
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<emphasis bold="true" box="[392,549,585,609]" italics="true" pageId="18" pageNumber="431">Brachystethus</emphasis>
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<emphasis bold="true" box="[555,650,585,609]" italics="true" pageId="18" pageNumber="431">vexillum</emphasis>
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<bibRefCitation author="BREDDIN, G." box="[656,816,585,609]" journalOrPublisher="Sber. Ges. naturf. Freunde Berl., Berlin" pageId="18" pageNumber="431" pagination="366 - 383" part="8" refId="ref20059" refString="BREDDIN, G. 1903. Beitrage zur Hemiptern-Fauna der Anden. Sber. Ges. naturf. Freunde Berl., Berlin, 8: 366 - 383." title="Beitrage zur Hemiptern-Fauna der Anden" type="journal article" year="1903">Breddin, 1903</bibRefCitation>
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</emphasis>
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</taxonomicName>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection box="[454,755,614,633]" pageId="18" pageNumber="431" type="description">
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<paragraph blockId="18.[392,816,585,633]" box="[454,755,614,633]" pageId="18" pageNumber="431">
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<heading box="[454,755,614,633]" pageId="18" pageNumber="431">
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(
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<figureCitation box="[461,528,614,633]" captionStart="Figs" captionStartId="5.[101,144,1441,1460]" captionTargetBox="[161,975,188,1422]" captionTargetId="figure@5.[161,975,188,1422]" captionText="Figs. 2-13. 2, 3, Brachystethus rubromaculatus Dallas, 1851; 4, 5, B. signoreti Stål; 1872; 6, B. cribrus (Fabricius, 1781); 7, B. schuhi sp. nov.; 8, B. tricolor Bolívar, 1879; 9, B. vexillum Breddin, 1903; 10, B. geniculatus (Fabricius, 1787); 11, B. coxalis Breddin, 1904; 12, B. improvisus Breddin, 1905; 13, B. vicinus Signoret, 1851. Magnification 1.63X." figureDoi="http://doi.org/10.5281/zenodo.3719471" httpUri="https://zenodo.org/record/3719471/files/figure.png" pageId="18" pageNumber="431">Figs. 9</figureCitation>
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,
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<figureCitation box="[540,564,614,633]" captionStart="Figs" captionStartId="7.[101,143,1361,1380]" captionText="Figs. 14-19. Pygophore, dorsal: 14, Brachystethus rubromaculatus; 15, B. signoreti; 16, B. cribrus; 17, B. schuhi; 18, B. tricolor; 19, B. vexillum (dor, dorsal rim; gcp, genital cup process; ldor, lobe of dorsal rim; pa, paramere; vr, ventral rim; X, tenth segment). Scale, 1 mm" pageId="18" pageNumber="431">19</figureCitation>
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,
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<figureCitation box="[577,602,614,633]" captionStart="Figs" captionStartId="12.[139,181,1149,1168]" captionTargetBox="[230,975,180,1114]" captionTargetId="figure@12.[222,987,179,1123]" captionTargetPageId="12" captionText="Figs. 24-29. Pygophore, caudal (tenth segment omitted): 24, Brachystethus rubromaculatus; 25, B. signoreti; 26, B. cribrus; 27, B. schuhi; 28, B. tricolor; 29, B. vexillum (dor, dorsal rim; gcp, genital cup process; pa, paramere; vr, ventral rim). Scale, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3719473" httpUri="https://zenodo.org/record/3719473/files/figure.png" pageId="18" pageNumber="431">29</figureCitation>
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,
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<figureCitation box="[614,639,614,633]" captionStart="Figs" captionStartId="17.[101,144,1245,1264]" captionTargetBox="[157,966,181,1208]" captionTargetId="figure@17.[152,974,177,1222]" captionTargetPageId="17" captionText="Figs. 34-43. Pygophore, ventral rim, ventral: 34, Brachystethus rubromaculatus; 35, B. signoreti; 36, B. cribrus; 37, B. schuhi; 38, B. tricolor; 39, B. vexillum; 40, B. geniculatus; 41, B. coxalis; 42, B. improvisus; 43, B. vicinus. Scale, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3719477" httpUri="https://zenodo.org/record/3719477/files/figure.png" pageId="18" pageNumber="431">39</figureCitation>
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,
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<figureCitation box="[651,676,614,633]" captionStart="Figs" captionStartId="20.[139,181,1131,1150]" captionTargetBox="[281,927,182,1105]" captionTargetId="figure@20.[275,935,178,1113]" captionTargetPageId="20" captionText="Figs. 44-52. Left paramere, posteroventral: 44, Brachystethus rubromaculatus; 45, B. signoreti; 46, B. cribrus; 47, B. schuhi; 48, B. tricolor; 49, B. vexillum; 50, B. geniculatus; 51, B. improvisus; 52, B. vicinus. Scale, 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3719479" httpUri="https://zenodo.org/record/3719479/files/figure.png" pageId="18" pageNumber="431">49</figureCitation>
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,
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<figureCitation box="[688,713,614,633]" captionStart="Figs" captionStartId="24.[139,184,1161,1180]" captionTargetBox="[176,1022,181,1137]" captionTargetId="figure@24.[168,1032,172,1151]" captionText="Figs.56-60. Genital plates, ventral: 56, Brachystethus rubromaculatus; 57, B. signoreti; 58, B. cribrus; 59, B. tricolor; 60, B. vexillum (gc8, gonocoxite 8; gc9, gonocoxites 9; la8, laterotergite 8; la9, laterotergite 9; VII, seventh segment; X, tenth segment). Scale, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3719483" httpUri="https://zenodo.org/record/3719483/files/figure.png" pageId="18" pageNumber="431">60</figureCitation>
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,
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<figureCitation box="[724,748,614,633]" captionStart="Figs" captionStartId="30.[139,183,1392,1411]" captionTargetBox="[159,1043,188,1362]" captionTargetId="figure@30.[154,1051,188,1372]" captionTargetPageId="30" captionText="Figs. 69-72. Laterotergites, gonocoxites and gonapophyses of ninth segment, and ectodermical genital ducts, ventral: 69, Brachystethus vexillum; 70, B. geniculatus; 71, B. improvisus; 72, B. vicinus (aac, anterior annular crest; ch, chitinellipsen; cs, capsula seminalis; dr, ductus receptaculi; g9, gonapophyses 9; gc9, gonocoxites 9; la9, laterotergites 9; mt, medial thickening of gonapophyses 9; or, orificium receptaculi; pac, posterior annular crest; pco, pars comunis; pi, pars intermedialis; tvi, thickening of vaginal intima; X, tenth segment). Scale, 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3719489" httpUri="https://zenodo.org/record/3719489/files/figure.png" pageId="18" pageNumber="431">69</figureCitation>
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)
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="18" pageNumber="431" type="reference_group">
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<paragraph blockId="18.[139,1070,662,706]" box="[139,1070,662,682]" pageId="18" pageNumber="431">
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<treatmentCitationGroup box="[139,1070,662,682]" pageId="18" pageNumber="431">
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<treatmentCitation author="BREDDIN" box="[139,517,662,682]" page="371" pageId="18" pageNumber="431" year="1903">
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<taxonomicName authority="BREDDIN, 1903: 371" authorityName="Breddin" authorityYear="1903" box="[139,517,662,682]" class="Insecta" family="Pentatomidae" genus="Brachystethus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="431" phylum="Arthropoda" rank="species" species="vexillum">
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<emphasis box="[139,341,663,682]" italics="true" pageId="18" pageNumber="431">Brachystethus vexillum</emphasis>
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<bibRefCitation author="BREDDIN, G." box="[349,517,662,681]" journalOrPublisher="Sber. Ges. naturf. Freunde Berl., Berlin" pageId="18" pageNumber="431" pagination="366 - 383" part="8" refId="ref20059" refString="BREDDIN, G. 1903. Beitrage zur Hemiptern-Fauna der Anden. Sber. Ges. naturf. Freunde Berl., Berlin, 8: 366 - 383." title="Beitrage zur Hemiptern-Fauna der Anden" type="journal article" year="1903">BREDDIN, 1903:371</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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;
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<treatmentCitation author="BERGROTH" box="[528,710,662,681]" page="178" pageId="18" pageNumber="431" year="1908">
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<bibRefCitation author="BERGROTH, E." box="[528,710,662,681]" journalOrPublisher="Mem. Soc. r. ent. Belg., Bruxelles" pageId="18" pageNumber="431" pagination="131 - 200" part="15" refId="ref19790" refString="BERGROTH, E. 1908. Enumeratio Pentatomidarum post catalogum Bruxellensem descriptarum. Mem. Soc. r. ent. Belg., Bruxelles, 15: 131 - 200." title="Enumeratio Pentatomidarum post catalogum Bruxellensem descriptarum" type="journal article" year="1908">BERGROTH, 1908:178</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation author="KIRKALDY, G. W." box="[721,909,662,681]" page="152" pageId="18" pageNumber="431" year="1909">
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<bibRefCitation author="KIRKALDY, G. W." bookContentInfo="392 p." box="[721,909,662,681]" journalOrPublisher="Berlin, Felix L. Dames" pageId="18" pageNumber="431" refId="ref20822" refString="KIRKALDY, G. W. 1909. Catalogue of the Hemiptera (Heteroptera). 1. Cimicidae. Berlin, Felix L. Dames. 392 p." title="Catalogue of the Hemiptera (Heteroptera). 1. Cimicidae" type="book" year="1909">KIRKALDY, 1909: 152</bibRefCitation>
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</treatmentCitation>
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;
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<treatmentCitation author="PIRÁN" box="[920,1066,662,681]" page="337" pageId="18" pageNumber="431" year="1955">
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<bibRefCitation author="PIRAN, A. P." box="[920,1066,662,681]" journalOrPublisher="Beitr. Ent., Berlin" pageId="18" pageNumber="431" pagination="337 - 338" part="5" refId="ref21103" refString="PIRAN, A. P. 1955. Dos notas de sinonimia sobre Hemiptera neotropicales. Beitr. Ent., Berlin, 5 (3 / 4): 337 - 338." title="Dos notas de sinonimia sobre Hemiptera neotropicales" type="journal article" year="1955">PIRÁN, 1955:337</bibRefCitation>
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</treatmentCitation>
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.
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</treatmentCitationGroup>
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</paragraph>
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<paragraph blockId="18.[139,1070,662,706]" box="[139,792,686,706]" pageId="18" pageNumber="431">
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<treatmentCitationGroup box="[139,792,686,706]" pageId="18" pageNumber="431">
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<treatmentCitation author="KORMILEV" box="[139,557,686,706]" page="38" pageId="18" pageNumber="431" year="1951">
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<taxonomicName authority="KORMILEV, 1951: 38" authorityName="Kormilev" authorityYear="1951" box="[139,557,686,706]" class="Insecta" family="Pentatomidae" genus="Daimonocoris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="414" phylum="Arthropoda" rank="species" species="bolivianus">
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<emphasis box="[139,368,687,706]" italics="true" pageId="18" pageNumber="431">Daimonocoris bolivianus</emphasis>
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<bibRefCitation author="KORMILEV, N. A." box="[380,557,686,705]" journalOrPublisher="Folia Univ. Cochabamba, Cochabamba" pageId="18" pageNumber="431" pagination="35 - 45" part="5" refId="ref20852" refString="KORMILEV, N. A. 1951. Notas sobre hemipteros bolivianos con la descripcion de un genero y tres especies nuevos. Folia Univ. Cochabamba, Cochabamba, 5: 35 - 45." title="Notas sobre hemipteros bolivianos con la descripcion de un genero y tres especies nuevos" type="journal article" year="1951">KORMILEV, 1951:38</bibRefCitation>
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</taxonomicName>
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</treatmentCitation>
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;
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<treatmentCitation author="PIRÁN" box="[572,723,686,705]" page="337" pageId="18" pageNumber="431" year="1955">
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<bibRefCitation author="PIRAN, A. P." box="[572,723,686,705]" journalOrPublisher="Beitr. Ent., Berlin" pageId="18" pageNumber="431" pagination="337 - 338" part="5" refId="ref21103" refString="PIRAN, A. P. 1955. Dos notas de sinonimia sobre Hemiptera neotropicales. Beitr. Ent., Berlin, 5 (3 / 4): 337 - 338." title="Dos notas de sinonimia sobre Hemiptera neotropicales" type="journal article" year="1955">PIRÁN, 1955:337</bibRefCitation>
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</treatmentCitation>
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(syn.).
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</treatmentCitationGroup>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="18" pageNumber="431" type="materials_examined">
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<paragraph blockId="18.[139,1070,710,1525]" pageId="18" pageNumber="431">
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Type material of
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<taxonomicName authorityName="Breddin" authorityYear="1903" box="[402,530,711,734]" class="Insecta" family="Pentatomidae" genus="Brachystethus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="431" phylum="Arthropoda" rank="species" species="vexillum">
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<emphasis box="[402,530,711,734]" italics="true" pageId="18" pageNumber="431">B. vexillum</emphasis>
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</taxonomicName>
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.
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<materialsCitation ID-GBIF-Occurrence="2579424832" collectionCode="DEIC" collectorName="Breddin" country="Bolivia" location="Brachystethus vexillum Bredd" municipality="Brachystethus vexillum Bredd" pageId="18" pageNumber="431" specimenCount="2" specimenCount-male="1" typeStatus="holotype">
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<typeStatus box="[548,651,710,733]" pageId="18" pageNumber="431">Holotype</typeStatus>
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, examined, labeled: a)
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<collectingCountry box="[947,1032,710,733]" name="Bolivia" pageId="18" pageNumber="431" value="Bolivia">Bolivia</collectingCountry>
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; b)
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<taxonomicName authority="Bredd" authorityName="Breddin" authorityYear="1903" box="[139,457,740,763]" class="Insecta" family="Pentatomidae" genus="Brachystethus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="431" phylum="Arthropoda" rank="species" species="vexillum">
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<collectingMunicipality box="[139,457,740,763]" pageId="18" pageNumber="431">
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<emphasis box="[139,384,740,763]" italics="true" pageId="18" pageNumber="431">Brachystethus vexillum</emphasis>
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Bredd
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</collectingMunicipality>
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</taxonomicName>
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; c) coll.
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<collectorName box="[543,632,740,763]" pageId="18" pageNumber="431">Breddin</collectorName>
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; d)
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<typeStatus box="[668,782,740,763]" pageId="18" pageNumber="431">Holotypus</typeStatus>
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; e) Dtsch. Entomol. Institut Berlin; f) HT
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<taxonomicName authorityName="Breddin" authorityYear="1903" box="[280,369,769,792]" class="Insecta" family="Pentatomidae" genus="Brachystethus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="18" pageNumber="431" phylum="Arthropoda" rank="species" species="vexillum">
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<emphasis box="[280,369,769,792]" italics="true" pageId="18" pageNumber="431">vexillum</emphasis>
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</taxonomicName>
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(
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<collectionCode box="[381,448,769,792]" collectionName="Germany, Muencheberg, Deutsches Entomologisches Institut im ZALF" pageId="18" pageNumber="431">DEIC</collectionCode>
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).
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</materialsCitation>
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Typelocality: Bolivia.
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</paragraph>
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</subSubSection>
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<subSubSection lastPageId="19" lastPageNumber="432" pageId="18" pageNumber="431" type="description">
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<paragraph blockId="18.[139,1070,710,1525]" pageId="18" pageNumber="431">
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Description. Dorsal surface black, with yellowish stripes on corium (
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<figureCitation box="[934,993,798,821]" captionStart="Figs" captionStartId="5.[101,144,1441,1460]" captionTargetBox="[161,975,188,1422]" captionTargetId="figure@5.[161,975,188,1422]" captionTargetPageId="5" captionText="Figs. 2-13. 2, 3, Brachystethus rubromaculatus Dallas, 1851; 4, 5, B. signoreti Stål; 1872; 6, B. cribrus (Fabricius, 1781); 7, B. schuhi sp. nov.; 8, B. tricolor Bolívar, 1879; 9, B. vexillum Breddin, 1903; 10, B. geniculatus (Fabricius, 1787); 11, B. coxalis Breddin, 1904; 12, B. improvisus Breddin, 1905; 13, B. vicinus Signoret, 1851. Magnification 1.63X." figureDoi="http://doi.org/10.5281/zenodo.3719471" httpUri="https://zenodo.org/record/3719471/files/figure.png" pageId="18" pageNumber="431">fig. 9</figureCitation>
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). Head black, with well-marked wrinkles. Punctures moderately distributed, restrict to base of jugae and tylus. Antennae black; fourth antennal segment cylindrical. First antennal segment about 3/4 longer than second, third slightly longer than second, fourth almost twice longer than third and about 3/4 the length of fifth, the longest. Pronotum black, with fine punctures regularly distributed and coarse punctures sparse behind cicatrices. Before cicatrices, medium-sized punctures densely distributed. Anterolateral angles yellowish at apex. Scutellum black, with coarse punctures sparse, more frequent beside lateral margins of basal half. Corium black, without callus, with two yellow stripes: one on clavus, adjacent to claval suture and another occupying 2/3 ectad to radial vein, becoming broader apically toward clavus. Legsdark reddish-castaneous to black. Ventral surface of thorax dark reddish-castaneous to black; medium-sized punctures, concolor. Ostiolar rugae short, reaching 1/3 of distance between inner margin of ostiolum and lateral margin of metapleura. Metasternal carina with conspicuous transversal sulcus; in profile, little elevatedanteriorly. Connexivum black, inner superior angle of each segment orange, segments finely punctured. In some specimens, the orange area extends along inner lateral margin of seventhsegment. Venter red, with black spots on lateral thirds, as follows: one near lateral margin of each segment, becoming broader posterad, other small, around spiracles. On sternites III to VI, also present, entad to others, a larger spot, sub-elliptical, close to spiracle. Coarse punctures more concentrated on lateral thirds, over these spots. Spiracles black.
|
|
</paragraph>
|
|
<paragraph blockId="18.[139,1070,710,1525]" pageId="18" pageNumber="431">
|
|
Male genitalia. Pygophore globose, apex and base equally wide. Lobes of dorsal rim conspicuous, strongly projected posterad, rounded at apex. Genital cup processes littlevisible dorsally (
|
|
<figureCitation box="[362,481,1473,1496]" pageId="18" pageNumber="431">figs. 19, 29</figureCitation>
|
|
). Parameres withacute apex, anteriorly directed (
|
|
<figureCitation box="[986,1057,1473,1496]" pageId="18" pageNumber="431">fig. 49</figureCitation>
|
|
). Ventralrimopenly V-like (
|
|
<figureCitation box="[412,481,1502,1525]" pageId="18" pageNumber="431">fig. 39</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph blockId="19.[101,1032,190,1059]" pageId="19" pageNumber="432">Measurements (n=3). Headlength 1.73 (1.68-1.76) ± 0.05, width 2.56 (2.52-2,60) ± 0.04; interoculardistance 1.49 (1.44-1.52) ± 0.05; anteocularlength 0.64; lengthof antennal segments: I - 0.85 (0.80-0.88) ± 0.05; II - 1.17 (1.12-1.20) ± 0.05; III - 1.44 (1.28-1.52) ± 0.14; IV - 2.59 (2.48-2.72) ± 0.12; V - 3.68; pronotallength 3.70 (3.59-3.84) ± 0.13, width 8.63 (8.52-8.85) ± 0.19; scutellarlength 5.87 (5.85-5.93) ± 0.05, width 5.29 (5.18-5.43) ± 0.13; coriumlength 8.24 (8.02-8.52) ± 0.26; abdominalwidth 10.08 (9.69-10.35) ± 0.35; total length 13.64 (13.03-14.28) ± 0.63.</paragraph>
|
|
<paragraph blockId="19.[101,1032,190,1059]" pageId="19" pageNumber="432">
|
|
Female genitalia. Sutural borders of gonocoxites 8 parallel, except on apical fifth. Posterior borders convex on outer half and sub-retilineous on inner half (fig. 60). Gonocoxites 9 almost three times wider across apex of arms than longmedially; anterior border openly U-like, moderately excavate at middle. Length of arms of gonocoxites 9 equal to 1/6 of plate width. Thickening of vaginal intima digitiform, elongated.
|
|
<emphasis italics="true" pageId="19" pageNumber="432">Ductus receptaculi</emphasis>
|
|
thin, one and a half longer before than after vesicular area.
|
|
<emphasis box="[837,1031,542,565]" italics="true" pageId="19" pageNumber="432">Capsula seminalis</emphasis>
|
|
digitiform. Annular crests sub-parallel, turned toward vesicular area (
|
|
<figureCitation box="[834,907,571,594]" pageId="19" pageNumber="432">fig. 69</figureCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph blockId="19.[101,1032,190,1059]" pageId="19" pageNumber="432">Measurements (n=9). Headlength 1.93 (1.80-2.08) ± 0.08, width 2.77 (2.72-2.84) ± 0.05; interoculardistance 1.60 (1.48-1.68) ± 0.06; anteocularlength 0.69 (0.64-0.72) ± 0.04; lengthof antennalsegments: I - 0.91 (0.84-0.96) ± 0.04; II - 1.24 (0.96-1.36) ± 0.13; III - 1.56 (1.36-1.88) ± 0.16; IV - 2.75 (2.60-2.96) ± 0.13; V - 3.58 (3.44-3.68) ± 0.09; pronotallength 4.12 (3.84-4.34) ± 0.15, width 9.65 (9.27-10.02) ± 0.25; scutellarlength 6.74 (6.51-7.01) ± 0.17, width 5.97 (5.68-6.43) ± 0.21; coriumlength 9.32 (8.93-9.85) ± 0.27; abdominalwidth 11.44 (11.19-11.52) ± 0.13; totallength 15.56 (14.95-16.70) ± 0.56.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="19" pageNumber="432" type="materials_examined">
|
|
<paragraph blockId="19.[101,1032,190,1059]" pageId="19" pageNumber="432">
|
|
Material examined.
|
|
<materialsCitation ID-GBIF-Occurrence="2579424854" collectingDate="1982-11-28" collectionCode="DARC" collectorName="E. R. Ross" country="Peru" location="Machu Picchu" municipality="Machu Picchu" pageId="19" pageNumber="432" specimenCount="1" specimenCount-female="1" stateProvince="Cuzco">
|
|
<collectingCountry box="[353,413,807,826]" name="Peru" pageId="19" pageNumber="432" value="Peru">PERU</collectingCountry>
|
|
,
|
|
<emphasis bold="true" box="[424,483,807,826]" pageId="19" pageNumber="432">
|
|
<collectingRegion box="[424,483,807,826]" country="Peru" name="Cusco" pageId="19" pageNumber="432">Cuzco</collectingRegion>
|
|
</emphasis>
|
|
:
|
|
<collectingMunicipality box="[499,634,807,826]" pageId="19" pageNumber="432">Machu Picchu</collectingMunicipality>
|
|
,,
|
|
<date box="[671,777,807,826]" pageId="19" pageNumber="432" value="1982-11-28">
|
|
<collectingDate box="[671,777,807,826]" pageId="19" pageNumber="432" value="1982-11-28">28.XI.1982</collectingDate>
|
|
</date>
|
|
,
|
|
<collectorName box="[789,882,807,826]" pageId="19" pageNumber="432">E.R. Ross</collectorName>
|
|
col., California Academy of Sciences (
|
|
<collectionCode box="[314,380,832,851]" collectionName="DARC" pageId="19" pageNumber="432">DARC</collectionCode>
|
|
)
|
|
</materialsCitation>
|
|
;
|
|
<materialsCitation ID-GBIF-Occurrence="2579424849" box="[398,1028,832,851]" collectingDate="1964-07-01" collectionCode="MZSP" collectorName="B. Malkin" country="Peru" elevation="2700" pageId="19" pageNumber="432" specimenCount="1" specimenCount-female="4" specimenCount-male="2" stateProvince="Cuzco">
|
|
(over ruins,
|
|
<quantity box="[512,629,832,851]" metricMagnitude="3" metricUnit="m" metricValue="2.7" metricValueMax="2.8" metricValueMin="2.6" pageId="19" pageNumber="432" unit="m" value="2700.0" valueMax="2800.0" valueMin="2600.0">
|
|
<elevation box="[512,629,832,851]" metricMagnitude="3" metricUnit="m" metricValue="2.7" metricValueMax="2.8" metricValueMin="2.6" pageId="19" pageNumber="432" unit="m" value="2700.0" valueMax="2800.0" valueMin="2600.0">2600-2800m</elevation>
|
|
</quantity>
|
|
), 2, 4,
|
|
<date box="[739,839,832,851]" pageId="19" pageNumber="432" value="1964-07-01">
|
|
<collectingDate box="[739,839,832,851]" pageId="19" pageNumber="432" value="1964-07-01">1.VII.1964</collectingDate>
|
|
</date>
|
|
,
|
|
<collectorName box="[851,944,832,851]" pageId="19" pageNumber="432">B. Malkin</collectorName>
|
|
(
|
|
<collectionCode box="[959,1022,832,851]" collectionName="Brazil, Sao Paulo, Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="19" pageNumber="432">MZSP</collectionCode>
|
|
)
|
|
</materialsCitation>
|
|
;
|
|
<materialsCitation ID-GBIF-Occurrence="2579424860" box="[112,379,858,877]" collectingDate="1965-05-07" collectionCode="MZSP" country="Peru" pageId="19" pageNumber="432" specimenCount="1" specimenCount-female="1" stateProvince="Cuzco">
|
|
,
|
|
<date box="[126,206,858,877]" pageId="19" pageNumber="432" value="1965-05-07">
|
|
<collectingDate box="[126,206,858,877]" pageId="19" pageNumber="432" value="1965-05-07">7.V.1965</collectingDate>
|
|
</date>
|
|
(arbusto) (
|
|
<collectionCode box="[310,373,858,877]" collectionName="Brazil, Sao Paulo, Sao Paulo, Museu de Zoologia da Universidade de Sao Paulo" pageId="19" pageNumber="432">MZSP</collectionCode>
|
|
)
|
|
</materialsCitation>
|
|
<materialsCitation ID-GBIF-Occurrence="2579424891" box="[380,654,858,877]" collectionCode="USNM" country="Peru" pageId="19" pageNumber="432" specimenCount="1" specimenCount-female="1" stateProvince="Cuzco">
|
|
;, 1942, Weyrauch (
|
|
<collectionCode box="[579,648,858,877]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" pageId="19" pageNumber="432">USNM</collectionCode>
|
|
)
|
|
</materialsCitation>
|
|
;
|
|
<materialsCitation ID-GBIF-Occurrence="2579424828" box="[666,1028,858,877]" country="Peru" elevation="2800" location="Pillahuata" pageId="19" pageNumber="432" specimenCount="1" specimenCount-female="2" specimenCount-male="1" stateProvince="Prov. Paucartambo">
|
|
<collectingRegion box="[666,840,858,877]" pageId="19" pageNumber="432">Prov. Paucartambo</collectingRegion>
|
|
,
|
|
<location LSID="urn:lsid:plazi:treatment:217887A7991F80224BF85C0BFE906036:AC0E606A991E802249235D18FC2F6778" box="[851,941,858,877]" country="Peru" name="Pillahuata" pageId="19" pageNumber="432" stateProvince="Prov. Paucartambo">Pillahuata</location>
|
|
(
|
|
<quantity box="[955,1022,858,877]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="19" pageNumber="432" unit="m" value="2800.0">
|
|
<elevation box="[955,1022,858,877]" metricMagnitude="3" metricUnit="m" metricValue="2.8" pageId="19" pageNumber="432" unit="m" value="2800.0">2800m</elevation>
|
|
</quantity>
|
|
)
|
|
</materialsCitation>
|
|
,
|
|
<materialsCitation ID-GBIF-Occurrence="2579424962" box="[115,516,882,901]" collectingDate="1952-12-19" collectionCode="USNM" collectorName="F. Woytkowski" country="Peru" pageId="19" pageNumber="432" specimenCount="1" stateProvince="Prov. Paucartambo">
|
|
, 2,
|
|
<date box="[173,281,882,901]" pageId="19" pageNumber="432" value="1952-12-19">
|
|
<collectingDate box="[173,281,882,901]" pageId="19" pageNumber="432" value="1952-12-19">19.XII.1952</collectingDate>
|
|
</date>
|
|
,
|
|
<collectorName box="[294,428,882,901]" pageId="19" pageNumber="432">F. Woytkowski</collectorName>
|
|
(
|
|
<collectionCode box="[442,510,882,901]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" pageId="19" pageNumber="432">USNM</collectionCode>
|
|
)
|
|
</materialsCitation>
|
|
;
|
|
<materialsCitation ID-GBIF-Occurrence="2579424968" box="[528,906,882,901]" collectionCode="DARC" country="Peru" location="Marcapata" municipality="Marcapata" pageId="19" pageNumber="432" specimenCount="1" stateProvince="Prov. Paucartambo">
|
|
<collectingMunicipality box="[528,623,882,901]" pageId="19" pageNumber="432">Marcapata</collectingMunicipality>
|
|
,, “Coll. I.R.Sc. N.B.” (
|
|
<collectionCode box="[835,901,882,901]" collectionName="DARC" pageId="19" pageNumber="432">DARC</collectionCode>
|
|
)
|
|
</materialsCitation>
|
|
.
|
|
<materialsCitation ID-GBIF-Occurrence="2579424871" collectionCode="USNM" collectorName="R. Zischka" country="Bolivia" elevation="2000" location="Yungas de Palmar" municipality="Yungas de Palmar" pageId="19" pageNumber="432" specimenCount="1">
|
|
<collectingCountry box="[917,1009,882,901]" name="Bolivia" pageId="19" pageNumber="432" value="Bolivia">BOLIVIA</collectingCountry>
|
|
,
|
|
<collectingMunicipality box="[101,276,907,926]" pageId="19" pageNumber="432">Yungas de Palmar</collectingMunicipality>
|
|
(
|
|
<quantity box="[291,361,907,926]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="19" pageNumber="432" unit="m" value="2000.0">
|
|
<elevation box="[291,361,907,926]" metricMagnitude="3" metricUnit="m" metricValue="2.0" pageId="19" pageNumber="432" unit="m" value="2000.0">2000m</elevation>
|
|
</quantity>
|
|
),,
|
|
<collectorName box="[407,511,907,926]" pageId="19" pageNumber="432">R. Zischka</collectorName>
|
|
(
|
|
<collectionCode box="[527,599,907,926]" collectionName="USA, Washington D.C., National Museum of Natural History, [formerly, United States National Museum]" pageId="19" pageNumber="432">USNM</collectionCode>
|
|
)
|
|
</materialsCitation>
|
|
;
|
|
<materialsCitation ID-GBIF-Occurrence="2579424848" collectingDate="1961-11" collectionCode="DARC" country="Bolivia" elevation="110" location="Chapare" municipality="Cochabamba" pageId="19" pageNumber="432" specimenCount="1" specimenCount-female="2" specimenCount-male="2" stateProvince="Cochabamba">
|
|
<emphasis bold="true" box="[618,748,906,925]" pageId="19" pageNumber="432">
|
|
<collectingMunicipality box="[618,748,906,925]" pageId="19" pageNumber="432">Cochabamba</collectingMunicipality>
|
|
</emphasis>
|
|
:
|
|
<location LSID="urn:lsid:plazi:treatment:217887A7991F80224BF85C0BFE906036:AC0E606A991E8022488C5DC9FCC8678B" box="[764,842,907,926]" country="Bolivia" municipality="Cochabamba" name="Chapare" pageId="19" pageNumber="432" stateProvince="Cochabamba">Chapare</location>
|
|
prov., Alto Palmar (
|
|
<quantity box="[108,163,932,951]" metricMagnitude="2" metricUnit="m" metricValue="1.1" pageId="19" pageNumber="432" unit="m" value="110.0">
|
|
<elevation box="[108,163,932,951]" metricMagnitude="2" metricUnit="m" metricValue="1.1" pageId="19" pageNumber="432" unit="m" value="110.0">110m</elevation>
|
|
</quantity>
|
|
), 2, 2,
|
|
<date box="[275,349,932,951]" pageId="19" pageNumber="432" value="1961-11">
|
|
<collectingDate box="[275,349,932,951]" pageId="19" pageNumber="432" value="1961-11">XI.1961</collectingDate>
|
|
</date>
|
|
(
|
|
<collectionCode box="[364,431,932,951]" collectionName="DARC" pageId="19" pageNumber="432">DARC</collectionCode>
|
|
)
|
|
</materialsCitation>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection box="[165,700,954,977]" pageId="19" pageNumber="432" type="distribution">
|
|
<paragraph blockId="19.[101,1032,190,1059]" box="[165,700,954,977]" pageId="19" pageNumber="432">
|
|
Distribution. Peru (Cuzco), Bolivia (
|
|
<collectingRegion box="[544,689,954,977]" country="Bolivia" name="Cochabamba" pageId="19" pageNumber="432">Cochabamba</collectingRegion>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="19" pageNumber="432" type="discussion">
|
|
<paragraph blockId="19.[101,1032,190,1059]" pageId="19" pageNumber="432">
|
|
Comments. Bra
|
|
<emphasis box="[336,548,981,1004]" italics="true" pageId="19" pageNumber="432">chystethus vexillum</emphasis>
|
|
is closely related to
|
|
<taxonomicName authorityName="Bolívar" authorityYear="1879" box="[783,898,981,1004]" class="Insecta" family="Pentatomidae" genus="Brachystethus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="432" phylum="Arthropoda" rank="species" species="tricolor">
|
|
<emphasis box="[783,898,981,1004]" italics="true" pageId="19" pageNumber="432">B. tricolor</emphasis>
|
|
</taxonomicName>
|
|
. The dorsal color pattern, with yellow stripes, and the globose pygophore distinguish this species from
|
|
<emphasis box="[159,274,1036,1059]" italics="true" pageId="19" pageNumber="432">
|
|
<taxonomicName authorityName="Bolívar" authorityYear="1879" box="[159,270,1036,1059]" class="Insecta" family="Pentatomidae" genus="Brachystethus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Hemiptera" pageId="19" pageNumber="432" phylum="Arthropoda" rank="species" species="tricolor">B. tricolor</taxonomicName>
|
|
.
|
|
</emphasis>
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |