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<document id="FD810201765407E40D7430C33C04E9A8" ID-CLB-Dataset="156940" ID-DOI="10.11646/zootaxa.5190.4.7" ID-GBIF-Dataset="39f670af-d883-4a0b-9956-d6e29af453c3" ID-ISSN="1175-5326" ID-Zenodo-Dep="7138572" ID-ZooBank="763C975C-7D97-497B-BC7C-8A94CD92FBD1" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1664793912077" checkinUser="plazi" docAuthor="Shavrin, Alexey V." docDate="2022" docId="03F14B1A5B40BA31FF24EA481D044F2C" docLanguage="en" docName="zootaxa.5190.4.7.pdf" docOrigin="Zootaxa 5190 (4)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Amphichroum discolor Shavrin 2022, sp.n." docType="treatment" docVersion="7" lastPageNumber="578" masterDocId="FFC833625B41BA32FFB3E9751C754B19" masterDocTitle="New species and records of Amphichroum Kraatz, 1857 from China and the Himalayan Region (Coleoptera: Staphylinidae: Omaliinae: Anthophagini)" masterLastPageNumber="583" masterPageNumber="575" pageNumber="576" updateTime="1699362515216" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="6E1E736978FFDCC856DC32736297EB79" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="8A3FB2F102CCEF7A826AEE25C9353A53">
<mods:title id="0E7560428B76AC272E004F8C06C48CE9">New species and records of Amphichroum Kraatz, 1857 from China and the Himalayan Region (Coleoptera: Staphylinidae: Omaliinae: Anthophagini)</mods:title>
</mods:titleInfo>
<mods:name id="AC69528BA954C9711DE8BACC1784E395" type="personal">
<mods:role id="14744BEC34917D437355E90FE6E8E9F3">
<mods:roleTerm id="E45DD6D3D3FD52BF6BC0DAC4A3F85F62">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A540FC224256DCAE314562130A22CE6A">Shavrin, Alexey V.</mods:namePart>
</mods:name>
<mods:typeOfResource id="80DAC23DE311C3206661DD3A54AC126F">text</mods:typeOfResource>
<mods:relatedItem id="5CC28E574DF3162241B835CE67410A16" type="host">
<mods:titleInfo id="98509297511B62B440B51F1DD2455C4C">
<mods:title id="C02A6CEF893AC274B244DB23BDE164D2">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="28CEFBC51823018F9CDF82D772F834BF">
<mods:date id="878636C031EB0204CDBCEBC32D86125D">2022</mods:date>
<mods:detail id="4F928B54313EB63E30DBCA1F20020B4D" type="pubDate">
<mods:number id="CF8BD5EFA3BB6897EC279BED13C7FADE">2022-09-30</mods:number>
</mods:detail>
<mods:detail id="C93E91DA475588411A0B3ED6DEB4DDF9" type="volume">
<mods:number id="847E43224228DFAE6A18CFB2FA6D6D1B">5190</mods:number>
</mods:detail>
<mods:detail id="2BB66F35F639EB78609E9802E5FABC33" type="issue">
<mods:number id="973CB1FDB308044E68685AD79EA60F66">4</mods:number>
</mods:detail>
<mods:extent id="8A9C9E942EF631856BE047D369692B05" unit="page">
<mods:start id="CD6E23FB22DFC537355D384C5A312440">575</mods:start>
<mods:end id="776E1F579A6C5620686B9FC382D31F65">583</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="3DDA9E9C8BFC4698802CC62F778F77A8">journal article</mods:classification>
<mods:identifier id="8E5617C6132BCB62E14377C9E35683CB" type="CLB-Dataset">156940</mods:identifier>
<mods:identifier id="C9DE81E13955502FB7E0536B3A428A28" type="DOI">10.11646/zootaxa.5190.4.7</mods:identifier>
<mods:identifier id="0088CE9A8F952B010CCE459D34FE7DEE" type="GBIF-Dataset">39f670af-d883-4a0b-9956-d6e29af453c3</mods:identifier>
<mods:identifier id="250ABC1CFE2BCAE136AB8BE7A402CE8F" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="CD5B58591541F1828EB0D11055DCE9BC" type="Zenodo-Dep">7138572</mods:identifier>
<mods:identifier id="3BE1D81EDB7212C91A9690493FFD0F61" type="ZooBank">763C975C-7D97-497B-BC7C-8A94CD92FBD1</mods:identifier>
</mods:mods>
<treatment id="03F14B1A5B40BA31FF24EA481D044F2C" ID-DOI="http://doi.org/10.5281/zenodo.7140338" ID-GBIF-Taxon="201582203" ID-Zenodo-Dep="7140338" LSID="urn:lsid:plazi:treatment:03F14B1A5B40BA31FF24EA481D044F2C" httpUri="http://treatment.plazi.org/id/03F14B1A5B40BA31FF24EA481D044F2C" lastPageId="3" lastPageNumber="578" pageId="1" pageNumber="576">
<subSubSection id="C342A9875B40BA33FF24EA481D84484E" box="[151,497,829,856]" pageId="1" pageNumber="576" type="nomenclature">
<paragraph id="8BE7FA0C5B40BA33FF24EA481D84484E" blockId="1.[151,497,829,892]" box="[151,497,829,856]" pageId="1" pageNumber="576">
<heading id="D0AF4D605B40BA33FF24EA481D84484E" bold="true" box="[151,497,829,856]" fontSize="11" level="1" pageId="1" pageNumber="576" reason="1">
<emphasis id="B92C261E5B40BA33FF24EA481D84484E" bold="true" box="[151,497,829,856]" pageId="1" pageNumber="576">
<taxonomicName id="4C58818F5B40BA33FF24EA481DDA4841" authority="Shavrin, 2022" authorityName="Shavrin" authorityYear="2022" box="[151,431,829,856]" class="Insecta" family="Staphylinidae" genus="Amphichroum" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="576" phylum="Arthropoda" rank="species" species="discolor" status="sp. nov.">
<emphasis id="B92C261E5B40BA33FF24EA481DDA4841" bold="true" box="[151,431,829,856]" italics="true" pageId="1" pageNumber="576">Amphichroum discolor</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A21F9B655B40BA33FE05EA481D84484E" box="[438,497,829,855]" pageId="1" pageNumber="576" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C342A9875B40BA33FF24EA141D674865" box="[151,274,865,892]" pageId="1" pageNumber="576" type="description">
<paragraph id="8BE7FA0C5B40BA33FF24EA141D674865" blockId="1.[151,497,829,892]" box="[151,274,865,892]" pageId="1" pageNumber="576">
(
<figureCitation id="1363E6895B40BA33FF2CEA141D7C4865" box="[159,265,865,892]" captionStart-0="FIGURES 1–2" captionStart-1="FIGURES 3–10" captionStartId-0="2.[151,264,1849,1873]" captionStartId-1="4.[151,264,1983,2007]" captionTargetBox-0="[170,1425,609,1812]" captionTargetBox-1="[161,1424,355,1952]" captionTargetId-0="figure-196@2.[151,1436,594,1823]" captionTargetId-1="figure-103@4.[151,1436,345,1959]" captionTargetPageId-0="2" captionTargetPageId-1="4" captionText-0="FIGURES 1–2. Habitus of Amphichroum discolor: 1–holotype, 2–paratype. Scale bar: 1.0 mm." captionText-1="FIGURES 3–10. Aedeagus of Amphichroum (Figs. 3–5: A. discolor; Fig. 6 (apical portion): A. rotundatum; Figs. 7–8: A. grandidentatum; Figs. 9–10: A. reticulatum): 3–4, 6–7, 9–parameral view, 5, 8, 10–lateral view. Scale bar: 0.1 mm." figureDoi-0="http://doi.org/10.5281/zenodo.7138574" figureDoi-1="http://doi.org/10.5281/zenodo.7138578" httpUri-0="https://zenodo.org/record/7138574/files/figure.png" httpUri-1="https://zenodo.org/record/7138578/files/figure.png" pageId="1" pageNumber="576">Figs. 1–5</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C342A9875B40BA33FF24EADC1DFE4F82" pageId="1" pageNumber="576" type="materials_examined">
<paragraph id="8BE7FA0C5B40BA33FF24EADC18BC4F36" blockId="1.[151,1437,937,2008]" pageId="1" pageNumber="576">
<emphasis id="B92C261E5B40BA33FF24EADC1DB048DA" bold="true" box="[151,453,937,963]" pageId="1" pageNumber="576">Type material examined.</emphasis>
<materialsCitation id="3B30F0515B40BA33FE7DEADC18B04F36" ID-GBIF-Occurrence="3920575307" collectingDate="2019-07-05" collectionCode="NMPC" collectorName="D. Kral &amp; J. Ruzicka" country="China" county="Gongshan Co." elevation="3900" latitude="28.075" location="Biluo Mts. Pass" longLatPrecision="123" longitude="98.76" municipality="Biluo Mts. Pass" pageId="1" pageNumber="576" specimenCount="1" specimenCount-male="1" stateProvince="YUNNAN PROv." typeStatus="holotype">
<typeStatus id="54E344AE5B40BA33FE7DEADC1E4348DA" box="[462,566,937,963]" pageId="1" pageNumber="576" type="holotype">Holotype</typeStatus>
<specimenCount id="9D5E31855B40BA33FDF3EADF1E2048DD" box="[576,597,938,964]" pageId="1" pageNumber="576" type="male">♂</specimenCount>
(
<figureCitation id="1363E6895B40BA33FDD4EADC1ED948DA" box="[615,684,937,963]" captionStart="FIGURES 1–2" captionStartId="2.[151,264,1849,1873]" captionTargetBox="[170,1425,609,1812]" captionTargetId="figure-196@2.[151,1436,594,1823]" captionTargetPageId="2" captionText="FIGURES 1–2. Habitus of Amphichroum discolor: 1–holotype, 2–paratype. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7138574" httpUri="https://zenodo.org/record/7138574/files/figure.png" pageId="1" pageNumber="576">Fig. 1</figureCitation>
): ‘
<collectingCountry id="F34FBA9C5B40BA33FD7FEADC1F5B48DA" box="[716,814,937,963]" name="China" pageId="1" pageNumber="576">CHINA</collectingCountry>
:
<collectingRegion id="499C34EE5B40BA33FC89EADF187548DA" box="[826,1024,937,964]" country="China" name="Yunnan" pageId="1" pageNumber="576">YUNNAN PROv.</collectingRegion>
, |
<collectingCounty id="628682805B40BA33FBAFEADC18B448DA" box="[1052,1217,937,963]" pageId="1" pageNumber="576">Gongshan Co.</collectingCounty>
, |
<collectingMunicipality id="6B8360765B40BA33FB6CEADC19E248DD" box="[1247,1431,937,964]" pageId="1" pageNumber="576">Biluo Mts. Pass</collectingMunicipality>
,
<quantity id="4CA057E95B40BA33FF24EAB81D4048F1" box="[151,309,973,1000]" metricMagnitude="3" metricUnit="m" metricValue="3.9" metricValueMax="3.91" metricValueMin="3.8899999999999997" pageId="1" pageNumber="576" unit="m" value="3900.0" valueMax="3910.0" valueMin="3890.0">
<elevation id="00751D3F5B40BA33FF24EAB81D4048F1" box="[151,309,973,1000]" metricMagnitude="3" metricUnit="m" metricValue="3.9" metricValueMax="3.91" metricValueMin="3.8899999999999997" pageId="1" pageNumber="576" unit="m" value="3900.0" valueMax="3910.0" valueMin="3890.0">3890–3910 m</elevation>
</quantity>
, |
<geoCoordinate id="EE6C9CCB5B40BA33FEFDEABA1DB648F3" box="[334,451,975,1002]" degrees="28" direction="north" minutes="04.5" orientation="latitude" pageId="1" pageNumber="576" precision="92" value="28.075">28°04.5′N</geoCoordinate>
,
<geoCoordinate id="EE6C9CCB5B40BA33FE7CEABA1E3A48F3" box="[463,591,975,1002]" degrees="098" direction="east" minutes="45.6" orientation="longitude" pageId="1" pageNumber="576" precision="92" value="98.76">098°45.6′E</geoCoordinate>
, |
<collectorName id="26AD9FDA5B40BA33FDDBEABB1F2348F1" box="[616,854,973,1000]" pageId="1" pageNumber="576">D. Král &amp; J. Růžička</collectorName>
leg.’ &lt;printed&gt;, ‘
<date id="FFE6DCCC5B40BA33FBA4EAB818FA48F1" box="[1047,1167,973,1000]" pageId="1" pageNumber="576" value="2019-07-05">
<collectingDate id="EFA225245B40BA33FBA4EAB818FA48F1" box="[1047,1167,973,1000]" pageId="1" pageNumber="576" value="2019-07-05">5.vii.2019</collectingDate>
</date>
, sift #06 [Y06], | mixed forest with dominant |
<taxonomicName id="4C58818F5B40BA33FE28EA841DAC4F12" box="[411,473,1009,1035]" class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" order="Pinales" pageId="1" pageNumber="576" phylum="Tracheophyta" rank="genus">
<emphasis id="B92C261E5B40BA33FE28EA841DAC4F12" box="[411,473,1009,1035]" italics="true" pageId="1" pageNumber="576">Abies</emphasis>
</taxonomicName>
and
<taxonomicName id="4C58818F5B40BA33FDA0EA841ECD4F12" box="[531,696,1009,1035]" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" kingdom="Plantae" order="Ericales" pageId="1" pageNumber="576" phylum="Tracheophyta" rank="genus">
<emphasis id="B92C261E5B40BA33FDA0EA841ECD4F12" box="[531,696,1009,1035]" italics="true" pageId="1" pageNumber="576">Rhododendron</emphasis>
</taxonomicName>
shrubs, | litter under shrubs and along | fallen trunks’ &lt;printed&gt;, ‘HOLOTYPE |
<taxonomicName id="4C58818F5B40BA33FEFBED601D934F36" authorityName="Kraatz" authorityYear="1857" box="[328,486,1045,1071]" class="Insecta" family="Staphylinidae" genus="Amphichroum" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="576" phylum="Arthropoda" rank="genus">
<emphasis id="B92C261E5B40BA33FEFBED601D934F36" box="[328,486,1045,1071]" italics="true" pageId="1" pageNumber="576">Amphichroum</emphasis>
</taxonomicName>
|
<taxonomicName id="4C58818F5B40BA33FE4AED601E204F36" authority="Shavrin, 2022" authorityName="Shavrin" authorityYear="2022" box="[505,597,1045,1071]" class="Insecta" family="Staphylinidae" genus="Amphichroum" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="576" phylum="Arthropoda" rank="species" species="discolor" status="sp. nov.">
<emphasis id="B92C261E5B40BA33FE4AED601E204F36" box="[505,597,1045,1071]" italics="true" pageId="1" pageNumber="576">discolor</emphasis>
</taxonomicName>
<emphasis id="B92C261E5B40BA33FDEFED601EE64F36" bold="true" box="[604,659,1045,1071]" pageId="1" pageNumber="576">
<taxonomicNameLabel id="A21F9B655B40BA33FDEFED601EE64F36" box="[604,659,1045,1071]" pageId="1" pageNumber="576" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
| Shavrin A.V. des. 2022’ &lt;red, printed&gt; (
<collectionCode id="ED4962C95B40BA33FBD4ED6018C84F36" box="[1127,1213,1045,1071]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="1" pageNumber="576">NMPC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8BE7FA0C5B40BA33FF74ED4F1DFE4F82" blockId="1.[151,1437,937,2008]" pageId="1" pageNumber="576">
<typeStatus id="54E344AE5B40BA33FF74ED4F1D4D4F4D" box="[199,312,1082,1108]" pageId="1" pageNumber="576" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D5E31855B40BA33FEF7ED4C1DE64F4D" box="[324,403,1081,1108]" pageId="1" pageNumber="576" type="male">24 ♂♂</specimenCount>
(
<specimenCount id="9D5E31855B40BA33FE10ED4C1E244F4A" box="[419,593,1081,1107]" pageId="1" pageNumber="576" type="generic">four specimens</specimenCount>
dissected),
<specimenCount id="9D5E31855B40BA33FD6AED4C1F5F4F4D" box="[729,810,1081,1108]" pageId="1" pageNumber="576" type="female">27 ♀♀</specimenCount>
: same data as the holotype, with additional red printed label: ‘
<typeStatus id="54E344AE5B40BA33FF52ED281D044F6E" box="[225,369,1117,1143]" pageId="1" pageNumber="576" type="paratype">PARATYPE</typeStatus>
|
<taxonomicName id="4C58818F5B40BA33FE35ED281E514F6E" authorityName="Kraatz" authorityYear="1857" box="[390,548,1117,1143]" class="Insecta" family="Staphylinidae" genus="Amphichroum" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="576" phylum="Arthropoda" rank="genus">
<emphasis id="B92C261E5B40BA33FE35ED281E514F6E" box="[390,548,1117,1143]" italics="true" pageId="1" pageNumber="576">Amphichroum</emphasis>
</taxonomicName>
|
<taxonomicName id="4C58818F5B40BA33FD8BED281EE14F6E" authority="Shavrin, 2022" authorityName="Shavrin" authorityYear="2022" box="[568,660,1117,1143]" class="Insecta" family="Staphylinidae" genus="Amphichroum" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="576" phylum="Arthropoda" rank="species" species="discolor" status="sp. nov.">
<emphasis id="B92C261E5B40BA33FD8BED281EE14F6E" box="[568,660,1117,1143]" italics="true" pageId="1" pageNumber="576">discolor</emphasis>
</taxonomicName>
<emphasis id="B92C261E5B40BA33FD28ED281EA74F6E" bold="true" box="[667,722,1117,1143]" pageId="1" pageNumber="576">
<taxonomicNameLabel id="A21F9B655B40BA33FD28ED281EA74F6E" box="[667,722,1117,1143]" pageId="1" pageNumber="576" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
| Shavrin A.
<collectionCode id="ED4962C95B40BA33FCEDED2B1F014F61" box="[862,884,1118,1144]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="1" pageNumber="576" type="Museum">V</collectionCode>
. des. 2022’ &lt;red, printed&gt; (
<specimenCount id="9D5E31855B40BA33FB18ED2818994F61" box="[1195,1260,1117,1144]" pageId="1" pageNumber="576" type="male">3 ♂♂</specimenCount>
,
<specimenCount id="9D5E31855B40BA33FB4BED28194F4F61" box="[1272,1338,1117,1144]" pageId="1" pageNumber="576" type="female">3 ♀♀</specimenCount>
: cSh;
<specimenCount id="9D5E31855B40BA33FA33ED281CB64F85" pageId="1" pageNumber="576" type="male">21 ♂♂</specimenCount>
,
<specimenCount id="9D5E31855B40BA33FF7CEDF41D6B4F85" box="[207,286,1153,1180]" pageId="1" pageNumber="576" type="female">24 ♀♀</specimenCount>
:
<collectionCode id="ED4962C95B40BA33FE98EDF41DF54F82" box="[299,384,1153,1179]" country="Czech Republic" httpUri="http://grbio.org/cool/hj12-2xm7" name="National Museum Prague" pageId="1" pageNumber="576">NMPC</collectionCode>
).
</paragraph>
</subSubSection>
<subSubSection id="C342A9875B40BA31FF74EDD01E674844" lastPageId="3" lastPageNumber="578" pageId="1" pageNumber="576" type="description">
<paragraph id="8BE7FA0C5B40BA33FF74EDD019354E1E" blockId="1.[151,1437,937,2008]" pageId="1" pageNumber="576">
<emphasis id="B92C261E5B40BA33FF74EDD01D2C4FA6" bold="true" box="[199,345,1189,1215]" pageId="1" pageNumber="576">Description.</emphasis>
Measurements (n=52): HW: 0.47–0.60; HL: 0.27–0.35; AL(
<typeStatus id="54E344AE5B40BA33FB94EDD018FB4FA6" box="[1063,1166,1189,1215]" pageId="1" pageNumber="576" type="holotype">holotype</typeStatus>
): 1.12; OL: 0.15–0.17; PL: 0.47– 0.65; PW: 0.75–0.90; ESL: 0.88–1.20; EW: 0.95–1.02; AW: 0.94–1.07; MTbL(
<typeStatus id="54E344AE5B40BA33FB50EDBC193C4FFA" box="[1251,1353,1225,1251]" pageId="1" pageNumber="576" type="holotype">holotype</typeStatus>
): 0.57; MTrL(
<typeStatus id="54E344AE5B40BA33FF54ED981D394E1E" box="[231,332,1261,1287]" pageId="1" pageNumber="576" type="holotype">holotype</typeStatus>
): 0.32 (MTrL 1–4: 0.15; MTrL 5: 0.17); AedL: 0.45–0.60; TL: 2.35–3.35 (
<typeStatus id="54E344AE5B40BA33FB3FED9818814E1E" box="[1164,1268,1261,1287]" pageId="1" pageNumber="576" type="holotype">holotype</typeStatus>
: 2.70).
</paragraph>
<paragraph id="8BE7FA0C5B40BA33FF74EC64195F4DC2" blockId="1.[151,1437,937,2008]" pageId="1" pageNumber="576">
Habitus as in
<figureCitation id="1363E6895B40BA33FED6EC641DA74E32" box="[357,466,1297,1323]" captionStart="FIGURES 1–2" captionStartId="2.[151,264,1849,1873]" captionTargetBox="[170,1425,609,1812]" captionTargetId="figure-196@2.[151,1436,594,1823]" captionTargetPageId="2" captionText="FIGURES 1–2. Habitus of Amphichroum discolor: 1–holotype, 2–paratype. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7138574" httpUri="https://zenodo.org/record/7138574/files/figure.png" pageId="1" pageNumber="576">Figs. 1–2</figureCitation>
. Body yellow-brown to brown; antennomeres 3–11 (brown specimens) or 5–11 (pale specimens) brownish; mouthparts, antennomeres 1–2 or 1–4 and legs yellow to yellow-brown (some specimens with yellow-brown forebody and narrow brown longitudinal spot along midline (
<figureCitation id="1363E6895B40BA33FC59EC2C185B4E6A" box="[1002,1070,1369,1395]" captionStart="FIGURES 1–2" captionStartId="2.[151,264,1849,1873]" captionTargetBox="[170,1425,609,1812]" captionTargetId="figure-196@2.[151,1436,594,1823]" captionTargetPageId="2" captionText="FIGURES 1–2. Habitus of Amphichroum discolor: 1–holotype, 2–paratype. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7138574" httpUri="https://zenodo.org/record/7138574/files/figure.png" pageId="1" pageNumber="576">Fig. 2</figureCitation>
), brown abdomen and yellowish paratergites; pronotum of specimens with brown body (
<figureCitation id="1363E6895B40BA33FC93EC081F114E8E" box="[800,868,1405,1431]" captionStart="FIGURES 1–2" captionStartId="2.[151,264,1849,1873]" captionTargetBox="[170,1425,609,1812]" captionTargetId="figure-196@2.[151,1436,594,1823]" captionTargetPageId="2" captionText="FIGURES 1–2. Habitus of Amphichroum discolor: 1–holotype, 2–paratype. Scale bar: 1.0 mm." figureDoi="http://doi.org/10.5281/zenodo.7138574" httpUri="https://zenodo.org/record/7138574/files/figure.png" pageId="1" pageNumber="576">Fig. 1</figureCitation>
) with yellow-brown lateral margins and slightly paler elytra). Body shiny; head with dense transverse microsculpture, denser and isodiametric on infraorbital ridges (some pale specimens without distinct meshes on infraorbital ridges); neck with distinct isodiametric or transverse microreticulation; pronotum with indistinct transverse microsculpure or without it; visible part of scutellum with very fine transverse meshes or without them; abdominal tergites with very dense isodiametric microsculpture. Head with several very fine and sparse punctures in middle; pronotum with moderately dense, large and deep punctation, sparser in middle and mediobasal portions; punctation of elytra distinctly larger, deeper and denser than that on pronotum, denser and coarser in mediobasal portion, finer and sparser along suture; abdominal tergites with indistinct and very sparse moderately fine punctation. Frontal portion of head with several long setae; pronotum and elytra with dense long setation; abdominal tergites with regular, short and moderately dense pubescence.
</paragraph>
<paragraph id="8BE7FA0C5B40BA30FF74EF9018144BE0" blockId="1.[151,1437,937,2008]" lastBlockId="2.[151,1437,151,573]" lastPageId="2" lastPageNumber="577" pageId="1" pageNumber="576">Head 1.7 times as broad as long, distinctly flattened in middle and slightly elevated on infraorbital ridges; anteocellar foveae (grooves in front of ocelli) narrow and very long, diagonally stretching toward antennal bases to about level of anterior margins of eyes. Ocelli large, located at level of posterior margins of eyes near occipital furrow, distance between ocelli about as long as distance between ocellus and posterior margin of eye or slightly longer. Eyes very large, convex. Apical segment of maxillary palpi 1.1–1.3 times as long as preceding segment, from basal portion gradually narrowed apicad. Antenna moderately long, reaching basal third of elytral length when reclined; basal antennomere moderately long, 1.4–1.6 times as long as broad, antennomere 2 about as broad as and distinctly shorter than basal antennomere, 3 about as long as and narrower than 2, 4 slightly shorter and broader than 3, 5–9 about as long as and distinctly broader than 4, 10 slightly shorter and broader than 9, apical antennomere about twice longer than 10, from apical third strongly narrowed toward subacute apex.</paragraph>
<paragraph id="8BE7FA0C5B43BA30FF74E876191B4A7C" blockId="2.[151,1437,151,573]" pageId="2" pageNumber="577">Pronotum convex, 1.3–1.5 times as broad as long, 1.5 times as broad as head, widest in middle, gradually or slightly more narrowed anteriad than posteriad; anterior angles rounded, distinctly protruded apicad; posterior angles widely rounded; lateral portions widely flattened and slightly explanate, gradually broadened posteriad.</paragraph>
<paragraph id="8BE7FA0C5B43BA30FF74E81A1FFA4AB4" blockId="2.[151,1437,151,573]" pageId="2" pageNumber="577">Elytra about as broad as long, distinctly broadened apicad, reaching apical margin of abdominal tergite IV or V, 1.8 times as long as pronotum, with widely rounded apical margins.</paragraph>
<paragraph id="8BE7FA0C5B43BA30FF74E8C21C854AEC" blockId="2.[151,1437,151,573]" pageId="2" pageNumber="577">Legs moderately long; metatibia 1.7 times as long as metatrasus, with long strong thorns on inner and outer surface.</paragraph>
<paragraph id="8BE7FA0C5B43BA30FF74E88A1E824924" blockId="2.[151,1437,151,573]" pageId="2" pageNumber="577">Abdomen about as broad as elytra, with a pair of small transverse tomentose spots in middle of tergite IV, with narrow palisade fringe on apical margin of tergite VII.</paragraph>
<caption id="DF27AA845B43BA30FF24EE4C18174C48" ID-DOI="http://doi.org/10.5281/zenodo.7138574" ID-Zenodo-Dep="7138574" box="[151,1122,1849,1873]" httpUri="https://zenodo.org/record/7138574/files/figure.png" pageId="2" pageNumber="577" startId="2.[151,264,1849,1873]" targetBox="[170,1425,609,1812]" targetPageId="2">
<paragraph id="8BE7FA0C5B43BA30FF24EE4C18174C48" blockId="2.[151,1122,1849,1873]" box="[151,1122,1849,1873]" pageId="2" pageNumber="577">
<emphasis id="B92C261E5B43BA30FF24EE4C1D4F4C48" bold="true" box="[151,314,1849,1873]" pageId="2" pageNumber="577">FIGURES 1–2.</emphasis>
Habitus of
<taxonomicName id="4C58818F5B43BA30FE02EE4C1EEC4C48" authorityName="Shavrin" authorityYear="2022" box="[433,665,1849,1873]" class="Insecta" family="Staphylinidae" genus="Amphichroum" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="577" phylum="Arthropoda" rank="species" species="discolor">
<emphasis id="B92C261E5B43BA30FE02EE4C1EEC4C48" box="[433,665,1849,1873]" italics="true" pageId="2" pageNumber="577">Amphichroum discolor</emphasis>
</taxonomicName>
: 1–holotype, 2–paratype. Scale bar: 1.0 mm.
</paragraph>
</caption>
<paragraph id="8BE7FA0C5B43BA31FF74EE0A1D6E4A04" blockId="2.[151,1436,1919,2018]" lastBlockId="3.[151,1437,151,609]" lastPageId="3" lastPageNumber="578" pageId="2" pageNumber="577">
Male. Protarsomeres 1–4 wide. Medial margin of apical half of protibia with two parallel rows of several short peg setae; mesotibia strongly curved mediad, with dense row of 17–21 very short thorns beginning from medial side of bend and stretching to apex of mesotibia. Apical margin of abdominal tergite VIII truncate. Apical margin of abdominal sternite VIII slightly concave. Aedeagus with very wide basal portion, strongly narrowed apicad toward very narrow and long apical portion of median lobe; parameres narrow, slightly exceeding apex of median lobe, with three short apical setae; internal sac long, with two fields of small thorns (
<figureCitation id="1363E6895B42BA31FC67E9AA18424BE0" box="[980,1079,223,249]" captionStart="FIGURES 3–10" captionStartId="4.[151,264,1983,2007]" captionTargetBox="[161,1424,355,1952]" captionTargetId="figure-103@4.[151,1436,345,1959]" captionTargetPageId="4" captionText="FIGURES 3–10. Aedeagus of Amphichroum (Figs. 3–5: A. discolor; Fig. 6 (apical portion): A. rotundatum; Figs. 7–8: A. grandidentatum; Figs. 9–10: A. reticulatum): 3–4, 6–7, 9–parameral view, 5, 8, 10–lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7138578" httpUri="https://zenodo.org/record/7138578/files/figure.png" pageId="3" pageNumber="578">Fig. 3–4</figureCitation>
). Lateral aspect of the aedeagus as in
<figureCitation id="1363E6895B42BA31FF61E8761D634A04" box="[210,278,258,285]" captionStart="FIGURES 3–10" captionStartId="4.[151,264,1983,2007]" captionTargetBox="[161,1424,355,1952]" captionTargetId="figure-103@4.[151,1436,345,1959]" captionTargetPageId="4" captionText="FIGURES 3–10. Aedeagus of Amphichroum (Figs. 3–5: A. discolor; Fig. 6 (apical portion): A. rotundatum; Figs. 7–8: A. grandidentatum; Figs. 9–10: A. reticulatum): 3–4, 6–7, 9–parameral view, 5, 8, 10–lateral view. Scale bar: 0.1 mm." figureDoi="http://doi.org/10.5281/zenodo.7138578" httpUri="https://zenodo.org/record/7138578/files/figure.png" pageId="3" pageNumber="578">Fig. 5</figureCitation>
.
</paragraph>
<paragraph id="8BE7FA0C5B42BA31FF74E8521C9B4A90" blockId="3.[151,1437,151,609]" pageId="3" pageNumber="578">Female.Protarsomeres 1–4narrow.Medial margin of protibia without peg setae; mesotibia without modifications, slender, gradually slightly widened apicad. Abdominal tergite VIII and sternite VIII with truncate or rounded apical margin.</paragraph>
<paragraph id="8BE7FA0C5B42BA31FF74E8E71D074978" blockId="3.[151,1437,151,609]" pageId="3" pageNumber="578">
<emphasis id="B92C261E5B42BA31FF74E8E71DC54AB4" bold="true" box="[199,432,402,429]" pageId="3" pageNumber="578">Comparative notes.</emphasis>
Based on the general shape of the body and apical portion of the median lobe, it is similar to Chinese
<taxonomicName id="4C58818F5B42BA31FEA8E8C21F194AC8" authority="Shavrin &amp; Smetana, 2018" authorityName="Shavrin &amp; Smetana" authorityYear="2018" box="[283,876,439,465]" class="Insecta" family="Staphylinidae" genus="Amphichroum" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="578" phylum="Arthropoda" rank="species" species="subaequale">
<emphasis id="B92C261E5B42BA31FEA8E8C21D594AC8" box="[283,300,439,465]" italics="true" pageId="3" pageNumber="578">A</emphasis>
.
<emphasis id="B92C261E5B42BA31FE8DE8C21DCB4AC8" box="[318,446,439,465]" italics="true" pageId="3" pageNumber="578">subaequale</emphasis>
<bibRefCitation id="EFC987FD5B42BA31FE74E8C21E874AC8" author="Shavrin, A. V. &amp; Smetana, A." box="[455,754,439,465]" pageId="3" pageNumber="578" pagination="377 - 402" refId="ref5394" refString="Shavrin, A. V. &amp; Smetana, A. (2018) New species of the genus Amphichroum Kraatz, 1857 (Coleoptera: Staphylinidae: Omaliinae: Anthophagini) from China. Zootaxa, 4508 (3), 377 - 402. https: // doi. org / 10.11646 / zootaxa. 4508.3.3" type="journal article" year="2018">Shavrin &amp; Smetana, 2018</bibRefCitation>
(Shaanxi)
</taxonomicName>
,
<taxonomicName id="4C58818F5B42BA31FCC9E8C2194C4AC8" authority="Shavrin, 2022" authorityName="Shavrin" authorityYear="2022" box="[890,1337,439,465]" class="Insecta" family="Staphylinidae" genus="Amphichroum" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="578" phylum="Arthropoda" rank="species" species="cuccodoroi">
<emphasis id="B92C261E5B42BA31FCC9E8C21FFE4AC8" box="[890,907,439,465]" italics="true" pageId="3" pageNumber="578">A</emphasis>
.
<emphasis id="B92C261E5B42BA31FC2EE8C2186F4AC8" box="[925,1050,439,465]" italics="true" pageId="3" pageNumber="578">cuccodoroi</emphasis>
<bibRefCitation id="EFC987FD5B42BA31FB97E8C218B14AC8" author="Shavrin, A. V." box="[1060,1220,439,465]" pageId="3" pageNumber="578" pagination="143 - 148" refId="ref5335" refString="Shavrin, A. V. (2022) Two new species and additional records of Amphichroum Kraatz, 1857 from Nepal, China and India (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa, 5104 (1), 143 - 148. https: // doi. org / 10.11646 / zootaxa. 5104.1.9" type="journal article" year="2022">Shavrin, 2022</bibRefCitation>
(Yunnan)
</taxonomicName>
, and the Himalayan
<taxonomicName id="4C58818F5B42BA31FEA4E8AE1E2F4AEC" authority="Cameron, 1941" authorityName="Cameron" authorityYear="1941" box="[279,602,475,501]" class="Insecta" family="Staphylinidae" genus="Amphichroum" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="578" phylum="Arthropoda" rank="species" species="altivagans">
<emphasis id="B92C261E5B42BA31FEA4E8AE1D5D4AEC" box="[279,296,475,501]" italics="true" pageId="3" pageNumber="578">A</emphasis>
.
<emphasis id="B92C261E5B42BA31FE86E8AE1DDF4AEC" box="[309,426,475,501]" italics="true" pageId="3" pageNumber="578">altivagans</emphasis>
Cameron, 1941
</taxonomicName>
(
<collectingCountry id="F34FBA9C5B42BA31FDD4E8AE1ED34AEC" box="[615,678,475,501]" name="India" pageId="3" pageNumber="578">India</collectingCountry>
: Kashmir) and
<taxonomicName id="4C58818F5B42BA31FCFFE8AE18C04AEC" authority="Shavrin, 2021" authorityName="Shavrin" authorityYear="2021" box="[844,1205,475,501]" class="Insecta" family="Staphylinidae" genus="Amphichroum" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="578" phylum="Arthropoda" rank="species" species="telnovi">
<emphasis id="B92C261E5B42BA31FCFFE8AE1F284AEC" box="[844,861,475,501]" italics="true" pageId="3" pageNumber="578">A</emphasis>
.
<emphasis id="B92C261E5B42BA31FCDAE8AE1FC04AEC" box="[873,949,475,501]" italics="true" pageId="3" pageNumber="578">telnovi</emphasis>
<bibRefCitation id="EFC987FD5B42BA31FC08E8AE18234AEC" author="Shavrin, A. V." box="[955,1110,475,501]" pageId="3" pageNumber="578" pagination="1 - 14" refId="ref5276" refString="Shavrin, A. V. (2021) New species and records of the genus Amphichroum Kraatz, 1857 from the Himalayan Region and Tibet (Coleoptera: Staphylinidae: Omaliinae: Anthophagini). Zootaxa, 5082 (1), 1 - 14. https: // doi. org / 10.11646 / zootaxa. 5082.1.1" type="journal article" year="2021">Shavrin, 2021</bibRefCitation>
(
<collectingCountry id="F34FBA9C5B42BA31FBD0E8AE18D94AEC" box="[1123,1196,475,501]" name="Nepal" pageId="3" pageNumber="578">Nepal</collectingCountry>
)
</taxonomicName>
. From
<taxonomicName id="4C58818F5B42BA31FB4CE8AE19E84AEC" authorityName="Shavrin &amp; Smetana" authorityYear="2018" box="[1279,1437,475,501]" class="Insecta" family="Staphylinidae" genus="Amphichroum" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="578" phylum="Arthropoda" rank="species" species="subaequale">
<emphasis id="B92C261E5B42BA31FB4CE8AE19654AEC" box="[1279,1296,475,501]" italics="true" pageId="3" pageNumber="578">A</emphasis>
.
<emphasis id="B92C261E5B42BA31FAAEE8AE19E84AEC" box="[1309,1437,475,501]" italics="true" pageId="3" pageNumber="578">subaequale</emphasis>
</taxonomicName>
it differs by the less transverse pronotum, and shorter and narrower parameres. From
<taxonomicName id="4C58818F5B42BA31FBEFE88A188C4900" authorityName="Shavrin" authorityYear="2022" box="[1116,1273,511,537]" class="Insecta" family="Staphylinidae" genus="Amphichroum" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="578" phylum="Arthropoda" rank="species" species="cuccodoroi">
<emphasis id="B92C261E5B42BA31FBEFE88A18184900" box="[1116,1133,511,537]" italics="true" pageId="3" pageNumber="578">A</emphasis>
.
<emphasis id="B92C261E5B42BA31FBCFE88A188C4900" box="[1148,1273,511,537]" italics="true" pageId="3" pageNumber="578">cuccodoroi</emphasis>
</taxonomicName>
,
<taxonomicName id="4C58818F5B42BA31FABBE88A19E84900" authorityName="Cameron" authorityYear="1941" box="[1288,1437,511,537]" class="Insecta" family="Staphylinidae" genus="Amphichroum" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="578" phylum="Arthropoda" rank="species" species="altivagans">
<emphasis id="B92C261E5B42BA31FABBE88A196C4900" box="[1288,1305,511,537]" italics="true" pageId="3" pageNumber="578">A</emphasis>
.
<emphasis id="B92C261E5B42BA31FA9BE88A19E84900" box="[1320,1437,511,537]" italics="true" pageId="3" pageNumber="578">altivagans</emphasis>
</taxonomicName>
and
<taxonomicName id="4C58818F5B42BA31FF79EB561D4D4924" authorityName="Shavrin" authorityYear="2021" box="[202,312,547,573]" class="Insecta" family="Staphylinidae" genus="Amphichroum" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="578" phylum="Arthropoda" rank="species" species="telnovi">
<emphasis id="B92C261E5B42BA31FF79EB561CAE4924" box="[202,219,547,573]" italics="true" pageId="3" pageNumber="578">A</emphasis>
.
<emphasis id="B92C261E5B42BA31FF5FEB561D4D4924" box="[236,312,547,573]" italics="true" pageId="3" pageNumber="578">telnovi</emphasis>
</taxonomicName>
it differs by the absence of the large sclerotized teeth in the internal sac. Additionally, it can be distinguished from:
</paragraph>
<paragraph id="8BE7FA0C5B42BA31FF74EBFA197C49B0" blockId="3.[199,1289,655,753]" box="[199,1289,655,681]" pageId="3" pageNumber="578">
<taxonomicName id="4C58818F5B42BA31FF74EBFA1D1649B0" authorityName="Shavrin" authorityYear="2022" box="[199,355,655,681]" class="Insecta" family="Staphylinidae" genus="Amphichroum" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="578" phylum="Arthropoda" rank="species" species="cuccodoroi">
<emphasis id="B92C261E5B42BA31FF74EBFA1CAD49B0" box="[199,216,655,681]" italics="true" pageId="3" pageNumber="578">A</emphasis>
.
<emphasis id="B92C261E5B42BA31FF55EBFA1D1649B0" box="[230,355,655,681]" italics="true" pageId="3" pageNumber="578">cuccodoroi</emphasis>
</taxonomicName>
by the less transverse pronotum, longer elytra, and significantly shorter parameres;
</paragraph>
<paragraph id="8BE7FA0C5B42BA31FF74EBC61F1E49D4" blockId="3.[199,1289,655,753]" box="[199,875,691,717]" pageId="3" pageNumber="578">
<taxonomicName id="4C58818F5B42BA31FF74EBC61D2E49D4" authorityName="Cameron" authorityYear="1941" box="[199,347,691,717]" class="Insecta" family="Staphylinidae" genus="Amphichroum" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="578" phylum="Arthropoda" rank="species" species="altivagans">
<emphasis id="B92C261E5B42BA31FF74EBC61CAD49D4" box="[199,216,691,717]" italics="true" pageId="3" pageNumber="578">A</emphasis>
.
<emphasis id="B92C261E5B42BA31FF55EBC61D2E49D4" box="[230,347,691,717]" italics="true" pageId="3" pageNumber="578">altivagans</emphasis>
</taxonomicName>
by the longer antennomeres and shorter elytra;
</paragraph>
<paragraph id="8BE7FA0C5B42BA31FF74EBA2188849E8" blockId="3.[199,1289,655,753]" box="[199,1277,727,753]" pageId="3" pageNumber="578">
<taxonomicName id="4C58818F5B42BA31FF74EBA21D4749E8" authorityName="Shavrin" authorityYear="2021" box="[199,306,727,753]" class="Insecta" family="Staphylinidae" genus="Amphichroum" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="578" phylum="Arthropoda" rank="species" species="telnovi">
<emphasis id="B92C261E5B42BA31FF74EBA21CAD49E8" box="[199,216,727,753]" italics="true" pageId="3" pageNumber="578">A</emphasis>
.
<emphasis id="B92C261E5B42BA31FF55EBA21D4749E8" box="[230,306,727,753]" italics="true" pageId="3" pageNumber="578">telnovi</emphasis>
</taxonomicName>
by the smaller body, longer antennomeres 6–10, slightly broader pronotum and elytra.
</paragraph>
<paragraph id="8BE7FA0C5B42BA31FF74EA6A1E674844" blockId="3.[151,1436,799,1077]" pageId="3" pageNumber="578">From all these species it can be distinguished by the darker elytra, and the shape of very narrow and elongate apical portion of the median lobe.</paragraph>
</subSubSection>
<subSubSection id="C342A9875B42BA31FF74EA131E4D48F4" pageId="3" pageNumber="578" type="distribution">
<paragraph id="8BE7FA0C5B42BA31FF74EA1319264898" blockId="3.[151,1436,799,1077]" box="[199,1363,870,897]" pageId="3" pageNumber="578">
<emphasis id="B92C261E5B42BA31FF74EA131D144899" bold="true" box="[199,353,870,896]" pageId="3" pageNumber="578">Distribution.</emphasis>
The species is at present known only from the
<typeStatus id="54E344AE5B42BA31FCC2EA121FD44898" box="[881,929,871,897]" pageId="3" pageNumber="578">type</typeStatus>
locality in Biluo Mts.,
<collectingRegion id="499C34EE5B42BA31FB17EA1218884898" box="[1188,1277,871,897]" country="China" name="Yunnan" pageId="3" pageNumber="578">Yunnan</collectingRegion>
,
<collectingCountry id="F34FBA9C5B42BA31FABBEA12193A4898" box="[1288,1359,871,897]" name="China" pageId="3" pageNumber="578">China</collectingCountry>
.
</paragraph>
<paragraph id="8BE7FA0C5B42BA31FF74EAFF1E4D48F4" blockId="3.[151,1436,799,1077]" pageId="3" pageNumber="578">
<emphasis id="B92C261E5B42BA31FF74EAFF1D3E48BD" bold="true" box="[199,331,906,932]" pageId="3" pageNumber="578">Bionomics.</emphasis>
Specimens were collected at elevation from
<quantity id="4CA057E95B42BA31FCFEEAFE187648BC" box="[845,1027,907,933]" metricMagnitude="3" metricUnit="m" metricValue="3.9" metricValueMax="3.91" metricValueMin="3.8899999999999997" pageId="3" pageNumber="578" unit="m" value="3900.0" valueMax="3910.0" valueMin="3890.0">3890 to 3910 m</quantity>
a.s.l. together with specimens of
<taxonomicName id="4C58818F5B42BA31FA37EAFE1D3F48D0" authority="Shavrin, 2022" authorityName="Shavrin" authorityYear="2022" class="Insecta" family="Staphylinidae" genus="Amphichroum" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="578" phylum="Arthropoda" rank="species" species="grandidentatum" status="sp. nov.">
<emphasis id="B92C261E5B42BA31FA37EAFE19E048BC" box="[1412,1429,907,933]" italics="true" pageId="3" pageNumber="578">A</emphasis>
.
<emphasis id="B92C261E5B42BA31FF24EADA1D3F48D0" box="[151,330,943,969]" italics="true" pageId="3" pageNumber="578">grandidentatum</emphasis>
</taxonomicName>
<emphasis id="B92C261E5B42BA31FEE1EADA1DFC48D0" bold="true" box="[338,393,943,969]" pageId="3" pageNumber="578">
<taxonomicNameLabel id="A21F9B655B42BA31FEE1EADA1DFC48D0" box="[338,393,943,969]" pageId="3" pageNumber="578" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
(see below). They were taken by sifting litter under shrubs and along fallen trunks in mixed forest with
<taxonomicName id="4C58818F5B42BA31FEA4EAA61D2048F4" box="[279,341,979,1005]" class="Pinopsida" family="Pinaceae" genus="Abies" kingdom="Plantae" order="Pinales" pageId="3" pageNumber="578" phylum="Tracheophyta" rank="genus">
<emphasis id="B92C261E5B42BA31FEA4EAA61D2048F4" box="[279,341,979,1005]" italics="true" pageId="3" pageNumber="578">Abies</emphasis>
</taxonomicName>
and
<taxonomicName id="4C58818F5B42BA31FE38EAA61E4548F4" box="[395,560,979,1005]" class="Magnoliopsida" family="Ericaceae" genus="Rhododendron" kingdom="Plantae" order="Ericales" pageId="3" pageNumber="578" phylum="Tracheophyta" rank="genus">
<emphasis id="B92C261E5B42BA31FE38EAA61E4548F4" box="[395,560,979,1005]" italics="true" pageId="3" pageNumber="578">Rhododendron</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C342A9875B42BA31FF74EA821D044F2C" pageId="3" pageNumber="578" type="etymology">
<paragraph id="8BE7FA0C5B42BA31FF74EA821D044F2C" blockId="3.[151,1436,799,1077]" pageId="3" pageNumber="578">
<emphasis id="B92C261E5B42BA31FF74EA821D3B4F08" bold="true" box="[199,334,1015,1041]" pageId="3" pageNumber="578">Etymology.</emphasis>
The specific epithet (Latin adjective: of different colours) refers to the different coloration of the body of specimens.
</paragraph>
</subSubSection>
</treatment>
</document>
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