429 lines
56 KiB
XML
429 lines
56 KiB
XML
<document id="DEAE53B74AF297C5091A2CC5A6675C34" ID-CLB-Dataset="26382" ID-DOI="10.11646/zootaxa.4624.1.6" ID-GBIF-Dataset="6d724005-a6af-4345-9407-5932133b3c4b" ID-ISSN="1175-5326" ID-Zenodo-Dep="3258482" ID-ZooBank="593D3E52-0DBC-4BCB-AABF-AE3D7B46BA65" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1561648504544" checkinUser="plazi" docAuthor="Bedoya-Roqueme, Edwin & Torres, Richard A." docDate="2019" docId="934D6B7A28148D4BFF3AFC55FD78FAE0" docLanguage="en" docName="zootaxa.4624.1.6.pdf" docOrigin="Zootaxa 4624 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tenebriochernes Bedoya-Roqueme & Torres 2019, gen. nov." docType="treatment" docVersion="14" lastPageNumber="90" masterDocId="6F74130228168D48FFADFFE4FF8DFF9B" masterDocTitle="Tenebriochernes, a new genus and species of Chernetidae (Arachnida: Pseudoscorpiones) from north-western Colombia, with ecological observations" masterLastPageNumber="107" masterPageNumber="87" pageNumber="89" updateTime="1698737322199" updateUser="ExternalLinkService">
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<mods:mods id="468C3658EB1864D99684DDAF5C22CCAB" xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo id="994002BC8D9A7BAE4DB4EA49648825F8">
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<mods:title id="045D2C7377ADDDD19A922C9497F7B597">Tenebriochernes, a new genus and species of Chernetidae (Arachnida: Pseudoscorpiones) from north-western Colombia, with ecological observations</mods:title>
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<mods:namePart id="42BAC9F16E837648BC8DD566AA13A0FB">Bedoya-Roqueme, Edwin</mods:namePart>
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<mods:roleTerm id="9D4C3463F0566BED0FC147107B6FB197">Author</mods:roleTerm>
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<mods:namePart id="7648B79090A4739076CEF8857FBC4458">Torres, Richard A.</mods:namePart>
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<mods:typeOfResource id="3190C87C54E72BD738BE333E9D3B33EF">text</mods:typeOfResource>
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<mods:title id="FFB1A48A2244BC39CA47C33CCAD716DE">Zootaxa</mods:title>
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<mods:part id="EBA3B87EE888A24163AAF93CA62196C5">
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<mods:date id="9CDF22363E6901294038C86049EAEA5B">2019</mods:date>
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<mods:number id="BA415506755299E83FD85603D90888D8">2019-06-27</mods:number>
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<mods:number id="39429F1A926FF7A70E621A671FFACC0B">4624</mods:number>
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<mods:detail id="8EC2087D9890B4FB8D4D0A069C368268" type="issue">
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<mods:identifier id="FE75D761AB8A6F95D7C9CEC78323F52F" type="DOI">10.11646/zootaxa.4624.1.6</mods:identifier>
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<treatment id="934D6B7A28148D4BFF3AFC55FD78FAE0" ID-DOI="http://doi.org/10.5281/zenodo.5922506" ID-GBIF-Taxon="158407556" ID-Zenodo-Dep="5922506" LSID="urn:lsid:plazi:treatment:934D6B7A28148D4BFF3AFC55FD78FAE0" httpUri="http://treatment.plazi.org/id/934D6B7A28148D4BFF3AFC55FD78FAE0" lastPageId="3" lastPageNumber="90" pageId="2" pageNumber="89">
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<subSubSection id="53FE89E728148D4AFF3AFC55FC48FC74" pageId="2" pageNumber="89" type="nomenclature">
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<paragraph id="1B5BDA6C28148D4AFF3AFC55FE58FC50" blockId="2.[151,965,945,1007]" box="[151,469,945,972]" pageId="2" pageNumber="89">
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<heading id="40136D0028148D4AFF3AFC55FE58FC50" bold="true" box="[151,469,945,972]" fontSize="11" level="1" pageId="2" pageNumber="89" reason="1">
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<emphasis id="2990067E28148D4AFF3AFC55FE58FC50" bold="true" box="[151,469,945,972]" pageId="2" pageNumber="89">
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<taxonomicName id="DCE4A1EF28148D4AFF3AFC55FEEDFC57" ID-CoL="7TRV" authority="Bedoya-Roqueme & Torres, 2019" authorityName="Bedoya-Roqueme & Torres" authorityYear="2019" box="[151,352,945,972]" class="Arachnida" family="Chernetidae" genus="Tenebriochernes" kingdom="Animalia" order="Pseudoscorpiones" pageId="2" pageNumber="89" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="2990067E28148D4AFF3AFC55FEEDFC57" bold="true" box="[151,352,945,972]" italics="true" pageId="2" pageNumber="89">Tenebriochernes</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="32A3BB0528148D4AFEC5FC55FE58FC50" box="[360,469,945,971]" pageId="2" pageNumber="89" rank="genus">gen. nov.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="1B5BDA6C28148D4AFF3AFC30FC48FC74" blockId="2.[151,965,945,1007]" box="[151,965,980,1007]" pageId="2" pageNumber="89">
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<uri id="6F75D66E28148D4AFF3AFC30FC48FC74" box="[151,965,980,1007]" pageId="2" pageNumber="89">
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urn:lsid:zoobank.org:act:
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<uuid id="6F42E0B928148D4AFE04FC30FC48FC74" box="[425,965,980,1007]" pageId="2" pageNumber="89">A4181424-D1A0-4A5A-8025-1EFC040DF730</uuid>
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</uri>
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</paragraph>
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</subSubSection>
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<subSubSection id="53FE89E728148D4AFF3AFBFDFDFAFBAA" box="[151,631,1048,1073]" pageId="2" pageNumber="89" type="materials_examined">
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<paragraph id="1B5BDA6C28148D4AFF3AFBFDFDFAFBAA" blockId="2.[151,631,1048,1073]" box="[151,631,1048,1073]" pageId="2" pageNumber="89">
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<emphasis id="2990067E28148D4AFF3AFBFDFE93FBAA" bold="true" box="[151,286,1048,1073]" pageId="2" pageNumber="89">
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<typeStatus id="C45F64CE28148D4AFF3AFBFDFF41FBAA" box="[151,204,1049,1073]" pageId="2" pageNumber="89">Type</typeStatus>
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species
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</emphasis>
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:
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<taxonomicName id="DCE4A1EF28148D4AFE87FBFDFDBAFBAA" authority="Bedoya-Roqueme & Torres, 2019" authorityName="Bedoya-Roqueme & Torres" authorityYear="2019" box="[298,567,1049,1073]" class="Arachnida" family="Chernetidae" genus="Tenebriochernes" kingdom="Animalia" order="Pseudoscorpiones" pageId="2" pageNumber="89" phylum="Arthropoda" rank="species" species="concavus" status="sp. nov.">
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<emphasis id="2990067E28148D4AFE87FBFDFDBAFBAA" box="[298,567,1049,1073]" italics="true" pageId="2" pageNumber="89">Tenebriochernes concavus</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="32A3BB0528148D4AFD90FBFDFDFAFBAA" box="[573,631,1049,1073]" pageId="2" pageNumber="89" rank="species">
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<emphasis id="2990067E28148D4AFD90FBFDFDFAFBAA" bold="true" box="[573,631,1049,1073]" pageId="2" pageNumber="89">n. sp.</emphasis>
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</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="53FE89E728148D4AFF3AFB84FF67FB5A" pageId="2" pageNumber="89" type="etymology">
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<paragraph id="1B5BDA6C28148D4AFF3AFB84FF67FB5A" blockId="2.[151,1437,1119,2010]" pageId="2" pageNumber="89">
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<emphasis id="2990067E28148D4AFF3AFB84FE95FBE1" bold="true" box="[151,280,1120,1146]" pageId="2" pageNumber="89">Etymology</emphasis>
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. The generic name refers to the presence of the
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<typeStatus id="C45F64CE28148D4AFC90FB84FCE0FBE1" box="[829,877,1120,1146]" pageId="2" pageNumber="89">type</typeStatus>
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species within caves (
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<emphasis id="2990067E28148D4AFBC3FB84FB46FBE1" box="[1134,1227,1120,1146]" italics="true" pageId="2" pageNumber="89">tenebrio</emphasis>
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, Latin, shunner of light, lover of darkness), which is combined with the genus name
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<taxonomicName id="DCE4A1EF28148D4AFC3AFB60FC79FB05" authorityName="Menge" authorityYear="1855" box="[919,1012,1156,1182]" class="Arachnida" family="Chernetidae" genus="Chernes" kingdom="Animalia" order="Pseudoscorpiones" pageId="2" pageNumber="89" phylum="Arthropoda" rank="genus">
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<emphasis id="2990067E28148D4AFC3AFB60FC79FB05" box="[919,1012,1156,1182]" italics="true" pageId="2" pageNumber="89">Chernes</emphasis>
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</taxonomicName>
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(Greek, labourer). It is masculine in gender.
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</paragraph>
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</subSubSection>
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<subSubSection id="53FE89E728148D4AFF6AFB2FFEFFFA79" pageId="2" pageNumber="89" type="diagnosis">
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<paragraph id="1B5BDA6C28148D4AFF6AFB2FFEFFFA79" blockId="2.[151,1437,1119,2010]" pageId="2" pageNumber="89">
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<emphasis id="2990067E28148D4AFF6AFB2FFEB7FB7D" bold="true" box="[199,314,1227,1254]" pageId="2" pageNumber="89">Diagnosis</emphasis>
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:
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<taxonomicName id="DCE4A1EF28148D4AFEE5FB28FD8FFB7D" authority="Bedoya-Roqueme & Torres, 2019" authorityName="Bedoya-Roqueme & Torres" authorityYear="2019" box="[328,514,1228,1254]" class="Arachnida" family="Chernetidae" genus="Tenebriochernes" kingdom="Animalia" order="Pseudoscorpiones" pageId="2" pageNumber="89" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="2990067E28148D4AFEE5FB28FD8FFB7D" box="[328,514,1228,1254]" italics="true" pageId="2" pageNumber="89">Tenebriochernes</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="32A3BB0528148D4AFDA5FB28FDDBFB7D" box="[520,598,1228,1254]" pageId="2" pageNumber="89" rank="genus">
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<emphasis id="2990067E28148D4AFDA5FB28FDDBFB7D" bold="true" box="[520,598,1228,1254]" pageId="2" pageNumber="89">n. gen.</emphasis>
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</taxonomicNameLabel>
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can be distinguished from all other chernetid genera by the following combination of characters: presence of 7 denticulate and thickened setae (
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<emphasis id="2990067E28148D4AFC35FB14FC39FA91" box="[920,948,1264,1290]" italics="true" pageId="2" pageNumber="89">Sd</emphasis>
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) proximal of the base of fixed chelal finger; the concavity on the retrolateral margin of the pedipalpal patella; the chelal hand shape and the strong concavity on the base of the movable chelal finger proximal of the condyle; trichobothrium
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<emphasis id="2990067E28148D4AFC52FADCFBE2FAC9" box="[1023,1135,1336,1362]" italics="true" pageId="2" pageNumber="89">ib, ist, esb</emphasis>
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and
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<emphasis id="2990067E28148D4AFB09FADCFB33FAC9" box="[1188,1214,1336,1362]" italics="true" pageId="2" pageNumber="89">eb</emphasis>
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grouped at the base of the fixed finger in a diagonal row, trichobothrium
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<taxonomicName id="DCE4A1EF28148D4AFD72FAB8FCFEFAEE" box="[735,883,1371,1397]" pageId="2" pageNumber="89" rank="subSpecies" subSpecies="equidistant">
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<emphasis id="2990067E28148D4AFD72FAB8FD7FFAEE" box="[735,754,1372,1397]" italics="true" pageId="2" pageNumber="89">st</emphasis>
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equidistant
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</taxonomicName>
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between
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<emphasis id="2990067E28148D4AFC71FAB8FC69FAEE" box="[988,996,1372,1397]" italics="true" pageId="2" pageNumber="89">t</emphasis>
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and
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<emphasis id="2990067E28148D4AFBB4FAB8FBBFFAED" box="[1049,1074,1372,1398]" italics="true" pageId="2" pageNumber="89">sb</emphasis>
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, proximal of nodus ramosus and approximately at the same level as
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<emphasis id="2990067E28148D4AFD8AFA64FDCBFA02" box="[551,582,1408,1433]" italics="true" pageId="2" pageNumber="89">est</emphasis>
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; females spermathecae with a distally expanded and less globose sacs with darkened tips, and two long tubes which not leading to a single medium chamber with a cribriform plate; tarsus IV without tactile seta.
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</paragraph>
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</subSubSection>
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<subSubSection id="53FE89E728148D4BFF6AFA08FD78FAE0" lastPageId="3" lastPageNumber="90" pageId="2" pageNumber="89" type="discussion">
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<paragraph id="1B5BDA6C28148D4BFF6AFA08FD5BFEFC" blockId="2.[151,1437,1119,2010]" lastBlockId="3.[151,1437,153,1403]" lastPageId="3" lastPageNumber="90" pageId="2" pageNumber="89">
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<emphasis id="2990067E28148D4AFF6AFA08FEB9F99D" bold="true" box="[199,308,1516,1542]" pageId="2" pageNumber="89">Remarks</emphasis>
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: The genus
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<taxonomicName id="DCE4A1EF28148D4AFE6EFA08FDF0F99D" authority="Bedoya-Roqueme & Torres, 2019" authorityName="Bedoya-Roqueme & Torres" authorityYear="2019" box="[451,637,1516,1542]" class="Arachnida" family="Chernetidae" genus="Tenebriochernes" kingdom="Animalia" order="Pseudoscorpiones" pageId="2" pageNumber="89" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="2990067E28148D4AFE6EFA08FDF0F99D" box="[451,637,1516,1542]" italics="true" pageId="2" pageNumber="89">Tenebriochernes</emphasis>
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</taxonomicName>
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is referred to the subfamily
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<taxonomicName id="DCE4A1EF28148D4AFC69FA0FFBC6F99E" authorityName="Menge" authorityYear="1855" box="[964,1099,1515,1541]" class="Arachnida" family="Chernetidae" kingdom="Animalia" order="Pseudoscorpiones" pageId="2" pageNumber="89" phylum="Arthropoda" rank="subFamily" subFamily="Chernetinae">Chernetinae</taxonomicName>
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by lacking T-shaped spermathecae as occurs in the
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<taxonomicName id="DCE4A1EF28148D4AFE32F9EBFDF4F9B2" authorityName="Beier" authorityYear="1932" box="[415,633,1551,1577]" class="Arachnida" family="Chernetidae" kingdom="Animalia" order="Pseudoscorpiones" pageId="2" pageNumber="89" phylum="Arthropoda" rank="subFamily" subFamily="Lamprochernetinae">Lamprochernetinae</taxonomicName>
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and the angular posterior margin of the carapace, exclusive of the
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<taxonomicName id="DCE4A1EF28148D4AFADCF9F4FECDF9D6" authorityName="Beier" authorityYear="1932" class="Arachnida" family="Chernetidae" kingdom="Animalia" order="Pseudoscorpiones" pageId="2" pageNumber="89" phylum="Arthropoda" rank="subFamily" subFamily="Goniochernetinae">Goniochernetinae</taxonomicName>
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. Like a number of other chernetine genera,
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<taxonomicName id="DCE4A1EF28148D4AFC85F9D0FC6FF9D5" authority="Bedoya-Roqueme & Torres, 2019" authorityName="Bedoya-Roqueme & Torres" authorityYear="2019" box="[808,994,1588,1614]" class="Arachnida" family="Chernetidae" genus="Tenebriochernes" kingdom="Animalia" order="Pseudoscorpiones" pageId="2" pageNumber="89" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="2990067E28148D4AFC85F9D0FC6FF9D5" box="[808,994,1588,1614]" italics="true" pageId="2" pageNumber="89">Tenebriochernes</emphasis>
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</taxonomicName>
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has four blades in the cheliceral rallum and lacks a tactile seta on tarsi III and IV.
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<taxonomicName id="DCE4A1EF28148D4AFDC1F9BCFCABF9E9" authority="Bedoya-Roqueme & Torres, 2019" authorityName="Bedoya-Roqueme & Torres" authorityYear="2019" box="[620,806,1624,1650]" class="Arachnida" family="Chernetidae" genus="Tenebriochernes" kingdom="Animalia" order="Pseudoscorpiones" pageId="2" pageNumber="89" phylum="Arthropoda" rank="genus" status="gen. nov.">
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<emphasis id="2990067E28148D4AFDC1F9BCFCABF9E9" box="[620,806,1624,1650]" italics="true" pageId="2" pageNumber="89">Tenebriochernes</emphasis>
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</taxonomicName>
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differs from the genera
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<taxonomicName id="DCE4A1EF28148D4AFB99F9BCFAD2F9E9" authority="Beier, 1955" authorityName="Beier" authorityYear="1955" box="[1076,1375,1623,1650]" class="Arachnida" family="Chernetidae" genus="Atherochernes" kingdom="Animalia" order="Pseudoscorpiones" pageId="2" pageNumber="89" phylum="Arthropoda" rank="genus">
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<emphasis id="2990067E28148D4AFB99F9BCFB5BF9E9" box="[1076,1238,1624,1650]" italics="true" pageId="2" pageNumber="89">Atherochernes</emphasis>
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Beier, 1955
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</taxonomicName>
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from
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<collectingCountry id="63F39AFC28148D4AFF3AF99FFE80F90E" box="[151,269,1659,1685]" name="Venezuela" pageId="2" pageNumber="89">Venezuela</collectingCountry>
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,
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<taxonomicName id="DCE4A1EF28148D4AFEBAF998FDB3F90D" authority="Beier, 1932" authorityName="Beier" authorityYear="1932" box="[279,574,1659,1686]" class="Arachnida" family="Chernetidae" genus="Austrochernes" kingdom="Animalia" order="Pseudoscorpiones" pageId="2" pageNumber="89" phylum="Arthropoda" rank="genus">
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<emphasis id="2990067E28148D4AFEBAF998FE35F90D" box="[279,440,1660,1686]" italics="true" pageId="2" pageNumber="89">Austrochernes</emphasis>
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<bibRefCitation id="7F75A79D28148D4AFE13F99FFDB3F90D" author="Beier, M." box="[446,574,1659,1686]" pageId="2" pageNumber="89" pagination="1 - 294" refId="ref11488" refString="Beier, M. (1932) Pseudoscorpionidea II. Subord. C. Cheliferinea. Tierreich, 58, 1 - 294." type="journal article" year="1932">Beier, 1932</bibRefCitation>
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</taxonomicName>
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and
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<emphasis id="2990067E28148D4AFDDFF998FC81F90D" box="[626,780,1660,1686]" italics="true" pageId="2" pageNumber="89">Balgachernes</emphasis>
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<bibRefCitation id="7F75A79D28148D4AFCBFF998FC2AF90D" author="Harvey, M. S." box="[786,935,1660,1686]" pageId="2" pageNumber="89" pagination="115 - 130" refId="ref12195" refString="Harvey, M. S. (2018) Balgachernes occultus, a new genus and species of pseudoscorpion (Pseudoscorpiones: Chernetidae) associated with balga (Xanthorrhoea preissii) in south-western Australia, with remarks on Austrochernes and Troglochernes. Records of the Western Australian Museum, 33 (1): 115 - 130." type="journal article" year="2018">Harvey, 2018</bibRefCitation>
|
|
from
|
|
<collectingCountry id="63F39AFC28148D4AFC4AF99FFBC3F90E" box="[999,1102,1659,1685]" name="Australia" pageId="2" pageNumber="89">Australia</collectingCountry>
|
|
(
|
|
<bibRefCitation id="7F75A79D28148D4AFBF6F998FB62F90D" author="Harvey, M. S." box="[1115,1263,1660,1686]" pageId="2" pageNumber="89" pagination="115 - 130" refId="ref12195" refString="Harvey, M. S. (2018) Balgachernes occultus, a new genus and species of pseudoscorpion (Pseudoscorpiones: Chernetidae) associated with balga (Xanthorrhoea preissii) in south-western Australia, with remarks on Austrochernes and Troglochernes. Records of the Western Australian Museum, 33 (1): 115 - 130." type="journal article" year="2018">Harvey 2018</bibRefCitation>
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|
) and
|
|
<emphasis id="2990067E28148D4AFA86F998FF63F921" italics="true" pageId="2" pageNumber="89">Eumechochernes</emphasis>
|
|
<bibRefCitation id="7F75A79D28148D4AFF58F97BFEFAF921" author="Beier, M." box="[245,375,1695,1722]" pageId="2" pageNumber="89" pagination="1 - 294" refId="ref11488" refString="Beier, M. (1932) Pseudoscorpionidea II. Subord. C. Cheliferinea. Tierreich, 58, 1 - 294." type="journal article" year="1932">Beier, 1932</bibRefCitation>
|
|
from Hawaii (
|
|
<bibRefCitation id="7F75A79D28148D4AFDB7F97BFD18F921" author="With, C. J." box="[538,661,1695,1722]" pageId="2" pageNumber="89" pagination="94 - 143" refId="ref13725" refString="With, C. J. (1905) On Chelonethi, chiefly from the Australian region, in the collection of the British Museum, with observations on the " coxal sac " and on some cases of abnormal segmentation. Annals and Magazine of Natural History, 15 (7), 94 - 143, 328." type="journal article" year="1905">With 1905</bibRefCitation>
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|
;
|
|
<bibRefCitation id="7F75A79D28148D4AFD0DF97BFC92F921" author="Beier, M." box="[672,799,1695,1722]" pageId="2" pageNumber="89" pagination="1 - 294" refId="ref11488" refString="Beier, M. (1932) Pseudoscorpionidea II. Subord. C. Cheliferinea. Tierreich, 58, 1 - 294." type="journal article" year="1932">Beier 1932</bibRefCitation>
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|
,
|
|
<bibRefCitation id="7F75A79D28148D4AFC87F97BFCE8F921" author="Beier, M." box="[810,869,1695,1722]" pageId="2" pageNumber="89" pagination="131 - 142" refId="ref11546" refString="Beier, M. (1954) Eine Pseudoscorpioniden-Ausbeute aus Venezuela. Memorie del Museo Civico di Storia Naturale di Verona, 4, 131 - 142." type="journal article" year="1954">1954</bibRefCitation>
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|
;
|
|
<bibRefCitation id="7F75A79D28148D4AFCDDF944FB89F921" author="Harvey, M. S." box="[880,1028,1696,1722]" pageId="2" pageNumber="89" refId="ref12147" refString="Harvey, M. S. (2013) Pseudoscorpions of the World, version 3.0. Western Australian Museum, Perth. Available from: http: // www. museum. wa. gov. au / catalogues-beta / pseudoscorpions (accessed 15 May 2018)" type="url" year="2013">Harvey 2013</bibRefCitation>
|
|
) by the pattern of trichobothrium
|
|
<emphasis id="2990067E28148D4AFAD2F944FF69F945" italics="true" pageId="2" pageNumber="89">ib, ist, esb</emphasis>
|
|
and
|
|
<emphasis id="2990067E28148D4AFEB7F920FEB9F945" box="[282,308,1732,1758]" italics="true" pageId="2" pageNumber="89">eb</emphasis>
|
|
grouped at the base of the fixed finger in a diagonal row; trichobothrium
|
|
<taxonomicName id="DCE4A1EF28148D4AFBCAF920FB76F946" box="[1127,1275,1731,1757]" pageId="2" pageNumber="89" rank="subSpecies" subSpecies="equidistant">
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|
<emphasis id="2990067E28148D4AFBCAF920FBF7F946" box="[1127,1146,1732,1757]" italics="true" pageId="2" pageNumber="89">st</emphasis>
|
|
equidistant
|
|
</taxonomicName>
|
|
between
|
|
<emphasis id="2990067E28148D4AFACBF920FAE3F946" box="[1382,1390,1732,1757]" italics="true" pageId="2" pageNumber="89">t</emphasis>
|
|
and
|
|
<emphasis id="2990067E28148D4AFF3AF90CFF3DF899" box="[151,176,1768,1794]" italics="true" pageId="2" pageNumber="89">sb</emphasis>
|
|
, trichobothrium
|
|
<taxonomicName id="DCE4A1EF28148D4AFEC7F90CFE40F89A" box="[362,461,1767,1793]" pageId="2" pageNumber="89" rank="subSpecies" subSpecies="almost">
|
|
<emphasis id="2990067E28148D4AFEC7F90CFEF0F89A" box="[362,381,1768,1793]" italics="true" pageId="2" pageNumber="89">st</emphasis>
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|
almost
|
|
</taxonomicName>
|
|
at the same level as
|
|
<emphasis id="2990067E28148D4AFD01F90CFD46F89A" box="[684,715,1768,1793]" italics="true" pageId="2" pageNumber="89">est</emphasis>
|
|
; venom duct long, nodus ramosus midway between trichobothria
|
|
<emphasis id="2990067E28148D4AFF3AF8E8FF12F8BE" box="[151,159,1804,1829]" italics="true" pageId="2" pageNumber="89">t</emphasis>
|
|
and
|
|
<emphasis id="2990067E28148D4AFF7EF8E8FF6BF8BE" box="[211,230,1804,1829]" italics="true" pageId="2" pageNumber="89">st</emphasis>
|
|
; the chelal hand shape and the strong concavity on the base of movable chelal finger proximal of the condyle, the robust pedipalps and the concavity on the retrolateral margin of the pedipalpal patella. It differs from the genus
|
|
<taxonomicName id="DCE4A1EF28148D4AFF3AF8B0FE41F8F5" authority="Harvey, 1992" authorityName="Harvey" authorityYear="1992" box="[151,460,1876,1902]" class="Arachnida" family="Chernetidae" genus="Marachernes" kingdom="Animalia" order="Pseudoscorpiones" pageId="2" pageNumber="89" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="2990067E28148D4AFF3AF8B0FEA1F8F5" box="[151,300,1876,1902]" italics="true" pageId="2" pageNumber="89">Marachernes</emphasis>
|
|
<bibRefCitation id="7F75A79D28148D4AFE98F8B0FE41F8F5" author="Harvey, M. S." box="[309,460,1876,1902]" pageId="2" pageNumber="89" pagination="1373 - 1435" refId="ref12063" refString="Harvey, M. S. (1992) The phylogeny and classification of the Pseudoscorpionida (Chelicerata). Invertebrate Taxonomy, 6, 1373 - 1435. https: // doi. org / 10.1071 / IT 992137 3" type="journal article" year="1992">Harvey, 1992</bibRefCitation>
|
|
</taxonomicName>
|
|
from
|
|
<collectingCountry id="63F39AFC28148D4AFDBCF8B7FDF5F8F6" box="[529,632,1875,1901]" name="Australia" pageId="2" pageNumber="89">Australia</collectingCountry>
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|
by the absence of an internolateral mound on the male movable chelal finger and the spermathecae with two long tubes and distally expanded sacs with darkened tips. It differs from the genera
|
|
<taxonomicName id="DCE4A1EF28148D4AFF44F878FD8CF82D" authority="Beier, 1930" authorityName="Beier" authorityYear="1930" box="[233,513,1947,1974]" class="Arachnida" family="Chernetidae" genus="Mirochernes" kingdom="Animalia" order="Pseudoscorpiones" pageId="2" pageNumber="89" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="2990067E28148D4AFF44F878FEFAF82D" box="[233,375,1948,1974]" italics="true" pageId="2" pageNumber="89">Mirochernes</emphasis>
|
|
<bibRefCitation id="7F75A79D28148D4AFED2F87FFD8CF82D" author="Beier, M." box="[383,513,1947,1974]" pageId="2" pageNumber="89" pagination="199 - 222" refId="ref11459" refString="Beier, M. (1930) Die Pseudoskorpione des Wiener Naturhistorischen Museums. III. Annalen des Naturhistorischen Museums in Wien, 44, 199 - 222." type="journal article" year="1930">Beier, 1930</bibRefCitation>
|
|
</taxonomicName>
|
|
from
|
|
<collectingCountry id="63F39AFC28148D4AFDEAF87FFD53F82D" box="[583,734,1947,1974]" name="United States of America" pageId="2" pageNumber="89">United States</collectingCountry>
|
|
and
|
|
<taxonomicName id="DCE4A1EF28148D4AFCBBF878FBCAF82D" authority="Beier, 1932" authorityName="Beier" authorityYear="1932" box="[790,1095,1947,1974]" class="Arachnida" family="Chernetidae" genus="Semeiochernes" kingdom="Animalia" order="Pseudoscorpiones" pageId="2" pageNumber="89" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="2990067E28148D4AFCBBF878FC33F82D" box="[790,958,1948,1974]" italics="true" pageId="2" pageNumber="89">Semeiochernes</emphasis>
|
|
<bibRefCitation id="7F75A79D28148D4AFC68F87FFBCAF82D" author="Beier, M." box="[965,1095,1947,1974]" pageId="2" pageNumber="89" pagination="1 - 294" refId="ref11488" refString="Beier, M. (1932) Pseudoscorpionidea II. Subord. C. Cheliferinea. Tierreich, 58, 1 - 294." type="journal article" year="1932">Beier, 1932</bibRefCitation>
|
|
</taxonomicName>
|
|
from
|
|
<collectingCountry id="63F39AFC28148D4AFB20F878FA85F82E" box="[1165,1288,1947,1974]" name="Costa Rica" pageId="2" pageNumber="89">Costa Rica</collectingCountry>
|
|
(
|
|
<bibRefCitation id="7F75A79D28148D4AFABAF87FFA15F82D" author="Beier, M." box="[1303,1432,1947,1974]" pageId="2" pageNumber="89" pagination="199 - 222" refId="ref11459" refString="Beier, M. (1930) Die Pseudoskorpione des Wiener Naturhistorischen Museums. III. Annalen des Naturhistorischen Museums in Wien, 44, 199 - 222." type="journal article" year="1930">Beier 1930</bibRefCitation>
|
|
,
|
|
<bibRefCitation id="7F75A79D28148D4AFF3AF824FF5FF841" author="Beier, M." box="[151,210,1984,2010]" pageId="2" pageNumber="89" pagination="1 - 294" refId="ref11488" refString="Beier, M. (1932) Pseudoscorpionidea II. Subord. C. Cheliferinea. Tierreich, 58, 1 - 294." type="journal article" year="1932">1932</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="7F75A79D28148D4AFF72F824FEF9F841" author="Harvey, M. S." box="[223,372,1984,2010]" pageId="2" pageNumber="89" refId="ref12147" refString="Harvey, M. S. (2013) Pseudoscorpions of the World, version 3.0. Western Australian Museum, Perth. Available from: http: // www. museum. wa. gov. au / catalogues-beta / pseudoscorpions (accessed 15 May 2018)" type="url" year="2013">Harvey 2013</bibRefCitation>
|
|
) by the robust and granulated pedipalps highly sexually dimorphic, the males possess greatly enlarged chelae and chelal peg-like processes, that are absent in females (Chrystophoryová
|
|
<emphasis id="2990067E28158D4BFB33FF7EFB5AFF28" box="[1182,1239,153,179]" italics="true" pageId="3" pageNumber="90">et al.</emphasis>
|
|
2018), as well as the distribution of the trichobothria, with
|
|
<emphasis id="2990067E28158D4BFDC9FF5AFDF3FF4C" box="[612,638,190,215]" italics="true" pageId="3" pageNumber="90">ist</emphasis>
|
|
distal of
|
|
<emphasis id="2990067E28158D4BFD4BFF5AFC88FF4C" box="[742,773,190,215]" italics="true" pageId="3" pageNumber="90">est</emphasis>
|
|
on the fixed chelal finger and
|
|
<taxonomicName id="DCE4A1EF28158D4BFBF5FF5AFB39FF4C" box="[1112,1204,189,215]" pageId="3" pageNumber="90" rank="subSpecies" subSpecies="closer">
|
|
<emphasis id="2990067E28158D4BFBF5FF5AFBE6FF4C" box="[1112,1131,190,215]" italics="true" pageId="3" pageNumber="90">st</emphasis>
|
|
closer
|
|
</taxonomicName>
|
|
to
|
|
<emphasis id="2990067E28158D4BFB7AFF5AFB52FF4C" box="[1239,1247,190,215]" italics="true" pageId="3" pageNumber="90">t</emphasis>
|
|
than to
|
|
<emphasis id="2990067E28158D4BFA95FF59FADCFF4C" box="[1336,1361,189,215]" italics="true" pageId="3" pageNumber="90">sb</emphasis>
|
|
on the movable chelal finger. It differs from the genera
|
|
<taxonomicName id="DCE4A1EF28158D4BFD68FF05FBA8FF60" authority="Chamberlin, 1929" authorityName="Chamberlin" authorityYear="1929" box="[709,1061,225,251]" class="Arachnida" family="Chernetidae" genus="Dinocheirus" kingdom="Animalia" order="Pseudoscorpiones" pageId="3" pageNumber="90" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="2990067E28158D4BFD68FF05FCC2FF60" box="[709,847,225,251]" italics="true" pageId="3" pageNumber="90">Dinocheirus</emphasis>
|
|
<bibRefCitation id="7F75A79D28158D4BFCF4FF05FBA8FF60" author="Chamberlin, J. C." box="[857,1061,225,251]" pageId="3" pageNumber="90" pagination="171 - 173" refId="ref11745" refString="Chamberlin, J. C. (1929) Dinocheirus tenoch, an hitherto undescribed genus and species of false scorpion from Mexico (Arachnida - Chelonethida). Pan-Pacific Entomologist, 5, 171 - 173." type="journal article" year="1929">Chamberlin, 1929</bibRefCitation>
|
|
</taxonomicName>
|
|
from American region, Asia and Europe, that bears only five setae on the cheliceral hand (
|
|
<emphasis id="2990067E28158D4BFCBCFEE1FC92FE84" box="[785,799,261,287]" italics="true" pageId="3" pageNumber="90">b</emphasis>
|
|
and
|
|
<emphasis id="2990067E28158D4BFCF9FEE2FCE6FE84" box="[852,875,262,287]" italics="true" pageId="3" pageNumber="90">es</emphasis>
|
|
being acuminate), tarsus with an acuminate tactile seta distal of midpoint, position of the trichobothrium
|
|
<emphasis id="2990067E28158D4BFD5DFECDFC8BFED8" box="[752,774,297,323]" italics="true" pageId="3" pageNumber="90">ib</emphasis>
|
|
at the same level of
|
|
<emphasis id="2990067E28158D4BFC44FECEFB85FED8" box="[1001,1032,298,323]" italics="true" pageId="3" pageNumber="90">est</emphasis>
|
|
, and trichobothrium
|
|
<emphasis id="2990067E28158D4BFB5EFECDFA99FED8" box="[1267,1300,297,323]" italics="true" pageId="3" pageNumber="90">isb</emphasis>
|
|
proximal of
|
|
<emphasis id="2990067E28158D4BFF3AFEAAFF2AFEFC" box="[151,167,334,359]" italics="true" pageId="3" pageNumber="90">it</emphasis>
|
|
(
|
|
<bibRefCitation id="7F75A79D28158D4BFF18FEA9FEF0FEFC" author="Chamberlin, J. C." box="[181,381,333,359]" pageId="3" pageNumber="90" pagination="171 - 173" refId="ref11745" refString="Chamberlin, J. C. (1929) Dinocheirus tenoch, an hitherto undescribed genus and species of false scorpion from Mexico (Arachnida - Chelonethida). Pan-Pacific Entomologist, 5, 171 - 173." type="journal article" year="1929">Chamberlin 1929</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="7F75A79D28158D4BFE24FEA9FE72FEFC" author="Hoff, C. C." box="[393,511,333,359]" pageId="3" pageNumber="90" pagination="471 - 550" refId="ref12474" refString="Hoff, C. C. (1947) The species of the pseudoscorpion genus Chelanops described by Banks. Bulletin of the Museum of Comparative Zoology, 98, 471 - 550." type="journal article" year="1947">Hoff 1947</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="7F75A79D28158D4BFDA6FEA9FD44FEFC" author="Muchmore, W. B." box="[523,713,333,359]" pageId="3" pageNumber="90" pagination="25 - 36" refId="ref13172" refString="Muchmore, W. B. (1974). Clarification of the genera Hesperochernes and Dinocheirus (Pseudoscorpionida, Chernetidae). Journal of Arachnology, 2, 25 - 36" type="journal article" year="1974">Muchmore 1974</bibRefCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph id="1B5BDA6C28158D4BFF6AFE95FD0FFCC4" blockId="3.[151,1437,153,1403]" pageId="3" pageNumber="90">
|
|
The morphology of the spermatheca of females has been found to provide reliable characters for the delimitation of genera in
|
|
<taxonomicName id="DCE4A1EF28158D4BFED9FE71FE76FE34" authorityName="Menge" authorityYear="1855" box="[372,507,405,431]" class="Arachnida" family="Chernetidae" kingdom="Animalia" order="Pseudoscorpiones" pageId="3" pageNumber="90" phylum="Arthropoda" rank="family">Chernetidae</taxonomicName>
|
|
(e.g.
|
|
<bibRefCitation id="7F75A79D28158D4BFD92FE71FD73FE34" author="Muchmore, W. B." box="[575,766,405,431]" pageId="3" pageNumber="90" pagination="25 - 36" refId="ref13172" refString="Muchmore, W. B. (1974). Clarification of the genera Hesperochernes and Dinocheirus (Pseudoscorpionida, Chernetidae). Journal of Arachnology, 2, 25 - 36" type="journal article" year="1974">Muchmore 1974</bibRefCitation>
|
|
,
|
|
<bibRefCitation id="7F75A79D28158D4BFCA1FE70FCCAFE34" author="Muchmore, W. B." box="[780,839,404,431]" pageId="3" pageNumber="90" pagination="17 - 20" refId="ref13204" refString="Muchmore, W. B. (1975) Use of the spermathecae in the taxonomy of chernetid pseudoscorpions. Proceedings of the International Arachnological Congress, Amsterdam, 6, 17 - 20." type="journal article" year="1975">1975</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="7F75A79D28158D4BFCF8FE71FB21FE34" author="Muchmore, W. B. & Hentschel, E." box="[853,1196,405,431]" pageId="3" pageNumber="90" pagination="41 - 45" refId="ref13389" refString="Muchmore, W. B. & Hentschel, E. (1982) Epichernes aztecus, a new genus and species of pseudoscorpion from Mexico (Pseudoscorpionida, Chernetidae). Journal of Arachnology, 10, 41 - 45." type="journal article" year="1982">Muchmore & Hentschel 1982</bibRefCitation>
|
|
,
|
|
<bibRefCitation id="7F75A79D28158D4BFB14FE71FE94FE48" author="Hentschel, E. & Muchmore, W. B." pageId="3" pageNumber="90" pagination="345 - 349" refId="ref12434" refString="Hentschel, E. & Muchmore, W. B. (1989) Cocinachernes foliosus, a new genus and species of pseudoscorpion (Chernetidae) from Mexico. Journal of Arachnology, 17, 345 - 349." type="journal article" year="1989">Hentschel & Muchmore, 1989</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="7F75A79D28158D4BFE88FE5DFE48FE48" author="Mahnert, V." box="[293,453,441,467]" pageId="3" pageNumber="90" pagination="719 - 810" refId="ref12766" refString="Mahnert, V. (1979) Pseudoskorpione (Arachnida) aus dem Amazonas-Gebiet (Brasilien). Revue Suisse de Zoologie, 86, 719 - 810." type="journal article" year="1979">Mahnert 1979</bibRefCitation>
|
|
,
|
|
<bibRefCitation id="7F75A79D28158D4BFE7DFE5CFD86FE48" author="Mahnert, V." box="[464,523,440,467]" pageId="3" pageNumber="90" pagination="17 - 28" refId="ref12795" refString="Mahnert, V. (1985) Pseudoscorpions (Arachnida} recoltes durant la mission speologique espagnole au Perou en 1977. Revue Arachnologique, 6, 17 - 28." type="journal article" year="1985">1985</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="7F75A79D28158D4BFDBBFE5DFD27FE48" author="Harvey, M. S." box="[534,682,440,467]" pageId="3" pageNumber="90" pagination="199 - 208" refId="ref12104" refString="Harvey, M. S. (1995) Barbaraella gen. nov. and Cacoxylus Beier CPseudoscorpionida: Chernetidae), two remarkable sexually dimorphic pseudoscorpions from Australasia. Records ofthe Western Australian Museum Supplement, 52, 199 - 208." type="journal article" year="1995">Harvey 1995</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="7F75A79D28158D4BFD1BFE5DFC6FFE48" author="Harvey, M. S. & Volschenk, E. S." box="[694,994,441,467]" pageId="3" pageNumber="90" pagination="238 - 277" refId="ref12390" refString="Harvey, M. S. & Volschenk, E. S. (2007) Review of some Australasian Chernetidae: Sundochernes, Troglochernes and a new genus (Pseudoscorpiones). The Journal of Arachnology, 35, 238 - 277." type="journal article" year="2007">Harvey & Volschenk 2007</bibRefCitation>
|
|
). The spermathecae of
|
|
<taxonomicName id="DCE4A1EF28158D4BFB4EFE5DFA10FE48" authority="Bedoya-Roqueme & Torres, 2019" authorityName="Bedoya-Roqueme & Torres" authorityYear="2019" box="[1251,1437,441,467]" class="Arachnida" family="Chernetidae" genus="Tenebriochernes" kingdom="Animalia" order="Pseudoscorpiones" pageId="3" pageNumber="89" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
|
<emphasis id="2990067E28158D4BFB4EFE5DFA10FE48" box="[1251,1437,441,467]" italics="true" pageId="3" pageNumber="90">Tenebriochernes</emphasis>
|
|
</taxonomicName>
|
|
have two long tubes, opening distally into expanded sacs with darkened tips, which distinguish it from the genera
|
|
<taxonomicName id="DCE4A1EF28158D4BFF3AFDE5FE1DFD80" authority="Menge, 1855" authorityName="Menge" authorityYear="1855" box="[151,400,512,539]" class="Arachnida" family="Chernetidae" genus="Chernes" kingdom="Animalia" order="Pseudoscorpiones" pageId="3" pageNumber="90" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="2990067E28158D4BFF3AFDE5FF79FD80" box="[151,244,513,539]" italics="true" pageId="3" pageNumber="90">Chernes</emphasis>
|
|
Menge,
|
|
<quantity id="DC1C778928158D4BFEF5FDE4FE1DFD80" box="[344,400,512,539]" metricMagnitude="1" metricUnit="m" metricValue="4.7116999999999996" pageId="3" pageNumber="90" unit="in" value="1855.0">1855</quantity>
|
|
</taxonomicName>
|
|
in the Holarctic region,
|
|
<taxonomicName id="DCE4A1EF28158D4BFD0EFDE5FC49FD80" authority="Beier, 1937" authorityName="Beier" authorityYear="1937" box="[675,964,513,539]" class="Arachnida" family="Chernetidae" genus="Ceriochernes" kingdom="Animalia" order="Pseudoscorpiones" pageId="3" pageNumber="90" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="2990067E28158D4BFD0EFDE5FCB7FD80" box="[675,826,513,539]" italics="true" pageId="3" pageNumber="90">Ceriochernes</emphasis>
|
|
Beier, 1937
|
|
</taxonomicName>
|
|
from Asia and South America,
|
|
<taxonomicName id="DCE4A1EF28158D4BFA85FDE5FDBAFDA4" authority="Hentschel & Muchmore, 1989" authorityName="Hentschel & Muchmore" authorityYear="1989" class="Arachnida" family="Chernetidae" genus="Cocinachernes" kingdom="Animalia" order="Pseudoscorpiones" pageId="3" pageNumber="90" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="2990067E28158D4BFA85FDE5FF59FDA4" italics="true" pageId="3" pageNumber="90">Cocinachernes</emphasis>
|
|
<bibRefCitation id="7F75A79D28158D4BFF70FDC1FDBAFDA4" author="Hentschel, E. & Muchmore, W. B." box="[221,567,549,575]" pageId="3" pageNumber="90" pagination="345 - 349" refId="ref12434" refString="Hentschel, E. & Muchmore, W. B. (1989) Cocinachernes foliosus, a new genus and species of pseudoscorpion (Chernetidae) from Mexico. Journal of Arachnology, 17, 345 - 349." type="journal article" year="1989">Hentschel & Muchmore, 1989</bibRefCitation>
|
|
</taxonomicName>
|
|
from
|
|
<collectingCountry id="63F39AFC28158D4BFDD2FDC1FD5AFDA4" box="[639,727,549,575]" name="Mexico" pageId="3" pageNumber="90">Mexico</collectingCountry>
|
|
,
|
|
<taxonomicName id="DCE4A1EF28158D4BFD48FDC1FC77FDA4" authority="Hoff, 1949" authorityName="Hoff" authorityYear="1949" box="[741,1018,549,575]" class="Arachnida" family="Chernetidae" genus="Illinichernes" kingdom="Animalia" order="Pseudoscorpiones" pageId="3" pageNumber="90" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="2990067E28158D4BFD48FDC1FCFEFDA4" box="[741,883,549,575]" italics="true" pageId="3" pageNumber="90">Illinichernes</emphasis>
|
|
<bibRefCitation id="7F75A79D28158D4BFCD6FDC1FC77FDA4" author="Hoff, C. C." box="[891,1018,549,575]" pageId="3" pageNumber="90" pagination="407 - 498" refId="ref12508" refString="Hoff, C. C. (1949) The pseudoscorpions of Illinois. Bulletin of the Illinois Natural History Survey, 24, 407 - 498." type="journal article" year="1949">Hoff, 1949</bibRefCitation>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="DCE4A1EF28158D4BFBAAFDC1FAB8FDA4" authority="Beier, 1964" authorityName="Beier" authorityYear="1964" box="[1031,1333,549,575]" class="Arachnida" family="Chernetidae" genus="Neochelanops" kingdom="Animalia" order="Pseudoscorpiones" pageId="3" pageNumber="90" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="2990067E28158D4BFBAAFDC1FB2BFDA4" box="[1031,1190,549,575]" italics="true" pageId="3" pageNumber="90">Neochelanops</emphasis>
|
|
<bibRefCitation id="7F75A79D28158D4BFB02FDC1FAB8FDA4" author="Beier, M." box="[1199,1333,549,575]" pageId="3" pageNumber="90" pagination="592 - 598" refId="ref11599" refString="Beier, M. (1964) Further records of Pseudoscorpionidea from the Solomon Islands. Pacific lnsects, 6, 592 - 598." type="journal article" year="1964">Beier, 1964</bibRefCitation>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="DCE4A1EF28158D4BFAEFFDC1FEF0FDF8" authority="Beier, 1932" authorityName="Beier" authorityYear="1932" class="Arachnida" family="Chernetidae" genus="Semeiochernes" kingdom="Animalia" order="Pseudoscorpiones" pageId="3" pageNumber="90" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="2990067E28158D4BFAEFFDC1FF63FDF8" italics="true" pageId="3" pageNumber="90">Semeiochernes</emphasis>
|
|
<bibRefCitation id="7F75A79D28158D4BFF5AFDADFEF0FDF8" author="Beier, M." box="[247,381,585,611]" pageId="3" pageNumber="90" pagination="1 - 294" refId="ref11488" refString="Beier, M. (1932) Pseudoscorpionidea II. Subord. C. Cheliferinea. Tierreich, 58, 1 - 294." type="journal article" year="1932">Beier, 1932</bibRefCitation>
|
|
</taxonomicName>
|
|
;
|
|
<taxonomicName id="DCE4A1EF28158D4BFE27FDADFC9EFDF8" authority="Chamberlin, 1924" authorityName="Chamberlin" authorityYear="1924" box="[394,787,585,611]" class="Arachnida" family="Chernetidae" genus="Hesperochernes" kingdom="Animalia" order="Pseudoscorpiones" pageId="3" pageNumber="90" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="2990067E28158D4BFE27FDADFDB2FDF8" box="[394,575,585,611]" italics="true" pageId="3" pageNumber="90">Hesperochernes</emphasis>
|
|
Chamberlin, 1924
|
|
</taxonomicName>
|
|
from the American region;
|
|
<taxonomicName id="DCE4A1EF28158D4BFBFEFDADFAD0FDF8" authority="Feio, 1960" authorityName="Feio" authorityYear="1960" box="[1107,1373,585,611]" class="Arachnida" family="Chernetidae" genus="Maxchernes" kingdom="Animalia" order="Pseudoscorpiones" pageId="3" pageNumber="90" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="2990067E28158D4BFBFEFDADFB51FDF8" box="[1107,1244,585,611]" italics="true" pageId="3" pageNumber="90">Maxchernes</emphasis>
|
|
Feio, 1960
|
|
</taxonomicName>
|
|
from South America;
|
|
<taxonomicName id="DCE4A1EF28158D4BFEE0FD89FDF4FD1C" authority="Beier, 1932" authorityName="Beier" authorityYear="1932" box="[333,633,621,647]" class="Arachnida" family="Chernetidae" genus="Austrochernes" kingdom="Animalia" order="Pseudoscorpiones" pageId="3" pageNumber="90" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="2990067E28158D4BFEE0FD89FE63FD1C" box="[333,494,621,647]" italics="true" pageId="3" pageNumber="90">Austrochernes</emphasis>
|
|
<bibRefCitation id="7F75A79D28158D4BFE58FD89FDF4FD1C" author="Beier, M." box="[501,633,621,647]" pageId="3" pageNumber="90" pagination="1 - 294" refId="ref11488" refString="Beier, M. (1932) Pseudoscorpionidea II. Subord. C. Cheliferinea. Tierreich, 58, 1 - 294." type="journal article" year="1932">Beier, 1932</bibRefCitation>
|
|
</taxonomicName>
|
|
,
|
|
<taxonomicName id="DCE4A1EF28158D4BFD29FD89FC55FD1C" authority="Beier, 1966" authorityName="Beier" authorityYear="1966" box="[644,984,621,647]" class="Arachnida" family="Chernetidae" genus="Paraustrochernes" kingdom="Animalia" order="Pseudoscorpiones" pageId="3" pageNumber="90" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="2990067E28158D4BFD29FD89FCC1FD1C" box="[644,844,621,647]" italics="true" pageId="3" pageNumber="90">Paraustrochernes</emphasis>
|
|
<bibRefCitation id="7F75A79D28158D4BFCFEFD89FC55FD1C" author="Beier, M." box="[851,984,621,647]" pageId="3" pageNumber="90" pagination="275 - 303" refId="ref11624" refString="Beier, M. (1966). On the Pseudoscorpionidea of Australia. Australian Journal of Zoology, 14, 275 - 303." type="journal article" year="1966">Beier, 1966</bibRefCitation>
|
|
</taxonomicName>
|
|
, and
|
|
<taxonomicName id="DCE4A1EF28158D4BFBBFFD89FF42FD30" authority="Harvey and Volschenk, 2007" authorityName="Harvey and Volschenk" authorityYear="2007" class="Arachnida" family="Chernetidae" genus="Satrapanus" kingdom="Animalia" order="Pseudoscorpiones" pageId="3" pageNumber="90" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="2990067E28158D4BFBBFFD89FB1FFD1C" box="[1042,1170,621,647]" italics="true" pageId="3" pageNumber="90">Satrapanus</emphasis>
|
|
<bibRefCitation id="7F75A79D28158D4BFB34FD89FF42FD30" author="Harvey, M. S. & Volschenk, E. S." pageId="3" pageNumber="90" pagination="238 - 277" refId="ref12390" refString="Harvey, M. S. & Volschenk, E. S. (2007) Review of some Australasian Chernetidae: Sundochernes, Troglochernes and a new genus (Pseudoscorpiones). The Journal of Arachnology, 35, 238 - 277." type="journal article" year="2007">Harvey and Volschenk, 2007</bibRefCitation>
|
|
</taxonomicName>
|
|
from
|
|
<collectingCountry id="63F39AFC28158D4BFEBFFD75FEF4FD30" box="[274,377,657,683]" name="Australia" pageId="3" pageNumber="90">Australia</collectingCountry>
|
|
(
|
|
<bibRefCitation id="7F75A79D28158D4BFE25FD75FD85FD30" author="Beier, M." box="[392,520,657,683]" pageId="3" pageNumber="90" pagination="1 - 294" refId="ref11488" refString="Beier, M. (1932) Pseudoscorpionidea II. Subord. C. Cheliferinea. Tierreich, 58, 1 - 294." type="journal article" year="1932">Beier 1932</bibRefCitation>
|
|
; 1937; 1966;
|
|
<bibRefCitation id="7F75A79D28158D4BFD0FFD75FC95FD30" author="Hoff, C. C." box="[674,792,657,683]" pageId="3" pageNumber="90" pagination="407 - 498" refId="ref12508" refString="Hoff, C. C. (1949) The pseudoscorpions of Illinois. Bulletin of the Illinois Natural History Survey, 24, 407 - 498." type="journal article" year="1949">Hoff 1949</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="7F75A79D28158D4BFC89FD75FBF8FD30" author="Hentschel, E. & Muchmore, W. B." box="[804,1141,657,683]" pageId="3" pageNumber="90" pagination="345 - 349" refId="ref12434" refString="Hentschel, E. & Muchmore, W. B. (1989) Cocinachernes foliosus, a new genus and species of pseudoscorpion (Chernetidae) from Mexico. Journal of Arachnology, 17, 345 - 349." type="journal article" year="1989">Hentschel & Muchmore 1989</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="7F75A79D28158D4BFB2CFD75FAB3FD30" author="Muchmore, W. B." box="[1153,1342,657,683]" pageId="3" pageNumber="90" pagination="103 - 111" refId="ref13359" refString="Muchmore, W. B. (1999) Redefinition of the genus Chelanops Gervais (Pseudoscorpionida: Chernetidae). Pan-Pacific Entomologist, 75, 103 - 111." type="journal article" year="1999">Muchmore 1999</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="7F75A79D28158D4BFAE7FD75FEE7FD54" author="Harvey, M. S. & Volschenk, E. S." pageId="3" pageNumber="90" pagination="238 - 277" refId="ref12390" refString="Harvey, M. S. & Volschenk, E. S. (2007) Review of some Australasian Chernetidae: Sundochernes, Troglochernes and a new genus (Pseudoscorpiones). The Journal of Arachnology, 35, 238 - 277." type="journal article" year="2007">Harvey & Volschenk 2007</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="7F75A79D28158D4BFED8FD51FD81FD54" author="Harvey, M. S." box="[373,524,693,719]" pageId="3" pageNumber="90" refId="ref12147" refString="Harvey, M. S. (2013) Pseudoscorpions of the World, version 3.0. Western Australian Museum, Perth. Available from: http: // www. museum. wa. gov. au / catalogues-beta / pseudoscorpions (accessed 15 May 2018)" type="url" year="2013">Harvey 2013</bibRefCitation>
|
|
). Unfortunately, the spermathecae of many chernetine genera remain undescribed and others have yet to be studied in detail, which hinders our understanding of the group. In this sense, a further review of the spermathecal morphology is necessary for each of the genera of
|
|
<taxonomicName id="DCE4A1EF28158D4BFC53FD19FB08FC8C" authorityName="Menge" authorityYear="1855" box="[1022,1157,765,791]" class="Arachnida" family="Chernetidae" kingdom="Animalia" order="Pseudoscorpiones" pageId="3" pageNumber="90" phylum="Arthropoda" rank="family">Chernetidae</taxonomicName>
|
|
family, to clarify and establish phylogenetic relationships within this large and complex family (e.g. Harvey 1991, 1995, 2013;
|
|
<bibRefCitation id="7F75A79D28158D4BFA8CFCC5FF5FFCC4" author="Muchmore, W. B." pageId="3" pageNumber="90" pagination="25 - 36" refId="ref13172" refString="Muchmore, W. B. (1974). Clarification of the genera Hesperochernes and Dinocheirus (Pseudoscorpionida, Chernetidae). Journal of Arachnology, 2, 25 - 36" type="journal article" year="1974">Muchmore 1974</bibRefCitation>
|
|
,
|
|
<bibRefCitation id="7F75A79D28158D4BFF70FCA0FE95FCC4" author="Muchmore, W. B." box="[221,280,836,863]" pageId="3" pageNumber="90" pagination="17 - 20" refId="ref13204" refString="Muchmore, W. B. (1975) Use of the spermathecae in the taxonomy of chernetid pseudoscorpions. Proceedings of the International Arachnological Congress, Amsterdam, 6, 17 - 20." type="journal article" year="1975">1975</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="7F75A79D28158D4BFE89FCA1FDFBFCC4" author="Hentschel, E. & Muchmore, W. B." box="[292,630,837,863]" pageId="3" pageNumber="90" pagination="345 - 349" refId="ref12434" refString="Hentschel, E. & Muchmore, W. B. (1989) Cocinachernes foliosus, a new genus and species of pseudoscorpion (Chernetidae) from Mexico. Journal of Arachnology, 17, 345 - 349." type="journal article" year="1989">Hentschel & Muchmore 1989</bibRefCitation>
|
|
).
|
|
</paragraph>
|
|
<paragraph id="1B5BDA6C28158D4BFF6AFC8DFD78FAE0" blockId="3.[151,1437,153,1403]" pageId="3" pageNumber="90">
|
|
Finally,
|
|
<taxonomicName id="DCE4A1EF28158D4BFE89FC8DFE53FC18" authority="Bedoya-Roqueme & Torres, 2019" authorityName="Bedoya-Roqueme & Torres" authorityYear="2019" box="[292,478,873,899]" class="Arachnida" family="Chernetidae" genus="Tenebriochernes" kingdom="Animalia" order="Pseudoscorpiones" pageId="3" pageNumber="89" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
|
<emphasis id="2990067E28158D4BFE89FC8DFE53FC18" box="[292,478,873,899]" italics="true" pageId="3" pageNumber="90">Tenebriochernes</emphasis>
|
|
</taxonomicName>
|
|
resembles
|
|
<taxonomicName id="DCE4A1EF28158D4BFDF3FC8DFC1FFC18" authority="Hoff, 1947" authorityName="Hoff" authorityYear="1947" box="[606,914,873,899]" class="Arachnida" family="Chernetidae" genus="Neoallochernes" kingdom="Animalia" order="Pseudoscorpiones" pageId="3" pageNumber="90" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="2990067E28158D4BFDF3FC8DFC83FC18" box="[606,782,873,899]" italics="true" pageId="3" pageNumber="90">Neoallochernes</emphasis>
|
|
<bibRefCitation id="7F75A79D28158D4BFCBBFC8DFC1FFC18" author="Hoff, C. C." box="[790,914,873,899]" pageId="3" pageNumber="90" pagination="471 - 550" refId="ref12474" refString="Hoff, C. C. (1947) The species of the pseudoscorpion genus Chelanops described by Banks. Bulletin of the Museum of Comparative Zoology, 98, 471 - 550." type="journal article" year="1947">Hoff, 1947</bibRefCitation>
|
|
</taxonomicName>
|
|
from the
|
|
<collectingCountry id="63F39AFC28158D4BFBA9FC8DFB12FC18" box="[1028,1183,873,899]" name="United States of America" pageId="3" pageNumber="90">United States</collectingCountry>
|
|
, Central America, and some Caribbean islands in having spermatheca with dark tips. The genus
|
|
<taxonomicName id="DCE4A1EF28158D4BFC4EFC69FB1EFC3C" authorityName="Hoff" authorityYear="1947" box="[995,1171,909,935]" class="Arachnida" family="Chernetidae" genus="Neoallochernes" kingdom="Animalia" order="Pseudoscorpiones" pageId="3" pageNumber="90" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="2990067E28158D4BFC4EFC69FB1EFC3C" box="[995,1171,909,935]" italics="true" pageId="3" pageNumber="90">Neoallochernes</emphasis>
|
|
</taxonomicName>
|
|
is characterized by the spermathecae in form of elongated sacs with short slender tubules leading into a single median chamber with a cribriform plate; in this sense, the unusual spermathecal morphology is clearly diagnostic for this genus (e.g.
|
|
<bibRefCitation id="7F75A79D28158D4BFAC5FC31FF5FFB88" author="Hoff, C. C." pageId="3" pageNumber="90" pagination="83 - 88" refId="ref12536" refString="Hoff, C. C. (1957) Tejachernes (Arachnida - Chelonethida, Chernetidae - Chernetinae), a new genus of pseudoscorpion based on Dinocheirus stercoreus. Southwestern Naturalist, 2, 83 - 88." type="journal article" year="1957">Hoff 1957</bibRefCitation>
|
|
;
|
|
<bibRefCitation id="7F75A79D28158D4BFF4DFC1DFE12FB88" author="Muchmore, W. B." box="[224,415,1016,1043]" pageId="3" pageNumber="90" pagination="17 - 20" refId="ref13204" refString="Muchmore, W. B. (1975) Use of the spermathecae in the taxonomy of chernetid pseudoscorpions. Proceedings of the International Arachnological Congress, Amsterdam, 6, 17 - 20." type="journal article" year="1975">Muchmore 1975</bibRefCitation>
|
|
,
|
|
<bibRefCitation id="7F75A79D28158D4BFE01FC1DFE6AFB88" author="Muchmore, W. B." box="[428,487,1017,1043]" pageId="3" pageNumber="90" pagination="127 - 153" refId="ref13239" refString="Muchmore, W. B. (1992) Cavernicolous pseudoscorpions from Texas and New Mexico (Arachnida: Pseudoscorpionida). Texas Memorial Museum, Speleological Monographs, 3, 127 - 153." type="journal article" year="1992">1992</bibRefCitation>
|
|
,
|
|
<bibRefCitation id="7F75A79D28158D4BFE59FC1DFDBDFB8F" author="Muchmore, W. B." box="[500,560,1017,1044]" pageId="3" pageNumber="90" pagination="387 - 389" refId="ref13326" refString="Muchmore, W. B. (1996) A new species of Neoallochernes from Antigua (Pseudoscorpionida: Chernetidae). Caribbean Journal of Science, 32, 387 - 389." type="journal article" year="1996">1996</bibRefCitation>
|
|
).
|
|
<taxonomicName id="DCE4A1EF28158D4BFDE8FC1DFD72FB88" authority="Bedoya-Roqueme & Torres, 2019" authorityName="Bedoya-Roqueme & Torres" authorityYear="2019" box="[581,767,1017,1043]" class="Arachnida" family="Chernetidae" genus="Tenebriochernes" kingdom="Animalia" order="Pseudoscorpiones" pageId="3" pageNumber="89" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
|
<emphasis id="2990067E28158D4BFDE8FC1DFD72FB88" box="[581,767,1017,1043]" italics="true" pageId="3" pageNumber="90">Tenebriochernes</emphasis>
|
|
</taxonomicName>
|
|
can be distinguished from
|
|
<taxonomicName id="DCE4A1EF28158D4BFB94FC1DFB64FB88" authorityName="Hoff" authorityYear="1947" box="[1081,1257,1017,1043]" class="Arachnida" family="Chernetidae" genus="Neoallochernes" kingdom="Animalia" order="Pseudoscorpiones" pageId="3" pageNumber="90" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="2990067E28158D4BFB94FC1DFB64FB88" box="[1081,1257,1017,1043]" italics="true" pageId="3" pageNumber="90">Neoallochernes</emphasis>
|
|
</taxonomicName>
|
|
by the spermathecae with distally expanded and less globose sacs, and two long tubes which are not leading to a single medium chamber with a cribriform plate. Likewise,
|
|
<taxonomicName id="DCE4A1EF28158D4BFD3CFBA5FCC6FBC0" authority="Bedoya-Roqueme & Torres, 2019" authorityName="Bedoya-Roqueme & Torres" authorityYear="2019" box="[657,843,1089,1115]" class="Arachnida" family="Chernetidae" genus="Tenebriochernes" kingdom="Animalia" order="Pseudoscorpiones" pageId="3" pageNumber="89" phylum="Arthropoda" rank="genus" status="gen. nov.">
|
|
<emphasis id="2990067E28158D4BFD3CFBA5FCC6FBC0" box="[657,843,1089,1115]" italics="true" pageId="3" pageNumber="90">Tenebriochernes</emphasis>
|
|
</taxonomicName>
|
|
has the setae
|
|
<emphasis id="2990067E28158D4BFC5EFBA5FB9AFBC0" box="[1011,1047,1089,1115]" italics="true" pageId="3" pageNumber="90">sbs</emphasis>
|
|
on the cheliceral hand (absent in
|
|
<taxonomicName id="DCE4A1EF28158D4BFF3AFB81FECAFBE4" authorityName="Hoff" authorityYear="1947" box="[151,327,1125,1151]" class="Arachnida" family="Chernetidae" genus="Neoallochernes" kingdom="Animalia" order="Pseudoscorpiones" pageId="3" pageNumber="90" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="2990067E28158D4BFF3AFB81FECAFBE4" box="[151,327,1125,1151]" italics="true" pageId="3" pageNumber="90">Neoallochernes</emphasis>
|
|
</taxonomicName>
|
|
), four blades on the rallum (three blades in
|
|
<taxonomicName id="DCE4A1EF28158D4BFC90FB81FC60FBE4" authorityName="Hoff" authorityYear="1947" box="[829,1005,1125,1151]" class="Arachnida" family="Chernetidae" genus="Neoallochernes" kingdom="Animalia" order="Pseudoscorpiones" pageId="3" pageNumber="90" phylum="Arthropoda" rank="genus">
|
|
<emphasis id="2990067E28158D4BFC90FB81FC60FBE4" box="[829,1005,1125,1151]" italics="true" pageId="3" pageNumber="90">Neoallochernes</emphasis>
|
|
</taxonomicName>
|
|
), and also differs by the concavity on the prolateral margin of the pedipalpal patella; further in the shape of the chelal hand: the strong concavity at the base of the movable chelal finger proximal of the condyle, and the distribution of trichobothria
|
|
<emphasis id="2990067E28158D4BFB78FB49FAC4FB5C" box="[1237,1353,1197,1223]" italics="true" pageId="3" pageNumber="90">ib, ist, esb</emphasis>
|
|
and
|
|
<emphasis id="2990067E28158D4BFA2FFB49FA11FB5C" box="[1410,1436,1197,1223]" italics="true" pageId="3" pageNumber="90">eb</emphasis>
|
|
which are grouped at the base of the fixed finger in a diagonal row. Due to these characters, the new genus can be distinguished from all other genera of the family
|
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<taxonomicName id="DCE4A1EF28158D4BFD7DFB11FCDAFA94" authorityName="Menge" authorityYear="1855" box="[720,855,1269,1295]" class="Arachnida" family="Chernetidae" kingdom="Animalia" order="Pseudoscorpiones" pageId="3" pageNumber="90" phylum="Arthropoda" rank="family">Chernetidae</taxonomicName>
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by the shape of the spermathecae, the chelal hand shape with a strong concavity near the base of the movable finger, the form of the retrolateral accessory teeth of the movable chelal finger, proximal of fingertip of fixed chelal finger, the position of the trichobothria, and the concavity on the retrolateral margin of the pedipalpal patella.
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</paragraph>
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</subSubSection>
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</treatment>
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</document> |