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<document id="0CD2162D51F5DBA88E8EA31793E9766C" ID-DOI="10.11646/zootaxa.5361.3.4" ID-ISSN="1175-5326" ID-Zenodo-Dep="10147202" ID-ZooBank="89A338C1-1DD5-439E-9CBE-74151FC5A4B5" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="GgImagineBatch" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1700199036336" checkinUser="plazi" docAuthor="Galindo-Malagón, Ximena Alejandra, Morales, Irina &amp; Moreira, Felipe Ferraz Figueiredo" docDate="2023" docId="587AC751B570A208FF42F97CBAF2FBED" docLanguage="en" docName="zootaxa.5361.3.4.pdf" docOrigin="Zootaxa 5361 (3)" docSource="https://www.mapress.com/zt/article/download/zootaxa.5361.3.4/52196" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Rhagovelia calceola Padilla-Gil 2011" docType="treatment" docVersion="2" lastPageNumber="374" masterDocId="A443BF29B576A20FFFD5FFDEB930FC24" masterDocTitle="A new species and updates on the Rhagovelia angustipes complex (Hemiptera, Heteroptera, Veliidae) in Colombia" masterLastPageNumber="389" masterPageNumber="367" pageNumber="373" updateTime="1700199459245" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="73FD9B4841F4B9BD85E45EF511068029">A new species and updates on the Rhagovelia angustipes complex (Hemiptera, Heteroptera, Veliidae) in Colombia</mods:title>
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<mods:namePart id="8B7E6DACB0947B50F42D3844142DA91A">Galindo-Malagón, Ximena Alejandra</mods:namePart>
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<mods:affiliation id="87EB5F1AF3DC88606690E11289B283C0">Red de Biodiversidad y Sistemática, Instituto de Ecología, A. C. (INECOL), Xalapa, Veracruz, 91073, Mexico &amp; Laboratorio de Entomología, Grupo de investigación Biodiversidad y Conservación, Universidad Pedagógica y Tecnológica de Colombia, Tunja, BY, Colombia &amp; ximena. galindo @ uptc. edu. co; https: // orcid. org / 0000 - 0003 - 0617 - 9403</mods:affiliation>
<mods:nameIdentifier id="6E60A3CABE2115FAE8BE3FF1FA22BA56" type="email">ximena.galindo@uptc.edu.co</mods:nameIdentifier>
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<mods:namePart id="680D4A488FC7E7CAAD72304D07F69075">Morales, Irina</mods:namePart>
<mods:nameIdentifier id="733C2F7D6CBB800FE60098266B3BA652" type="ORCID">0000-0003-2456-5674</mods:nameIdentifier>
<mods:affiliation id="C9FD3CDA1A619C6C9708E0A330B5AA63">Laboratorio de Entomología, Grupo de investigación Biodiversidad y Conservación, Universidad Pedagógica y Tecnológica de Colombia, Tunja, BY, Colombia &amp; irina. morales @ uptc. edu. co; https: // orcid. org / 0000 - 0003 - 2456 - 5674</mods:affiliation>
<mods:nameIdentifier id="BFF8D1AC3D09A0FED8EAED4696A11095" type="email">irina.morales@uptc.edu.co</mods:nameIdentifier>
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<mods:namePart id="081994893CD2088A24105019D758A199">Moreira, Felipe Ferraz Figueiredo</mods:namePart>
<mods:nameIdentifier id="E1152554127C601593C3DB6942E5C5FD" type="ORCID">0000-0002-6692-0323</mods:nameIdentifier>
<mods:affiliation id="7E9DB348A9B893E9B505EB04F902A525">Laboratório de Entomologia, Instituto Oswaldo Cruz, Fundação Oswaldo Cruz, Rio de Janeiro, RJ, Brasil ppmeiameiameia @ gmail. com; https: // orcid. org / 0000 - 0002 - 6692 - 0323</mods:affiliation>
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<treatment id="587AC751B570A208FF42F97CBAF2FBED" LSID="urn:lsid:plazi:treatment:587AC751B570A208FF42F97CBAF2FBED" httpUri="http://treatment.plazi.org/id/587AC751B570A208FF42F97CBAF2FBED" lastPageId="7" lastPageNumber="374" pageId="6" pageNumber="373">
<subSubSection id="98C925CCB570A209FF42F97CBB53FA98" box="[151,611,1698,1725]" pageId="6" pageNumber="373" type="nomenclature">
<paragraph id="D06C7647B570A209FF42F97CBB53FA98" blockId="6.[151,611,1698,1761]" box="[151,611,1698,1725]" pageId="6" pageNumber="373">
<heading id="8B24C12BB570A209FF42F97CBB53FA98" bold="true" box="[151,611,1698,1725]" fontSize="11" level="1" pageId="6" pageNumber="373" reason="1">
<taxonomicName id="17D30DC4B570A209FF42F97CBB53FA98" ID-CoL="4RZ9W" authority="Padilla-Gil, 2011" authorityName="Padilla-Gil" authorityYear="2011" box="[151,611,1698,1725]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="6" pageNumber="373" phylum="Arthropoda" rank="species" species="calceola">
<emphasis id="E2A7AA55B570A209FF42F97CBB53FA98" bold="true" box="[151,611,1698,1725]" pageId="6" pageNumber="373">
<emphasis id="E2A7AA55B570A209FF42F97CB8BAFA99" bold="true" box="[151,394,1698,1725]" italics="true" pageId="6" pageNumber="373">Rhagovelia calceola</emphasis>
Padilla-Gil, 2011
</emphasis>
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</paragraph>
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<subSubSection id="98C925CCB570A209FF42F919B9CEFAC5" box="[151,254,1735,1761]" pageId="6" pageNumber="373" type="description">
<paragraph id="D06C7647B570A209FF42F919B9CEFAC5" blockId="6.[151,611,1698,1761]" box="[151,254,1735,1761]" pageId="6" pageNumber="373">
(
<figureCitation id="48E86AC2B570A209FF4AF919B9C6FAC5" box="[159,246,1735,1761]" captionStart="FIGURE 4" captionStartId="7.[152,255,1716,1740]" captionTargetBox="[402,1181,191,1681]" captionTargetId="figure-17@7.[397,1191,181,1693]" captionTargetPageId="7" captionText="FIGURE 4. Geographic distribution, including new records: A. Rhagovelia gastrotricha; B. R. calceola; C. R. molanoi." figureDoi="http://doi.org/10.5281/zenodo.10147212" httpUri="https://zenodo.org/record/10147212/files/figure.png" pageId="6" pageNumber="373">Fig. 4B</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="98C925CCB570A209FF42F8D0BC6EFBB1" pageId="6" pageNumber="373" type="materials_examined">
<paragraph id="D06C7647B570A209FF42F8D0BC6EFBB1" blockId="6.[151,1436,1806,1943]" pageId="6" pageNumber="373">
<emphasis id="E2A7AA55B570A209FF42F8D0B8B4FB0C" bold="true" box="[151,388,1806,1832]" pageId="6" pageNumber="373">Material examined.</emphasis>
<materialsCitation id="60BB7C1AB570A209FE58F8D1BC6AFBB1" collectingDate="2021-06-05" collectorName="Morales-Alba, A. F. &amp; Martinez, D. C." country="Colombia" elevation="431" latitude="6.0761113" location="Finca Los Mangos" longLatPrecision="1" longitude="-74.21083" municipality="Vereda Locacion" pageId="6" pageNumber="373" specimenCount="2" specimenCount-female="2" stateProvince="Santander">
<collectingCountry id="A8C436D7B570A209FE58F8D1BB16FB0D" box="[397,550,1807,1833]" name="Colombia" pageId="6" pageNumber="373">COLOMBIA</collectingCountry>
.
<emphasis id="E2A7AA55B570A209FDE6F8D0BB85FB0C" bold="true" box="[563,693,1806,1832]" pageId="6" pageNumber="373">
<collectingRegion id="1217B8A5B570A209FDE6F8D0BB9FFB0C" box="[563,687,1806,1832]" country="Colombia" name="Santander" pageId="6" pageNumber="373">Santander</collectingRegion>
.
</emphasis>
<collectingRegion id="1217B8A5B570A209FD6BF8D1BA24FB0D" box="[702,788,1807,1833]" country="Colombia" name="Bolivar" pageId="6" pageNumber="373">Bolívar</collectingRegion>
,
<collectingMunicipality id="3008EC3DB570A209FCF4F8D1BAD4FB0D" box="[801,996,1807,1833]" pageId="6" pageNumber="373">Vereda Locación</collectingMunicipality>
,
<location id="D50C209CB570A209FC27F8D1BDFBFB0D" LSID="urn:lsid:plazi:treatment:587AC751B570A208FF42F97CBAF2FBED:D50C209CB570A209FC27F8D1BDFBFB0D" box="[1010,1227,1807,1833]" country="Colombia" latitude="6.0761113" longLatPrecision="1" longitude="-74.21083" municipality="Vereda Locacion" name="Finca Los Mangos" pageId="6" pageNumber="373" stateProvince="Santander">Finca Los Mangos</location>
, quebrada interior bosque,
<geoCoordinate id="B5E71080B570A209FF21F8ECB874FB69" box="[244,324,1842,1869]" degrees="6.0745" direction="north" orientation="latitude" pageId="6" pageNumber="373" precision="5" value="6.0745">6.0745</geoCoordinate>
,
<geoCoordinate id="B5E71080B570A209FE9BF8EDB885FB69" box="[334,437,1843,1869]" degrees="74.2136" direction="west" orientation="longitude" pageId="6" pageNumber="373" precision="5" value="-74.2136">-74.2136</geoCoordinate>
,
<quantity id="172BDBA2B570A209FE15F8EDBB38FB69" box="[448,520,1843,1869]" metricMagnitude="2" metricUnit="m" metricValue="4.31" pageId="6" pageNumber="373" unit="m" value="431.0">
<elevation id="5BFE9174B570A209FE15F8EDBB38FB69" box="[448,520,1843,1869]" metricMagnitude="2" metricUnit="m" metricValue="4.31" pageId="6" pageNumber="373" unit="m" value="431.0">431 m</elevation>
</quantity>
,
<date id="A46D5087B570A209FDC6F8ECBBA7FB69" box="[531,663,1842,1869]" pageId="6" pageNumber="373" value="2021-06-05">
<collectingDate id="B429A96FB570A209FDC6F8ECBBA7FB69" box="[531,663,1842,1869]" pageId="6" pageNumber="373" value="2021-06-05">2021-VI-05</collectingDate>
</date>
(
<collectorName id="7D261391B570A209FD70F8EDBA4EFB69" box="[677,894,1843,1869]" pageId="6" pageNumber="373">Morales-Alba, A.F.</collectorName>
&amp;
<collectorName id="7D261391B570A209FC75F8EDBD77FB69" box="[928,1095,1843,1869]" pageId="6" pageNumber="373">Martínez, D.C.</collectorName>
leg.): 1 apterous ♁, 1 apterous
<specimenCount id="C6D5BDCEB570A209FF42F889B99CFB55" box="[151,172,1879,1905]" count="1" pageId="6" pageNumber="373" type="female">♀</specimenCount>
(UPTC-In-22651). Vereda Locación, Finca Los Mangos, quebrada interior bosque,
<geoCoordinate id="B5E71080B570A209FB80F889BDFCFB55" box="[1109,1228,1879,1905]" degrees="6.0761111" direction="north" orientation="latitude" pageId="6" pageNumber="373" precision="1" value="6.0761113">6.0761111</geoCoordinate>
,
<geoCoordinate id="B5E71080B570A209FB03F889BC58FB55" box="[1238,1384,1879,1905]" degrees="74.2108333" direction="west" orientation="longitude" pageId="6" pageNumber="373" precision="1" value="-74.21083">-74.2108333</geoCoordinate>
,
<quantity id="172BDBA2B570A209FAA7F889B99FFBB1" metricMagnitude="2" metricUnit="m" metricValue="3.9299999999999997" pageId="6" pageNumber="373" unit="m" value="393.0">
<elevation id="5BFE9174B570A209FAA7F889B99FFBB1" metricMagnitude="2" metricUnit="m" metricValue="3.9299999999999997" pageId="6" pageNumber="373" unit="m" value="393.0">393 m</elevation>
</quantity>
,
<date id="A46D5087B570A209FF6EF8A4B80FFBB1" box="[187,319,1914,1941]" pageId="6" pageNumber="373" value="2021-06-05">
<collectingDate id="B429A96FB570A209FF6EF8A4B80FFBB1" box="[187,319,1914,1941]" pageId="6" pageNumber="373" value="2021-06-05">2021-VI-05</collectingDate>
</date>
(
<collectorName id="7D261391B570A209FE9BF8A5BB17FBB1" box="[334,551,1915,1941]" pageId="6" pageNumber="373">Morales-Alba, A.F.</collectorName>
&amp;
<collectorName id="7D261391B570A209FD9EF8A5BBC3FBB1" box="[587,755,1915,1941]" pageId="6" pageNumber="373">Martínez, D.C.</collectorName>
leg.): 23 apterous ♁, 21 apterous
<specimenCount id="C6D5BDCEB570A209FBA5F8A5BDB5FBB1" box="[1136,1157,1915,1941]" count="1" pageId="6" pageNumber="373" type="female">♀</specimenCount>
(UPTC-In-22652)
</materialsCitation>
.
</paragraph>
</subSubSection>
<caption id="84AC26CFB571A208FF4DF96ABC6FFAE8" ID-DOI="http://doi.org/10.5281/zenodo.10147212" ID-Zenodo-Dep="10147212" box="[152,1375,1716,1741]" httpUri="https://zenodo.org/record/10147212/files/figure.png" pageId="7" pageNumber="374" startId="7.[152,255,1716,1740]" targetBox="[402,1181,191,1681]" targetPageId="7" targetType="figure">
<paragraph id="D06C7647B571A208FF4DF96ABC6FFAE8" blockId="7.[152,1375,1716,1741]" box="[152,1375,1716,1741]" pageId="7" pageNumber="374">
<emphasis id="E2A7AA55B571A208FF4DF96AB828FAE9" bold="true" box="[152,280,1716,1741]" pageId="7" pageNumber="374">FIGURE 4.</emphasis>
Geographic distribution, including new records:
<emphasis id="E2A7AA55B571A208FCDEF96ABA14FAE8" bold="true" box="[779,804,1716,1740]" pageId="7" pageNumber="374">A.</emphasis>
<taxonomicName id="17D30DC4B571A208FCFFF96BBD10FAE9" authorityName="Padilla-Gil" authorityYear="2011" box="[810,1056,1717,1741]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="374" phylum="Arthropoda" rank="species" species="gastrotricha">
<emphasis id="E2A7AA55B571A208FCFFF96BBD10FAE9" box="[810,1056,1717,1741]" italics="true" pageId="7" pageNumber="374">Rhagovelia gastrotricha</emphasis>
</taxonomicName>
;
<emphasis id="E2A7AA55B571A208FBFBF96BBD75FAE9" bold="true" box="[1070,1093,1717,1741]" pageId="7" pageNumber="374">B.</emphasis>
<taxonomicName id="17D30DC4B571A208FB9EF96BBD8DFAE9" authorityName="Padilla-Gil" authorityYear="2011" box="[1099,1213,1717,1741]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="374" phylum="Arthropoda" rank="species" species="calceola">
<emphasis id="E2A7AA55B571A208FB9EF96BBD8DFAE9" box="[1099,1213,1717,1741]" italics="true" pageId="7" pageNumber="374">R. calceola</emphasis>
</taxonomicName>
;
<emphasis id="E2A7AA55B571A208FB1FF96ABDD3FAE8" bold="true" box="[1226,1251,1716,1740]" pageId="7" pageNumber="374">C.</emphasis>
<taxonomicName id="17D30DC4B571A208FB3CF96BBC69FAE9" authorityName="Galindo-Malagon, Morales &amp; Moreira" authorityYear="2021" box="[1257,1369,1717,1741]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="374" phylum="Arthropoda" rank="species" species="molanoi">
<emphasis id="E2A7AA55B571A208FB3CF96BBC69FAE9" box="[1257,1369,1717,1741]" italics="true" pageId="7" pageNumber="374">R. molanoi</emphasis>
</taxonomicName>
.
</paragraph>
</caption>
<subSubSection id="98C925CCB571A208FF12F8C1BAF2FBED" pageId="7" pageNumber="374" type="discussion">
<paragraph id="D06C7647B571A208FF12F8C1BAF2FBED" blockId="7.[151,1436,1823,1993]" pageId="7" pageNumber="374">
<emphasis id="E2A7AA55B571A208FF12F8C1B87FFB1D" bold="true" box="[199,335,1823,1849]" pageId="7" pageNumber="374">Comments.</emphasis>
Following the procedure established by D.
<bibRefCitation id="B4420BB6B571A208FC82F8C1BD2DFB1D" author="Polhemus, D. A." box="[855,1053,1823,1849]" pageId="7" pageNumber="374" refId="ref12676" refString="Polhemus, D. A. (1997) Systematics of the genus Rhagovelia Mayr (Heteroptera: Velliidae) in the Western Hemisphere (exclusive of the angustipes complex). Entomological Society of America, Lanham, Maryland, 386 pp. https: // doi. org / 10.4182 / JCRD 5977" type="book" year="1997">Polhemus (1997)</bibRefCitation>
, Padilla-Gil (2011) provisionally assigned
<taxonomicName id="17D30DC4B571A208FED5F89AB84FFB79" authorityName="Padilla-Gil" authorityYear="2011" box="[256,383,1859,1885]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="374" phylum="Arthropoda" rank="species" species="calceola">
<emphasis id="E2A7AA55B571A208FED5F89AB84FFB79" box="[256,383,1859,1885]" italics="true" pageId="7" pageNumber="374">R. calceola</emphasis>
</taxonomicName>
to the
<taxonomicName id="17D30DC4B571A208FE1AF89DBB07FB79" authorityName="Bacon" authorityYear="1948" box="[463,567,1859,1885]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="374" phylum="Arthropoda" rank="species" species="bisignata">
<emphasis id="E2A7AA55B571A208FE1AF89DBB07FB79" box="[463,567,1859,1885]" italics="true" pageId="7" pageNumber="374">bisignata</emphasis>
</taxonomicName>
group due to the lack of macropterous specimens. Subsequently,
<bibRefCitation id="B4420BB6B571A208FAF4F89DB9D2FBA6" author="Padilla-Gil, D. N." pageId="7" pageNumber="374" pagination="71 - 95" refId="ref11980" refString="Padilla-Gil, D. N. (2015) Ten new species of Rhagovelia in the angustipes complex (Hemiptera: Heteroptera: Veliidae) from Colombia, with a key to the Colombian species. Zootaxa, 4059 (1), 71 - 95. https: // doi. org / 10.11646 / zootaxa. 4059.1.4" type="journal article" year="2015">Padilla-Gil (2015)</bibRefCitation>
described
<taxonomicName id="17D30DC4B571A208FE89F8B6B8E2FBA5" authorityName="Padilla-Gil" authorityYear="2015" box="[348,466,1896,1921]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="374" phylum="Arthropoda" rank="species" species="tricoma">
<emphasis id="E2A7AA55B571A208FE89F8B6B8E2FBA5" box="[348,466,1896,1921]" italics="true" pageId="7" pageNumber="374">R. tricoma</emphasis>
</taxonomicName>
in the
<emphasis id="E2A7AA55B571A208FDF4F8B9BBAFFBA5" box="[545,671,1895,1921]" italics="true" pageId="7" pageNumber="374">hambletoni</emphasis>
group, because the macropterous forms displayed three closed cells on the forewings. Because
<bibRefCitation id="B4420BB6B571A208FE14F855BA25FB81" author="Galindo-Malagon, X. A. &amp; Morales, I. &amp; Moreira, F. F. F." box="[449,789,1931,1957]" pageId="7" pageNumber="374" pagination="167 - 225" refId="ref11045" refString="Galindo-Malagon, X. A., Morales, I. &amp; Moreira, F. F. F. (2021) Revision of the Rhagovelia angustipes complex (Insecta: Hemiptera: Veliidae) from Colombia. Zootaxa, 4958 (1), 167 - 225. https: // doi. org / 10.11646 / zootaxa. 4958.1.11" type="journal article" year="2021">
Galindo-Malagón
<emphasis id="E2A7AA55B571A208FD5BF852BBF5FB81" box="[654,709,1931,1957]" italics="true" pageId="7" pageNumber="374">et al.</emphasis>
(2021)
</bibRefCitation>
synonymized both species,
<taxonomicName id="17D30DC4B571A208FB99F852BDF8FB81" authorityName="Padilla-Gil" authorityYear="2011" box="[1100,1224,1931,1957]" class="Insecta" family="Veliidae" genus="Rhagovelia" kingdom="Animalia" order="Hemiptera" pageId="7" pageNumber="374" phylum="Arthropoda" rank="species" species="calceola">
<emphasis id="E2A7AA55B571A208FB99F852BDF8FB81" box="[1100,1224,1931,1957]" italics="true" pageId="7" pageNumber="374">R. calceola</emphasis>
</taxonomicName>
can be definitively placed in the
<emphasis id="E2A7AA55B571A208FEF8F871B89BFBED" box="[301,427,1967,1993]" italics="true" pageId="7" pageNumber="374">hambletoni</emphasis>
group, which was overlooked by these authors.
</paragraph>
</subSubSection>
</treatment>
</document>
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