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<document id="676990614531B1858DCF9CCF2C7C90ED" ID-DOI="10.5281/zenodo.12584736" ID-ISSN="2064-2474" ID-Zenodo-Dep="12584736" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1719592945761" checkinUser="felipe" docAuthor="Brake, I., Papp, L. &amp; Bd, Uk." docDate="2009" docId="03EF87E31215E412FD1AFE2C4924BAF5" docLanguage="en" docName="ActaZoolAcadSciH.55.3.227-234.pdf" docOrigin="Acta Zoologica Academiae Scientiarum Hungaricae 55 (3)" docStyle="DocumentStyle:4FD749C6DF7B87D8940F4D8C7B0A0514.3:ActaZoolAcadSciH.2002-2014.journal_article" docStyleId="4FD749C6DF7B87D8940F4D8C7B0A0514" docStyleName="ActaZoolAcadSciH.2002-2014.journal_article" docStyleVersion="3" docTitle="Madiza africana Brake, Papp &amp; Bd, 2009, sp. n." docType="treatment" docVersion="2" lastPageNumber="231" masterDocId="FFD6FF9B1214E414FF95FF9B4B6FBE5C" masterDocTitle="A Peculiar New Species Of Madiza (Diptera: Milichiidae) From Tanzania" masterLastPageNumber="234" masterPageNumber="227" pageNumber="228" updateTime="1719594409504" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-4.0">
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<mods:title id="BBC4FE0497409AD5B55F37776F5F928E">A Peculiar New Species Of Madiza (Diptera: Milichiidae) From Tanzania</mods:title>
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<mods:name id="0C9F4C23ED1E25E2AA3E740727722C34" type="personal">
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<mods:namePart id="B0B848B504398B200225C567C4A1F246">Brake, I.</mods:namePart>
<mods:affiliation id="D22AF756FB62C7A3AED663CB57A547B3">and P, L. &amp; Department of Entomology, The Natural History Museum Cromwell Road, London SW 7 5 BD, UK. E-mail: i. brake @ nhm. ac. uk</mods:affiliation>
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<mods:namePart id="DA8B6E90D24C719C71BFDE8198BDABAE">Papp, L.</mods:namePart>
<mods:affiliation id="7CCF932F0832762674E212DD71A728B6">Department of Zoology, Hungarian Natural History Museum and Animal Ecology Research Group of the Hungarian Academy of Sciences PO. Box 137, H- 1431 Budapest, Hungary E-mail: lpapp @ nhmus. hu</mods:affiliation>
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<mods:namePart id="509168F753D3EBE51025169972F06479">Bd, Uk.</mods:namePart>
<mods:affiliation id="16DE5D0ED61F81772769183A891BB50C">Department of Zoology, Hungarian Natural History Museum and Animal Ecology Research Group of the Hungarian Academy of Sciences PO. Box 137, H- 1431 Budapest, Hungary E-mail: lpapp @ nhmus. hu</mods:affiliation>
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<mods:title id="8FD7DC82C2A1CD63314148E2D38D24A6">Acta Zoologica Academiae Scientiarum Hungaricae</mods:title>
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<mods:date id="EB78B1E44236E827414B243AD3B406B4">2009</mods:date>
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<treatment id="03EF87E31215E412FD1AFE2C4924BAF5" LSID="urn:lsid:plazi:treatment:03EF87E31215E412FD1AFE2C4924BAF5" httpUri="http://treatment.plazi.org/id/03EF87E31215E412FD1AFE2C4924BAF5" lastPageId="6" lastPageNumber="231" pageId="1" pageNumber="228">
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<taxonomicName id="4C464D761215E415FD1AFE2C4832BF8D" authorityName="Brake &amp; Papp &amp; Bd" authorityYear="2009" box="[655,861,439,465]" class="Insecta" family="Milichiidae" genus="Madiza" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="228" phylum="Arthropoda" rank="species" species="africana" status="sp. nov.">
<emphasis id="B932EAE71215E415FD1AFE2C4832BF8D" bold="true" box="[655,861,439,465]" pageId="1" pageNumber="228">Madiza africana</emphasis>
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<taxonomicNameLabel id="A201579C1215E415FCF0FE2C48CCBF8D" box="[869,931,439,465]" pageId="1" pageNumber="228" rank="species">sp. n.</taxonomicNameLabel>
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(
<figureCitation id="137D2A701215E415FD48FE40483BBFA9" box="[733,852,475,501]" captionStart-0="Figs 13" captionStart-1="Figs 48" captionStart-2="Figs 910" captionStart-3="Figs 1113" captionStartId-0="2.[317,359,1774,1795]" captionStartId-1="3.[317,360,1721,1742]" captionStartId-2="4.[317,360,1777,1798]" captionStartId-3="5.[317,360,1730,1751]" captionTargetBox-0="[392,1197,425,1742]" captionTargetBox-1="[322,1267,454,1664]" captionTargetBox-2="[322,1269,1185,1748]" captionTargetBox-3="[384,1204,424,1702]" captionTargetId-0="figure-18@2.[389,1197,425,1746]" captionTargetId-1="figure-13@3.[317,1269,450,1667]" captionTargetId-2="figure-353@4.[317,1269,1174,1748]" captionTargetId-3="figure-13@5.[380,1207,424,1702]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionTargetPageId-3="5" captionText-0="Figs 13. Madiza africana sp. n., paratype male, abdomen. 1 = sternites 1 to 3, ventral view, 2 = sternites 4 and 5,ventral view, 3 =mid anterior part oftergite 5. Scale:0.2 mm forFigs 12, 0.1 mm forFig.3" captionText-1="Figs 48. Madiza africana sp. n., paratype male: 4 = genital complex with epandrium and cercus, lateral view; 5 = epandrium, surstyli and subepandrial sclerite, ventral view; 6 = phallus, basiphallus in ventral view, knob of distiphallus in lateral view; 7 = syntergosternite 78 with spiracular openings," captionText-2="Figs 910. Madiza africana sp. n., paratype male.910 = hypandrial complex: 9 = ventral view, 10 = lateral view. Scale 0.1 mm" captionText-3="Figs 1113. Madiza africana sp. n., paratype female:11 = postabdomen with sternite 5 and margin of tergite 5, ventral view, segment 8 twisted, hypoproct, epiproct and cerci in lateral view; 12 = postabdominal sclerites in higher magnification, dorsal view, 13 = terminalia, ventral view, cf. Fig. 12. Scales: 0.4 mm for Figs 11 &amp; 13, 0.2 mm for Fig. 12" figureDoi-0="http://doi.org/10.5281/zenodo.12584738" figureDoi-1="http://doi.org/10.5281/zenodo.12584740" figureDoi-2="http://doi.org/10.5281/zenodo.12584742" figureDoi-3="http://doi.org/10.5281/zenodo.12584744" httpUri-0="https://zenodo.org/record/12584738/files/figure.png" httpUri-1="https://zenodo.org/record/12584740/files/figure.png" httpUri-2="https://zenodo.org/record/12584742/files/figure.png" httpUri-3="https://zenodo.org/record/12584744/files/figure.png" pageId="1" pageNumber="228">Figs 113</figureCitation>
)
</heading>
</paragraph>
<paragraph id="8BF936F51215E415FEEBFD854F99BC6F" blockId="1.[317,1270,542,592]" box="[382,1270,542,563]" pageId="1" pageNumber="228">
<materialsCitation id="3B2E3CA81215E415FEEBFD854F2DBC6E" box="[382,1090,542,563]" collectingDate="1988-02-06" collectionCode="HNHM" country="Tanzania" location="Lake Natron" pageId="1" pageNumber="228" specimenCount="1" typeStatus="holotype">
<typeStatus id="54FD88571215E415FEEBFD854AB8BC6F" box="[382,471,542,563]" pageId="1" pageNumber="228" type="holotype">Holotype</typeStatus>
male (
<collectionCode id="ED57AE301215E415FD83FD854909BC6F" box="[534,614,542,563]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="1" pageNumber="228">HNHM</collectionCode>
):
<collectingCountry id="F35176651215E415FDE1FD8549A1BC6F" box="[628,718,542,563]" name="Tanzania" pageId="1" pageNumber="228">Tanzánia</collectingCountry>
,
<location id="8E99602E1215E415FD42FD85483EBC6F" LSID="urn:lsid:plazi:treatment:03EF87E31215E412FD1AFE2C4924BAF5:8E99602E1215E415FD42FD85483EBC6F" box="[727,849,542,563]" country="Tanzania" name="Lake Natron" pageId="1" pageNumber="228">Lake Natron</location>
,
<date id="FFF810351215E415FCCCFD8548ADBC6F" box="[857,962,542,563]" pageId="1" pageNumber="228" value="1988-02-06">
<collectingDate id="EFBCE9DD1215E415FCCCFD8548ADBC6F" box="[857,962,542,563]" pageId="1" pageNumber="228" value="1988-02-06">1988. II. 6.</collectingDate>
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, leg.
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.
<typeStatus id="54FD88571215E415FBD8FD854FC1BC6F" box="[1101,1198,542,563]" pageId="1" pageNumber="228" type="paratype">Paratypes</typeStatus>
:
<specimenCount id="9D40FD7C1215E415FB23FD854F99BC6F" box="[1206,1270,542,563]" count="1" pageId="1" pageNumber="228" type="male">1 male</specimenCount>
</paragraph>
<paragraph id="8BF936F51215E415FEA8FDA048DDBC0C" blockId="1.[317,1270,542,592]" box="[317,946,571,592]" pageId="1" pageNumber="228">
<materialsCitation id="3B2E3CA81215E415FEA8FDA048C1BC0C" box="[317,942,571,592]" collectionCode="HNHM, NHM" pageId="1" pageNumber="228" specimenCount="5" specimenCount-female="5">
<specimenCount id="9D40FD7C1215E415FEA8FDA04AFDBC0C" box="[317,402,571,592]" count="4" pageId="1" pageNumber="228" type="female">4 female</specimenCount>
(
<collectionCode id="ED57AE301215E415FE35FDA04A82BC0C" box="[416,493,571,592]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34991" lsid="urn:lsid:biocol.org:col:34991" name="Hungarian Natural History Museum (Termeszettudomanyi Muzeum)" pageId="1" pageNumber="228">HNHM</collectionCode>
,
<specimenCount id="9D40FD7C1215E415FE6DFDA04922BC0C" box="[504,589,571,592]" count="1" pageId="1" pageNumber="228" type="female">1 female</specimenCount>
in
<collectionCode id="ED57AE301215E415FDFAFDA049C3BC0C" box="[623,684,571,592]" country="United Kingdom" lsid="urn:lsid:biocol.org:col:14908" name="University of Nottingham" pageId="1" pageNumber="228" type="Herbarium">NHM</collectionCode>
): with the same label data
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.
</paragraph>
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Measurements in mm: body length 2.20 (
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), 2.48 (
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male), 2.913.52 (
<typeStatus id="54FD88571215E415FB34FDEF4F9ABCD5" box="[1185,1269,628,649]" pageId="1" pageNumber="228" type="paratype">paratype</typeStatus>
females), wing length 2.20 (
<typeStatus id="54FD88571215E415FDC1FD0A49C2BCFA" box="[596,685,657,678]" pageId="1" pageNumber="228" type="holotype">holotype</typeStatus>
), 2.59 (
<typeStatus id="54FD88571215E415FD6DFD0A4823BCFA" box="[760,844,657,678]" pageId="1" pageNumber="228" type="paratype">paratype</typeStatus>
male), 3.033.52 (
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females), wing width 0.95 (
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), 1.03 (
<typeStatus id="54FD88571215E415FDC3FD3549C5BC9F" box="[598,682,686,707]" pageId="1" pageNumber="228" type="paratype">paratype</typeStatus>
male), 1.291.37 (
<typeStatus id="54FD88571215E415FCF1FD3548D7BC9F" box="[868,952,686,707]" pageId="1" pageNumber="228" type="paratype">paratype</typeStatus>
females).
</paragraph>
<paragraph id="8BF936F51215E415FEEBFD504F9CBCBC" blockId="1.[317,1270,628,1744]" box="[382,1267,715,736]" pageId="1" pageNumber="228">
A comparatively large-bodied fly of
<taxonomicName id="4C464D761215E415FD73FD504821BCBC" box="[742,846,715,736]" class="Insecta" family="Milichiidae" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="228" phylum="Arthropoda" rank="subFamily" subFamily="Madizinae">Madizinae</taxonomicName>
. Body and legs dark grey microtomentose.
</paragraph>
<paragraph id="8BF936F51215E415FEEBFD734F2DBD45" blockId="1.[317,1270,628,1744]" pageId="1" pageNumber="228">
Head. Cheek, prefrons and pregena greyish yellow. Palpus mainly dirty yellow, with apical and ventral margins grey, on outer surface this colour occupies
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.
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<paragraph id="8BF936F51215E415FEEBFCBA4F11BA0A" blockId="1.[317,1270,628,1744]" pageId="1" pageNumber="228">
Two pairs of mesoclinate frontal setae anteriorly, 2 lateroreclinate anterior orbital setae and a slightly mesoclinate very long posterior orbital seta (
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long). A very large (
<quantity id="4CBE9B101215E415FBF1FCA54FD4BD0F" box="[1124,1211,830,851]" metricMagnitude="-4" metricUnit="m" metricValue="2.0" pageId="1" pageNumber="228" unit="mm" value="0.2">0.20 mm</quantity>
long) parallel postocellar pair present. Ocellar, outer and inner vertical pairs rather long. Some small additional setulae lateral to frontal setae down to cheeks. Frons without interfrontal stripes, i.e. without an M shaped area. 4 pairs of medium-long interfrontals, and an additional similarly long seta present between anterior interfrontal and frontal seta. Lunule not extended ventrally as a flat plate between antennae, with 1 pair of distinct setae. Prefrons concave, below antennae not much more deepened; no carina. Clypeus very small, but Π- shaped rather than U-shaped anteriorly, slightly protruding before the protruded prefrons. Vibrissa inserted at level of lower eye margin, 2 large upcurved peristomals, other 2 large ventrally directed peristomals and some shorter peristomals. Gena below eye
<quantity id="4CBE9B101215E415FB35FBBF4F9ABA65" box="[1184,1269,1060,1081]" metricMagnitude="-5" metricUnit="m" metricValue="8.0" pageId="1" pageNumber="228" unit="mm" value="0.08">0.08 mm</quantity>
broad. Genal setae on postgenal border (or on postgena) only. Postocular setae short.
</paragraph>
<paragraph id="8BF936F51215E415FEEBFBC54AF3BA95" blockId="1.[317,1270,628,1744]" pageId="1" pageNumber="228">
Antenna short, pedicel with a long dorsal apical seta. First flagellomere globular, arista c.
<quantity id="4CBE9B101215E415FB42FBC54A0DBACC" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="1" pageNumber="228" unit="mm" value="0.4">0.4 mm</quantity>
long, basal aristomeres thickened. Aristal cilia somewhat longer than
<quantity id="4CBE9B101215E415FB81FBE04F02BACC" box="[1044,1133,1147,1168]" metricMagnitude="-5" metricUnit="m" metricValue="1.0" pageId="1" pageNumber="228" unit="mm" value="0.01">0.01 mm</quantity>
. Palpus short, broad with several medium-long but thick setae. Proboscis without labella
<quantity id="4CBE9B101215E415FB80FB034F01BAF1" box="[1045,1134,1176,1197]" metricMagnitude="-4" metricUnit="m" metricValue="3.3" pageId="1" pageNumber="228" unit="mm" value="0.33">0.33 mm</quantity>
, labella
<quantity id="4CBE9B101215E415FB55FB034A0DBA95" metricMagnitude="-4" metricUnit="m" metricValue="2.75" pageId="1" pageNumber="228" unit="mm" value="0.275">0.275 mm</quantity>
long.
</paragraph>
<paragraph id="8BF936F51215E415FEEBFB4A49B6BB7C" blockId="1.[317,1270,628,1744]" pageId="1" pageNumber="228">
Thorax. Basisternum broadly trapezoid, more similar to that of
<taxonomicName id="4C464D761215E415FC4CFB4A4F4EBABA" authorityName="FALLEN" authorityYear="1810" box="[985,1057,1233,1254]" class="Insecta" family="Milichiidae" genus="Madiza" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="228" phylum="Arthropoda" rank="genus">
<emphasis id="B932EAE71215E415FC4CFB4A4F4EBABA" box="[985,1057,1233,1254]" italics="true" pageId="1" pageNumber="228">Madiza</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFD74B041215E415FBB8FB4A4FC5BABA" author="BRAKE, I." box="[1069,1194,1233,1254]" pageId="1" pageNumber="228" pagination="1 - 120" refId="ref2227" refString="BRAKE, I. (2000) Phylogenetic systematics of the Milichiidae (Diptera, Schizophora). Entomologica Scandinavica (Insect Systematics &amp; Evolution) Supplement 57: 1 - 120." type="journal article" year="2000">
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2000
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: Pl. 8B) than to basisternum of
<taxonomicName id="4C464D761215E415FD82FB7549CFBB5F" box="[535,672,1262,1283]" class="Insecta" family="Milichiidae" genus="Desmometopa" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="228" phylum="Arthropoda" rank="genus">
<emphasis id="B932EAE71215E415FD82FB7549CFBB5F" box="[535,672,1262,1283]" italics="true" pageId="1" pageNumber="228">Desmometopa</emphasis>
</taxonomicName>
(
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2000
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: Pl. 8A). A pair of small lateral secondary sclerites (
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broad) also present.
</paragraph>
<paragraph id="8BF936F51215E415FEEBFAB348E4B85A" blockId="1.[317,1270,628,1744]" pageId="1" pageNumber="228">
Thoracic chaetotaxy: 1 large postpronotal, 2 notopleural, 1 presutural, 1 short prealar, 1 supra-alar (or, 2 supra-alar pairs), 1 extremely long postalar (
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), 1 less long medial postalar, 0+3 dorsocentral pairs, anterior pair shorter and aligned with posterior notopleural, 1 prescutellar acrostichal pair. Anepisternum and anepimeron bare. 1 minute supracoxal, 2 long katepisternals plus some short katepisternals anteriorly; no other seta on pleura. Anepisternum anteriorly and ventrally with an obliquely quadratic shiny black spot:
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long and
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broad (
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), on a female
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0.33 ×
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. Both ampullae below wing base comparatively large, dorsal one longish, oblique, proclinate, ventral (“greater”) one bulbous.
</paragraph>
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Wing. Wing membrane light brown, veins slightly darker but still light brown. Fringe of calypter dark grey.
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R
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emargination narrow. Veins
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R
<subScript id="17C234B01215E415FCBAF9AC4859B81F" attach="left" box="[815,822,1591,1603]" fontSize="5" pageId="1" pageNumber="228">2</subScript>
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+3,
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R
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+5 and M
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parallel, stronger costal fringe of 45 thornlets occupying only 1/5 of third costal section. Inter-crossvein section of M
<subScript id="17C234B01215E415FB1CF9CF4FFFB83C" attach="left" box="[1161,1168,1620,1632]" fontSize="5" pageId="1" pageNumber="228">1</subScript>
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, dm-cu
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, terminal section of CuA
<subScript id="17C234B01215E415FD40F9EA49B3B821" attach="left" box="[725,732,1649,1661]" fontSize="5" pageId="1" pageNumber="228">1</subScript>
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. Costagial setae: dorsal
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, ventral
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(
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) up to
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and
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(
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female). A large costal seta over vein h
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(
<typeStatus id="54FD88571215E415FEFBF9054AA7B8EF" box="[366,456,1694,1715]" pageId="1" pageNumber="228" type="holotype">holotype</typeStatus>
). Anal lobe liguliform,
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broad at base. No anal vein distal to anal cell: a rather thick fold running close to wing margin discernible. Haltere brownish grey
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.
</paragraph>
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<paragraph id="8BF936F51216E416FEA8F9754F9BB97D" blockId="2.[317,1270,1774,1825]" pageId="2" pageNumber="229">
<emphasis id="B932EAE71216E416FEA8F9754AE0B95F" bold="true" box="[317,399,1774,1795]" pageId="2" pageNumber="229">Figs 13</emphasis>
.
<taxonomicName id="4C464D761216E416FE0FF974495AB958" authorityName="Brake &amp; Papp &amp; Bd" authorityYear="2009" box="[410,565,1775,1796]" class="Insecta" family="Milichiidae" genus="Madiza" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="229" phylum="Arthropoda" rank="species" species="africana" status="sp. nov.">
<emphasis id="B932EAE71216E416FE0FF974495AB958" box="[410,565,1775,1796]" italics="true" pageId="2" pageNumber="229">Madiza africana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A201579C1216E416FDAFF9744902B958" box="[570,621,1775,1796]" pageId="2" pageNumber="229" rank="species">sp. n.</taxonomicNameLabel>
, paratype male, abdomen. 1 = sternites 1 to 3, ventral view, 2 = sternites 4 and 5, ventral view, 3 = mid anterior part of tergite 5. Scale:0.2 mm for Figs 12, 0.1 mm for Fig. 3
</paragraph>
</caption>
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<paragraph id="8BF936F51217E417FEA8F9224F99B95B" blockId="3.[317,1270,1721,1828]" pageId="3" pageNumber="230">
<emphasis id="B932EAE71217E417FEA8F9224AFFB892" bold="true" box="[317,400,1721,1742]" pageId="3" pageNumber="230">Figs 48</emphasis>
.
<taxonomicName id="4C464D761217E417FE0EF9224956B892" authorityName="Brake &amp; Papp &amp; Bd" authorityYear="2009" box="[411,569,1721,1742]" class="Insecta" family="Milichiidae" genus="Madiza" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="230" phylum="Arthropoda" rank="species" species="africana" status="sp. nov.">
<emphasis id="B932EAE71217E417FE0EF9224956B892" box="[411,569,1721,1742]" italics="true" pageId="3" pageNumber="230">Madiza africana</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A201579C1217E417FDABF922491EB892" box="[574,625,1721,1742]" pageId="3" pageNumber="230" rank="species">sp. n.</taxonomicNameLabel>
, paratype male: 4 = genital complex with epandrium and cercus, lateral view; 5 = epandrium, surstyli and subepandrial sclerite, ventral view; 6 = phallus, basiphallus in ventral view, knob of distiphallus in lateral view; 7 = syntergosternite 78 with spiracular openings,
</paragraph>
</caption>
<paragraph id="8BF936F51217E417FE11F8944FC0B978" blockId="3.[317,1270,1721,1828]" box="[388,1199,1807,1828]" pageId="3" pageNumber="230">
lateral view; 8 = ejaculatory apodeme, broadest (ventral) view. Scale:
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for all
</paragraph>
<paragraph id="8BF936F51210E410FEEBFE2F4978BC5F" blockId="4.[317,1271,436,1120]" pageId="4" pageNumber="231">Legs. Mid tibia with a strong ventroapical seta, no other characteristic setae on legs. Hind tibia without tibial organ. Male hind tibia not enlarged. First tarsomere of hind leg without a posterior comb of ventral setae.</paragraph>
<paragraph id="8BF936F51210E410FEEBFD904F43BCB5" blockId="4.[317,1271,436,1120]" pageId="4" pageNumber="231">
Abdomen. Abdominal tergites of male with dark grey microtomentum. Tergites 1 and 2 with narrow membranous area between them laterally, wholly fused on lateral parts of dorsal surface and with a
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broad membranous area medially. Male tergites 4 and 5 with socketed warts anteriorly (
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). Length of normally setose caudal part of tergite 5
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, section with warts medially
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, laterally
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mm
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long. Tergite 4 with similarly broad warts zone and with
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long normal surface. Male sternite
<quantity id="4CBE9B101210E410FD2EFD00498FBCEC" box="[699,736,667,688]" metricMagnitude="-2" metricUnit="m" metricValue="2.54" pageId="4" pageNumber="231" unit="in" value="1.0">1 in</quantity>
2 parts, microtrichose only (
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), sternite 2 quadrate with medium-long setae laterally and with 2 pairs of longer setae caudally. Sternite 3 longer than broad. Sternite 4 and 5 much longer than broad with scattered setae (
<figureCitation id="137D2A701210E410FC76FD4F4F70BCB5" box="[995,1055,724,745]" captionStart="Figs 13" captionStartId="2.[317,359,1774,1795]" captionTargetBox="[392,1197,425,1742]" captionTargetId="figure-18@2.[389,1197,425,1746]" captionTargetPageId="2" captionText="Figs 13. Madiza africana sp. n., paratype male, abdomen. 1 = sternites 1 to 3, ventral view, 2 = sternites 4 and 5,ventral view, 3 =mid anterior part oftergite 5. Scale:0.2 mm forFigs 12, 0.1 mm forFig.3" figureDoi="http://doi.org/10.5281/zenodo.12584738" httpUri="https://zenodo.org/record/12584738/files/figure.png" pageId="4" pageNumber="231">Fig. 2</figureCitation>
).
</paragraph>
<paragraph id="8BF936F51210E410FEEBFD6A4A18BA55" blockId="4.[317,1271,436,1120]" pageId="4" pageNumber="231">
Postabdomen and genitalia. Male syn(tergo)sternite 78 not completely symmetrical (
<figureCitation id="137D2A701210E410FB3AFD6A4F85BD5A" box="[1199,1258,753,774]" captionStart="Figs 48" captionStartId="3.[317,360,1721,1742]" captionTargetBox="[322,1267,454,1664]" captionTargetId="figure-13@3.[317,1269,450,1667]" captionTargetPageId="3" captionText="Figs 48. Madiza africana sp. n., paratype male: 4 = genital complex with epandrium and cercus, lateral view; 5 = epandrium, surstyli and subepandrial sclerite, ventral view; 6 = phallus, basiphallus in ventral view, knob of distiphallus in lateral view; 7 = syntergosternite 78 with spiracular openings," figureDoi="http://doi.org/10.5281/zenodo.12584740" httpUri="https://zenodo.org/record/12584740/files/figure.png" pageId="4" pageNumber="231">Fig. 7</figureCitation>
). Epandrium (
<figureCitation id="137D2A701210E410FE2DFC954A9BBD7F" box="[440,500,782,803]" captionStart="Figs 48" captionStartId="3.[317,360,1721,1742]" captionTargetBox="[322,1267,454,1664]" captionTargetId="figure-13@3.[317,1269,450,1667]" captionTargetPageId="3" captionText="Figs 48. Madiza africana sp. n., paratype male: 4 = genital complex with epandrium and cercus, lateral view; 5 = epandrium, surstyli and subepandrial sclerite, ventral view; 6 = phallus, basiphallus in ventral view, knob of distiphallus in lateral view; 7 = syntergosternite 78 with spiracular openings," figureDoi="http://doi.org/10.5281/zenodo.12584740" httpUri="https://zenodo.org/record/12584740/files/figure.png" pageId="4" pageNumber="231">Fig. 4</figureCitation>
) saddle-shaped dorsally, antero-laterally with a pair of large, well-sclerotised lobes (
<figureCitation id="137D2A701210E410FEEEFCB04AA0BD1C" box="[379,463,811,832]" captionStart="Figs 48" captionStartId="3.[317,360,1721,1742]" captionTargetBox="[322,1267,454,1664]" captionTargetId="figure-13@3.[317,1269,450,1667]" captionTargetPageId="3" captionText="Figs 48. Madiza africana sp. n., paratype male: 4 = genital complex with epandrium and cercus, lateral view; 5 = epandrium, surstyli and subepandrial sclerite, ventral view; 6 = phallus, basiphallus in ventral view, knob of distiphallus in lateral view; 7 = syntergosternite 78 with spiracular openings," figureDoi="http://doi.org/10.5281/zenodo.12584740" httpUri="https://zenodo.org/record/12584740/files/figure.png" pageId="4" pageNumber="231">Figs 45</figureCitation>
) and with robust but not particularly long setae, rather far from medio-caudal margin. Surstylus very large, completely fused to epandrium, with fine setae only (
<figureCitation id="137D2A701210E410FB80FCD34F01BD01" box="[1045,1134,840,861]" captionStart="Figs 48" captionStartId="3.[317,360,1721,1742]" captionTargetBox="[322,1267,454,1664]" captionTargetId="figure-13@3.[317,1269,450,1667]" captionTargetPageId="3" captionText="Figs 48. Madiza africana sp. n., paratype male: 4 = genital complex with epandrium and cercus, lateral view; 5 = epandrium, surstyli and subepandrial sclerite, ventral view; 6 = phallus, basiphallus in ventral view, knob of distiphallus in lateral view; 7 = syntergosternite 78 with spiracular openings," figureDoi="http://doi.org/10.5281/zenodo.12584740" httpUri="https://zenodo.org/record/12584740/files/figure.png" pageId="4" pageNumber="231">Figs 45</figureCitation>
), medial edge with fine indentations (
<figureCitation id="137D2A701210E410FDB3FCFF490ABD25" box="[550,613,868,889]" captionStart="Figs 48" captionStartId="3.[317,360,1721,1742]" captionTargetBox="[322,1267,454,1664]" captionTargetId="figure-13@3.[317,1269,450,1667]" captionTargetPageId="3" captionText="Figs 48. Madiza africana sp. n., paratype male: 4 = genital complex with epandrium and cercus, lateral view; 5 = epandrium, surstyli and subepandrial sclerite, ventral view; 6 = phallus, basiphallus in ventral view, knob of distiphallus in lateral view; 7 = syntergosternite 78 with spiracular openings," figureDoi="http://doi.org/10.5281/zenodo.12584740" httpUri="https://zenodo.org/record/12584740/files/figure.png" pageId="4" pageNumber="231">Fig. 5</figureCitation>
). Cercus large but weakly sclerotised. Subepandrial sclerite short sagittally (
<figureCitation id="137D2A701210E410FE34FC1A4AB4BDCA" box="[417,475,897,918]" captionStart="Figs 48" captionStartId="3.[317,360,1721,1742]" captionTargetBox="[322,1267,454,1664]" captionTargetId="figure-13@3.[317,1269,450,1667]" captionTargetPageId="3" captionText="Figs 48. Madiza africana sp. n., paratype male: 4 = genital complex with epandrium and cercus, lateral view; 5 = epandrium, surstyli and subepandrial sclerite, ventral view; 6 = phallus, basiphallus in ventral view, knob of distiphallus in lateral view; 7 = syntergosternite 78 with spiracular openings," figureDoi="http://doi.org/10.5281/zenodo.12584740" httpUri="https://zenodo.org/record/12584740/files/figure.png" pageId="4" pageNumber="231">Fig. 5</figureCitation>
). Hypandrial complex (
<figureCitation id="137D2A701210E410FD29FC1A4872BDCA" box="[700,797,897,918]" captionStart="Figs 910" captionStartId="4.[317,360,1777,1798]" captionTargetBox="[322,1269,1185,1748]" captionTargetId="figure-353@4.[317,1269,1174,1748]" captionTargetPageId="4" captionText="Figs 910. Madiza africana sp. n., paratype male.910 = hypandrial complex: 9 = ventral view, 10 = lateral view. Scale 0.1 mm" figureDoi="http://doi.org/10.5281/zenodo.12584742" httpUri="https://zenodo.org/record/12584742/files/figure.png" pageId="4" pageNumber="231">Figs 910</figureCitation>
) not much longer than broad, hypandrial arms robust, pregonites rather flat (in the horizontal plane). Basiphallus better sclerotised (
<figureCitation id="137D2A701210E410FBE6FC054FDFBDEF" box="[1139,1200,926,947]" captionStart="Figs 48" captionStartId="3.[317,360,1721,1742]" captionTargetBox="[322,1267,454,1664]" captionTargetId="figure-13@3.[317,1269,450,1667]" captionTargetPageId="3" captionText="Figs 48. Madiza africana sp. n., paratype male: 4 = genital complex with epandrium and cercus, lateral view; 5 = epandrium, surstyli and subepandrial sclerite, ventral view; 6 = phallus, basiphallus in ventral view, knob of distiphallus in lateral view; 7 = syntergosternite 78 with spiracular openings," figureDoi="http://doi.org/10.5281/zenodo.12584740" httpUri="https://zenodo.org/record/12584740/files/figure.png" pageId="4" pageNumber="231">Fig. 6</figureCitation>
), distiphallus bulbous membranous. Ejaculatory apodeme (
<figureCitation id="137D2A701210E410FCA3FC204801BD8C" box="[822,878,955,976]" captionStart="Figs 48" captionStartId="3.[317,360,1721,1742]" captionTargetBox="[322,1267,454,1664]" captionTargetId="figure-13@3.[317,1269,450,1667]" captionTargetPageId="3" captionText="Figs 48. Madiza africana sp. n., paratype male: 4 = genital complex with epandrium and cercus, lateral view; 5 = epandrium, surstyli and subepandrial sclerite, ventral view; 6 = phallus, basiphallus in ventral view, knob of distiphallus in lateral view; 7 = syntergosternite 78 with spiracular openings," figureDoi="http://doi.org/10.5281/zenodo.12584740" httpUri="https://zenodo.org/record/12584740/files/figure.png" pageId="4" pageNumber="231">Fig. 8</figureCitation>
) positioned in the main axis of abdomen, long slightly asymmetrical, in ventral view broadened sub-medially; that extended part with fine holes.
</paragraph>
<paragraph id="8BF936F51210E412FEEBFB8A4924BAF5" blockId="4.[317,1271,436,1120]" lastBlockId="6.[317,1269,1115,1193]" lastPageId="6" lastPageNumber="233" pageId="4" pageNumber="231">
Female sternite 5 much longer than broad, with a pair of particularly strong caudal appendages. Female tergites 4 and 5 without socketed warts anteriorly. Female ovipositor rather long (
<figureCitation id="137D2A701210E410FB5AFBB54A39BA3C" captionStart="Figs 1113" captionStartId="5.[317,360,1730,1751]" captionTargetBox="[384,1204,424,1702]" captionTargetId="figure-13@5.[380,1207,424,1702]" captionTargetPageId="5" captionText="Figs 1113. Madiza africana sp. n., paratype female:11 = postabdomen with sternite 5 and margin of tergite 5, ventral view, segment 8 twisted, hypoproct, epiproct and cerci in lateral view; 12 = postabdominal sclerites in higher magnification, dorsal view, 13 = terminalia, ventral view, cf. Fig. 12. Scales: 0.4 mm for Figs 11 &amp; 13, 0.2 mm for Fig. 12" figureDoi="http://doi.org/10.5281/zenodo.12584744" httpUri="https://zenodo.org/record/12584744/files/figure.png" pageId="4" pageNumber="231">Fig. 11</figureCitation>
), sternites 5 and 6 with long introsuscepted caudal apodemes (serving insertion points for muscles when postabdomen inverted). Tergite 8 not divided, sternite
<quantity id="4CBE9B101212E412FCEDFBC048F4BA2C" box="[888,923,1115,1136]" metricMagnitude="-1" metricUnit="m" metricValue="2.032" pageId="6" pageNumber="233" unit="in" value="8.0">8 in</quantity>
3 parts (
<figureCitation id="137D2A701212E412FC7FFBC04F36BA2C" box="[1002,1113,1115,1136]" captionStart="Figs 1113" captionStartId="5.[317,360,1730,1751]" captionTargetBox="[384,1204,424,1702]" captionTargetId="figure-13@5.[380,1207,424,1702]" captionTargetPageId="5" captionText="Figs 1113. Madiza africana sp. n., paratype female:11 = postabdomen with sternite 5 and margin of tergite 5, ventral view, segment 8 twisted, hypoproct, epiproct and cerci in lateral view; 12 = postabdominal sclerites in higher magnification, dorsal view, 13 = terminalia, ventral view, cf. Fig. 12. Scales: 0.4 mm for Figs 11 &amp; 13, 0.2 mm for Fig. 12" figureDoi="http://doi.org/10.5281/zenodo.12584744" httpUri="https://zenodo.org/record/12584744/files/figure.png" pageId="6" pageNumber="233">Figs 1113</figureCitation>
). Cerci not elongated, with medium-long setae. Spermathecal ducts tightly coiled together, similar to
<taxonomicName id="4C464D761212E412FBFCFBE34F9ABAD1" box="[1129,1269,1144,1165]" class="Insecta" family="Milichiidae" genus="Madiza" kingdom="Animalia" order="Diptera" pageId="6" pageNumber="233" phylum="Arthropoda" rank="species" species="glabra">
<emphasis id="B932EAE71212E412FBFCFBE34F9ABAD1" box="[1129,1269,1144,1165]" italics="true" pageId="6" pageNumber="233">Madiza glabra</emphasis>
</taxonomicName>
(see
<bibRefCitation id="EFD74B041212E412FEFEFB0F4A85BAF5" author="BRAKE, I." box="[363,490,1172,1193]" pageId="6" pageNumber="233" pagination="1 - 120" refId="ref2227" refString="BRAKE, I. (2000) Phylogenetic systematics of the Milichiidae (Diptera, Schizophora). Entomologica Scandinavica (Insect Systematics &amp; Evolution) Supplement 57: 1 - 120." type="journal article" year="2000">
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2000
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: Pl. 24E).
</paragraph>
</subSubSection>
</treatment>
</document>