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<document id="253A86D11A60FB887A55E085639A6898" ID-DOI="10.11646/phytotaxa.229.1.1" ID-ISSN="1179-3163" ID-Zenodo-Dep="13632375" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatmentCitations_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1725316854097" checkinUser="felipe" docAuthor="Chen, Xin, He, Hai &amp; Zhang, Li-Bing" docDate="2015" docId="03F887E9FFE9C76DFF03FE2AFA373956" docLanguage="en" docName="phytotaxa.229.1.1.pdf" docOrigin="Phytotaxa 229 (1)" docSource="http://dx.doi.org/10.11646/phytotaxa.229.1.1" docStyle="DocumentStyle:F08184CE06D8A97EA3E6DE35D99648B0.4:Phytotaxa.2014-.monograph" docStyleId="F08184CE06D8A97EA3E6DE35D99648B0" docStyleName="Phytotaxa.2014-.monograph" docStyleVersion="4" docTitle="Anisophyllea beccariana Baillon 1875" docType="treatment" docVersion="2" lastPageNumber="450" masterDocId="FFC1FF91FFFBC77EFF94FFF6FFAA3111" masterDocTitle="A monograph of the Anisophylleaceae (Cucurbitales) with description of 18 new species of Anisophyllea" masterLastPageNumber="450" masterPageNumber="448" pageNumber="448" updateTime="1725318148254" updateUser="ExternalLinkService">
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<mods:title id="AE699A6CC1FEC643A4D734A1835E465F">A monograph of the Anisophylleaceae (Cucurbitales) with description of 18 new species of Anisophyllea</mods:title>
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<mods:name id="580534FE2AAA338BE01C28479CADD62A" type="personal">
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<mods:namePart id="4909F47BB1502106BD204D7490DC0AC8">Chen, Xin</mods:namePart>
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<mods:namePart id="AFD5C181EB9041E6FD8AEAB5228B183B">He, Hai</mods:namePart>
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<mods:name id="E62165967A568D5D15F730B4408426DE" type="personal">
<mods:role id="404BABCB951E2FFACC6E41B4B608E467">
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<mods:namePart id="49ABADB6EF8883B9AAD39DCF9F004233">Zhang, Li-Bing</mods:namePart>
<mods:affiliation id="9B2B20DA1C4EEF3F1B250F1348D2966C">1 Department of Botany, College of Boology and the Environment, Nanjing Forestry University, 159 Longpan Rd., Xuanwu Qu, Nanjing, 210037, P. R. China 2 College of Life Sciences, Chongqing Normal University, Shapingba, Chongqing 400047, P. R. China 3 Missouri Botanical Garden, P. O. Box 299, St. Louis, Missouri 63166 - 0299, U. S. A. and Chengdu Institute of Biology, Chinese Academy of Sciences, P. O. Box 416, Chengdu, Sichuan 610041, P. R. China * Authors for correspondence: e-mails: hehaicq @ yahoo. com; libing. zhang @ mobot. org</mods:affiliation>
<mods:nameIdentifier id="B01083C6E57C4FFE7CDD3D8EA3EBD674" type="email">hehaicq@yahoo.com</mods:nameIdentifier>
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<mods:title id="BEC32C07CFD89C757A08F9DEE9721DEB">Phytotaxa</mods:title>
</mods:titleInfo>
<mods:part id="0D46CACF6AA59A2821B28B622A763C73">
<mods:date id="9F6A8C0CA1E41B982D900A3FFC972A8E">2015</mods:date>
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<mods:number id="4D39750D1AA0D6C1741F5C9229992124">2015-10-02</mods:number>
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<mods:number id="7CC83DEF4EA654D61FBC4421D900A236">229</mods:number>
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<mods:classification id="6B1BF68BE3F6B9AA3F906C8BE8D51FFF">journal article</mods:classification>
<mods:identifier id="3AD0DF062F385849A17B3AB7A1AAE01A" type="DOI">10.11646/phytotaxa.229.1.1</mods:identifier>
<mods:identifier id="780438B2AF2565ECA1E8E6BFDB8BC052" type="ISSN">1179-3163</mods:identifier>
<mods:identifier id="E1D5AE1CDD91F6830AF7F9BBB9C58F53" type="Zenodo-Dep">13632375</mods:identifier>
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<treatment id="03F887E9FFE9C76DFF03FE2AFA373956" LSID="urn:lsid:plazi:treatment:03F887E9FFE9C76DFF03FE2AFA373956" httpUri="http://treatment.plazi.org/id/03F887E9FFE9C76DFF03FE2AFA373956" lastPageId="19" pageId="18">
<subSubSection id="C34B6574FFE9C76CFF03FE2AFC8130E4" box="[151,811,475,501]" pageId="18" type="nomenclature">
<paragraph id="8BEE36FFFFE9C76CFF03FE2AFC8130E4" blockId="18.[151,811,475,501]" box="[151,811,475,501]" pageId="18">
<emphasis id="B925EAEDFFE9C76CFF03FE2AFF0630E4" bold="true" box="[151,172,476,501]" pageId="18">3.</emphasis>
<taxonomicName id="4C514D7CFFE9C76CFF27FE2DFD1930E4" ID-CoL="673FL" authority="Baillon (1875: 311)" authorityName="Baillon" authorityPageNumber="311" authorityYear="1875" box="[179,691,475,501]" class="Magnoliopsida" family="Anisophylleaceae" genus="Anisophyllea" kingdom="Plantae" order="Cucurbitales" pageId="18" phylum="Tracheophyta" rank="species" species="beccariana">
<emphasis id="B925EAEDFFE9C76CFF27FE2DFE7B30E5" bold="true" box="[179,465,475,500]" italics="true" pageId="18">Anisophyllea beccariana</emphasis>
<bibRefCitation id="EFC04B0EFFE9C76CFE43FE2AFD1930E4" author="Baillon, H. E." box="[471,691,476,501]" pageId="18" pagination="292 - 312" refId="ref96116" refString="Baillon, H. E. (1875) Stirpes exoticae novae. Adansonia 11: 292 - 312." type="journal article" year="1875">Baillon (1875: 311)</bibRefCitation>
</taxonomicName>
(
<figureCitation id="136A2A7AFFE9C76CFD55FE2AFC8830E4" box="[705,802,476,501]" captionStart="FIGURE 6" captionStartId="19.[151,244,1782,1803]" captionTargetBox="[151,1421,194,1753]" captionTargetId="figure-105@19.[151,1436,179,1775]" captionTargetPageId="19" captionText="FIGURE 6. Anisophyllea beccariana Baillon. Flowering branch. (O. Beccari 1001; lectotype, K-000493121)." figureDoi="http://doi.org/10.5281/zenodo.13632391" httpUri="https://zenodo.org/record/13632391/files/figure.png" pageId="18">Figure 6</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C34B6574FFE9C76CFF03FDD5FB3D34EC" pageId="18" type="materials_examined">
<paragraph id="8BEE36FFFFE9C76CFF03FDD5FC303346" blockId="18.[151,1436,547,599]" pageId="18">
<materialsCitation id="3B393CA2FFE9C76CFF03FDD5FC3C3346" collectionCode="O" country="Indonesia" location="Borneo" municipality="Type" pageId="18" specimenCount="2" stateProvince="Kalimantan Barat" typeStatus="lectotype">
<collectingMunicipality id="6B8AAC85FFE9C76CFF03FDD5FF633329" box="[151,201,547,568]" pageId="18">Type</collectingMunicipality>
:—
<collectingCountry id="F346766FFFE9C76CFF70FDD5FEC03329" box="[228,362,547,568]" name="Indonesia" pageId="18">INDONESIA</collectingCountry>
.?
<collectingRegion id="4995F81DFFE9C76CFEEAFDD5FD873329" box="[382,557,547,568]" country="Indonesia" name="Kalimantan Barat" pageId="18">West Kalimantan</collectingRegion>
:
<location id="8E8E6024FFE9C76CFDACFDD5FD2B3329" LSID="urn:lsid:plazi:treatment:03F887E9FFE9C76DFF03FE2AFA373956:8E8E6024FFE9C76CFDACFDD5FD2B3329" box="[568,641,547,568]" country="Indonesia" municipality="Type" name="Borneo" pageId="18" stateProvince="Kalimantan Barat">Borneo</location>
, 18651868,
<emphasis id="B925EAEDFFE9C76CFC9CFDD5FC013329" box="[776,939,547,568]" italics="true" pageId="18">
<collectionCode id="ED40AE3AFFE9C76CFC9CFDD5FCB63329" box="[776,796,547,568]" country="Norway" lsid="urn:lsid:biocol.org:col:13093" name="Botanical Museum - University of Oslo" pageId="18" type="Herbarium">O</collectionCode>
. Beccari 1001
</emphasis>
[
<typeStatus id="54EA885DFFE9C76CFC2DFDD5FBBE3329" box="[953,1044,547,568]" pageId="18" type="lectotype">lectotype</typeStatus>
K-000493121!, designated by
<bibRefCitation id="EFC04B0EFFE9C76CFADCFDD5FED73346" author="Wong, K. M. &amp; Madani, L." pageId="18" refId="ref101455" refString="Wong, K. M. &amp; Madani, L. (1995) Anisophylleaceae. In: Soepadmo, E. &amp; Wong, K. M. (Eds.) Tree Flora of Sabah and Sarawak. Vol. 1. Ampang Press Sdn. Bhd., Kuala Lumpur, pp. 15 - 26." type="book" year="1995">Wong &amp; Madani (1995: 18)</bibRefCitation>
,
<typeStatus id="54EA885DFFE9C76CFE12FDB4FDA93346" box="[390,515,578,599]" pageId="18" type="isolectotype">isolectotypes</typeStatus>
FI-008732!, G-00441118!, P-00641657!]
</materialsCitation>
.
</paragraph>
<paragraph id="8BEE36FFFFE9C76CFF03FD73FD1C34C9" blockId="18.[151,1437,644,1965]" pageId="18">
<emphasis id="B925EAEDFFE9C76CFF03FD73FF7E338D" box="[151,212,645,668]" italics="true" pageId="18">Trees</emphasis>
to
<quantity id="4CA99B1AFFE9C76CFF6EFD72FE9E338A" box="[250,308,644,669]" metricMagnitude="1" metricUnit="m" metricValue="2.5" pageId="18" unit="m" value="25.0">25 m</quantity>
tall, trunk slender; bark rolled or scaly, inner bark brownish; sapwood yellowish; young branches terete, glabrous; buds glabrous.
<emphasis id="B925EAEDFFE9C76CFE68FD5FFDE033D1" box="[508,586,681,704]" italics="true" pageId="18">Leaves</emphasis>
dimorphic, internodes between similar
<typeStatus id="54EA885DFFE9C76CFB93FD5EFBE833D1" box="[1031,1090,680,704]" pageId="18">types</typeStatus>
of leaves
<quantity id="4CA99B1AFFE9C76CFB20FD51FA9E33AE" box="[1204,1332,679,704]" metricMagnitude="-2" metricUnit="m" metricValue="1.85" metricValueMax="2.5" metricValueMin="1.2" pageId="18" unit="cm" value="1.85" valueMax="2.5" valueMin="1.2">1.22.5 cm</quantity>
, between two adjacent different
<typeStatus id="54EA885DFFE9C76CFE08FD3AFE7233F5" box="[412,472,716,740]" pageId="18">types</typeStatus>
of leaves
<quantity id="4CA99B1AFFE9C76CFDC4FD3AFD1933F2" box="[592,691,716,741]" metricMagnitude="-3" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="18" unit="mm" value="4.0" valueMax="5.0" valueMin="3.0">35 mm</quantity>
;
<emphasis id="B925EAEDFFE9C76CFD2BFD3AFCE733F4" box="[703,845,716,741]" italics="true" pageId="18">small leaves</emphasis>
caducous, only leaving small scars on twigs;
<emphasis id="B925EAEDFFE9C76CFAF5FD3AFF753219" italics="true" pageId="18">large leaves</emphasis>
petiolate, petiole
<quantity id="4CA99B1AFFE9C76CFE24FD19FDBA3216" box="[432,528,751,776]" metricMagnitude="-3" metricUnit="m" metricValue="6.0" metricValueMax="8.0" metricValueMin="4.0" pageId="18" unit="mm" value="6.0" valueMax="8.0" valueMin="4.0">48 mm</quantity>
long,
<quantity id="4CA99B1AFFE9C76CFDCFFD19FD4F3216" box="[603,741,751,776]" metricMagnitude="-3" metricUnit="m" metricValue="1.5499999999999998" metricValueMax="1.7" metricValueMin="1.4" pageId="18" unit="mm" value="1.5499999999999998" valueMax="1.7" valueMin="1.4">1.41.7 mm</quantity>
in diam., glabrous; leaf blade elliptic to ovate, 6.511.0 cm long,
<quantity id="4CA99B1AFFE9C76CFF4DFCE2FE84323A" box="[217,302,788,813]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="18" unit="cm" value="4.0" valueMax="5.0" valueMin="3.0">35 cm</quantity>
wide, base slightly oblique, acute to obtuse, apex acuminate to caudate, 0.82.0 cm long, margins often slightly revolute, thinly coriaceous, lustrous and bright green adaxially, dull and pale green abaxially, glabrous on both surfaces; main longitudinal veins 56, springing from blade base or ± from midrib a litter distance above blade base, midrib straight and bold, impressed adaxially and raised abaxially, lateral longitudinal main veins rather fine, outmost two lateral veins very close to blade margins and one often disappearing into blade margin when 6 main veins present, slightly prominent on both surfaces; transverse veins irregular, some of those from midrib and curved toward blade apex rather conspicuous and forming a pinnate-like venation; veinlets reticulate, slightly prominent on both surfaces.
<emphasis id="B925EAEDFFE9C76CFD33FBF9FC973539" box="[679,829,1039,1064]" italics="true" pageId="18">Inflorescence</emphasis>
axillary or slightly supra-axillary, in serials, simple or proximally branched, with female and male flowers; rachis short, tenuous,
<quantity id="4CA99B1AFFE9C76CFC70FBC2FBC8355A" box="[996,1122,1076,1101]" metricMagnitude="-2" metricUnit="m" metricValue="3.0" metricValueMax="4.5" metricValueMin="1.5" pageId="18" unit="cm" value="3.0" valueMax="4.5" valueMin="1.5">1.54.5 cm</quantity>
long,
<quantity id="4CA99B1AFFE9C76CFB3EFBC2FA98355A" box="[1194,1330,1076,1101]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" metricValueMax="6.0" metricValueMin="4.0" pageId="18" unit="mm" value="0.5" valueMax="0.6" valueMin="0.4">0.40.6 mm</quantity>
in diam., pannose with yellowish hairs
<quantity id="4CA99B1AFFE9C76CFE61FBA1FD2B357E" box="[501,641,1111,1136]" metricMagnitude="-4" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="18" unit="mm" value="0.15" valueMax="0.2" valueMin="0.1">0.10.2 mm</quantity>
long, remotely flowered with floral internodes
<quantity id="4CA99B1AFFE9C76CFB26FBA1FA94357E" box="[1202,1342,1111,1136]" metricMagnitude="-3" metricUnit="m" metricValue="1.1" metricValueMax="1.6" metricValueMin="0.6" pageId="18" unit="mm" value="1.1" valueMax="1.6" valueMin="0.6">0.61.6 mm</quantity>
distant; bracts deltoid, 0.51.0 mm long, ca.
<quantity id="4CA99B1AFFE9C76CFDAFFB8AFD393582" box="[571,659,1148,1173]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="18" unit="mm" value="0.5">0.5 mm</quantity>
wide, glabrous adaxially, pannose abaxially, margins ciliate; flower buds pannose;
<emphasis id="B925EAEDFFE9C76CFEA8FB69FE2535A9" box="[316,399,1183,1208]" italics="true" pageId="18">flowers</emphasis>
unisexual, 4-merous, brightly yellow;
<emphasis id="B925EAEDFFE9C76CFCD4FB69FC4E35A9" box="[832,996,1183,1208]" italics="true" pageId="18">female flowers</emphasis>
2.0
<quantity id="4CA99B1AFFE9C76CFB88FB69FBDB35A6" box="[1052,1137,1183,1208]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="18" unit="mm" value="2.6">2.6 mm</quantity>
long, sessile or subsessile; receptacle obovate,
<quantity id="4CA99B1AFFE9C76CFEE9FB32FD8E35CA" box="[381,548,1220,1245]" metricMagnitude="-3" metricUnit="m" metricValue="1.44" metricValueMax="1.6" metricValueMin="1.28" pageId="18" unit="mm" value="1.44" valueMax="1.6" valueMin="1.28">1.281.60 mm</quantity>
long, ca.
<quantity id="4CA99B1AFFE9C76CFD03FB32FD7335CA" box="[663,729,1220,1245]" metricMagnitude="-3" metricUnit="m" metricValue="1.0" pageId="18" unit="mm" value="1.0">1 mm</quantity>
in diam., sparsely pannose; sepals deltoid, 0.61.0 mm long, nearly glabrous; petals oblong, ca.
<quantity id="4CA99B1AFFE9C76CFDD4FB11FD3035EE" box="[576,666,1255,1280]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="18" unit="mm" value="0.6">0.6 mm</quantity>
long, slightly emarginate or shortly 2-lobed, margins incurved; staminodes 8, ca.
<quantity id="4CA99B1AFFE9C76CFEF4FAFAFE1D3432" box="[352,439,1292,1317]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="18" unit="mm" value="0.5">0.5 mm</quantity>
long; styles 4, free, ca.
<quantity id="4CA99B1AFFE9C76CFD51FAFAFCB63432" box="[709,796,1292,1317]" metricMagnitude="-4" metricUnit="m" metricValue="6.0" pageId="18" unit="mm" value="0.6">0.6 mm</quantity>
long, base conical ca.
<quantity id="4CA99B1AFFE9C76CFB88FAFAFB2B3432" box="[1052,1153,1292,1317]" metricMagnitude="-4" metricUnit="m" metricValue="2.5" pageId="18" unit="mm" value="0.25">0.25 mm</quantity>
in diam., pubescent with brownish hairs, distally attenuate, apex recurved;
<emphasis id="B925EAEDFFE9C76CFD40FAD9FCCD3459" box="[724,871,1327,1352]" italics="true" pageId="18">male flowers</emphasis>
to
<quantity id="4CA99B1AFFE9C76CFC19FAD9FC4F3456" box="[909,997,1327,1352]" metricMagnitude="-3" metricUnit="m" metricValue="1.5" pageId="18" unit="mm" value="1.5">1.5 mm</quantity>
long, hairy as female flowers; shortly pedicelled, pedicel
<quantity id="4CA99B1AFFE9C76CFEFBFAA2FDBB347A" box="[367,529,1364,1389]" metricMagnitude="-4" metricUnit="m" metricValue="5.75" metricValueMax="6.5" metricValueMin="5.0" pageId="18" unit="mm" value="0.575" valueMax="0.65" valueMin="0.5">0.500.65 mm</quantity>
long; receptacle cup-shaped; sepals and petals similar to female flowers; stamens 8, episepalous 4 fertile, ca.
<quantity id="4CA99B1AFFE9C76CFE5CFA81FD86349E" box="[456,556,1399,1424]" metricMagnitude="-4" metricUnit="m" metricValue="6.5" pageId="18" unit="mm" value="0.65">0.65 mm</quantity>
long, filaments compressed, anthers ca.
<quantity id="4CA99B1AFFE9C76CFC66FA81FBE2349E" box="[1010,1096,1399,1424]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="18" unit="mm" value="0.2">0.2 mm</quantity>
long, epipetalous 4 sterile and much shorter, ca.
<quantity id="4CA99B1AFFE9C76CFECBFA6AFE1C34A2" box="[351,438,1436,1461]" metricMagnitude="-4" metricUnit="m" metricValue="5.0" pageId="18" unit="mm" value="0.5">0.5 mm</quantity>
long, glandular at apex; disk obscure; pistil rudimentary; styles 4, free, shortly subulate, ca.
<quantity id="4CA99B1AFFE9C76CFF2AFA49FEBE34C6" box="[190,276,1471,1496]" metricMagnitude="-4" metricUnit="m" metricValue="3.0" pageId="18" unit="mm" value="0.3">0.3 mm</quantity>
long, apex recurved.
<emphasis id="B925EAEDFFE9C76CFD92FA37FDEA34C9" box="[518,576,1473,1496]" italics="true" pageId="18">Fruit</emphasis>
unknown.
</paragraph>
<paragraph id="8BEE36FFFFE9C76CFF52FA12FB3D34EC" blockId="18.[151,1437,644,1965]" box="[198,1175,1508,1533]" pageId="18">
<emphasis id="B925EAEDFFE9C76CFF52FA12FE4A34EC" bold="true" box="[198,480,1508,1533]" pageId="18">Flowering and fruiting:</emphasis>
—Flowering in October to March, fruiting in April to August.
</paragraph>
</subSubSection>
<subSubSection id="C34B6574FFE9C76CFF52F9F1FA0A395E" pageId="18" type="description">
<paragraph id="8BEE36FFFFE9C76CFF52F9F1FDAA3779" blockId="18.[151,1437,644,1965]" pageId="18">
<emphasis id="B925EAEDFFE9C76CFF52F9F1FDAA3731" bold="true" box="[198,512,1543,1568]" pageId="18">Habitat and distribution:</emphasis>
—In lowland forests, especially mixed with
<taxonomicName id="4C514D7CFFE9C76CFB81F9FEFAFE3731" authority="C. F. Gaertner" authorityName="C. F. Gaertner" box="[1045,1364,1543,1568]" class="Magnoliopsida" family="Dipterocarpaceae" genus="Dipterocarpus" kingdom="Plantae" order="Malvales" pageId="18" phylum="Tracheophyta" rank="genus">
<emphasis id="B925EAEDFFE9C76CFB81F9FEFB1D370E" box="[1045,1207,1544,1567]" italics="true" pageId="18">Dipterocarpus</emphasis>
C.F.Gaertner
</taxonomicName>
or in
<emphasis id="B925EAEDFFE9C76CFF03F9DAFEAE3754" box="[151,260,1580,1605]" italics="true" pageId="18">kerangas</emphasis>
(heath) forests; below
<specimenCount id="9D57FD76FFE9C76CFD89F9DAFDDA3752" box="[541,624,1580,1605]" count="600" pageId="18" type="generic">
<quantity id="4CA99B1AFFE9C76CFD89F9DAFDC63752" box="[541,620,1580,1605]" metricMagnitude="2" metricUnit="m" metricValue="6.0" pageId="18" unit="m" value="600.0">600 m</quantity>
.
</specimenCount>
<materialsCitation id="3B393CA2FFE9C76CFDEDF9DAFE563779" country="Brunei" location="West Kalimantan, Sumatra" pageId="18" specimenCount="1" stateProvince="West Kalimantan, Sumatra">
<collectingCountry id="F346766FFFE9C76CFDEDF9DAFD673754" box="[633,717,1580,1605]" name="Brunei" pageId="18">Brunei</collectingCountry>
;
<collectingCountry id="F346766FFFE9C76CFD4FF9DAFCE53754" box="[731,847,1580,1605]" name="Indonesia" pageId="18">Indonesia</collectingCountry>
(?
<collectingRegion id="4995F81DFFE9C76CFCFFF9DAFB1C3754" box="[875,1206,1580,1605]" isEnumeration="true" pageId="18">West Kalimantan, Sumatra</collectingRegion>
);
<collectingCountry id="F346766FFFE9C76CFB59F9DAFA933754" box="[1229,1337,1580,1605]" name="Malaysia" pageId="18">Malaysia</collectingCountry>
(
<collectingRegion id="4995F81DFFE9C76CFADFF9DAFA323754" box="[1355,1432,1580,1605]" country="Malaysia" name="Sabah" pageId="18">Sabah</collectingRegion>
, Sarawak);
<collectingCountry id="F346766FFFE9C76CFE9BF9B9FE2B3779" box="[271,385,1615,1640]" name="Singapore" pageId="18">Singapore</collectingCountry>
(
<figureCitation id="136A2A7AFFE9C76CFE1BF9B9FE5B3779" box="[399,497,1615,1640]" captionStart="FIGURE 7" captionStartId="20.[151,244,1056,1077]" captionTargetBox="[151,1436,191,1031]" captionTargetId="figure-440@20.[151,1436,191,1031]" captionTargetPageId="20" captionText="FIGURE 7. Geographical distribution of Anisophyllea beccariana Baillon." figureDoi="http://doi.org/10.5281/zenodo.13632393" httpUri="https://zenodo.org/record/13632393/files/figure.png" pageId="18">Figure 7</figureCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BEE36FFFFE9C76CFF52F982FD69379C" blockId="18.[151,1437,644,1965]" box="[198,707,1652,1677]" pageId="18">
<emphasis id="B925EAEDFFE9C76CFF52F982FE6E379C" bold="true" box="[198,452,1652,1677]" pageId="18">Vernacular names:—</emphasis>
<emphasis id="B925EAEDFFE9C76CFE51F983FE40379D" box="[453,490,1653,1676]" italics="true" pageId="18">Pei</emphasis>
(Kayan,
<collectingCountry id="F346766FFFE9C76CFDC6F982FD12379C" box="[594,696,1652,1677]" name="Malaysia" pageId="18">Sarawak</collectingCountry>
).
</paragraph>
<paragraph id="8BEE36FFFFE9C76CFF52F961FC1C36BC" blockId="18.[151,1437,644,1965]" pageId="18">
<emphasis id="B925EAEDFFE9C76CFF52F961FE3037A1" bold="true" box="[198,410,1687,1712]" pageId="18">Taxonomic notes:</emphasis>
—Although
<taxonomicName id="4C514D7CFFE9C76CFDBDF961FCE937A1" box="[553,835,1687,1712]" class="Magnoliopsida" family="Anisophylleaceae" genus="Anisophyllea" kingdom="Plantae" order="Cucurbitales" pageId="18" phylum="Tracheophyta" rank="species" species="beccariana">
<emphasis id="B925EAEDFFE9C76CFDBDF961FCE937A1" box="[553,835,1687,1712]" italics="true" pageId="18">Anisophyllea beccariana</emphasis>
</taxonomicName>
has obvious longitudinal lateral veins, its transverse veins arising from the midrib are so prominent, which makes the venation somewhat pinnate-like.
<bibRefCitation id="EFC04B0EFFE9C76CFB63F94AFA3637C4" author="Baillon, H. E." box="[1271,1436,1724,1749]" pageId="18" pagination="292 - 312" refId="ref96116" refString="Baillon, H. E. (1875) Stirpes exoticae novae. Adansonia 11: 292 - 312." type="journal article" year="1875">Baillon (1875)</bibRefCitation>
was quite right in that he compared this species with
<taxonomicName id="4C514D7CFFE9C76CFD67F916FCDB37E9" box="[755,881,1759,1784]" class="Magnoliopsida" family="Anisophylleaceae" genus="Anisophyllea" kingdom="Plantae" order="Cucurbitales" pageId="18" phylum="Tracheophyta" rank="species" species="griffithii">
<emphasis id="B925EAEDFFE9C76CFD67F916FCDB37E9" box="[755,881,1759,1784]" italics="true" pageId="18">A. griffithii</emphasis>
</taxonomicName>
, but he did not attach importance of the pinnate-like venation of this species. Neither
<bibRefCitation id="EFC04B0EFFE9C76CFDAFF8F2FD57360C" author="Ding Hou" box="[571,765,1796,1821]" pageId="18" pagination="429 - 493" refId="ref97361" refString="Ding Hou (1958) Rhizophoraceae. In: Van Steenis, C. G. G .. J. (Ed.) Flora Malesiana. Series 1, vol. 5. Noordhoff-kolff N. V., Djakarta, pp. 429 - 493. http: // dx. doi. org / 10.5962 / bhl. title. 40744" type="book chapter" year="1958">Ding Hou (1958)</bibRefCitation>
nor
<bibRefCitation id="EFC04B0EFFE9C76CFCA6F8F2FBEB360C" author="Wong, K. M. &amp; Madani, L." box="[818,1089,1796,1821]" pageId="18" refId="ref101455" refString="Wong, K. M. &amp; Madani, L. (1995) Anisophylleaceae. In: Soepadmo, E. &amp; Wong, K. M. (Eds.) Tree Flora of Sabah and Sarawak. Vol. 1. Ampang Press Sdn. Bhd., Kuala Lumpur, pp. 15 - 26." type="book" year="1995">Wong &amp; Madani (1995)</bibRefCitation>
used the character of venation in their keys to species. Morphologically this species differs from
<taxonomicName id="4C514D7CFFE9C76CFC10F8DEFBA83651" box="[900,1026,1831,1856]" class="Magnoliopsida" family="Anisophylleaceae" genus="Anisophyllea" kingdom="Plantae" order="Cucurbitales" pageId="18" phylum="Tracheophyta" rank="species" species="griffithii">
<emphasis id="B925EAEDFFE9C76CFC10F8DEFBA83651" box="[900,1026,1831,1856]" italics="true" pageId="18">A. griffithii</emphasis>
</taxonomicName>
in having longer petioles, unisexual flowers, and glabrous young branches. In addition, this species often has longer and broader leaves than
<taxonomicName id="4C514D7CFFE9C76CFA17F8BAFF5F3699" class="Magnoliopsida" family="Anisophylleaceae" genus="Anisophyllea" kingdom="Plantae" order="Cucurbitales" pageId="18" phylum="Tracheophyta" rank="species" species="griffithii">
<emphasis id="B925EAEDFFE9C76CFA17F8BAFF5F3699" italics="true" pageId="18">A. griffithii</emphasis>
</taxonomicName>
. Its distribution in
<collectingCountry id="F346766FFFE9C76CFE45F899FDEE3699" box="[465,580,1903,1928]" name="Singapore" pageId="18">Singapore</collectingCountry>
is based on our examination of specimens cited below and that in Sumatra (unspecified province in CE Sumatra) follows Wong &amp; Madnai (1995).
</paragraph>
<paragraph id="8BEE36FFFFE9C76CFF01F7C0FDE5395A" blockId="18.[149,591,2102,2123]" box="[149,591,2102,2123]" pageId="18">
MONOGRAPH OF
<taxonomicName id="4C514D7CFFE9C76CFECEF7C0FDE5395A" authorityName="Ridley" authorityYear="1922" box="[346,591,2102,2123]" class="Magnoliopsida" family="Anisophylleaceae" kingdom="Plantae" order="Cucurbitales" pageId="18" phylum="Tracheophyta" rank="family">ANISOPHYLLEACEAE</taxonomicName>
</paragraph>
<paragraph id="8BEE36FFFFE9C76CFC28F7C1FA0A395E" blockId="18.[956,1440,2098,2127]" box="[956,1440,2098,2127]" pageId="18">
<emphasis id="B925EAEDFFE9C76CFC28F7C1FBB7395D" box="[956,1053,2103,2124]" italics="true" pageId="18">Phytotaxa</emphasis>
229 (1) © 2015 Magnolia Press •
<emphasis id="B925EAEDFFE9C76CFA14F7C4FA0A395E" bold="true" box="[1408,1440,2098,2127]" pageId="18">19</emphasis>
</paragraph>
</subSubSection>
<caption id="DF2E6677FFE8C76DFF03F900FB10361A" ID-DOI="http://doi.org/10.5281/zenodo.13632391" ID-Zenodo-Dep="13632391" box="[151,1210,1782,1803]" httpUri="https://zenodo.org/record/13632391/files/figure.png" pageId="19" startId="19.[151,244,1782,1803]" targetBox="[151,1421,194,1753]" targetPageId="19" targetType="figure">
<paragraph id="8BEE36FFFFE8C76DFF03F900FB10361A" blockId="19.[151,1210,1782,1803]" box="[151,1210,1782,1803]" pageId="19">
<emphasis id="B925EAEDFFE8C76DFF03F900FEA6361A" bold="true" box="[151,268,1782,1803]" pageId="19">FIGURE 6.</emphasis>
<taxonomicName id="4C514D7CFFE8C76DFE86F900FD17361A" authority="Baillon. Flowering" authorityName="Baillon. Flowering" box="[274,701,1782,1803]" class="Magnoliopsida" family="Anisophylleaceae" genus="Anisophyllea" kingdom="Plantae" order="Cucurbitales" pageId="19" phylum="Tracheophyta" rank="species" species="beccariana">
<emphasis id="B925EAEDFFE8C76DFE86F900FDAB361A" box="[274,513,1782,1803]" italics="true" pageId="19">Anisophyllea beccariana</emphasis>
Baillon. Flowering
</taxonomicName>
branch. (
<emphasis id="B925EAEDFFE8C76DFC83F900FC1F361A" box="[791,949,1782,1803]" italics="true" pageId="19">O. Beccari 1001</emphasis>
; lectotype, K-000493121).
</paragraph>
</caption>
<subSubSection id="C34B6574FFE8C76DFF52F8CCFDD63956" pageId="19" type="materials_examined">
<paragraph id="8BEE36FFFFE8C76DFF52F8CCFCC336F2" blockId="19.[151,1436,1850,2019]" pageId="19">
<emphasis id="B925EAEDFFE8C76DFF52F8CCFDE33642" bold="true" box="[198,585,1850,1875]" pageId="19">Additional specimens examined:</emphasis>
<materialsCitation id="3B393CA2FFE8C76DFDF2F8CCFA32368A" collectingDate="1960-08-12" collectingDateMax="1996-10-21" collectingDateMin="1960-08-12" collectionCode="K, J, L, NY, F, R, S" collectorName="Sinclair &amp; Kadim bin Tassim" country="Malaysia" county="Kampung Nalumad" location="Baka National Park" municipality="Daim Andau" pageId="19" specimenCount="1" stateProvince="Sabah">
<collectingCountry id="F346766FFFE8C76DFDF2F8CCFD573642" box="[614,765,1850,1875]" name="Malaysia" pageId="19">MALAYSIA</collectingCountry>
.
<collectingRegion id="4995F81DFFE8C76DFC93F8CCFCFA3642" box="[775,848,1850,1875]" country="Malaysia" name="Sabah" pageId="19">Sabah</collectingRegion>
:
<collectingCounty id="628F4E73FFE8C76DFCCEF8CDFB933642" box="[858,1081,1850,1875]" pageId="19">Kampung Nalumad</collectingCounty>
,
<date id="FFEF103FFFE8C76DFBD0F8CCFAA93642" box="[1092,1283,1850,1875]" pageId="19" value="1996-10-21">
<collectingDate id="EFABE9D7FFE8C76DFBD0F8CCFAA93642" box="[1092,1283,1850,1875]" pageId="19" value="1996-10-21">21 October 1996</collectingDate>
</date>
,
<emphasis id="B925EAEDFFE8C76DFA9AF8CDFF6B3666" italics="true" pageId="19">
<collectingMunicipality id="6B8AAC85FFE8C76DFA9AF8CDFA363642" box="[1294,1436,1850,1875]" pageId="19">Daim Andau</collectingMunicipality>
877
</emphasis>
(
<collectionCode id="ED40AE3AFFE8C76DFF45F8A8FF433666" box="[209,233,1886,1911]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="19" type="Herbarium">K</collectionCode>
).
<collectingRegion id="4995F81DFFE8C76DFF69F8A8FEC83666" box="[253,354,1886,1911]" country="Malaysia" name="Sarawak" pageId="19">Sarawak</collectingRegion>
:
<location id="8E8E6024FFE8C76DFEF9F8A8FDE13666" LSID="urn:lsid:plazi:treatment:03F887E9FFE9C76DFF03FE2AFA373956:8E8E6024FFE8C76DFEF9F8A8FDE13666" box="[365,587,1886,1911]" country="Malaysia" county="Kampung Nalumad" municipality="Daim Andau" name="Baka National Park" pageId="19" stateProvince="Sabah">Baka National Park</location>
, Lintang path (north end),
<date id="FFEF103FFFE8C76DFCEDF8A8FB843666" box="[889,1070,1886,1911]" pageId="19" value="1960-08-12">
<collectingDate id="EFABE9D7FFE8C76DFCEDF8A8FB843666" box="[889,1070,1886,1911]" pageId="19" value="1960-08-12">12 August 1960</collectingDate>
</date>
,
<emphasis id="B925EAEDFFE8C76DFBAEF8A9FF77368A" italics="true" pageId="19">
<collectionCode id="ED40AE3AFFE8C76DFBAEF8A9FBE03667" box="[1082,1098,1887,1910]" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="19" type="Herbarium">J</collectionCode>
.
<collectorName id="26A45329FFE8C76DFBC1F8A8FB043666" box="[1109,1198,1886,1911]" pageId="19">Sinclair</collectorName>
&amp;
<collectorName id="26A45329FFE8C76DFB46F8A8FA363667" box="[1234,1436,1886,1911]" pageId="19">Kadim bin Tassim</collectorName>
10331
</emphasis>
(
<collectionCode id="ED40AE3AFFE8C76DFF78F874FEAA368A" box="[236,256,1922,1947]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="19" type="Herbarium">L</collectionCode>
,
<collectionCode id="ED40AE3AFFE8C76DFE9FF874FE92368A" box="[267,312,1922,1947]" country="USA" lsid="urn:lsid:biocol.org:col:15556" name="William and Lynda Steere Herbarium of the New York Botanical Garden" pageId="19" type="Herbarium">NY</collectionCode>
); Kuching, Semengoh
<collectionCode id="ED40AE3AFFE8C76DFDA1F875FDED368B" box="[565,583,1923,1946]" country="USA" lsid="urn:lsid:biocol.org:col:15707" name="Field Museum of Natural History, Botany Department" pageId="19" type="Herbarium">F</collectionCode>
.
<collectionCode id="ED40AE3AFFE8C76DFDD8F875FDC8368B" box="[588,610,1923,1946]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="19">R</collectionCode>
.,
<locationDeviation id="9A5C2FDDFFE8C76DFDE5F874FC3A368A" box="[625,912,1922,1947]" location="Penrissen Road" pageId="19">
<quantity id="4CA99B1AFFE8C76DFDE5F874FD70368A" box="[625,730,1922,1947]" metricMagnitude="4" metricUnit="m" metricValue="2.01168" pageId="19" unit="mi" value="12.5">12.5 mile</quantity>
Penrissen Road
</locationDeviation>
,
<date id="FFEF103FFFE8C76DFC0FF874FBE2368A" box="[923,1096,1922,1947]" pageId="19" value="1973-03-13">
<collectingDate id="EFABE9D7FFE8C76DFC0FF874FBE2368A" box="[923,1096,1922,1947]" pageId="19" value="1973-03-13">13 March 1973</collectingDate>
</date>
,
<emphasis id="B925EAEDFFE8C76DFBC6F874FAC1368A" box="[1106,1387,1922,1947]" italics="true" pageId="19">
Othman Ismawi
<collectionCode id="ED40AE3AFFE8C76DFA9DF875FAB0368B" box="[1289,1306,1923,1946]" country="Sweden" lsid="urn:lsid:biocol.org:col:15668" name="Department of Botany, Swedish Museum of Natural History" pageId="19" type="Herbarium">S</collectionCode>
. 32992
</emphasis>
(
<collectionCode id="ED40AE3AFFE8C76DFAEEF874FA25368A" box="[1402,1423,1922,1947]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="19" type="Herbarium">L</collectionCode>
)
</materialsCitation>
.
<materialsCitation id="3B393CA2FFE8C76DFF03F850FCCF36F2" collectingDate="1953-04-24" collectingDateMax="1954-02-20" collectingDateMin="1953-04-24" collectionCode="J, K, L" country="Singapore" location="Off" pageId="19" specimenCount="1">
<collectingCountry id="F346766FFFE8C76DFF03F850FEE836AE" box="[151,322,1958,1983]" name="Singapore" pageId="19">SINGAPORE</collectingCountry>
.
<location id="8E8E6024FFE8C76DFEDBF850FED236AE" LSID="urn:lsid:plazi:treatment:03F887E9FFE9C76DFF03FE2AFA373956:8E8E6024FFE8C76DFEDBF850FED236AE" box="[335,376,1958,1983]" country="Singapore" name="Off" pageId="19">Off</location>
<locationDeviation id="9A5C2FDDFFE8C76DFE15F850FD1A36AE" box="[385,688,1958,1983]" location="Mandai Road" pageId="19">
<quantity id="4CA99B1AFFE8C76DFE15F850FE5536AE" box="[385,511,1958,1983]" metricMagnitude="4" metricUnit="m" metricValue="1.8507456" pageId="19" unit="mi" value="11.5">11.5 miles</quantity>
Mandai Road
</locationDeviation>
,
<date id="FFEF103FFFE8C76DFD2BF850FC3136AE" box="[703,923,1958,1983]" pageId="19" value="1954-02-20">
<collectingDate id="EFABE9D7FFE8C76DFD2BF850FC3136AE" box="[703,923,1958,1983]" pageId="19" value="1954-02-20">20 February 1954</collectingDate>
</date>
,
<emphasis id="B925EAEDFFE8C76DFC3DF851FBD636AE" box="[937,1148,1958,1983]" italics="true" pageId="19">
<collectionCode id="ED40AE3AFFE8C76DFC3DF851FC1036AF" box="[937,954,1959,1982]" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="19" type="Herbarium">J</collectionCode>
. Sinclair 40205
</emphasis>
(
<collectionCode id="ED40AE3AFFE8C76DFB1BF850FB0236AE" box="[1167,1192,1958,1983]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:15867" name="Royal Botanic Gardens" pageId="19" type="Herbarium">K</collectionCode>
,
<collectionCode id="ED40AE3AFFE8C76DFB22F851FB6036AF" box="[1206,1226,1959,1982]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="19" type="Herbarium">L</collectionCode>
, US); South side MacRitchie Reservoir,
<date id="FFEF103FFFE8C76DFE0EF83CFD9136F2" box="[410,571,1994,2019]" pageId="19" value="1953-04-24">
<collectingDate id="EFABE9D7FFE8C76DFE0EF83CFD9136F2" box="[410,571,1994,2019]" pageId="19" value="1953-04-24">24 April 1953</collectingDate>
</date>
,
<emphasis id="B925EAEDFFE8C76DFDD1F83DFCAF36F2" box="[581,773,1994,2019]" italics="true" pageId="19">
<collectionCode id="ED40AE3AFFE8C76DFDD1F83DFDFF36F3" box="[581,597,1995,2018]" country="South Africa" lsid="urn:lsid:biocol.org:col:14647" name="University of the Witwatersrand" pageId="19" type="Herbarium">J</collectionCode>
. Sinclair 39586
</emphasis>
(
<collectionCode id="ED40AE3AFFE8C76DFC81F83CFC8336F2" box="[789,809,1994,2019]" country="Netherlands" lsid="urn:lsid:biocol.org:col:15678" name="Nationaal Herbarium Nederland, Leiden University branch" pageId="19" type="Herbarium">L</collectionCode>
,
<collectingCountry id="F346766FFFE8C76DFCA0F83CFCF636F2" box="[820,860,1994,2019]" name="United States of America" pageId="19">US</collectingCountry>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BEE36FFFFE8C76DFF03F7DAFDD63956" blockId="19.[151,636,2092,2121]" box="[151,636,2092,2121]" pageId="19">
<emphasis id="B925EAEDFFE8C76DFF03F7DAFF1D3958" bold="true" box="[151,183,2092,2121]" pageId="19">20</emphasis>
<emphasis id="B925EAEDFFE8C76DFF75F7C4FEE83956" box="[225,322,2098,2119]" italics="true" pageId="19">Phytotaxa</emphasis>
229 (1) © 2015 Magnolia Press
</paragraph>
</subSubSection>
<subSubSection id="C34B6574FFE8C76DFA85F7C4FA373956" box="[1297,1437,2098,2119]" pageId="19" type="description">
<paragraph id="8BEE36FFFFE8C76DFA85F7C4FA373956" blockId="19.[1297,1437,2098,2119]" box="[1297,1437,2098,2119]" pageId="19">
CHEN
<emphasis id="B925EAEDFFE8C76DFACCF7C4FA373956" box="[1368,1437,2098,2119]" italics="true" pageId="19">
<collectingRegion id="4995F81DFFE8C76DFACCF7C4FADE3956" box="[1368,1396,2098,2119]" country="Cameroon" name="East" pageId="19">ET</collectingRegion>
AL.
</emphasis>
</paragraph>
</subSubSection>
</treatment>
</document>