137 lines
18 KiB
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137 lines
18 KiB
XML
<document id="92F0FA1696D9C5FAB02DFA89D5480D5D" ID-CLB-Dataset="27842" ID-DOI="10.11646/zootaxa.4527.3.9" ID-GBIF-Dataset="195c5a0b-ee6c-4865-ae6e-52403a5b0bc5" ID-ISSN="1175-5326" ID-Zenodo-Dep="2612371" ID-ZooBank="0BAB4687-ED83-416D-9573-8668E05F6E66" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1553698805300" checkinUser="plazi" docAuthor="Fiemapong, Armand Richard Nzoko & Enghoff, Henrik" docDate="2018" docId="03EF87FDFF9FB75AFF1016E1FBCCDEBC" docLanguage="en" docName="zootaxa.4527.3.9.pdf" docOrigin="Zootaxa 4527 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Pelmatojulus Saussure 1860" docType="treatment" docVersion="7" lastPageNumber="404" masterDocId="FFD6FF85FF9EB75BFF871528FFF2DB14" masterDocTitle="A giant African millipede re-discovered, re-located and re-classified after 120 years (Diplopoda, Spirobolida, Pachybolidae)" masterLastPageNumber="413" masterPageNumber="403" pageNumber="404" updateTime="1698714523762" updateUser="ExternalLinkService">
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<mods:title id="93E38DF362201118AABE51340C20DAC9">A giant African millipede re-discovered, re-located and re-classified after 120 years (Diplopoda, Spirobolida, Pachybolidae)</mods:title>
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<mods:namePart id="AADEB5C13A0F6A51F92999BE23750C44">Fiemapong, Armand Richard Nzoko</mods:namePart>
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<mods:namePart id="CA9F089E27DF338EB214DB106C7CFF47">Enghoff, Henrik</mods:namePart>
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<mods:title id="33C0D9519B36EFBC25A30454FFD15B62">Zootaxa</mods:title>
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<mods:part id="70199D17178975ECDFFE580F35B91C48">
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<mods:date id="EFC70B608D239DA81CFC82CE0BD22262">2018</mods:date>
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<mods:number id="C5ED4CF48D0C1AE1A39AA783B2A73385">2018-12-06</mods:number>
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<mods:number id="1FFF63370F5539C6B2AD44EBDB30BD65">4527</mods:number>
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<treatment id="03EF87FDFF9FB75AFF1016E1FBCCDEBC" ID-DOI="http://doi.org/10.5281/zenodo.5990957" ID-GBIF-Taxon="154508268" ID-Zenodo-Dep="5990957" LSID="urn:lsid:plazi:treatment:03EF87FDFF9FB75AFF1016E1FBCCDEBC" httpUri="http://treatment.plazi.org/id/03EF87FDFF9FB75AFF1016E1FBCCDEBC" lastPageNumber="404" pageId="1" pageNumber="404">
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<subSubSection id="C35C6560FF9FB75AFF1016E1FDBCD8F7" box="[151,590,968,995]" pageId="1" pageNumber="404" type="nomenclature">
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<paragraph id="8BF936EBFF9FB75AFF1016E1FDBCD8F7" blockId="1.[151,590,968,995]" box="[151,590,968,995]" pageId="1" pageNumber="404">
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<heading id="D0B18187FF9FB75AFF1016E1FDBCD8F7" bold="true" box="[151,590,968,995]" fontSize="11" level="1" pageId="1" pageNumber="404" reason="1">
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<emphasis id="B932EAF9FF9FB75AFF1016E1FDBCD8F7" bold="true" box="[151,590,968,995]" pageId="1" pageNumber="404">
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Genus
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<taxonomicName id="4C464D68FF9FB75AFF6916E0FDBCD8F7" ID-CoL="7PSWD" authority="Saussure, 1860" authorityName="Saussure" authorityYear="1860" box="[238,590,968,995]" class="Diplopoda" family="Pachybolidae" genus="Pelmatojulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirobolida" pageId="1" pageNumber="404" phylum="Arthropoda" rank="genus">
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<emphasis id="B932EAF9FF9FB75AFF6916E0FE7FD8F6" bold="true" box="[238,397,968,994]" italics="true" pageId="1" pageNumber="404">Pelmatojulus</emphasis>
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<bibRefCitation id="EFD74B1AFF9FB75AFE1216E1FDBCD8F7" author="Saussure, H. L. F. de" box="[405,590,969,995]" pageId="1" pageNumber="404" pagination="1 - 135" refId="ref4047" refString="Saussure, H. L. F. de (1860). Essai d`une faune des Myriapodes du Mexique avec la description de quelques especes des autres parties de l'Amerique. Memoires de la Societe de Physiques et d'Histoire naturelle de Geneve, 15 (2), 1 - 135." type="journal article" year="1860">Saussure, 1860</bibRefCitation>
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</taxonomicName>
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</emphasis>
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</heading>
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</paragraph>
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</subSubSection>
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<subSubSection id="C35C6560FF9FB75AFF101127FBCCDEBC" pageId="1" pageNumber="404" type="reference_group">
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<paragraph id="8BF936EBFF9FB75AFF101127FB15DF57" blockId="1.[151,1369,1038,1091]" pageId="1" pageNumber="404">
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<taxonomicName id="4C464D68FF9FB75AFF101127FE4DDF30" ID-CoL="7PSWD" authority="Saussure, 1860" authorityName="Saussure" authorityYear="1860" box="[151,447,1038,1060]" class="Diplopoda" family="Pachybolidae" genus="Pelmatojulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirobolida" pageId="1" pageNumber="404" phylum="Arthropoda" rank="genus">
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<emphasis id="B932EAF9FF9FB75AFF101127FEEFDF30" box="[151,285,1039,1060]" italics="true" pageId="1" pageNumber="404">Pelmatojulus</emphasis>
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<bibRefCitation id="EFD74B1AFF9FB75AFEA41126FE4DDF30" author="Saussure, H. L. F. de" box="[291,447,1038,1060]" pageId="1" pageNumber="404" pagination="1 - 135" refId="ref4047" refString="Saussure, H. L. F. de (1860). Essai d`une faune des Myriapodes du Mexique avec la description de quelques especes des autres parties de l'Amerique. Memoires de la Societe de Physiques et d'Histoire naturelle de Geneve, 15 (2), 1 - 135." type="journal article" year="1860">Saussure, 1860</bibRefCitation>
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</taxonomicName>
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.
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<typeStatus id="54FD8849FF9FB75AFE4E1127FE09DF30" box="[457,507,1039,1060]" pageId="1" pageNumber="404">Type</typeStatus>
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species
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<taxonomicName id="4C464D68FF9FB75AFDD61127FC8FDF30" authority="Saussure, 1859" authorityName="Saussure" authorityYear="1859" box="[593,893,1038,1060]" class="Diplopoda" family="Julidae" genus="Julus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Julida" pageId="1" pageNumber="404" phylum="Arthropoda" rank="species" species="insignis">
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<emphasis id="B932EAF9FF9FB75AFDD61127FD29DF30" box="[593,731,1039,1060]" italics="true" pageId="1" pageNumber="404">Julus insignis</emphasis>
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<bibRefCitation id="EFD74B1AFF9FB75AFD651126FC8FDF30" author="Saussure, H. L. F. de" box="[738,893,1038,1060]" pageId="1" pageNumber="404" pagination="328 - 332" refId="ref4021" refString="Saussure, H. L. F. de (1859) Diagnose de divers Myriapodes nouveaux. Linnaea entomologica, 13, 328 - 332" type="journal article" year="1859">Saussure, 1859</bibRefCitation>
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</taxonomicName>
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, by subsequent designation of
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<bibRefCitation id="EFD74B1AFF9FB75AFB331126FAABDF30" author="Hoffman, R. L." box="[1204,1369,1038,1060]" pageId="1" pageNumber="404" pagination="759 - 783" refId="ref3723" refString="Hoffman, R. L. (1962) Studies on spiroboloid millipeds IV. Systematic and nomenclatorial notes on the family Pachybolidae. Revue Suisse de Zoologie, 69, 759 - 783. https: // doi. org / 10.5962 / bhl. part. 75592" type="journal article" year="1962">Hoffman (1962)</bibRefCitation>
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<taxonomicName id="4C464D68FF9FB75AFF101105FE7FDF57" ID-CoL="7PRBK" authority="Cook, 1897" authorityName="Cook" authorityYear="1897" box="[151,397,1069,1091]" class="Diplopoda" family="Pachybolidae" genus="Pachybolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirobolida" pageId="1" pageNumber="404" phylum="Arthropoda" rank="genus">
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<emphasis id="B932EAF9FF9FB75AFF101105FEFFDF56" box="[151,269,1069,1090]" italics="true" pageId="1" pageNumber="404">Pachybolus</emphasis>
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<bibRefCitation id="EFD74B1AFF9FB75AFE941105FE7FDF57" author="Cook, O. F." box="[275,397,1069,1091]" pageId="1" pageNumber="404" pagination="73 - 75" refId="ref3291" refString="Cook, O. F. (1897) New relatives of Spirobolus giganteus. Brandtia, 18, 73 - 75." type="journal article" year="1897">Cook, 1897</bibRefCitation>
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</taxonomicName>
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.
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<typeStatus id="54FD8849FF9FB75AFE111106FE3ADF57" box="[406,456,1070,1091]" pageId="1" pageNumber="404">Type</typeStatus>
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species
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<taxonomicName id="4C464D68FF9FB75AFD991105FCA6DF57" ID-CoL="4BQ4T" authority="Cook, 1897" authorityName="Cook" authorityYear="1897" box="[542,852,1069,1091]" class="Diplopoda" family="Pachybolidae" genus="Pachybolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirobolida" pageId="1" pageNumber="404" phylum="Arthropoda" rank="species" species="tectus">
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<emphasis id="B932EAF9FF9FB75AFD991105FD24DF57" box="[542,726,1069,1091]" italics="true" pageId="1" pageNumber="404">Pachybolus tectus</emphasis>
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<bibRefCitation id="EFD74B1AFF9FB75AFD5B1105FCA6DF57" author="Cook, O. F." box="[732,852,1069,1091]" pageId="1" pageNumber="404" pagination="73 - 75" refId="ref3291" refString="Cook, O. F. (1897) New relatives of Spirobolus giganteus. Brandtia, 18, 73 - 75." type="journal article" year="1897">Cook, 1897</bibRefCitation>
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</taxonomicName>
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, by original designation,
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<emphasis id="B932EAF9FF9FB75AFBD31106FB15DF57" bold="true" box="[1108,1255,1070,1091]" pageId="1" pageNumber="404">new synonym</emphasis>
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</paragraph>
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<paragraph id="8BF936EBFF9FB75AFF101158FB1ADE91" blockId="1.[151,1437,1135,1448]" pageId="1" pageNumber="404">
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<taxonomicName id="4C464D68FF9FB75AFF101158FED9DF9C" ID-CoL="7PSWD" authorityName="Saussure" authorityYear="1860" box="[151,299,1136,1160]" class="Diplopoda" family="Pachybolidae" genus="Pelmatojulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirobolida" pageId="1" pageNumber="404" phylum="Arthropoda" rank="genus">
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<emphasis id="B932EAF9FF9FB75AFF101158FED9DF9C" box="[151,299,1136,1160]" italics="true" pageId="1" pageNumber="404">Pelmatojulus</emphasis>
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</taxonomicName>
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belongs to the tribe
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<taxonomicName id="4C464D68FF9FB75AFDB51147FBE5DF9C" ID-CoL="845B9" authority="sensu Wesener et al. (2008)" authorityName="Wesener" authorityYear="2008" box="[562,1047,1135,1160]" class="Diplopoda" family="Pachybolidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirobolida" pageId="1" pageNumber="404" phylum="Arthropoda" rank="tribe" tribe="Pachybolini">
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Pachybolini
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<emphasis id="B932EAF9FF9FB75AFD411159FCF6DF9C" box="[710,772,1137,1160]" italics="true" pageId="1" pageNumber="404">sensu</emphasis>
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<bibRefCitation id="EFD74B1AFF9FB75AFC951158FBE5DF9C" author="Wesener, T. & Enghoff, H. & Wagele, J. - W." box="[786,1047,1136,1160]" pageId="1" pageNumber="404" pagination="37 - 53" refId="ref4305" refString="Wesener, T., Enghoff, H. & Wagele, J. - W. (2008) Pachybolini-a tribe of giant Afrotropical millipedes. With arguments for monophyly and the description of a new genus from Madagascar (Diplopoda: Spirobolida: Pachybolidae). Invertebrate Systematics, 22, 37 - 53. https: // doi. org / 10.1071 / IS 07008" type="journal article" year="2008">
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Wesener
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<emphasis id="B932EAF9FF9FB75AFCF81159FC4DDF9C" box="[895,959,1136,1160]" italics="true" pageId="1" pageNumber="404">et al.</emphasis>
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(2008)
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</bibRefCitation>
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</taxonomicName>
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and
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<bibRefCitation id="EFD74B1AFF9FB75AFBDC1147FAE3DF9C" author="Enghoff, H." box="[1115,1297,1135,1160]" pageId="1" pageNumber="404" pagination="1 - 41" refId="ref3692" refString="Enghoff, H. (2011) East African giant millipedes of the tribe Pachybolini (Diplopoda, Spirobolida, Pachybolidae), Zootaxa, 2753, 1 - 41." type="journal article" year="2011">Enghoff (2011)</bibRefCitation>
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. Diagnostic apomorphies of
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<taxonomicName id="4C464D68FF9FB75AFECA11BCFE21DFB9" ID-CoL="845B9" box="[333,467,1172,1197]" class="Diplopoda" family="Pachybolidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirobolida" pageId="1" pageNumber="404" phylum="Arthropoda" rank="tribe" tribe="Pachybolini">Pachybolini</taxonomicName>
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include a closed incisura lateralis on the head capsule, two or more setae placed in an apical transverse row on each lamella lingualis of the gnathochilarium; kidney- to crescent-shaped vulvae with the valves meeting in a more or less sinuous fissure.
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<taxonomicName id="4C464D68FF9FB75AFD4211F4FCABDFE0" ID-CoL="7PSWD" authorityName="Saussure" authorityYear="1860" box="[709,857,1244,1268]" class="Diplopoda" family="Pachybolidae" genus="Pelmatojulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirobolida" pageId="1" pageNumber="404" phylum="Arthropoda" rank="genus">
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<emphasis id="B932EAF9FF9FB75AFD4211F4FCABDFE0" box="[709,857,1244,1268]" italics="true" pageId="1" pageNumber="404">Pelmatojulus</emphasis>
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</taxonomicName>
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differs from all other genera of
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<taxonomicName id="4C464D68FF9FB75AFB4F11F4FABCDFE1" ID-CoL="845B9" box="[1224,1358,1244,1269]" class="Diplopoda" family="Pachybolidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirobolida" pageId="1" pageNumber="404" phylum="Arthropoda" rank="tribe" tribe="Pachybolini">Pachybolini</taxonomicName>
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by the apomorphic, much enlarged number (15 or more) of apical antennal sensilla (the other genera have four such sensilla). A further, semi-diagnostic character is provided by the abovementioned setae of the lamellae linguales: in
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<taxonomicName id="4C464D68FF9FB75AFF101060FED9DE74" ID-CoL="7PSWD" authorityName="Saussure" authorityYear="1860" box="[151,299,1352,1376]" class="Diplopoda" family="Pachybolidae" genus="Pelmatojulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirobolida" pageId="1" pageNumber="404" phylum="Arthropoda" rank="genus">
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<emphasis id="B932EAF9FF9FB75AFF101060FED9DE74" box="[151,299,1352,1376]" italics="true" pageId="1" pageNumber="404">Pelmatojulus</emphasis>
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</taxonomicName>
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there are usually two such setae per lamella, a condition shared with the likewise West African genus
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<taxonomicName id="4C464D68FF9FB75AFF101044FE53DE90" ID-CoL="7PGCY" authority="Enghoff, 2008" authorityName="Enghoff" authorityYear="2008" box="[151,417,1388,1413]" class="Diplopoda" family="Pachybolidae" genus="Gabolus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirobolida" pageId="1" pageNumber="404" phylum="Arthropoda" rank="genus">
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<emphasis id="B932EAF9FF9FB75AFF101044FF04DE90" box="[151,246,1388,1412]" italics="true" pageId="1" pageNumber="404">Gabolus</emphasis>
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<bibRefCitation id="EFD74B1AFF9FB75AFF7A1044FE53DE90" author="Wesener, T. & Enghoff, H. & Wagele, J. - W." box="[253,417,1388,1413]" pageId="1" pageNumber="404" pagination="37 - 53" refId="ref4305" refString="Wesener, T., Enghoff, H. & Wagele, J. - W. (2008) Pachybolini-a tribe of giant Afrotropical millipedes. With arguments for monophyly and the description of a new genus from Madagascar (Diplopoda: Spirobolida: Pachybolidae). Invertebrate Systematics, 22, 37 - 53. https: // doi. org / 10.1071 / IS 07008" type="journal article" year="2008">Enghoff, 2008</bibRefCitation>
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</taxonomicName>
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, whereas the other genera in the tribe have three or more setae per lamella.
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</paragraph>
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<paragraph id="8BF936EBFF9FB75AFF4010A7FBCCDEBC" blockId="1.[151,1437,1135,1448]" box="[199,1086,1423,1448]" pageId="1" pageNumber="404">
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The species of
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<taxonomicName id="4C464D68FF9FB75AFEE810B8FDF1DEBC" ID-CoL="7PSWD" authorityName="Saussure" authorityYear="1860" box="[367,515,1424,1448]" class="Diplopoda" family="Pachybolidae" genus="Pelmatojulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Spirobolida" pageId="1" pageNumber="404" phylum="Arthropoda" rank="genus">
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<emphasis id="B932EAF9FF9FB75AFEE810B8FDF1DEBC" box="[367,515,1424,1448]" italics="true" pageId="1" pageNumber="404">Pelmatojulus</emphasis>
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</taxonomicName>
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and their known distributions are listed in
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<tableCitation id="C6C40350FF9FB75AFC6310A7FBCADEBC" box="[996,1080,1423,1448]" captionStart="TABLE 1" captionStartId="8.[147,232,593,615]" captionText="TABLE 1. Species of Pelmatojulus and their geographical distribution (countries arranged roughly from West to East). Based on Aouti (1980), Attems (1935), Billah et al. (2015), Bourdanné (1997), Carl (1905), Chamberlin (1927), Cook (1899), Demange (1965, 1971a, b, 1972, 1981), Demange & Gasc (1972), Demange & Mauriès (1995), Hoffman & Mahsberg (1996), Pesewu et al. (2015) and Toye (1970). The record of P. tectus from Tanzania (Zanzibar) is not included, nor are unsubstantiated records from various websites. * =new record. Details of new records from Ghana, see footnotes. The records of P. ligulatus from Ghana by Billah et al. (2015) Pesewu et al. (2015) might be doubted because the identification was made in a non-taxonomic framework; the records are, however, backed up by material studied by us1." pageId="1" pageNumber="404">Table 1</tableCitation>
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.
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|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |