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<document id="C1762CB2C83E2B41CA4628A8CA476CCE" ID-DOI="10.1111/j.1096-3642.2009.00557.x" ID-ISSN="0024-4082" ID-Zenodo-Dep="5438222" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630440985089" checkinUser="valdenar" docAuthor="Bonato, Lucio, Dányi, László &amp; Minelli, Alessandro" docDate="2010" docId="0398B8502F0F524E1BAC8DFBFDA2FA13" docLanguage="en" docName="j.1096-3642.2009.00557.x.pdf" docOrigin="Zoological Journal of the Linnean Society 158 (3)" docSource="http://dx.doi.org/10.1111/j.1096-3642.2009.00557.x" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Dicellophilus anomalus" docType="treatment" docVersion="3" lastPageNumber="520" masterDocId="FFA1C0282F00525D1B138B0DFF91FF81" masterDocTitle="Morphology and phylogeny of Dicellophilus, a centipede genus with a highly disjunct distribution (Chilopoda: Mecistocephalidae)" masterLastPageNumber="532" masterPageNumber="501" pageNumber="516" updateTime="1699271895951" updateUser="plazi">
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<mods:title id="0083349EE80415F459788AF99DDCD4F9">Morphology and phylogeny of Dicellophilus, a centipede genus with a highly disjunct distribution (Chilopoda: Mecistocephalidae)</mods:title>
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<mods:namePart id="DB9367439BB61F07E45E3F859EEACF4C">Bonato, Lucio</mods:namePart>
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<mods:namePart id="A4A74BA2D6F07F263B500FA08FE7315E">Dányi, László</mods:namePart>
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<mods:namePart id="BC50A6B267DE3F63E02D55C83F531224">Minelli, Alessandro</mods:namePart>
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<treatment id="0398B8502F0F524E1BAC8DFBFDA2FA13" LSID="urn:lsid:plazi:treatment:0398B8502F0F524E1BAC8DFBFDA2FA13" httpUri="http://treatment.plazi.org/id/0398B8502F0F524E1BAC8DFBFDA2FA13" lastPageId="19" lastPageNumber="520" pageId="15" pageNumber="516">
<subSubSection id="C32B5ACD2F0F52521BAC8DFBFD61F88A" box="[191,752,1781,1803]" pageId="15" pageNumber="516" type="nomenclature">
<paragraph id="8B8E09462F0F52521BAC8DFBFD61F88A" blockId="15.[191,752,1781,1803]" box="[191,752,1781,1803]" pageId="15" pageNumber="516">
<heading id="D0C6BE2A2F0F52521BAC8DFBFD61F88A" box="[191,752,1781,1803]" centered="true" fontSize="8" level="2" pageId="15" pageNumber="516" reason="8">
<taxonomicName id="4C3172C52F0F52521BAC8DFBFD61F88A" ID-CoL="35J7R" authority="(CHAMBERLIN, 1904)" baseAuthorityName="Chamberlin" baseAuthorityYear="1904" box="[191,752,1781,1803]" class="Chilopoda" family="Mecistocephalidae" genus="Dicellophilus" kingdom="Animalia" order="Geophilomorpha" pageId="15" pageNumber="516" phylum="Arthropoda" rank="species" species="anomalus">
<emphasis id="B945D5542F0F52521BAC8DFBFE66F888" box="[191,503,1782,1803]" italics="true" pageId="15" pageNumber="516">DICELLOPHILUS ANOMALUS</emphasis>
(
<bibRefCitation id="EFA074B72F0F5252191B8DF8FD79F88A" author="Chamberlin RV" box="[520,744,1781,1803]" pageId="15" pageNumber="516" pagination="651 - 657" refId="ref21072" refString="Chamberlin RV. 1904. New chilopods. Proceedings of the Academy of Natural Sciences of Philadelphia 56: 651 - 657." type="journal article" year="1904">CHAMBERLIN, 1904</bibRefCitation>
)
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C32B5ACD2F0F52521BB78C11FAEDFE1F" pageId="15" pageNumber="516" type="reference_group">
<paragraph id="8B8E09462F0F52521BB78C11FBF4FE98" blockId="15.[164,779,1820,1903]" lastBlockId="15.[826,1441,197,281]" pageId="15" pageNumber="516">
<taxonomicName id="4C3172C52F0F52521BB78C11FD29F8B0" authority="Chamberlin, 1904" authorityName="Chamberlin" authorityYear="1904" box="[164,696,1820,1842]" class="Chilopoda" family="Mecistocephalidae" genus="Mecistocephalus" kingdom="Animalia" order="Geophilomorpha" pageId="15" pageNumber="516" phylum="Arthropoda" rank="species" species="anomalus">
<emphasis id="B945D5542F0F52521BB78C11FE49F8B0" box="[164,472,1820,1841]" italics="true" pageId="15" pageNumber="516">Mecistocephalus anomalus</emphasis>
<bibRefCitation id="EFA074B72F0F52521AF78C11FD29F8B0" author="Chamberlin RV" box="[484,696,1820,1842]" pageId="15" pageNumber="516" pagination="651 - 657" refId="ref21072" refString="Chamberlin RV. 1904. New chilopods. Proceedings of the Academy of Natural Sciences of Philadelphia 56: 651 - 657." type="journal article" year="1904">Chamberlin, 1904</bibRefCitation>
</taxonomicName>
. Original description:
<bibRefCitation id="EFA074B72F0F52521A4D8C37FDF7F8D1" author="Chamberlin RV" box="[350,614,1850,1872]" pageId="15" pageNumber="516" pagination="651 - 657" refId="ref21072" refString="Chamberlin RV. 1904. New chilopods. Proceedings of the Academy of Natural Sciences of Philadelphia 56: 651 - 657." type="journal article" year="1904">Chamberlin, 1904: 655</bibRefCitation>
. Type locality: Pacific Grove, California [= Pacific Grove (
<collectingCountry id="F32649D62F0F5252199C8C54FD59F8EF" box="[655,712,1881,1902]" name="United States of America" pageId="15" pageNumber="516">USA</collectingCountry>
,
<collectingRegion id="49F5C7A42F0F525219C08C54FC12FF5D" box="[723,899,197,1903]" country="United States of America" name="California" pageId="15" pageNumber="516">California</collectingRegion>
)]. Type material:
<typeStatus id="548AB7E42F0F52521F598BCBFB3AFF5D" box="[1098,1195,198,220]" pageId="15" pageNumber="516" type="holotype">holotype</typeStatus>
male,
<quantity id="4CC9A4A32F0F52521FE48BCBFAD4FF5A" box="[1271,1349,198,219]" metricMagnitude="-2" metricUnit="m" metricValue="2.2" pageId="15" pageNumber="516" unit="mm" value="22.0">22 mm</quantity>
long, in the Museum of Comparative Zoology, Cambridge,
<collectingCountry id="F32649D62F0F525218298A0EFCFEFE99" box="[826,879,259,280]" name="United States of America" pageId="15" pageNumber="516">USA</collectingCountry>
(code number 1042).
</paragraph>
<paragraph id="8B8E09462F0F525218298A46FAEDFE1F" blockId="15.[826,1441,330,414]" pageId="15" pageNumber="516">
<emphasis id="B945D5542F0F525218298A46FB28FEDE" box="[826,1209,330,352]" italics="true" pageId="15" pageNumber="516">Main references for morphology:</emphasis>
<bibRefCitation id="EFA074B72F0F52521FD48A47FCF2FEFE" author="Chamberlin RV" pageId="15" pageNumber="516" pagination="651 - 657" refId="ref21072" refString="Chamberlin RV. 1904. New chilopods. Proceedings of the Academy of Natural Sciences of Philadelphia 56: 651 - 657." type="journal article" year="1904">Chamberlin, 1904: 655</bibRefCitation>
(original description);
<bibRefCitation id="EFA074B72F0F52521F948A64FA30FEFE" author="Chamberlin RV" box="[1159,1441,361,383]" pageId="15" pageNumber="516" pagination="651 - 672" refId="ref21095" refString="Chamberlin RV. 1912. The Chilopoda of California III. Pomona Journal of Entomology 4: 651 - 672." type="journal article" year="1912">Chamberlin, 1912: 653</bibRefCitation>
(redescription);
<bibRefCitation id="EFA074B72F0F525218E28A85FB55FE1C" author="Silvestri F" box="[1009,1220,392,414]" pageId="15" pageNumber="516" pagination="45 - 107" refId="ref23270" refString="Silvestri F. 1919. Contributions to a knowledge of the Chilopoda Geophilomorpha of India. Records of the Indian Museum 16: 45 - 107." type="journal article" year="1919">Silvestri, 1919: 83</bibRefCitation>
(redescription).
</paragraph>
</subSubSection>
<subSubSection id="C32B5ACD2F0F524E18298ADDFDA2FA13" lastPageId="19" lastPageNumber="520" pageId="15" pageNumber="516" type="diagnosis">
<paragraph id="8B8E09462F0F525218298ADDFC06F90F" blockId="15.[826,1442,463,1740]" pageId="15" pageNumber="516">
<emphasis id="B945D5542F0F525218298ADDFC20FE64" box="[826,945,464,485]" italics="true" pageId="15" pageNumber="516">Diagnosis:</emphasis>
A
<taxonomicName id="4C3172C52F0F525218C38AC2FB2DFE64" box="[976,1212,463,485]" class="Chilopoda" family="Mecistocephalidae" genus="Dicellophilus" kingdom="Animalia" order="Geophilomorpha" pageId="15" pageNumber="516" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B945D5542F0F525218C38AC2FBF4FE65" box="[976,1125,463,484]" italics="true" pageId="15" pageNumber="516">Dicellophilus</emphasis>
species
</taxonomicName>
with 41 leg-bearing segments. Antennae about four times as long as the maximum width of the head, strongly tapering (width of the terminal article less than half of the width of article I); terminal article more than 2 times as long as wide and less than 1.5 times as long as article XIII. The longest setae on the antennae are ~300-Mm long (
<figureCitation id="130A15C32F0F5252185189A8FC06FD3A" box="[834,919,677,699]" captionStart="Figure 2" captionStartId="3.[164,243,1217,1236]" captionTargetBox="[164,1443,195,1188]" captionTargetId="figure-207@3.[164,1446,195,1188]" captionTargetPageId="3" captionText="Figure 2. Dicellophilus anomalus. A, head, dorsal view. B, right part of the maxillary complex, ventral view (detached from head). C, anterior part of head, dorsal view. D, clypeus and labrum, ventral view (maxillae removed). E, left antennal articles IIIV, dorsal view. F, internal margin of left forcipule, dorsal view. G, last leg-bearing segment of an adult female, ventral view. H, left telopodite and medial projection of maxillae I, dorsal view (detached from head). I, left condylar process of forcipular coxosternum, dorsal view (head removed). Microscopic photographs and line drawings: ♀, 45-mm long, from Pacific Grove, California; the detailed data for the specimen are presented in the Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5438226" httpUri="https://zenodo.org/record/5438226/files/figure.png" pageId="15" pageNumber="516">Fig. 2E</figureCitation>
). Sensilla on the antennal tip usually with a crown-like projection at about mid-length. Head (
<figureCitation id="130A15C32F0F5252185189EFFC07FD78" box="[834,918,738,761]" captionStart="Figure 2" captionStartId="3.[164,243,1217,1236]" captionTargetBox="[164,1443,195,1188]" captionTargetId="figure-207@3.[164,1446,195,1188]" captionTargetPageId="3" captionText="Figure 2. Dicellophilus anomalus. A, head, dorsal view. B, right part of the maxillary complex, ventral view (detached from head). C, anterior part of head, dorsal view. D, clypeus and labrum, ventral view (maxillae removed). E, left antennal articles IIIV, dorsal view. F, internal margin of left forcipule, dorsal view. G, last leg-bearing segment of an adult female, ventral view. H, left telopodite and medial projection of maxillae I, dorsal view (detached from head). I, left condylar process of forcipular coxosternum, dorsal view (head removed). Microscopic photographs and line drawings: ♀, 45-mm long, from Pacific Grove, California; the detailed data for the specimen are presented in the Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5438226" httpUri="https://zenodo.org/record/5438226/files/figure.png" pageId="15" pageNumber="516">Fig. 2A</figureCitation>
) ~1.51.6 times as long as wide. Lateral margins of the cephalic plate almost straight, evidently converging for most of their length. Frontal line (
<figureCitation id="130A15C32F0F5252186F8833FC5EFCD5" box="[892,975,830,852]" captionStart="Figure 2" captionStartId="3.[164,243,1217,1236]" captionTargetBox="[164,1443,195,1188]" captionTargetId="figure-207@3.[164,1446,195,1188]" captionTargetPageId="3" captionText="Figure 2. Dicellophilus anomalus. A, head, dorsal view. B, right part of the maxillary complex, ventral view (detached from head). C, anterior part of head, dorsal view. D, clypeus and labrum, ventral view (maxillae removed). E, left antennal articles IIIV, dorsal view. F, internal margin of left forcipule, dorsal view. G, last leg-bearing segment of an adult female, ventral view. H, left telopodite and medial projection of maxillae I, dorsal view (detached from head). I, left condylar process of forcipular coxosternum, dorsal view (head removed). Microscopic photographs and line drawings: ♀, 45-mm long, from Pacific Grove, California; the detailed data for the specimen are presented in the Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5438226" httpUri="https://zenodo.org/record/5438226/files/figure.png" pageId="15" pageNumber="516">Fig. 2C</figureCitation>
) forming an angle pointing forwards. Clypeus (
<figureCitation id="130A15C32F0F525218B98850FB93FCF2" box="[938,1026,861,883]" captionStart="Figure 2" captionStartId="3.[164,243,1217,1236]" captionTargetBox="[164,1443,195,1188]" captionTargetId="figure-207@3.[164,1446,195,1188]" captionTargetPageId="3" captionText="Figure 2. Dicellophilus anomalus. A, head, dorsal view. B, right part of the maxillary complex, ventral view (detached from head). C, anterior part of head, dorsal view. D, clypeus and labrum, ventral view (maxillae removed). E, left antennal articles IIIV, dorsal view. F, internal margin of left forcipule, dorsal view. G, last leg-bearing segment of an adult female, ventral view. H, left telopodite and medial projection of maxillae I, dorsal view (detached from head). I, left condylar process of forcipular coxosternum, dorsal view (head removed). Microscopic photographs and line drawings: ♀, 45-mm long, from Pacific Grove, California; the detailed data for the specimen are presented in the Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5438226" httpUri="https://zenodo.org/record/5438226/files/figure.png" pageId="15" pageNumber="516">Fig. 2D</figureCitation>
): lateral margins converging backwards; a wide subtriangular posteromedial area without setae, but with a distinctly isolated posteromedian pair of setae. Posterior margin of each labral side piece usually also concave on its medial part. Non-areolate part of the buccae (
<figureCitation id="130A15C32F0F52521FDB88FBFA8CFB8D" box="[1224,1309,1014,1036]" captionStart="Figure 2" captionStartId="3.[164,243,1217,1236]" captionTargetBox="[164,1443,195,1188]" captionTargetId="figure-207@3.[164,1446,195,1188]" captionTargetPageId="3" captionText="Figure 2. Dicellophilus anomalus. A, head, dorsal view. B, right part of the maxillary complex, ventral view (detached from head). C, anterior part of head, dorsal view. D, clypeus and labrum, ventral view (maxillae removed). E, left antennal articles IIIV, dorsal view. F, internal margin of left forcipule, dorsal view. G, last leg-bearing segment of an adult female, ventral view. H, left telopodite and medial projection of maxillae I, dorsal view (detached from head). I, left condylar process of forcipular coxosternum, dorsal view (head removed). Microscopic photographs and line drawings: ♀, 45-mm long, from Pacific Grove, California; the detailed data for the specimen are presented in the Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5438226" httpUri="https://zenodo.org/record/5438226/files/figure.png" pageId="15" pageNumber="516">Fig. 2D</figureCitation>
) extending forwards distinctly beyond the labrum. Mandible with between five and seven pectinate lamellae. Coxosternum of maxillae I (
<figureCitation id="130A15C32F0F52521F388F5FFBEFFBE9" box="[1067,1150,1106,1128]" captionStart="Figure 2" captionStartId="3.[164,243,1217,1236]" captionTargetBox="[164,1443,195,1188]" captionTargetId="figure-207@3.[164,1446,195,1188]" captionTargetPageId="3" captionText="Figure 2. Dicellophilus anomalus. A, head, dorsal view. B, right part of the maxillary complex, ventral view (detached from head). C, anterior part of head, dorsal view. D, clypeus and labrum, ventral view (maxillae removed). E, left antennal articles IIIV, dorsal view. F, internal margin of left forcipule, dorsal view. G, last leg-bearing segment of an adult female, ventral view. H, left telopodite and medial projection of maxillae I, dorsal view (detached from head). I, left condylar process of forcipular coxosternum, dorsal view (head removed). Microscopic photographs and line drawings: ♀, 45-mm long, from Pacific Grove, California; the detailed data for the specimen are presented in the Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5438226" httpUri="https://zenodo.org/record/5438226/files/figure.png" pageId="15" pageNumber="516">Fig. 2B</figureCitation>
) less than 3.0 times as wide as medially long. Medial projection of maxillae I more than 2.0 times as long as wide; distal hyaline part enlarged, subtriangular; hyaline scales present on the dorsal surface (
<figureCitation id="130A15C32F0F52521F2F8FC1FB05FB63" box="[1084,1172,1228,1250]" captionStart="Figure 2" captionStartId="3.[164,243,1217,1236]" captionTargetBox="[164,1443,195,1188]" captionTargetId="figure-207@3.[164,1446,195,1188]" captionTargetPageId="3" captionText="Figure 2. Dicellophilus anomalus. A, head, dorsal view. B, right part of the maxillary complex, ventral view (detached from head). C, anterior part of head, dorsal view. D, clypeus and labrum, ventral view (maxillae removed). E, left antennal articles IIIV, dorsal view. F, internal margin of left forcipule, dorsal view. G, last leg-bearing segment of an adult female, ventral view. H, left telopodite and medial projection of maxillae I, dorsal view (detached from head). I, left condylar process of forcipular coxosternum, dorsal view (head removed). Microscopic photographs and line drawings: ♀, 45-mm long, from Pacific Grove, California; the detailed data for the specimen are presented in the Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5438226" httpUri="https://zenodo.org/record/5438226/files/figure.png" pageId="15" pageNumber="516">Fig. 2H</figureCitation>
). Distal hyaline part of the telopodite of maxillae I distinctly longer than the basal chitinous part. Distal article of the telopodite of maxillae II subconical, tapering gradually along most of its length, ~1.5 times as long as wide (
<figureCitation id="130A15C32F0F52521E2C8E4BFA05FADD" box="[1343,1428,1350,1372]" captionStart="Figure 2" captionStartId="3.[164,243,1217,1236]" captionTargetBox="[164,1443,195,1188]" captionTargetId="figure-207@3.[164,1446,195,1188]" captionTargetPageId="3" captionText="Figure 2. Dicellophilus anomalus. A, head, dorsal view. B, right part of the maxillary complex, ventral view (detached from head). C, anterior part of head, dorsal view. D, clypeus and labrum, ventral view (maxillae removed). E, left antennal articles IIIV, dorsal view. F, internal margin of left forcipule, dorsal view. G, last leg-bearing segment of an adult female, ventral view. H, left telopodite and medial projection of maxillae I, dorsal view (detached from head). I, left condylar process of forcipular coxosternum, dorsal view (head removed). Microscopic photographs and line drawings: ♀, 45-mm long, from Pacific Grove, California; the detailed data for the specimen are presented in the Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5438226" httpUri="https://zenodo.org/record/5438226/files/figure.png" pageId="15" pageNumber="516">Fig. 2B</figureCitation>
). Condylar processes of the forcipular coxosternum (
<figureCitation id="130A15C32F0F525218518E89FC1CFA1B" box="[834,909,1412,1434]" captionStart="Figure 2" captionStartId="3.[164,243,1217,1236]" captionTargetBox="[164,1443,195,1188]" captionTargetId="figure-207@3.[164,1446,195,1188]" captionTargetPageId="3" captionText="Figure 2. Dicellophilus anomalus. A, head, dorsal view. B, right part of the maxillary complex, ventral view (detached from head). C, anterior part of head, dorsal view. D, clypeus and labrum, ventral view (maxillae removed). E, left antennal articles IIIV, dorsal view. F, internal margin of left forcipule, dorsal view. G, last leg-bearing segment of an adult female, ventral view. H, left telopodite and medial projection of maxillae I, dorsal view (detached from head). I, left condylar process of forcipular coxosternum, dorsal view (head removed). Microscopic photographs and line drawings: ♀, 45-mm long, from Pacific Grove, California; the detailed data for the specimen are presented in the Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5438226" httpUri="https://zenodo.org/record/5438226/files/figure.png" pageId="15" pageNumber="516">Fig. 2I</figureCitation>
) well developed. Forcipular article I 1.21.3 times as long as wide, the tubercle relatively large, usually projecting beyond the distal margin of the article (
<figureCitation id="130A15C32F0F525218818ED2FC77FA74" box="[914,998,1503,1525]" captionStart="Figure 2" captionStartId="3.[164,243,1217,1236]" captionTargetBox="[164,1443,195,1188]" captionTargetId="figure-207@3.[164,1446,195,1188]" captionTargetPageId="3" captionText="Figure 2. Dicellophilus anomalus. A, head, dorsal view. B, right part of the maxillary complex, ventral view (detached from head). C, anterior part of head, dorsal view. D, clypeus and labrum, ventral view (maxillae removed). E, left antennal articles IIIV, dorsal view. F, internal margin of left forcipule, dorsal view. G, last leg-bearing segment of an adult female, ventral view. H, left telopodite and medial projection of maxillae I, dorsal view (detached from head). I, left condylar process of forcipular coxosternum, dorsal view (head removed). Microscopic photographs and line drawings: ♀, 45-mm long, from Pacific Grove, California; the detailed data for the specimen are presented in the Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5438226" httpUri="https://zenodo.org/record/5438226/files/figure.png" pageId="15" pageNumber="516">Fig. 2F</figureCitation>
). Forcipular tarsungulum with evident basal tubercle, the dorso-internal margin only weakly crenulated, with uniformly spaced notches. Sternum of the last leg-bearing segment (
<figureCitation id="130A15C32F0F52521FA98D36FA9EF9D0" box="[1210,1295,1595,1617]" captionStart="Figure 2" captionStartId="3.[164,243,1217,1236]" captionTargetBox="[164,1443,195,1188]" captionTargetId="figure-207@3.[164,1446,195,1188]" captionTargetPageId="3" captionText="Figure 2. Dicellophilus anomalus. A, head, dorsal view. B, right part of the maxillary complex, ventral view (detached from head). C, anterior part of head, dorsal view. D, clypeus and labrum, ventral view (maxillae removed). E, left antennal articles IIIV, dorsal view. F, internal margin of left forcipule, dorsal view. G, last leg-bearing segment of an adult female, ventral view. H, left telopodite and medial projection of maxillae I, dorsal view (detached from head). I, left condylar process of forcipular coxosternum, dorsal view (head removed). Microscopic photographs and line drawings: ♀, 45-mm long, from Pacific Grove, California; the detailed data for the specimen are presented in the Material and methods." figureDoi="http://doi.org/10.5281/zenodo.5438226" httpUri="https://zenodo.org/record/5438226/files/figure.png" pageId="15" pageNumber="516">Fig. 2G</figureCitation>
) wider than long. Posterior end of coxopleura usually distinctly pointed.
</paragraph>
<paragraph id="8B8E09462F0F525218408D9AFAFBF94A" blockId="15.[826,1442,463,1740]" pageId="15" pageNumber="516">
The main differential characters in respect to the other
<taxonomicName id="4C3172C52F0F525218938DBBFBFEF94D" box="[896,1135,1718,1740]" class="Chilopoda" family="Mecistocephalidae" genus="Dicellophilus" kingdom="Animalia" order="Geophilomorpha" pageId="15" pageNumber="516" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B945D5542F0F525218938DBBFB84F94A" box="[896,1045,1718,1739]" italics="true" pageId="15" pageNumber="516">Dicellophilus</emphasis>
species
</taxonomicName>
are given in
<tableCitation id="C6B33CFD2F0F52521E038DBBFAF4F94A" box="[1296,1381,1718,1740]" captionStart="Table 3" captionStartId="17.[144,208,525,546]" captionText="Table 3. Differential characters of the Dicellophilus species" pageId="15" pageNumber="516">Table 3</tableCitation>
.
</paragraph>
<paragraph id="8B8E09462F0F524D18298DF3FD61F936" blockId="15.[826,1441,1789,1903]" lastBlockId="16.[144,759,1666,1902]" lastPageId="16" lastPageNumber="517" pageId="15" pageNumber="516">
<emphasis id="B945D5542F0F525218298DF3FB8DF892" box="[826,1052,1790,1811]" italics="true" pageId="15" pageNumber="516">Notes on taxonomy:</emphasis>
Originally included in the genus
<taxonomicName id="4C3172C52F0F525218298C11FB3DF8B0" authority="Newport, 1843" authorityName="Newport" authorityYear="1843" box="[826,1196,1820,1841]" class="Chilopoda" family="Mecistocephalidae" genus="Mecistocephalus" kingdom="Animalia" order="Geophilomorpha" pageId="15" pageNumber="516" phylum="Arthropoda" rank="genus">
<emphasis id="B945D5542F0F525218298C11FC63F8B0" box="[826,1010,1820,1841]" italics="true" pageId="15" pageNumber="516">Mecistocephalus</emphasis>
<collectingRegion id="49F5C7A42F0F525218EC8C11FBF5F8B0" box="[1023,1124,1820,1841]" country="United Kingdom" name="Newport" pageId="15" pageNumber="516">Newport</collectingRegion>
, 1843
</taxonomicName>
(
<bibRefCitation id="EFA074B72F0F52521FD38C11FA04F8B3" author="Chamberlin RV" box="[1216,1429,1820,1842]" pageId="15" pageNumber="516" pagination="651 - 657" refId="ref21072" refString="Chamberlin RV. 1904. New chilopods. Proceedings of the Academy of Natural Sciences of Philadelphia 56: 651 - 657." type="journal article" year="1904">Chamberlin, 1904</bibRefCitation>
), the species was moved to
<taxonomicName id="4C3172C52F0F52521F6B8C37FA9CF8CE" authorityName="Cook" authorityYear="1896" box="[1144,1293,1850,1871]" class="Chilopoda" family="Mecistocephalidae" genus="Dicellophilus" kingdom="Animalia" order="Geophilomorpha" pageId="15" pageNumber="516" phylum="Arthropoda" rank="genus">
<emphasis id="B945D5542F0F52521F6B8C37FA9CF8CE" box="[1144,1293,1850,1871]" italics="true" pageId="15" pageNumber="516">Dicellophilus</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFA074B72F0F52521E528C37FC15F8EE" author="Silvestri F" pageId="15" pageNumber="516" pagination="45 - 107" refId="ref23270" refString="Silvestri F. 1919. Contributions to a knowledge of the Chilopoda Geophilomorpha of India. Records of the Indian Museum 16: 45 - 107." type="journal article" year="1919">Silvestri (1919)</bibRefCitation>
, followed by subsequent authors. After the original description, additional specimens were only recorded by
<bibRefCitation id="EFA074B72F10524D1A338DACFE68F936" author="Chamberlin RV" box="[288,505,1697,1719]" pageId="16" pageNumber="517" pagination="651 - 672" refId="ref21095" refString="Chamberlin RV. 1912. The Chilopoda of California III. Pomona Journal of Entomology 4: 651 - 672." type="journal article" year="1912">Chamberlin (1912)</bibRefCitation>
and
<bibRefCitation id="EFA074B72F10524D19248DACFD7DF936" author="Silvestri F" box="[567,748,1697,1719]" pageId="16" pageNumber="517" pagination="45 - 107" refId="ref23270" refString="Silvestri F. 1919. Contributions to a knowledge of the Chilopoda Geophilomorpha of India. Records of the Indian Museum 16: 45 - 107." type="journal article" year="1919">Silvestri (1919)</bibRefCitation>
.
</paragraph>
<caption id="DF4E59CE2F10524D1B838EEEFE32F9B0" ID-DOI="http://doi.org/10.5281/zenodo.5438242" ID-Zenodo-Dep="5438242" httpUri="https://zenodo.org/record/5438242/files/figure.png" pageId="16" pageNumber="517" startId="16.[144,223,1507,1526]" targetBox="[149,1426,200,1472]" targetPageId="16">
<paragraph id="8B8E09462F10524D1B838EEEFE32F9B0" blockId="16.[144,1421,1507,1585]" pageId="16" pageNumber="517">
<emphasis id="B945D5542F10524D1B838EEEFE94FA77" bold="true" box="[144,261,1507,1526]" pageId="16" pageNumber="517">Figure 10.</emphasis>
Geographical distribution and phylogenetic relationships of the species of
<taxonomicName id="4C3172C52F10524D1F268EEEFB2FFA77" authorityName="Cook" authorityYear="1896" box="[1077,1214,1507,1526]" class="Chilopoda" family="Mecistocephalidae" genus="Dicellophilus" kingdom="Animalia" order="Geophilomorpha" pageId="16" pageNumber="517" phylum="Arthropoda" rank="genus">
<emphasis id="B945D5542F10524D1F268EEEFB2FFA77" box="[1077,1214,1507,1526]" italics="true" pageId="16" pageNumber="517">Dicellophilus</emphasis>
</taxonomicName>
. The shaded areas indicate the established populations; stars indicate either occasional or dubious records. The dotted line indicates the 40 °N parallel of latitude.
</paragraph>
</caption>
<paragraph id="8B8E09462F10524D1BBB8DCDFB4DF937" blockId="16.[144,759,1666,1902]" lastBlockId="16.[806,1421,1666,1902]" pageId="16" pageNumber="517">
Even though
<taxonomicName id="4C3172C52F10524D1A5C8DCDFE79F955" authorityName="Chamberlin" authorityYear="1904" box="[335,488,1727,1749]" class="Chilopoda" family="Mecistocephalidae" genus="Mecistocephalus" kingdom="Animalia" order="Geophilomorpha" pageId="16" pageNumber="517" phylum="Arthropoda" rank="species" species="anomalus">
<emphasis id="B945D5542F10524D1A5C8DCDFE79F955" box="[335,488,1727,1749]" italics="true" pageId="16" pageNumber="517">D. anomalus</emphasis>
</taxonomicName>
was maintained as a valid species by most authors, it was cited as a junior synonym of
<taxonomicName id="4C3172C52F10524D1A0B8DF0FE50F890" authorityName="C. L. Koch" authorityYear="1847" box="[280,449,1788,1810]" class="Chilopoda" family="Geophilidae" genus="Clinopodes" kingdom="Animalia" order="Geophilomorpha" pageId="16" pageNumber="517" phylum="Arthropoda" rank="species" species="carniolensis">
<emphasis id="B945D5542F10524D1A0B8DF0FE50F890" box="[280,449,1788,1810]" italics="true" pageId="16" pageNumber="517">D. carniolensis</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFA074B72F10524D1AF98DF0FD1CF893" author="Attems CG" box="[490,653,1788,1810]" pageId="16" pageNumber="517" refId="ref20647" refString="Attems CG. 1929. Myriapoda I: Geophilomorpha. Das Tierreich 52. Berlin: De Gruyter." type="book" year="1929">Attems (1929)</bibRefCitation>
, followed by other authors (
<bibRefCitation id="EFA074B72F10524D1A7F8C16FE68F8B1" author="Matic Z" box="[364,505,1819,1841]" pageId="16" pageNumber="517" pagination="2" refId="ref22620" refString="Matic Z. 1972. Clasa Chilopoda, Subclasa Epimorpha. Fauna Republicii Socialiste Romania 6 (2). Bucharest: Academiei Republicii Socialiste Romania." type="journal article" year="1972">Matic, 1972</bibRefCitation>
;
<bibRefCitation id="EFA074B72F10524D19198C16FD31F8B1" author="Kevan DKMcE" box="[522,672,1819,1840]" pageId="16" pageNumber="517" pagination="2938 - 2955" refId="ref22292" refString="Kevan DKMcE. 1983. A preliminary survey of known and potentially Canadian and Alaskan centipedes (Chilopoda). Canadian Journal of Zoology 61: 2938 - 2955." type="journal article" year="1983">Kevan, 1983</bibRefCitation>
;
<bibRefCitation id="EFA074B72F10524D19A28C16FF5BF8D1" author="Mikos MZ" pageId="16" pageNumber="517" pagination="9 - 18" refId="ref22788" refString="Mikos MZ. 1991. Dicellophilus carniolensis (C. L. Koch, 1847) kot predstavnik druzine Mecistocephalidae (superfam. Geophilomorpha). Bioloski Vestnik, Ljubljana 39: 9 - 18." type="journal article" year="1991">Mikoš 1991</bibRefCitation>
). Most probably Attems relied on the influential redescription provided by
<bibRefCitation id="EFA074B72F10524D1AD38C55FDE5F8EF" author="Silvestri F" box="[448,628,1880,1902]" pageId="16" pageNumber="517" pagination="45 - 107" refId="ref23270" refString="Silvestri F. 1919. Contributions to a knowledge of the Chilopoda Geophilomorpha of India. Records of the Indian Museum 16: 45 - 107." type="journal article" year="1919">Silvestri (1919)</bibRefCitation>
, where the number of leg-bearing segments was given erroneously as 43 instead of 41 (see below).
</paragraph>
<paragraph id="8B8E09462F10524D182C8DB2FAB2F8EF" blockId="16.[806,1421,1666,1902]" pageId="16" pageNumber="517">
<materialsCitation id="3B59031B2F10524D182C8DB2FA8EF8EF" collectionCode="R, V" collectorName="Chamberlin &amp; French &amp; H. Ribaut. The &amp; L. Leibensperger" location="Museum of Comparative Zoology" municipality="Cambridge" pageId="16" pageNumber="517" specimenCount="1" typeStatus="holotype">
Worth noting is that the
<specimenCount id="9D37C2CF2F10524D1F788DCDFA85F954" box="[1131,1300,1727,1749]" pageId="16" pageNumber="517" type="generic">two specimens</specimenCount>
examined by us (see Material and methods) are topotypic, and were sent by
<collectionCode id="ED2091832F10524D18D18DF0FC44F893" box="[962,981,1789,1810]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="16" pageNumber="517">R</collectionCode>
.
<collectionCode id="ED2091832F10524D18CB8DF0FC7DF893" box="[984,1004,1789,1810]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="16" pageNumber="517" type="Museum">V</collectionCode>
.
<collectorName id="26C46C902F10524D18EA8DF1FB10F893" box="[1017,1153,1788,1810]" pageId="16" pageNumber="517">Chamberlin</collectorName>
as a gift to the
<collectorName id="26C46C902F10524D1E2F8DF1FA1CF893" box="[1340,1421,1788,1810]" pageId="16" pageNumber="517">French</collectorName>
myriapodologist
<collectorName id="26C46C902F10524D18F58C16FB05F8B0" box="[998,1172,1819,1841]" pageId="16" pageNumber="517">H. Ribaut. The</collectorName>
<typeStatus id="548AB7E42F10524D1F8E8C16FB6FF8B0" box="[1181,1278,1819,1841]" pageId="16" pageNumber="517" type="holotype">holotype</typeStatus>
is currently preserved at the
<location id="8EEE5F9D2F10524D18EE8C37FA1BF8CE" LSID="urn:lsid:plazi:treatment:0398B8502F0F524E1BAC8DFBFDA2FA13:8EEE5F9D2F10524D18EE8C37FA1BF8CE" box="[1021,1418,1849,1871]" municipality="Cambridge" name="Museum of Comparative Zoology" pageId="16" pageNumber="517">Museum of Comparative Zoology</location>
,
<collectingMunicipality id="6BEA933C2F10524D18358C55FC32F8EF" box="[806,931,1880,1902]" pageId="16" pageNumber="517">Cambridge</collectingMunicipality>
(
<collectorName id="26C46C902F10524D18A58C55FBEAF8EF" box="[950,1147,1880,1902]" pageId="16" pageNumber="517">L. Leibensperger</collectorName>
, pers. comm.)
</materialsCitation>
.
</paragraph>
<caption id="DF4E59CE2F11524C1B838900FC83FDA3" box="[144,786,525,546]" pageId="17" pageNumber="518" startId="17.[144,208,525,546]">
<paragraph id="8B8E09462F11524C1B838900FC83FDA3" blockId="17.[144,786,525,546]" box="[144,786,525,546]" pageId="17" pageNumber="518">
<emphasis id="B945D5542F11524C1B838900FF78FDA3" bold="true" box="[144,233,525,546]" pageId="17" pageNumber="518">Table 3.</emphasis>
Differential characters of the
<taxonomicName id="4C3172C52F11524C19258900FC83FDA3" box="[566,786,525,546]" class="Chilopoda" family="Mecistocephalidae" genus="Dicellophilus" kingdom="Animalia" order="Geophilomorpha" pageId="17" pageNumber="518" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="B945D5542F11524C19258900FD2EFDA3" box="[566,703,525,546]" italics="true" pageId="17" pageNumber="518">Dicellophilus</emphasis>
species
</taxonomicName>
</paragraph>
</caption>
<paragraph id="8B8E09462F11524C1942894AF910FDDD" blockId="17.[593,729,583,604]" box="[593,1665,583,604]" lastBlockId="17.[1554,1665,583,604]" pageId="17" pageNumber="518">
<emphasis id="B945D5542F11524C1942894AF910FDDD" box="[593,1665,583,604]" italics="true" pageId="17" pageNumber="518">
<taxonomicName id="4C3172C52F11524C1942894AFD48FDDD" authorityName="Chamberlin" authorityYear="1904" box="[593,729,583,604]" class="Chilopoda" family="Mecistocephalidae" genus="Mecistocephalus" kingdom="Animalia" order="Geophilomorpha" pageId="17" pageNumber="518" phylum="Arthropoda" rank="species" species="anomalus">D. anomalus</taxonomicName>
<taxonomicName id="4C3172C52F11524C1888894AFBA8FDDD" authorityName="C. L. Koch" authorityYear="1847" box="[923,1081,583,604]" class="Chilopoda" family="Geophilidae" genus="Clinopodes" kingdom="Animalia" order="Geophilomorpha" pageId="17" pageNumber="518" phylum="Arthropoda" rank="species" species="carniolensis">D. carniolensis</taxonomicName>
<taxonomicName id="4C3172C52F11524C1FDB894AFAA8FDDD" authorityName="Wood" authorityYear="1862" box="[1224,1337,583,604]" class="Chilopoda" family="Mecistocephalidae" genus="Mecistocephalus" kingdom="Animalia" order="Geophilomorpha" pageId="17" pageNumber="518" phylum="Arthropoda" rank="species" species="limatus">D. limatus</taxonomicName>
<taxonomicName id="4C3172C52F11524C1D01894AF910FDDD" baseAuthorityName="Kishida" baseAuthorityYear="1928" box="[1554,1665,583,604]" class="Chilopoda" family="Mecistocephalidae" genus="Dicellophilus" kingdom="Animalia" order="Geophilomorpha" pageId="17" pageNumber="518" phylum="Arthropoda" rank="species" species="pulcher">D. pulcher</taxonomicName>
</emphasis>
</paragraph>
<paragraph id="8B8E09462F11524C1B83898CFB73FD17" blockId="17.[144,1514,640,1569]" box="[144,1250,640,662]" pageId="17" pageNumber="518">Antenna length/head width ~4 ~3 ~4</paragraph>
<paragraph id="8B8E09462F11524C1B838993FB6BFD32" blockId="17.[144,1514,640,1569]" box="[144,1274,664,691]" pageId="17" pageNumber="518">
Antenna, width of article XIV/width
<emphasis id="B945D5542F11524C19428995FDCCFD30" box="[593,605,664,689]" italics="true" pageId="17" pageNumber="518">&lt;</emphasis>
0.5 ~0.5
<emphasis id="B945D5542F11524C1FDB8995FB45FD30" box="[1224,1236,664,689]" italics="true" pageId="17" pageNumber="518">&lt;</emphasis>
0.5
</paragraph>
<paragraph id="8B8E09462F11524C1BB589B6FE86FD51" blockId="17.[144,1514,640,1569]" box="[166,279,699,720]" pageId="17" pageNumber="518">of article I</paragraph>
<paragraph id="8B8E09462F11524C1B8389D4FA85FD6F" blockId="17.[144,1514,640,1569]" box="[144,1300,729,750]" pageId="17" pageNumber="518">Antenna, article XIV, length/width 2.22.4 1.72.0 2.22.4</paragraph>
<paragraph id="8B8E09462F11524C1B8389FBFB6BFC8A" blockId="17.[144,1514,640,1569]" box="[144,1274,751,779]" pageId="17" pageNumber="518">
Antenna, length of article XIV/length
<emphasis id="B945D5542F11524C194289E2FDCCFC89" box="[593,605,751,776]" italics="true" pageId="17" pageNumber="518">&lt;</emphasis>
1.5
<emphasis id="B945D5542F11524C188889E2FC36FC89" box="[923,935,751,776]" italics="true" pageId="17" pageNumber="518">&gt;</emphasis>
1.5
<emphasis id="B945D5542F11524C1FDB89E2FB45FC89" box="[1224,1236,751,776]" italics="true" pageId="17" pageNumber="518">&lt;</emphasis>
1.5
</paragraph>
<paragraph id="8B8E09462F11524C1BB5881EFEABFCA9" blockId="17.[144,1514,640,1569]" box="[166,314,787,808]" pageId="17" pageNumber="518">of article XIII</paragraph>
<paragraph id="8B8E09462F11524C1B83883CFAB3FCC7" blockId="17.[144,1514,640,1569]" box="[144,1314,816,838]" pageId="17" pageNumber="518">Antenna, setae, maximum length ~300 Mm 150200 Mm ~300 Mm</paragraph>
<paragraph id="8B8E09462F11524C1B838843FE9CFC01" blockId="17.[144,1514,640,1569]" pageId="17" pageNumber="518">Antenna, apical sensilla, crown-like Usually present Absent Usually present projection</paragraph>
<paragraph id="8B8E09462F11524C1B838885FA85FC1C" blockId="17.[144,1514,640,1569]" box="[144,1300,904,925]" pageId="17" pageNumber="518">Cephalic plate, length/width 1.51.6 1.21.4 1.51.6</paragraph>
<paragraph id="8B8E09462F11524C1B8388ABFC4CFB93" blockId="17.[144,1514,640,1569]" pageId="17" pageNumber="518">Cephalic plate, lateral margins, shape Almost straight, evidently Markedly convex, Almost straight, evidently converging for most of evidently converging converging for most of the length only for the posterior the length part</paragraph>
<paragraph id="8B8E09462F11524C1B838F16FAC1FBEB" blockId="17.[144,1514,640,1569]" pageId="17" pageNumber="518">Cephalic plate, frontal line, shape Angle pointing forwards Uniformly rounded Angle pointing forwards Clypeus, lateral margins, shape Evidently converging Only slightly converging Evidently converging posteriorly posteriorly posteriorly</paragraph>
<paragraph id="8B8E09462F11524C1B838F7FFF44FB24" blockId="17.[144,1514,640,1569]" pageId="17" pageNumber="518">Clypeus, setae on a central triangular Absent Present Absent area</paragraph>
<paragraph id="8B8E09462F11524C1B838FA0FE5EFB5E" blockId="17.[144,1514,640,1569]" pageId="17" pageNumber="518">Clypeus, distinctly isolated Present Absent Present posteromedian pair of setae</paragraph>
<paragraph id="8B8E09462F11524C1B838FE5FC6CFAB6" blockId="17.[144,1514,640,1569]" pageId="17" pageNumber="518">Bucca, non-areolate part, anterior Evidently overreaching the Not or only slightly Evidently overreaching the extent labrum overreaching the labrum labrum</paragraph>
<paragraph id="8B8E09462F11524C1B838E32FB7FFAD5" blockId="17.[144,1514,640,1569]" box="[144,1262,1343,1364]" pageId="17" pageNumber="518">Mandible, pectinate lamellae, number 57 35 57</paragraph>
<paragraph id="8B8E09462F11524C1B838E50FA85FAF3" blockId="17.[144,1514,640,1569]" box="[144,1300,1373,1394]" pageId="17" pageNumber="518">maxillae I, coxosternum, 2.63.0 3.33.5 2.63.0</paragraph>
<paragraph id="8B8E09462F11524C1BB58E77FEEDFA0E" blockId="17.[144,1514,640,1569]" box="[166,380,1402,1423]" pageId="17" pageNumber="518">width/medial length</paragraph>
<paragraph id="8B8E09462F11524C1B838E9AFA85FA2D" blockId="17.[144,1514,640,1569]" box="[144,1300,1431,1452]" pageId="17" pageNumber="518">Maxillae I, medial projection, length/ 2.02.5 0.51.5 2.02.5</paragraph>
<paragraph id="8B8E09462F11524C1BB58EB9FF73FA48" blockId="17.[144,1514,640,1569]" box="[166,226,1460,1481]" pageId="17" pageNumber="518">width</paragraph>
<paragraph id="8B8E09462F11524C1B838EDFFBD3F9A0" blockId="17.[144,1514,640,1569]" pageId="17" pageNumber="518">Maxillae I, medial projection, distal Evidently enlarged Only slightly enlarged Evidently enlarged hyaline part, shape distally, subtriangular distally, not distally, subtriangular subtriangular</paragraph>
<paragraph id="8B8E09462F11524C1D01898DF9BAFD17" blockId="17.[1554,1604,640,691]" box="[1554,1579,640,662]" pageId="17" pageNumber="518">~3</paragraph>
<paragraph id="8B8E09462F11524C1D018995F9D5FD32" blockId="17.[1554,1604,640,691]" box="[1554,1604,664,691]" pageId="17" pageNumber="518">
<emphasis id="B945D5542F11524C1D018995F98FFD30" box="[1554,1566,664,689]" italics="true" pageId="17" pageNumber="518">&lt;</emphasis>
0.5
</paragraph>
<paragraph id="8B8E09462F11524C1D0189D4F9CCFD6F" blockId="17.[1554,1629,729,779]" box="[1554,1629,729,750]" pageId="17" pageNumber="518">2.22.4</paragraph>
<paragraph id="8B8E09462F11524C1D0189E2F9D5FC8A" blockId="17.[1554,1629,729,779]" box="[1554,1604,751,779]" pageId="17" pageNumber="518">
<emphasis id="B945D5542F11524C1D0189E2F98FFC89" box="[1554,1566,751,776]" italics="true" pageId="17" pageNumber="518">&lt;</emphasis>
1.5
</paragraph>
<paragraph id="8B8E09462F11524C1D01883CF900FCC7" blockId="17.[1554,1724,816,867]" box="[1554,1681,816,838]" pageId="17" pageNumber="518">150200 Mm</paragraph>
<paragraph id="8B8E09462F11524C1D018843F92DFCE2" blockId="17.[1554,1724,816,867]" box="[1554,1724,846,867]" pageId="17" pageNumber="518">Usually present</paragraph>
<paragraph id="8B8E09462F11524C1D018885F9CCFC1C" blockId="17.[1554,1804,904,1159]" box="[1554,1629,904,925]" pageId="17" pageNumber="518">1.21.4</paragraph>
<paragraph id="8B8E09462F11524C1D0188ABF9C5FB93" blockId="17.[1554,1804,904,1159]" pageId="17" pageNumber="518">Markedly convex, evidently converging only for the posterior part</paragraph>
<paragraph id="8B8E09462F11524C1D018F16F94DFBB1" blockId="17.[1554,1804,904,1159]" box="[1554,1756,1051,1072]" pageId="17" pageNumber="518">Uniformly rounded</paragraph>
<paragraph id="8B8E09462F11524C1D018F35F908FBEB" blockId="17.[1554,1804,904,1159]" pageId="17" pageNumber="518">Evidently converging posteriorly</paragraph>
<paragraph id="8B8E09462F11524C1D018F7FF9CAFB06" blockId="17.[1554,1804,904,1159]" box="[1554,1627,1138,1159]" pageId="17" pageNumber="518">Absent</paragraph>
<paragraph id="8B8E09462F11524C1D018FA0F9CAFB43" blockId="17.[1554,1627,1197,1218]" box="[1554,1627,1197,1218]" pageId="17" pageNumber="518">Absent</paragraph>
<paragraph id="8B8E09462F11524C1D018FE5F9E5FA9B" blockId="17.[1554,1843,1256,1306]" pageId="17" pageNumber="518">Evidently overreaching the labrum</paragraph>
<paragraph id="8B8E09462F11524C1D018E32F9A6FAD5" blockId="17.[1554,1629,1343,1394]" box="[1554,1591,1343,1364]" pageId="17" pageNumber="518">57</paragraph>
<paragraph id="8B8E09462F11524C1D018E50F9CCFAF3" blockId="17.[1554,1629,1343,1394]" box="[1554,1629,1373,1394]" pageId="17" pageNumber="518">2.63.0</paragraph>
<paragraph id="8B8E09462F11524C1D018E9AF9CFFA2D" blockId="17.[1554,1630,1431,1452]" box="[1554,1630,1431,1452]" pageId="17" pageNumber="518">2.02.5</paragraph>
<paragraph id="8B8E09462F11524F1D018EDFFEAAFD39" blockId="17.[1554,1813,1490,1540]" lastBlockId="18.[144,1501,645,1457]" lastPageId="18" lastPageNumber="519" pageId="17" pageNumber="518">Evidently enlarged distally, subtriangular Maxillae I, medial projection, dorsal Present Absent Present hyaline scales</paragraph>
<paragraph id="8B8E09462F12524F1B8389CDFB65FD54" blockId="18.[144,1501,645,1457]" box="[144,1268,703,725]" pageId="18" pageNumber="519">Maxillae I, telopodite, length of the ~1.5 ~1.0 ~1.5</paragraph>
<paragraph id="8B8E09462F12524F1BB589D0FEEDFC8E" blockId="18.[144,1501,645,1457]" pageId="18" pageNumber="519">distal hyaline part/length of the basal chitinous part</paragraph>
<paragraph id="8B8E09462F12524F1B838815FBDCFC05" blockId="18.[144,1501,645,1457]" pageId="18" pageNumber="519">Maxillae II, telopodite, distal article, Subconical, tapering Subcylindrical for most Subconical, tapering shape gradually along most of of the length, gradually along most of the length tapering abruptly in the length the distal part</paragraph>
<paragraph id="8B8E09462F12524F1B838880FA85FC23" blockId="18.[144,1501,645,1457]" box="[144,1300,909,930]" pageId="18" pageNumber="519">Maxillae II, telopodite, distal article, 1.31.6 1.83.5 1.31.6</paragraph>
<paragraph id="8B8E09462F12524F1BB588A7FEBDFC3E" blockId="18.[144,1501,645,1457]" box="[166,300,938,959]" pageId="18" pageNumber="519">length/width</paragraph>
<paragraph id="8B8E09462F12524F1B8388CAFEFAFC7B" blockId="18.[144,1501,645,1457]" pageId="18" pageNumber="519">Forcipular segment, coxosternum, Well developed Virtually lacking Well developed condylar processes</paragraph>
<paragraph id="8B8E09462F12524F1B838F0FFB01FB28" blockId="18.[144,1501,645,1457]" pageId="18" pageNumber="519">Forcipular segment, 1.21.3 1.21.3 1.21.3 trochanteropraefemur, length/width forcipular segment, Large, greatly projecting Small, not or only Large, greatly projecting trochanteropraefemur, distal tubercle, beyond the distal slightly projecting beyond the distal relative size in respect to the article margin of the article beyond the distal margin of the article margin of the article</paragraph>
<paragraph id="8B8E09462F12524F1B838FBCFF6AFB65" blockId="18.[144,1501,645,1457]" pageId="18" pageNumber="519">Forcipular segment, tarsungulum, basal Well developed Virtually lacking Well developed tubercle</paragraph>
<paragraph id="8B8E09462F12524F1B838FE1FABEFA9F" blockId="18.[144,1501,645,1457]" pageId="18" pageNumber="519">Forcipular segment, tarsungulum, Weak, uniformly spaced Weak, uniformly spaced Weak, uniformly spaced marginal crenulation, shape notches notches notches</paragraph>
<paragraph id="8B8E09462F12524F1B838E2BFB70FABA" blockId="18.[144,1501,645,1457]" box="[144,1249,1318,1339]" pageId="18" pageNumber="519">Leg-bearing segments, number 41 43 45</paragraph>
<paragraph id="8B8E09462F12524F1B838E49FA82FAD8" blockId="18.[144,1501,645,1457]" box="[144,1299,1348,1369]" pageId="18" pageNumber="519">Last leg-bearing segment, sternum, 0.80.9 1.01.2 0.80.9</paragraph>
<paragraph id="8B8E09462F12524F1BB58E6CFEBDFAF7" blockId="18.[144,1501,645,1457]" box="[166,300,1377,1398]" pageId="18" pageNumber="519">length/width</paragraph>
<paragraph id="8B8E09462F12524F1B838E73FEEEFA30" blockId="18.[144,1501,645,1457]" pageId="18" pageNumber="519">Last leg-bearing segment, coxopleuron, Usually distinctly pointed Rounded Usually slightly pointed posterior end, shape</paragraph>
<paragraph id="8B8E09462F12524F1D018988F9F2FD1B" blockId="18.[1554,1635,645,666]" box="[1554,1635,645,666]" pageId="18" pageNumber="519">Present</paragraph>
<paragraph id="8B8E09462F12524F1D0189B2F9ACFD54" blockId="18.[1554,1597,703,725]" box="[1554,1597,703,725]" pageId="18" pageNumber="519">~1.5</paragraph>
<paragraph id="8B8E09462F12524F1D018815F907FCE6" blockId="18.[1554,1831,792,871]" pageId="18" pageNumber="519">Subconical, tapering gradually along most of the length</paragraph>
<paragraph id="8B8E09462F12524F1D018880F9CCFC23" blockId="18.[1554,1629,909,930]" box="[1554,1629,909,930]" pageId="18" pageNumber="519">1.31.6</paragraph>
<paragraph id="8B8E09462F12524F1D0188CAF921FC5D" blockId="18.[1554,1712,967,988]" box="[1554,1712,967,988]" pageId="18" pageNumber="519">Well developed</paragraph>
<paragraph id="8B8E09462F12524F1D018F0FF9A0FB96" blockId="18.[1554,1585,1026,1047]" box="[1554,1585,1026,1047]" pageId="18" pageNumber="519">1.0</paragraph>
<paragraph id="8B8E09462F12524F1D018F31F9FAFB28" blockId="18.[1554,1836,1084,1222]" pageId="18" pageNumber="519">Small, not or only slightly projecting beyond the distal margin of the article</paragraph>
<paragraph id="8B8E09462F12524F1D018FBCF921FB47" blockId="18.[1554,1836,1084,1222]" box="[1554,1712,1201,1222]" pageId="18" pageNumber="519">Well developed</paragraph>
<paragraph id="8B8E09462F12524F1D018FE1F9E9FA9F" blockId="18.[1554,1817,1260,1369]" pageId="18" pageNumber="519">Evident, variably spaced notches</paragraph>
<paragraph id="8B8E09462F12524F1D018E2BF9BAFABA" blockId="18.[1554,1817,1260,1369]" box="[1554,1579,1318,1339]" pageId="18" pageNumber="519">41</paragraph>
<paragraph id="8B8E09462F12524F1D018E49F9CCFAD8" blockId="18.[1554,1817,1260,1369]" box="[1554,1629,1348,1369]" pageId="18" pageNumber="519">1.01.2</paragraph>
<paragraph id="8B8E09462F12524E1D018E73FF7BFE6E" blockId="18.[1554,1810,1406,1427]" lastBlockId="19.[164,779,198,1107]" lastPageId="19" lastPageNumber="520" pageId="18" pageNumber="519">
Usually slightly pointed
<emphasis id="B945D5542F13524E1BB78BCAFE0BFF5A" box="[164,410,198,220]" italics="true" pageId="19" pageNumber="520">Notes on morphology:</emphasis>
The number of leg-bearing segments was reported as 43 by
<bibRefCitation id="EFA074B72F13524E1AE78BE9FD34FF7A" author="Silvestri F" box="[500,677,228,251]" pageId="19" pageNumber="520" pagination="45 - 107" refId="ref23270" refString="Silvestri F. 1919. Contributions to a knowledge of the Chilopoda Geophilomorpha of India. Records of the Indian Museum 16: 45 - 107." type="journal article" year="1919">Silvestri (1919)</bibRefCitation>
, but this was probably a mistake, as suggested by the fact that
<bibRefCitation id="EFA074B72F13524E1BB78A2FFEC6FEB9" author="Silvestri F" box="[164,343,290,312]" pageId="19" pageNumber="520" pagination="45 - 107" refId="ref23270" refString="Silvestri F. 1919. Contributions to a knowledge of the Chilopoda Geophilomorpha of India. Records of the Indian Museum 16: 45 - 107." type="journal article" year="1919">Silvestri (1919)</bibRefCitation>
did not comment upon this discrepancy, even though the number had been reported consistently as being invariantly 41 for the species (
<bibRefCitation id="EFA074B72F13524E1BB88A70FE13FE12" author="Chamberlin RV" box="[171,386,381,403]" pageId="19" pageNumber="520" pagination="651 - 657" refId="ref21072" refString="Chamberlin RV. 1904. New chilopods. Proceedings of the Academy of Natural Sciences of Philadelphia 56: 651 - 657." type="journal article" year="1904">Chamberlin, 1904</bibRefCitation>
,
<bibRefCitation id="EFA074B72F13524E1A858A73FE41FE15" author="Chamberlin RV" box="[406,464,382,404]" pageId="19" pageNumber="520" pagination="651 - 672" refId="ref21095" refString="Chamberlin RV. 1912. The Chilopoda of California III. Pomona Journal of Entomology 4: 651 - 672." type="journal article" year="1912">1912</bibRefCitation>
), and intraspecific invariance in the number was taken for granted for all mecistocephalid species at that time (Bonato
<emphasis id="B945D5542F13524E19DF8AB6FC95FE51" box="[716,772,443,464]" italics="true" pageId="19" pageNumber="520">et al.</emphasis>
, 2002).
</paragraph>
<paragraph id="8B8E09462F13524E1BAF8AF5FDC5FBD2" blockId="19.[164,779,198,1107]" pageId="19" pageNumber="520">
Apart from the number of trunk segments, two other characters were listed as being different between
<taxonomicName id="4C3172C52F13524E1A1D893BFE35FDCB" authorityName="Chamberlin" authorityYear="1904" box="[270,420,565,587]" class="Chilopoda" family="Mecistocephalidae" genus="Mecistocephalus" kingdom="Animalia" order="Geophilomorpha" pageId="19" pageNumber="520" phylum="Arthropoda" rank="species" species="anomalus">
<emphasis id="B945D5542F13524E1A1D893BFE35FDCB" box="[270,420,565,587]" italics="true" pageId="19" pageNumber="520">D. anomalus</emphasis>
</taxonomicName>
and
<taxonomicName id="4C3172C52F13524E1AF5893BFDF2FDCB" authorityName="Wood" authorityYear="1862" box="[486,611,565,587]" class="Chilopoda" family="Mecistocephalidae" genus="Mecistocephalus" kingdom="Animalia" order="Geophilomorpha" pageId="19" pageNumber="520" phylum="Arthropoda" rank="species" species="limatus">
<emphasis id="B945D5542F13524E1AF5893BFDF2FDCB" box="[486,611,565,587]" italics="true" pageId="19" pageNumber="520">D. limatus</emphasis>
</taxonomicName>
, namely body size (
<taxonomicName id="4C3172C52F13524E1BF58959FEEFFDE8" authorityName="Chamberlin" authorityYear="1904" box="[230,382,596,617]" class="Chilopoda" family="Mecistocephalidae" genus="Mecistocephalus" kingdom="Animalia" order="Geophilomorpha" pageId="19" pageNumber="520" phylum="Arthropoda" rank="species" species="anomalus">
<emphasis id="B945D5542F13524E1BF58959FEEFFDE8" box="[230,382,596,617]" italics="true" pageId="19" pageNumber="520">D. anomalus</emphasis>
</taxonomicName>
was described as being smaller than
<taxonomicName id="4C3172C52F13524E1BF1897EFECDFD06" authorityName="Wood" authorityYear="1862" box="[226,348,626,648]" class="Chilopoda" family="Mecistocephalidae" genus="Mecistocephalus" kingdom="Animalia" order="Geophilomorpha" pageId="19" pageNumber="520" phylum="Arthropoda" rank="species" species="limatus">
<emphasis id="B945D5542F13524E1BF1897EFECDFD06" box="[226,348,626,648]" italics="true" pageId="19" pageNumber="520">D. limatus</emphasis>
</taxonomicName>
, according to
<bibRefCitation id="EFA074B72F13524E1913897FFD5EFD09" author="Chamberlin RV" box="[512,719,626,648]" pageId="19" pageNumber="520" pagination="651 - 657" refId="ref21072" refString="Chamberlin RV. 1904. New chilopods. Proceedings of the Academy of Natural Sciences of Philadelphia 56: 651 - 657." type="journal article" year="1904">Chamberlin, 1904</bibRefCitation>
) and antennal length (shorter in
<taxonomicName id="4C3172C52F13524E1AE3899CFD15FD27" authorityName="Chamberlin" authorityYear="1904" box="[496,644,657,678]" class="Chilopoda" family="Mecistocephalidae" genus="Mecistocephalus" kingdom="Animalia" order="Geophilomorpha" pageId="19" pageNumber="520" phylum="Arthropoda" rank="species" species="anomalus">
<emphasis id="B945D5542F13524E1AE3899CFD15FD27" box="[496,644,657,678]" italics="true" pageId="19" pageNumber="520">D. anomalus</emphasis>
</taxonomicName>
than in
<taxonomicName id="4C3172C52F13524E19E2899CFF6AFD45" authorityName="Wood" authorityYear="1862" class="Chilopoda" family="Mecistocephalidae" genus="Mecistocephalus" kingdom="Animalia" order="Geophilomorpha" pageId="19" pageNumber="520" phylum="Arthropoda" rank="species" species="limatus">
<emphasis id="B945D5542F13524E19E2899CFF6AFD45" italics="true" pageId="19" pageNumber="520">D. limatus</emphasis>
</taxonomicName>
, according to
<bibRefCitation id="EFA074B72F13524E1AA389A2FD16FD47" author="Chamberlin RV" box="[432,647,687,710]" pageId="19" pageNumber="520" pagination="651 - 672" refId="ref21095" refString="Chamberlin RV. 1912. The Chilopoda of California III. Pomona Journal of Entomology 4: 651 - 672." type="journal article" year="1912">Chamberlin, 1912</bibRefCitation>
). However, Chamberlins evaluation of the body size of
<taxonomicName id="4C3172C52F13524E198089C2FC9AFD62" authorityName="Wood" authorityYear="1862" box="[659,779,718,740]" class="Chilopoda" family="Mecistocephalidae" genus="Mecistocephalus" kingdom="Animalia" order="Geophilomorpha" pageId="19" pageNumber="520" phylum="Arthropoda" rank="species" species="limatus">
<emphasis id="B945D5542F13524E198089C2FC9AFD62" box="[659,779,718,740]" italics="true" pageId="19" pageNumber="520">D. limatus</emphasis>
</taxonomicName>
was obviously based on a single specimen that was not fully grown, as suggested by the pattern of coxal pores described. Furthermore, our morphometric analyses of representative specimens of both species did not confirm any significant difference in the elongation of the antennae in respect to the head, nor in the elongation of the articles (
<tableCitation id="C6B33CFD2F13524E1AED88A9FDC2FC3B" box="[510,595,932,954]" captionStart="Table 3" captionStartId="17.[144,208,525,546]" captionText="Table 3. Differential characters of the Dicellophilus species" pageId="19" pageNumber="520">Table 3</tableCitation>
). Also, the body colour was described as slightly different between the species (
<bibRefCitation id="EFA074B72F13524E1A1088ECFE43FC76" author="Chamberlin RV" box="[259,466,993,1015]" pageId="19" pageNumber="520" pagination="651 - 657" refId="ref21072" refString="Chamberlin RV. 1904. New chilopods. Proceedings of the Academy of Natural Sciences of Philadelphia 56: 651 - 657." type="journal article" year="1904">Chamberlin, 1904</bibRefCitation>
,
<bibRefCitation id="EFA074B72F13524E1ACD88EFFD89FC79" author="Chamberlin RV" box="[478,536,994,1016]" pageId="19" pageNumber="520" pagination="651 - 672" refId="ref21095" refString="Chamberlin RV. 1912. The Chilopoda of California III. Pomona Journal of Entomology 4: 651 - 672." type="journal article" year="1912">1912</bibRefCitation>
), but it is expected to be obviously affected by the condition of preservation and the treatment of the specimens, as acknowledged for
<taxonomicName id="4C3172C52F13524E1BDE8F33FED9FBD3" authorityName="Wood" authorityYear="1862" box="[205,328,1085,1107]" class="Chilopoda" family="Mecistocephalidae" genus="Mecistocephalus" kingdom="Animalia" order="Geophilomorpha" pageId="19" pageNumber="520" phylum="Arthropoda" rank="species" species="limatus">
<emphasis id="B945D5542F13524E1BDE8F33FED9FBD3" box="[205,328,1085,1107]" italics="true" pageId="19" pageNumber="520">D. limatus</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFA074B72F13524E1A658F30FD91FBD2" author="Wood HC" box="[374,512,1085,1107]" pageId="19" pageNumber="520" pagination="5 - 52" refId="ref23857" refString="Wood HC. 1862. On the Chilopoda of North America with a catalogue of all the specimens in the collection of the Smithsonian Institution. Journal of the Academy of Natural Sciences of Philadelphia 5: 5 - 52." type="journal article" year="1862">Wood (1862</bibRefCitation>
,
<bibRefCitation id="EFA074B72F13524E191E8F30FDD6FBD2" author="Wood HC" box="[525,583,1085,1107]" pageId="19" pageNumber="520" pagination="137 - 248" refId="ref23898" refString="Wood HC. 1865. The Myriopoda of North America. Transactions of the American Philosophical Society (N. S.), 13: 137 - 248." type="journal article" year="1865">1865</bibRefCitation>
).
</paragraph>
<paragraph id="8B8E09462F13524E1BB78F8AFDA2FA13" blockId="19.[164,779,1159,1426]" pageId="19" pageNumber="520">
<emphasis id="B945D5542F13524E1BB78F8AFEA4FB1D" box="[164,309,1159,1180]" italics="true" pageId="19" pageNumber="520">Distribution:</emphasis>
The species occurs in a restricted area in the south-western part of North America, between the Pacific Coast Ranges and the Cascade Range and Sierra Nevada, from central California to Oregon (
<figureCitation id="130A15C32F13524E1BBF8E0CFF6CFA96" box="[172,253,1281,1303]" captionStart="Figure 10" captionStartId="16.[144,223,1507,1526]" captionTargetBox="[149,1426,200,1472]" captionTargetId="figure-125@16.[144,1426,195,1477]" captionTargetPageId="16" captionText="Figure 10. Geographical distribution and phylogenetic relationships of the species of Dicellophilus. The shaded areas indicate the established populations; stars indicate either occasional or dubious records. The dotted line indicates the 40 °N parallel of latitude." figureDoi="http://doi.org/10.5281/zenodo.5438242" httpUri="https://zenodo.org/record/5438242/files/figure.png" pageId="19" pageNumber="520">Fig. 10</figureCitation>
). It has been recorded in only a few localities, in particular around Monterey Bay, near Pacific Grove, California (
<bibRefCitation id="EFA074B72F13524E1A8A8E33FDE8FAD2" author="Chamberlin RV" box="[409,633,1342,1364]" pageId="19" pageNumber="520" pagination="651 - 657" refId="ref21072" refString="Chamberlin RV. 1904. New chilopods. Proceedings of the Academy of Natural Sciences of Philadelphia 56: 651 - 657." type="journal article" year="1904">Chamberlin, 1904</bibRefCitation>
,
<bibRefCitation id="EFA074B72F13524E19858E33FD41FAD5" author="Chamberlin RV" box="[662,720,1342,1364]" pageId="19" pageNumber="520" pagination="651 - 672" refId="ref21095" refString="Chamberlin RV. 1912. The Chilopoda of California III. Pomona Journal of Entomology 4: 651 - 672." type="journal article" year="1912">1912</bibRefCitation>
), at Oroville, California (
<bibRefCitation id="EFA074B72F13524E1ABD8E50FD1EFAF2" author="Chamberlin RV" box="[430,655,1373,1395]" pageId="19" pageNumber="520" pagination="651 - 672" refId="ref21095" refString="Chamberlin RV. 1912. The Chilopoda of California III. Pomona Journal of Entomology 4: 651 - 672." type="journal article" year="1912">Chamberlin, 1912</bibRefCitation>
), and at
<collectingCountry id="F32649D62F13524E1BB78E76FE98FA10" box="[164,265,1403,1425]" name="Lebanon" pageId="19" pageNumber="520">Lebanon</collectingCountry>
, Oregon (
<bibRefCitation id="EFA074B72F13524E1A6F8E76FDB6FA13" author="Silvestri F" box="[380,551,1403,1426]" pageId="19" pageNumber="520" pagination="45 - 107" refId="ref23270" refString="Silvestri F. 1919. Contributions to a knowledge of the Chilopoda Geophilomorpha of India. Records of the Indian Museum 16: 45 - 107." type="journal article" year="1919">Silvestri, 1919</bibRefCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>