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<document id="A778BACF1B265315FAB29D4557ACA123" ID-DOI="10.24349/ynmN-vroH" ID-ISSN="2107-7207" ID-Zenodo-Dep="7159408" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1665193151235" checkinUser="felipe" docAuthor="Ermilov, Sergey G., Subias, Luis S., Shtanchaeva, Umukusum Ya. &amp; Friedrich, Stefan" docDate="2021" docId="03F1DC592632FFAEFE21802C1DCBC820" docLanguage="en" docName="Acarologia.61.3.591-601.pdf.pdf" docOrigin="Acarologia 61 (3)" docSource="http://www1.montpellier.inrae.fr/CBGP/acarologia/article.php?id=4457" docStyle="DocumentStyle:0387F17C4AF9F8D6336E46CC5E5664D0.6:Acarologia.2018-.journal_article" docStyleId="0387F17C4AF9F8D6336E46CC5E5664D0" docStyleName="Acarologia.2018-.journal_article" docStyleVersion="6" docTitle="Pilobates wachteli Ermilov &amp; Subias &amp; Shtanchaeva &amp; Friedrich 2021, n. sp." docType="treatment" docVersion="3" lastPageNumber="597" masterDocId="FFC8A4212636FFA9FFCA82411868CB69" masterDocTitle="New faunistical and taxonomic data on oribatid mites (Acari: Oribatida) of Ethiopia, with description of two new species of the superfamily Oripodoidea" masterLastPageNumber="601" masterPageNumber="591" pageNumber="594" updateTime="1699364423098" updateUser="plazi" zenodo-license-document="CC-BY-4.0">
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<mods:title id="1BD93E78FA5992BC3722E1AED6F9EDE9">New faunistical and taxonomic data on oribatid mites (Acari: Oribatida) of Ethiopia, with description of two new species of the superfamily Oripodoidea</mods:title>
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<mods:namePart id="8FAC4E638A36CAC31A3C70E30F2311E8">Ermilov, Sergey G.</mods:namePart>
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<mods:namePart id="7536EBB2767D9685F113BCEA5495DC90">Subias, Luis S.</mods:namePart>
<mods:affiliation id="8DDD3949E07AD3DCE3928F46096C44DD">Complutense University, Madrid, Spain.</mods:affiliation>
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<mods:namePart id="CF25C61E9D2BF4D41DB7469EB6CA8557">Shtanchaeva, Umukusum Ya.</mods:namePart>
<mods:affiliation id="082BF6A8061EF447AA031A7716897882">Complutense University, Madrid, Spain.</mods:affiliation>
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<mods:namePart id="1567C9852ECE193E81B0084D7A2CCB8A">Friedrich, Stefan</mods:namePart>
<mods:affiliation id="307E77A5C19EFFD758FDD641B7983A51">SNSB-Bavarian State Collection of Zoology, Munich, Germany. &amp; Institute of Environmental and Agricultural Biology (X-BIO), Tyumen State University, Tyumen, Russia.</mods:affiliation>
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<mods:title id="2301AE8AF06E44AE2448383082E0FE54">Acarologia</mods:title>
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<mods:date id="F12AA058389D86B76F0C759661C3DA80">2021</mods:date>
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<mods:number id="42BE3E572C4619D0C13410F4CB1B5EF1">61</mods:number>
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<heading id="D0AFDA232632FFADFE21802C1B57C9E0" bold="true" box="[491,831,621,649]" fontSize="12" level="2" pageId="4" pageNumber="594" reason="6">
<emphasis id="B92CB15D2632FFADFE21802C1B57C9E0" bold="true" box="[491,831,621,649]" pageId="4" pageNumber="594">
<taxonomicName id="4C5816CC2632FFADFE21802C1A85C9E0" ID-CoL="8T9GS" authority="Ermilov &amp; Subias &amp; Shtanchaeva &amp; Friedrich, 2021" authorityName="Ermilov &amp; Subias &amp; Shtanchaeva &amp; Friedrich" authorityYear="2021" box="[491,749,621,649]" class="Arachnida" family="Haplozetidae" genus="Pilobates" kingdom="Animalia" order="Sarcoptiformes" pageId="4" pageNumber="594" phylum="Arthropoda" rank="species" species="wachteli" status="sp. nov.">
<emphasis id="B92CB15D2632FFADFE21802C1A85C9E0" bold="true" box="[491,749,621,649]" italics="true" pageId="4" pageNumber="594">Pilobates wachteli</emphasis>
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<taxonomicNameLabel id="A21F0C262632FFADFD3F802C1B57C9E0" box="[757,831,621,649]" pageId="4" pageNumber="594" rank="species">n. sp.</taxonomicNameLabel>
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Zoobank:
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<subSubSection id="C3423EC42632FFADFDD980871AF7C9B7" box="[531,671,710,734]" pageId="4" pageNumber="594" type="description">
<paragraph id="8BE76D4F2632FFADFDD980871AF7C9B7" blockId="4.[490,1548,679,2045]" box="[531,671,710,734]" pageId="4" pageNumber="594">
(
<figureCitation id="136371CA2632FFADFDD180871AEBC9B7" box="[539,643,710,734]" captionStart="Figure 2" captionStartId="5.[119,186,1944,1965]" captionTargetBox="[151,1459,271,1872]" captionTargetId="figure-2@5.[103,1494,221,1890]" captionTargetPageId="5" captionText="Figure 2 Pilobates wachtelin. sp., adult: a dorsal view (legs omitted); b ventral view (gnathosoma and legs omitted); c lateral view (gnathosoma and legs omitted). Scale bar 100 μm." figureDoi="http://doi.org/10.5281/zenodo.7159412" httpUri="https://zenodo.org/record/7159412/files/figure.png" pageId="4" pageNumber="594">Figures 2</figureCitation>
,
<figureCitation id="136371CA2632FFADFD4780871AF0C9B7" box="[653,664,710,734]" captionStart="Figure 3" captionStartId="6.[119,186,1910,1931]" captionTargetBox="[118,1488,238,1820]" captionTargetId="figure-2@6.[103,1494,228,1856]" captionTargetPageId="6" captionText="Figure 3 Pilobates wachtelin. sp., adult: a subcapitulum, ventral view; b lip with adoral setae, left, ventral view; c palp, left, antiaxial" figureDoi="http://doi.org/10.5281/zenodo.7159414" httpUri="https://zenodo.org/record/7159414/files/figure.png" pageId="4" pageNumber="594">3</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3423EC42632FFADFDD980A81BABC8A6" pageId="4" pageNumber="594" type="diagnosis">
<paragraph id="8BE76D4F2632FFADFDD980A81DB0C84A" blockId="4.[490,1548,679,2045]" pageId="4" pageNumber="594">
<emphasis id="B92CB15D2632FFADFDD980A81AE8C868" bold="true" box="[531,640,745,769]" pageId="4" pageNumber="594">Diagnosis</emphasis>
— Adult. Body size: 630713 × 365431. Body surface densely microgranulate and sparsely foveolate. Rostral, lamellar and interlamellar setae setiform, barbed;
<emphasis id="B92CB15D2632FFADFA91814D1D1AC84A" box="[1371,1394,780,803]" italics="true" pageId="4" pageNumber="594">ro</emphasis>
shortest,
</paragraph>
<paragraph id="8BE76D4F2632FFADFE21816F1BABC8A6" blockId="4.[490,1548,679,2045]" pageId="4" pageNumber="594">
<emphasis id="B92CB15D2632FFADFE21816F1A68C82C" box="[491,512,814,837]" italics="true" pageId="4" pageNumber="594">in</emphasis>
longest. Bothridial seta long, lanceolate, barbed. Tutorium of medium length. With 15 pairs of setiform, barbed notogastral setae,
<emphasis id="B92CB15D2632FFADFC7081101BAEC801" box="[954,966,849,872]" italics="true" pageId="4" pageNumber="594">c</emphasis>
<subScript id="17DC6F0A2632FFADFC0F811D1BA3C804" attach="left" box="[965,971,860,877]" fontSize="7" pageId="4" pageNumber="594">2</subScript>
comparatively short, others long. Epimeral and anogenital setae setiform, barbed; anal and adanal
<emphasis id="B92CB15D2632FFADFBC681331C4FC8E3" box="[1036,1063,882,906]" italics="true" pageId="4" pageNumber="594">ad</emphasis>
(
<subScript id="17DC6F0A2632FFADFBE2813F1C59C8E6" attach="none" box="[1064,1073,894,911]" fontSize="7" pageId="4" pageNumber="594">1</subScript>
,
<emphasis id="B92CB15D2632FFADFB8A81331C33C8E3" box="[1088,1115,882,906]" italics="true" pageId="4" pageNumber="594">ad</emphasis>
<subScript id="17DC6F0A2632FFADFB91813F1C09C8E6" attach="left" box="[1115,1121,894,911]" fontSize="7" pageId="4" pageNumber="594">2</subScript>
) setae long,
<emphasis id="B92CB15D2632FFADFB2781331D60C8E3" box="[1261,1288,882,906]" italics="true" pageId="4" pageNumber="594">ad</emphasis>
<subScript id="17DC6F0A2632FFADFAC3813F1D67C8E6" attach="left" box="[1289,1295,894,911]" fontSize="7" pageId="4" pageNumber="594">3</subScript>
of medium length. With six pairs of genital and three pairs of aggenital setae. All leg tarsi with three claws. Trochanter IV without tooth distodorsally.
</paragraph>
</subSubSection>
<subSubSection id="C3423EC42632FFAEFDD9819B1A24CAA4" lastPageId="7" lastPageNumber="597" pageId="4" pageNumber="594" type="description">
<paragraph id="8BE76D4F2632FFADFDD9819B1CE5CF7D" blockId="4.[490,1548,679,2045]" pageId="4" pageNumber="594">
<emphasis id="B92CB15D2632FFADFDD9819B1A90C89B" bold="true" box="[531,760,986,1010]" pageId="4" pageNumber="594">Description of adult</emphasis>
— Measurements Body length: 630 (
<typeStatus id="54E3D3ED2632FFADFB7B819B1D7DC89B" box="[1201,1301,986,1010]" pageId="4" pageNumber="594" type="holotype">holotype</typeStatus>
: male), 713 (
<specimenCount id="9D5EA6C62632FFADFA56819B1A2ECF7C" pageId="4" pageNumber="594" type="generic" typeStatus="paratype">one paratype</specimenCount>
: female); body width: 365 (
<typeStatus id="54E3D3ED2632FFADFCB681BD1B88CF7D" box="[892,992,1020,1044]" pageId="4" pageNumber="594" type="holotype">holotype</typeStatus>
), 431 (
<typeStatus id="54E3D3ED2632FFADFBEE81BD1CEACF7D" box="[1060,1154,1020,1044]" pageId="4" pageNumber="594" type="paratype">paratype</typeStatus>
).
</paragraph>
<paragraph id="8BE76D4F2632FFADFDD986611DBECF30" blockId="4.[490,1548,679,2045]" pageId="4" pageNumber="594">
Integument (
<figureCitation id="136371CA2632FFADFD55865E1A8ECF5E" box="[671,742,1055,1079]" captionStart="Figure 2" captionStartId="5.[119,186,1944,1965]" captionTargetBox="[151,1459,271,1872]" captionTargetId="figure-2@5.[103,1494,221,1890]" captionTargetPageId="5" captionText="Figure 2 Pilobates wachtelin. sp., adult: a dorsal view (legs omitted); b ventral view (gnathosoma and legs omitted); c lateral view (gnathosoma and legs omitted). Scale bar 100 μm." figureDoi="http://doi.org/10.5281/zenodo.7159412" httpUri="https://zenodo.org/record/7159412/files/figure.png" pageId="4" pageNumber="594">Figs 2</figureCitation>
a-c, 3a) Body color light brown. Body surface densely microgranulate and (except median part of epimeral region) sparsely foveolate (diameter of foveola up to 4).
</paragraph>
<paragraph id="8BE76D4F2632FFADFDD986251D84CFF6" blockId="4.[490,1548,679,2045]" pageId="4" pageNumber="594">
Prodorsum (
<figureCitation id="136371CA2632FFADFD5786251B7DCF15" box="[669,789,1124,1148]" captionStart="Figure 2" captionStartId="5.[119,186,1944,1965]" captionTargetBox="[151,1459,271,1872]" captionTargetId="figure-2@5.[103,1494,221,1890]" captionTargetPageId="5" captionText="Figure 2 Pilobates wachtelin. sp., adult: a dorsal view (legs omitted); b ventral view (gnathosoma and legs omitted); c lateral view (gnathosoma and legs omitted). Scale bar 100 μm." figureDoi="http://doi.org/10.5281/zenodo.7159412" httpUri="https://zenodo.org/record/7159412/files/figure.png" pageId="4" pageNumber="594">Figs 2a, 2c</figureCitation>
) Rostrum rounded. Lamella about 1/2 length of prodorsum. Prolamella absent. Sublamella slightly shorter than lamella. Sublamellar porose area oval (20
</paragraph>
<paragraph id="8BE76D4F2632FFADFE2186E81A53CE22" blockId="4.[490,1548,679,2045]" pageId="4" pageNumber="594">
24 × 1216), located ventral to sublamella. Tutorium similar to lamella in length, ridge-like, distinctly not reaching rostral margin. Rostral (7377), lamellar (131143), interlamellar (151 164), and exobothridial (5357) setae setiform, barbed;
<emphasis id="B92CB15D2632FFADFB8A86AE1C30CE6F" box="[1088,1112,1263,1286]" italics="true" pageId="4" pageNumber="594">ex</emphasis>
thinnest. Bothridial seta (123135) lanceolate, barbed, with long stalk and short head. Dorsosejugal porose area band-like, poorly visible.
</paragraph>
<paragraph id="8BE76D4F2632FFADFDD987171C3FCEBC" blockId="4.[490,1548,679,2045]" pageId="4" pageNumber="594">
Notogaster (
<figureCitation id="136371CA2632FFADFD5487171B7CCE07" box="[670,788,1366,1390]" captionStart="Figure 2" captionStartId="5.[119,186,1944,1965]" captionTargetBox="[151,1459,271,1872]" captionTargetId="figure-2@5.[103,1494,221,1890]" captionTargetPageId="5" captionText="Figure 2 Pilobates wachtelin. sp., adult: a dorsal view (legs omitted); b ventral view (gnathosoma and legs omitted); c lateral view (gnathosoma and legs omitted). Scale bar 100 μm." figureDoi="http://doi.org/10.5281/zenodo.7159412" httpUri="https://zenodo.org/record/7159412/files/figure.png" pageId="4" pageNumber="594">Figs 2a, 2c</figureCitation>
) Anterior notogastral margin slightly convex medially. Pteromorph subtriangular, rounded laterally, with well-developed hinge. Fifteen pairs of notogastral setae (
<emphasis id="B92CB15D2632FFADFDE587DD1A53CEDA" box="[559,571,1436,1459]" italics="true" pageId="4" pageNumber="594">c</emphasis>
<subScript id="17DC6F0A2632FFADFDF187E61A29CED1" attach="left" box="[571,577,1447,1464]" fontSize="7" pageId="4" pageNumber="594">2</subScript>
: 3641; others: 135147) setiform, barbed;
<emphasis id="B92CB15D2632FFADFBD187DD1C4FCEDA" box="[1051,1063,1436,1459]" italics="true" pageId="4" pageNumber="594">c</emphasis>
<subScript id="17DC6F0A2632FFADFBED87E61C45CED1" attach="left" box="[1063,1069,1447,1464]" fontSize="7" pageId="4" pageNumber="594">2</subScript>
thinnest. Four pairs of sacculi drop-like. Opisthonotal gland opening and all lyrifissures distinct.
</paragraph>
<paragraph id="8BE76D4F2632FFADFDD987A11DD0CD36" blockId="4.[490,1548,679,2045]" pageId="4" pageNumber="594">
Gnathosoma (
<figureCitation id="136371CA2632FFADFD7A87A11A91CE91" box="[688,761,1504,1528]" captionStart="Figure 3" captionStartId="6.[119,186,1910,1931]" captionTargetBox="[118,1488,238,1820]" captionTargetId="figure-2@6.[103,1494,228,1856]" captionTargetPageId="6" captionText="Figure 3 Pilobates wachtelin. sp., adult: a subcapitulum, ventral view; b lip with adoral setae, left, ventral view; c palp, left, antiaxial" figureDoi="http://doi.org/10.5281/zenodo.7159414" httpUri="https://zenodo.org/record/7159414/files/figure.png" pageId="4" pageNumber="594">Figs 3</figureCitation>
a-d) Subcapitulum size: 147151 × 98106. Subcapitular setae setiform,
<emphasis id="B92CB15D2632FFADFD9984421A08CD73" box="[595,608,1539,1562]" italics="true" pageId="4" pageNumber="594">a</emphasis>
and
<emphasis id="B92CB15D2632FFADFD5D84431ACCCD73" box="[663,676,1538,1562]" italics="true" pageId="4" pageNumber="594">h</emphasis>
(2428) barbed,
<emphasis id="B92CB15D2632FFADFC9584421B1ACD73" box="[863,882,1539,1562]" italics="true" pageId="4" pageNumber="594">m</emphasis>
(2024) slightly barbed, thinnest. Adoral seta (1620) setiform, barbed. Palp (6165) with setation 02139(+ω). Postpalpal seta (8) spiniform, roughened. Chelicera (159164) with two setiform, barbed setae
<emphasis id="B92CB15D2632FFADFB6184061CB9CD36" box="[1195,1233,1607,1631]" italics="true" pageId="4" pageNumber="594">cha</emphasis>
(: 5761;
<emphasis id="B92CB15D2632FFADFAF884061D30CD36" box="[1330,1368,1607,1631]" italics="true" pageId="4" pageNumber="594">chb</emphasis>
: 3641).
</paragraph>
<paragraph id="8BE76D4F2632FFADFDD9842B1C00CDAE" blockId="4.[490,1548,679,2045]" pageId="4" pageNumber="594">
Epimeral and lateral podosomal regions (
<figureCitation id="136371CA2632FFADFC15842B1C3ECDEB" box="[991,1110,1642,1666]" captionStart="Figure 2" captionStartId="5.[119,186,1944,1965]" captionTargetBox="[151,1459,271,1872]" captionTargetId="figure-2@5.[103,1494,221,1890]" captionTargetPageId="5" captionText="Figure 2 Pilobates wachtelin. sp., adult: a dorsal view (legs omitted); b ventral view (gnathosoma and legs omitted); c lateral view (gnathosoma and legs omitted). Scale bar 100 μm." figureDoi="http://doi.org/10.5281/zenodo.7159412" httpUri="https://zenodo.org/record/7159412/files/figure.png" pageId="4" pageNumber="594">Figs 2b, 2c</figureCitation>
) Epimeral setal formula: 3133. Setae (
<emphasis id="B92CB15D2632FFADFDFF84CD1A27CDCD" box="[565,591,1676,1700]" italics="true" pageId="4" pageNumber="594">1a</emphasis>
,
<emphasis id="B92CB15D2632FFADFD9784CD1AF6CDCD" box="[605,670,1676,1700]" italics="true" pageId="4" pageNumber="594">1c, 2a</emphasis>
,
<emphasis id="B92CB15D2632FFADFD6684CD1AAECDCD" box="[684,710,1676,1700]" italics="true" pageId="4" pageNumber="594">3a</emphasis>
: 2024;
<emphasis id="B92CB15D2632FFADFCE284CD1B29CDCD" box="[808,833,1676,1700]" italics="true" pageId="4" pageNumber="594">3c</emphasis>
: 4953; others: 3641) setiform, barbed. Circumpedal carina long, directed to pedotectum II. Discidium subtriangular.
</paragraph>
<paragraph id="8BE76D4F2632FFADFDD984901B26CC38" blockId="4.[490,1548,679,2045]" pageId="4" pageNumber="594">
Anogenital region (
<figureCitation id="136371CA2632FFADFD3884901B04CD80" box="[754,876,1745,1769]" captionStart="Figure 2" captionStartId="5.[119,186,1944,1965]" captionTargetBox="[151,1459,271,1872]" captionTargetId="figure-2@5.[103,1494,221,1890]" captionTargetPageId="5" captionText="Figure 2 Pilobates wachtelin. sp., adult: a dorsal view (legs omitted); b ventral view (gnathosoma and legs omitted); c lateral view (gnathosoma and legs omitted). Scale bar 100 μm." figureDoi="http://doi.org/10.5281/zenodo.7159412" httpUri="https://zenodo.org/record/7159412/files/figure.png" pageId="4" pageNumber="594">Figs 2b, 2c</figureCitation>
) Six pairs of genital (2024), three pairs of aggenital (2024), two pairs of anal (7382), and three pairs of adanal (
<emphasis id="B92CB15D2632FFADFBB484B51CF1CC65" box="[1150,1177,1780,1804]" italics="true" pageId="4" pageNumber="594">ad</emphasis>
<subScript id="17DC6F0A2632FFADFB5085411CCBCC78" attach="left" box="[1178,1187,1792,1809]" fontSize="7" pageId="4" pageNumber="594">1</subScript>
,
<emphasis id="B92CB15D2632FFADFB7884B51CA5CC65" box="[1202,1229,1780,1804]" italics="true" pageId="4" pageNumber="594">ad</emphasis>
<subScript id="17DC6F0A2632FFADFB0785411CBBCC78" attach="left" box="[1229,1235,1792,1809]" fontSize="7" pageId="4" pageNumber="594">2</subScript>
: 114121;
<emphasis id="B92CB15D2632FFADFA9984B51D06CC65" box="[1363,1390,1780,1804]" italics="true" pageId="4" pageNumber="594">ad</emphasis>
<subScript id="17DC6F0A2632FFADFAA585411D1DCC78" attach="left" box="[1391,1397,1792,1809]" fontSize="7" pageId="4" pageNumber="594">3</subScript>
: 4553) setae setiform, barbed. Adanal lyrifissure located close and parallel to anal plate. Marginal porose area complete, band-like.
</paragraph>
<paragraph id="8BE76D4F2632FFADFDD9851D1DC3CCF2" blockId="4.[490,1548,679,2045]" pageId="4" pageNumber="594">
Legs (
<figureCitation id="136371CA2632FFADFD94851A1ACFCC1A" box="[606,679,1883,1907]" captionStart="Figure 3" captionStartId="6.[119,186,1910,1931]" captionTargetBox="[118,1488,238,1820]" captionTargetId="figure-2@6.[103,1494,228,1856]" captionTargetPageId="6" captionText="Figure 3 Pilobates wachtelin. sp., adult: a subcapitulum, ventral view; b lip with adoral setae, left, ventral view; c palp, left, antiaxial" figureDoi="http://doi.org/10.5281/zenodo.7159414" httpUri="https://zenodo.org/record/7159414/files/figure.png" pageId="4" pageNumber="594">Figs 3</figureCitation>
e-h) Tridactylous. Median claw distinctly thicker than lateral claws, all slightly barbed on dorsal side. Tibia I with small dorsodistal apophysis bearing solenidiaandφ
<subScript id="17DC6F0A2632FFADFA6885CB1DC3CCF2" attach="both" box="[1442,1451,1930,1947]" fontSize="7" pageId="4" pageNumber="594">1</subScript>
</paragraph>
<paragraph id="8BE76D4F2632FFAFFE2185E01D24CCD1" blockId="4.[490,1548,679,2045]" lastBlockId="6.[118,1553,1909,1976]" lastPageId="6" lastPageNumber="596" pageId="4" pageNumber="594">
φ
<subScript id="17DC6F0A2632FFADFE3185ED1A69CCD4" attach="left" box="[507,513,1964,1981]" fontSize="7" pageId="4" pageNumber="594">2</subScript>
. Tibiae I and II with triangular tubercle proximoventrally. All femora rounded distoventrally. Trochanter IV rounded distodorsally (without tooth). Dorsoparaxial porose area on femora IIV and on trochanters III, IV well visible; proximoventral porose area on tarsi IIV and view; d chelicera, right, antiaxial view; e leg I, without trochanter, right, antiaxial view; f leg II, without trochanter and tarsus, right, antiaxial view; g leg III, without tarsus, left, antiaxial view; h leg IV, left, antiaxial view. Scale bars 20 μm (a, b; c; d; e-h).
</paragraph>
<caption id="DF273DC72633FFACFFBD85D91A1ECCAD" ID-DOI="http://doi.org/10.5281/zenodo.7159412" ID-Zenodo-Dep="7159412" httpUri="https://zenodo.org/record/7159412/files/figure.png" pageId="5" pageNumber="595" startId="5.[119,186,1944,1965]" targetBox="[151,1459,271,1872]" targetPageId="5">
<paragraph id="8BE76D4F2633FFACFFBD85D91A1ECCAD" blockId="5.[118,1525,1944,1988]" pageId="5" pageNumber="595">
<emphasis id="B92CB15D2633FFACFFBD85D918A6CCC4" bold="true" box="[119,206,1944,1965]" pageId="5" pageNumber="595">Figure 2</emphasis>
<taxonomicName id="4C5816CC2633FFACFF1685D919FCCCC7" authority="Ermilov &amp; Subias &amp; Shtanchaeva &amp; Friedrich, 2021" authorityName="Ermilov &amp; Subias &amp; Shtanchaeva &amp; Friedrich" authorityYear="2021" box="[220,404,1944,1966]" class="Arachnida" family="Haplozetidae" genus="Pilobates" kingdom="Animalia" order="Sarcoptiformes" pageId="5" pageNumber="595" phylum="Arthropoda" rank="species" species="wachteli" status="sp. nov.">
<emphasis id="B92CB15D2633FFACFF1685D919FCCCC7" box="[220,404,1944,1966]" italics="true" pageId="5" pageNumber="595">Pilobates wachteli</emphasis>
</taxonomicName>
<emphasis id="B92CB15D2633FFACFE5C85D919A6CCC4" bold="true" box="[406,462,1944,1965]" pageId="5" pageNumber="595">
<taxonomicNameLabel id="A21F0C262633FFACFE5C85D919A6CCC4" box="[406,462,1944,1965]" pageId="5" pageNumber="595" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, adult: a dorsal view (legs omitted); b ventral view (gnathosoma and legs omitted); c lateral view (gnathosoma and legs omitted). Scale bar 100 μm.
</paragraph>
</caption>
<caption id="DF273DC72630FFAFFFBD85371DB9CCE2" ID-DOI="http://doi.org/10.5281/zenodo.7159414" ID-Zenodo-Dep="7159414" box="[119,1489,1909,1931]" httpUri="https://zenodo.org/record/7159414/files/figure.png" pageId="6" pageNumber="596" startId="6.[119,186,1910,1931]" targetBox="[118,1488,238,1820]" targetPageId="6">
<paragraph id="8BE76D4F2630FFAFFFBD85371DB9CCE2" blockId="6.[118,1553,1909,1976]" box="[119,1489,1909,1931]" pageId="6" pageNumber="596">
<emphasis id="B92CB15D2630FFAFFFBD853718A5CCE2" bold="true" box="[119,205,1910,1931]" pageId="6" pageNumber="596">Figure 3</emphasis>
<taxonomicName id="4C5816CC2630FFAFFF10853419F9CCE2" authority="Ermilov &amp; Subias &amp; Shtanchaeva &amp; Friedrich, 2021" authorityName="Ermilov &amp; Subias &amp; Shtanchaeva &amp; Friedrich" authorityYear="2021" box="[218,401,1909,1931]" class="Arachnida" family="Haplozetidae" genus="Pilobates" kingdom="Animalia" order="Sarcoptiformes" pageId="6" pageNumber="596" phylum="Arthropoda" rank="species" species="wachteli" status="sp. nov.">
<emphasis id="B92CB15D2630FFAFFF10853419F9CCE2" box="[218,401,1909,1931]" italics="true" pageId="6" pageNumber="596">Pilobates wachteli</emphasis>
</taxonomicName>
<emphasis id="B92CB15D2630FFAFFE58853719A0CCE2" bold="true" box="[402,456,1910,1931]" pageId="6" pageNumber="596">
<taxonomicNameLabel id="A21F0C262630FFAFFE58853719A0CCE2" box="[402,456,1910,1931]" pageId="6" pageNumber="596" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, adult: a subcapitulum, ventral view; b lip with adoral setae, left, ventral view; c palp, left, antiaxial
</paragraph>
</caption>
<paragraph id="8BE76D4F2631FFAEFE2182A41DC8CA49" blockId="7.[491,1542,229,841]" pageId="7" pageNumber="597">distoventral porose area on tibiae IIV not observed. Formulas of leg setation and solenidia: I (153418) [122], II (153415) [112], III (231315) [110], IV (122312)</paragraph>
<paragraph id="8BE76D4F2631FFAEFE21836A1DAECA0C" blockId="7.[491,1542,229,841]" pageId="7" pageNumber="597">
[010]; homology of setae and solenidia indicated in
<tableCitation id="C6DA58F42631FFAEFB94836A1CD9CA2A" box="[1118,1201,299,323]" captionStart="Table 1" captionStartId="7.[119,175,1667,1688]" captionText="Table 1 Leg setation and solenidia of adultPilobates wachtelin. sp. andZetorchella robertbeckin. sp." pageId="7" pageNumber="597">Table 1</tableCitation>
. Famulus on tarsus I short, erect, slightly swollen distally, inserted between solenidion
<subScript id="17DC6F0A2631FFAEFBB483181CECCA03" attach="right" box="[1150,1156,345,362]" fontSize="7" pageId="7" pageNumber="597">2</subScript>
ωand seta
<emphasis id="B92CB15D2631FFAEFB3A830C1D66CA0C" box="[1264,1294,333,357]" italics="true" pageId="7" pageNumber="597">ft”</emphasis>
. Seta
<emphasis id="B92CB15D2631FFAEFA9F830F1D37CA0C" box="[1365,1375,334,357]" italics="true" pageId="7" pageNumber="597">s</emphasis>
on tarsus
</paragraph>
<paragraph id="8BE76D4F2631FFAEFE2183311A24CAA4" blockId="7.[491,1542,229,841]" pageId="7" pageNumber="597">
I eupathidial, located before setae
<emphasis id="B92CB15D2631FFAEFC9583311B1DCAEE" box="[863,885,368,391]" italics="true" pageId="7" pageNumber="597">a</emphasis>
and
<emphasis id="B92CB15D2631FFAEFC6683311BA0CAEE" box="[940,968,368,391]" italics="true" pageId="7" pageNumber="597">a”</emphasis>
. Seta
<emphasis id="B92CB15D2631FFAEFBDB83311C4FCAE1" box="[1041,1063,368,392]" italics="true" pageId="7" pageNumber="597">l”</emphasis>
on genu I located on small apophysis. Solenidion ω
<subScript id="17DC6F0A2631FFAEFDBD83DF1AE8CAC6" attach="left" box="[631,640,414,431]" fontSize="7" pageId="7" pageNumber="597">1</subScript>
on tarsi I, ω
<subScript id="17DC6F0A2631FFAEFCC283DF1B79CAC6" attach="left" box="[776,785,414,431]" fontSize="7" pageId="7" pageNumber="597">1</subScript>
and ω
<subScript id="17DC6F0A2631FFAEFC9383DF1B37CAC6" attach="left" box="[857,863,414,431]" fontSize="7" pageId="7" pageNumber="597">2</subScript>
on tarsus II and σ on genu III bacilliform, other solenidia setiform.
</paragraph>
</subSubSection>
<subSubSection id="C3423EC42631FFAEFDD983991BB7C97B" pageId="7" pageNumber="597" type="materials_examined">
<paragraph id="8BE76D4F2631FFAEFDD983991BB7C97B" blockId="7.[491,1542,229,841]" pageId="7" pageNumber="597">
<emphasis id="B92CB15D2631FFAEFDD983991AAFCA99" bold="true" box="[531,711,472,496]" pageId="7" pageNumber="597">Type deposition</emphasis>
— The
<typeStatus id="54E3D3ED2631FFAEFCEC83961BECCA86" box="[806,900,471,495]" pageId="7" pageNumber="597" type="holotype">holotype</typeStatus>
is deposited in the collection of the
<collectionCode id="ED49F58A2631FFAEFAEB83961D33CA86" box="[1313,1371,471,495]" country="Germany" httpUri="http://grbio.org/cool/4b7r-7sec" name="Bavarian State Collection of Zoology" pageId="7" pageNumber="597" type="Museum">ZSM</collectionCode>
;
<specimenCount id="9D5EA6C62631FFAEFAAC83991D83CA99" box="[1382,1515,472,496]" pageId="7" pageNumber="597" type="generic" typeStatus="paratype">one paratype</specimenCount>
is deposited in the collection of the
<collectionCode id="ED49F58A2631FFAEFCB483BB1BB4C97B" box="[894,988,506,530]" pageId="7" pageNumber="597">TSUMZ</collectionCode>
.
</paragraph>
</subSubSection>
<subSubSection id="C3423EC42631FFAEFDD9805C1BBAC910" pageId="7" pageNumber="597" type="etymology">
<paragraph id="8BE76D4F2631FFAEFDD9805C1BBAC910" blockId="7.[491,1542,229,841]" pageId="7" pageNumber="597">
<emphasis id="B92CB15D2631FFAEFDD9805C1AE6C95C" bold="true" box="[531,654,541,565]" pageId="7" pageNumber="597">Etymology</emphasis>
— The specific name is dedicated to our friend and colleague, the coleopterologist and all-around biologist Franz Wachtel (Grünwald,
<collectingCountry id="F34F2DDF2631FFAEFB4A807E1C88C93E" box="[1152,1248,575,599]" name="Germany" pageId="7" pageNumber="597">Germany</collectingCountry>
) for his expertise and assistance by collecting the Ethiopian mites.
</paragraph>
</subSubSection>
<subSubSection id="C3423EC42631FFAEFDD980C51DCBC820" pageId="7" pageNumber="597" type="discussion">
<paragraph id="8BE76D4F2631FFAEFDD980C51DA3C86D" blockId="7.[491,1542,229,841]" pageId="7" pageNumber="597">
<emphasis id="B92CB15D2631FFAEFDD980C51A12C9F5" bold="true" box="[531,634,644,668]" pageId="7" pageNumber="597">Remarks</emphasis>
— In having tridactylous legs,
<taxonomicName id="4C5816CC2631FFAEFC0280C51CFBC9F5" authority="Ermilov &amp; Subias &amp; Shtanchaeva &amp; Friedrich, 2021" authorityName="Ermilov &amp; Subias &amp; Shtanchaeva &amp; Friedrich" authorityYear="2021" box="[968,1171,644,668]" class="Arachnida" family="Haplozetidae" genus="Pilobates" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="597" phylum="Arthropoda" rank="species" species="wachteli" status="sp. nov.">
<emphasis id="B92CB15D2631FFAEFC0280C51CFBC9F5" box="[968,1171,644,668]" italics="true" pageId="7" pageNumber="597">Pilobates wachteli</emphasis>
</taxonomicName>
<emphasis id="B92CB15D2631FFAEFB5980C51CA6C9F5" bold="true" box="[1171,1230,644,668]" pageId="7" pageNumber="597">
<taxonomicNameLabel id="A21F0C262631FFAEFB5980C51CA6C9F5" box="[1171,1230,644,668]" pageId="7" pageNumber="597" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
is morphologically most similar to
<taxonomicName id="4C5816CC2631FFAEFD9F80E71BD1C9D7" authority="(Berlese, 1916)" baseAuthorityName="Berlese" baseAuthorityYear="1916" box="[597,953,678,702]" class="Arachnida" family="Haplozetidae" genus="Pilobates" kingdom="Animalia" order="Sarcoptiformes" pageId="7" pageNumber="597" phylum="Arthropoda" rank="species" species="incisura">
<emphasis id="B92CB15D2631FFAEFD9F80E71B73C9D7" box="[597,795,678,702]" italics="true" pageId="7" pageNumber="597">Pilobates incisura</emphasis>
(Berlese, 1916)
</taxonomicName>
from
<collectingCountry id="F34F2DDF2631FFAEFC3D80E71C26C9D7" box="[1015,1102,678,702]" name="Somalia" pageId="7" pageNumber="597">Somalia</collectingCountry>
(see
<bibRefCitation id="EFC910BE2631FFAEFB4E80E71D73C9D7" author="Berlese A." box="[1156,1307,678,702]" pageId="7" pageNumber="597" pagination="289 - 338" refId="ref4942" refString="Berlese A. 1916 b. Centuria terza di Acari nuovi. Redia, 12: 289 - 338." type="journal article" year="1916">Berlese 1916b</bibRefCitation>
;
<bibRefCitation id="EFC910BE2631FFAEFAEF80E71DA9C9D7" author="Mahunka S." box="[1317,1473,678,702]" pageId="7" pageNumber="597" pagination="29 - 49" refId="ref5311" refString="Mahunka S. 1994. Further notes, additions and redescriptions of the oribatid species preserved in the Berlese collection (Acari, Oribatida) I. Acta Zool. Acad. Sci. Hung., 40 (1): 29 - 49." type="journal article" year="1994">Mahunka 1994</bibRefCitation>
), however, the new species differs from the latter by the presence of 15 pairs of notogastral setae including
<emphasis id="B92CB15D2631FFAEFD9280AC1A0CC86D" box="[600,612,749,772]" italics="true" pageId="7" pageNumber="597">c</emphasis>
<subScript id="17DC6F0A2631FFAEFDAE80B91A02C860" attach="left" box="[612,618,760,777]" fontSize="7" pageId="7" pageNumber="597">3</subScript>
(versus 14 pairs,
<emphasis id="B92CB15D2631FFAEFCFA80AC1B54C86D" box="[816,828,749,772]" italics="true" pageId="7" pageNumber="597">c</emphasis>
<subScript id="17DC6F0A2631FFAEFCF680B91B2AC860" attach="left" box="[828,834,760,777]" fontSize="7" pageId="7" pageNumber="597">3</subScript>
absent), very long notogastral (except comparatively short
</paragraph>
<paragraph id="8BE76D4F2631FFAEFE21814E1DCBC820" blockId="7.[491,1542,229,841]" pageId="7" pageNumber="597">
<emphasis id="B92CB15D2631FFAEFE21814E199FC84F" box="[491,503,783,806]" italics="true" pageId="7" pageNumber="597">c</emphasis>
<subScript id="17DC6F0A2631FFAEFE3D815B1995C842" attach="left" box="[503,509,794,811]" fontSize="7" pageId="7" pageNumber="597">2</subScript>
), anal and adanal
<emphasis id="B92CB15D2631FFAEFD18814F1A85C84F" box="[722,749,782,806]" italics="true" pageId="7" pageNumber="597">ad</emphasis>
(
<subScript id="17DC6F0A2631FFAEFD24815B1A9FC842" attach="left" box="[750,759,794,811]" fontSize="7" pageId="7" pageNumber="597">1</subScript>
,
<emphasis id="B92CB15D2631FFAEFCCD814F1B4AC84F" box="[775,802,782,806]" italics="true" pageId="7" pageNumber="597">ad</emphasis>
<subScript id="17DC6F0A2631FFAEFCE9815B1B41C842" attach="left" box="[803,809,794,811]" fontSize="7" pageId="7" pageNumber="597">2</subScript>
) setae (versus all these setae comparatively short), lanceolate bothridial seta (versus setiform), and foveolate body integument (versus not foveolate).
</paragraph>
</subSubSection>
</treatment>
</document>