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<document id="61258D60D31DF6DA8DC05F9138B9910A" ID-CLB-Dataset="135067" ID-DOI="10.11646/zootaxa.5182.1.3" ID-GBIF-Dataset="8807b216-9d4c-49f2-bb19-7f0ae3e4d1b9" ID-ISSN="1175-5326" ID-Zenodo-Dep="7042806" ID-ZooBank="D36CE632-720E-4D9C-A7D8-654F62322509" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1662105380701" checkinUser="plazi" docAuthor="Kluge, Nikita" docDate="2022" docId="F26287EBE876AA6C38CEF9AB0210C5E6" docLanguage="en" docName="zootaxa.5182.1.3.pdf" docOrigin="Zootaxa 5182 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Waynokiops pentacanthus Kluge 2022, sp.n." docType="treatment" docVersion="6" lastPageNumber="44" masterDocId="0E5BFF93E875AA6F3859FFFB0050C248" masterDocTitle="Two new species of Waynokiops Hill et al. 2010 (Ephemeroptera: Baetidae) from the Oriental Region" masterLastPageNumber="63" masterPageNumber="41" pageNumber="44" updateTime="1699374775996" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="00CFDFA62CA77A2F56DAEB8B3B68F892">Two new species of Waynokiops Hill et al. 2010 (Ephemeroptera: Baetidae) from the Oriental Region</mods:title>
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<mods:namePart id="1E1E1507A4BAF2F616DC8470F62CA797">Kluge, Nikita</mods:namePart>
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<mods:date id="A063FDBC5441A268131BDE831B720406">2022</mods:date>
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<mods:number id="ADA754EB98FAEFBF9F6227972A3367BD">2022-09-02</mods:number>
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<treatment id="F26287EBE876AA6C38CEF9AB0210C5E6" ID-DOI="http://doi.org/10.5281/zenodo.7046947" ID-GBIF-Taxon="199919402" ID-Zenodo-Dep="7046947" LSID="urn:lsid:plazi:treatment:F26287EBE876AA6C38CEF9AB0210C5E6" httpUri="http://treatment.plazi.org/id/F26287EBE876AA6C38CEF9AB0210C5E6" lastPageNumber="44" pageId="3" pageNumber="44">
<subSubSection id="32D16576E876AA6C38CEF9AB0246C423" box="[151,534,1616,1643]" pageId="3" pageNumber="44" type="nomenclature">
<paragraph id="7A7436FDE876AA6C38CEF9AB0246C423" blockId="3.[151,534,1616,1678]" box="[151,534,1616,1643]" pageId="3" pageNumber="44">
<heading id="213C8191E876AA6C38CEF9AB0246C423" bold="true" box="[151,534,1616,1643]" fontSize="11" level="1" pageId="3" pageNumber="44" reason="1">
<emphasis id="48BFEAEFE876AA6C38CEF9AB0246C423" bold="true" box="[151,534,1616,1643]" pageId="3" pageNumber="44">
<taxonomicName id="BDCB4D7EE876AA6C38CEF9AB0184C423" authority="Kluge, 2022" authorityName="Kluge" authorityYear="2022" box="[151,468,1616,1643]" class="Insecta" family="Baetidae" genus="Waynokiops" kingdom="Animalia" order="Ephemeroptera" pageId="3" pageNumber="44" phylum="Arthropoda" rank="species" species="pentacanthus" status="sp. nov.">
<emphasis id="48BFEAEFE876AA6C38CEF9AB0184C423" bold="true" box="[151,468,1616,1643]" italics="true" pageId="3" pageNumber="44">Waynokiops pentacanthus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="538C5794E876AA6C3982F9AA0246C423" box="[475,534,1617,1643]" pageId="3" pageNumber="44" rank="species">sp.n.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
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<subSubSection id="32D16576E876AA6C38CEF98F0149C4C6" box="[151,281,1652,1678]" pageId="3" pageNumber="44" type="description">
<paragraph id="7A7436FDE876AA6C38CEF98F0149C4C6" blockId="3.[151,534,1616,1678]" box="[151,281,1652,1678]" pageId="3" pageNumber="44">
(
<figureCitation id="E2F02A78E876AA6C38C6F98F0141C4C6" box="[159,273,1652,1678]" captionStart-0="FIGURES 110" captionStart-1="FIGURES 1120" captionStart-2="FIGURES 2127" captionStart-3="FIGURES 2834" captionStart-4="FIGURES 3538" captionStart-5="FIGURES 3942" captionStart-6="FIGURES 4352" captionStart-7="FIGURES 5355" captionStart-8="FIGURES 5659" captionStart-9="FIGURES 6063" captionStartId-0="4.[151,264,1841,1865]" captionStartId-1="5.[151,264,1831,1855]" captionStartId-2="6.[151,264,1563,1587]" captionStartId-3="7.[151,264,1841,1865]" captionStartId-4="8.[151,264,1831,1855]" captionStartId-5="9.[151,264,1282,1306]" captionStartId-6="10.[151,264,1770,1794]" captionStartId-7="11.[151,264,1841,1865]" captionStartId-8="12.[151,264,1710,1734]" captionStartId-9="13.[151,264,1176,1200]" captionTargetBox-0="[151,1436,179,1812]" captionTargetBox-1="[151,1436,181,1808]" captionTargetBox-2="[153,1436,181,1539]" captionTargetBox-3="[172,1419,189,1816]" captionTargetBox-4="[151,1436,181,1808]" captionTargetBox-5="[151,1436,197,1258]" captionTargetBox-6="[151,1436,181,1746]" captionTargetBox-7="[151,1436,181,1818]" captionTargetBox-8="[152,1420,183,1678]" captionTargetBox-9="[151,1436,181,1152]" captionTargetId-0="figure-18@4.[151,1436,179,1815]" captionTargetId-1="figure-16@5.[151,1436,181,1808]" captionTargetId-2="figure-18@6.[151,1436,181,1539]" captionTargetId-3="figure-16@7.[151,1436,181,1817]" captionTargetId-4="figure-18@8.[151,1436,181,1808]" captionTargetId-5="figure-16@9.[151,1436,181,1258]" captionTargetId-6="figure-18@10.[151,1436,181,1746]" captionTargetId-7="figure-16@11.[151,1436,181,1818]" captionTargetId-8="figure-18@12.[151,1436,181,1687]" captionTargetId-9="figure-16@13.[151,1436,181,1152]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="8" captionTargetPageId-5="9" captionTargetPageId-6="10" captionTargetPageId-7="11" captionTargetPageId-8="12" captionTargetPageId-9="13" captionText-0="FIGURES 110. Waynokiops pentacanthus sp. n., larvae. 12, female larvae of last instar; 38, median part of abdominal terga II, VI, VII, VIII, IX and X; 9, median part of sternum VII; 10, thorax and abdomen of young larva." captionText-1="FIGURES 1120. Waynokiops pentacanthus sp. n., larval exuviae (1118 with same magnification). 11, abdomen; 1214, half of pronotum and mesonotum (arrows show small proruberances); 15, head; 1618, fore, middle and hind legs; 19, fragment of cercus (dorsal view); 20, paraproct (11, 14, 1619, holotype)." captionText-2="FIGURES 2127. Waynokiops pentacanthus sp. n., larvae. 2122, left and right mandibles; 23, maxilla; 24, labium (focus on ventral side); 25, hypopharynx with superlinguae; 26, labrum (focus on dorsal side); 27, the same, middle of anterior margin (focus on ventral side)." captionText-3="FIGURES 2834. Waynokiops pentacanthus sp. n., larvae. 28, abdomen without tergalii IIV; 2932, tergalii IIV; 3334, apices of right and left mandibles." captionText-4="FIGURES 3538. Waynokiops pentacanthus sp. n., exuviae of larval fore legs. 3536, two fragments of tibia, tarsus and claw, ventral view; 37, femur; 38, apex of femur and base of tibia. Abbreviation: pts, patella-tibial suture crossing ventral side of tibia." captionText-5="FIGURES 3942. Waynokiops pentacanthus sp. n., patella-tibial suture, anterior view. 3940, fore and middle legs of larva; 4142, fore and middle legs of female subimago." captionText-6="FIGURES 4352. Waynokiops pentacanthus sp. n. 4345, male imagines; 46, fore leg of male imago; 47, fore leg of female imago; 48, its subimaginal exuviae; 49, middle leg of female imago; 50, female imago; 51, subimaginal exuviae of meso- and metapleuron; 52, subimaginal exuviae of mesonotum (4446, holotype)." captionText-7="FIGURES 5355. Waynokiops pentacanthus sp. n. 53, genitalia of male imago, focus on dorsal side; 54, their subimaginal exuviae; 55, genitalia of male imago, focus on ventral side (holotype)." captionText-8="FIGURES 5659. Waynokiops pentacanthus sp. n. 56, precursors of subimaginal gonostyli starting to develop under larval cuticle; 57, subimaginal exuviae of gonostylus; 58, genitalia of male imago; 59, wing. Abbreviations: g1g3, segments of gonostylus; gd, gonoduct; m.IX-X, areas of anterior attachment of paired intersegmental muscles going to paraprocts base; m.s, median sterno-styligeral muscle." captionText-9="FIGURES 6063. Waynokiops spp., eggs. 6061, W. pentacanthus sp. n.; 6263, W. palifer sp. n." figureDoi-0="http://doi.org/10.5281/zenodo.7042808" figureDoi-1="http://doi.org/10.5281/zenodo.7042810" figureDoi-2="http://doi.org/10.5281/zenodo.7042812" figureDoi-3="http://doi.org/10.5281/zenodo.7042814" figureDoi-4="http://doi.org/10.5281/zenodo.7042816" figureDoi-5="http://doi.org/10.5281/zenodo.7042818" figureDoi-6="http://doi.org/10.5281/zenodo.7042820" figureDoi-7="http://doi.org/10.5281/zenodo.7042822" figureDoi-8="http://doi.org/10.5281/zenodo.7042826" figureDoi-9="http://doi.org/10.5281/zenodo.7042828" httpUri-0="https://zenodo.org/record/7042808/files/figure.png" httpUri-1="https://zenodo.org/record/7042810/files/figure.png" httpUri-2="https://zenodo.org/record/7042812/files/figure.png" httpUri-3="https://zenodo.org/record/7042814/files/figure.png" httpUri-4="https://zenodo.org/record/7042816/files/figure.png" httpUri-5="https://zenodo.org/record/7042818/files/figure.png" httpUri-6="https://zenodo.org/record/7042820/files/figure.png" httpUri-7="https://zenodo.org/record/7042822/files/figure.png" httpUri-8="https://zenodo.org/record/7042826/files/figure.png" httpUri-9="https://zenodo.org/record/7042828/files/figure.png" pageId="3" pageNumber="44">Figs 161</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="32D16576E876AA6C38CEF9460281C4B3" pageId="3" pageNumber="44" type="etymology">
<paragraph id="7A7436FDE876AA6C38CEF9460281C4B3" blockId="3.[151,1437,1724,1967]" pageId="3" pageNumber="44">
<emphasis id="48BFEAEFE876AA6C38CEF946014EC49F" bold="true" box="[151,286,1725,1751]" pageId="3" pageNumber="44">Etymology.</emphasis>
From.
<emphasis id="48BFEAEFE876AA6C3937F94601F7C49E" box="[366,423,1725,1750]" italics="true" pageId="3" pageNumber="44">πεντ-</emphasis>
(Gr.)—five-, and
<emphasis id="48BFEAEFE876AA6C3A36F9460293C49F" box="[623,707,1725,1751]" italics="true" pageId="3" pageNumber="44">ακανθα</emphasis>
(Gr.)—thorn; allusion to five especially long protrusions of equal shape on abdominal terga IIVII of larva (
<figureCitation id="E2F02A78E876AA6C3A28F91B0294C4B3" box="[625,708,1760,1787]" captionStart="FIGURES 110" captionStartId="4.[151,264,1841,1865]" captionTargetBox="[151,1436,179,1812]" captionTargetId="figure-18@4.[151,1436,179,1815]" captionTargetPageId="4" captionText="FIGURES 110. Waynokiops pentacanthus sp. n., larvae. 12, female larvae of last instar; 38, median part of abdominal terga II, VI, VII, VIII, IX and X; 9, median part of sternum VII; 10, thorax and abdomen of young larva." figureDoi="http://doi.org/10.5281/zenodo.7042808" httpUri="https://zenodo.org/record/7042808/files/figure.png" pageId="3" pageNumber="44">Fig. 10</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="32D16576E876AA6C389EF8FF0210C5E6" pageId="3" pageNumber="44" type="materials_examined">
<paragraph id="7A7436FDE876AA6C389EF8FF0210C5E6" blockId="3.[151,1437,1724,1967]" pageId="3" pageNumber="44">
<emphasis id="48BFEAEFE876AA6C389EF8FF01E2C556" bold="true" box="[199,434,1796,1822]" pageId="3" pageNumber="44">Material examined.</emphasis>
<materialsCitation id="CAA33CA0E876AA6C39E3F8FE0447C50A" ID-GBIF-Occurrence="3897738304" collectingDate="2015-02-09" collectorName="N. Kluge &amp; L. Sheyko." country="Thailand" county="Tan Jet Ton village" location="Pai" municipality="river Pai" pageId="3" pageNumber="44" specimenCount="1" specimenCount-male="1" typeStatus="holotype">
<emphasis id="48BFEAEFE876AA6C39E3F8FE0261C557" bold="true" box="[442,561,1797,1823]" pageId="3" pageNumber="44">
<typeStatus id="A570885FE876AA6C39E3F8FE027BC557" box="[442,555,1797,1823]" pageId="3" pageNumber="44" type="holotype">Holotype</typeStatus>
:
</emphasis>
L-S-I
<specimenCount id="6CCDFD74E876AA6C3A21F8FE02DBC557" box="[632,651,1797,1823]" pageId="3" pageNumber="44" type="male"></specimenCount>
{specimen [XIII](2) 2015},
<collectingCountry id="02DC766DE876AA6C3B8AF8FE0437C557" box="[979,1127,1797,1823]" name="Thailand" pageId="3" pageNumber="44">THAILAND</collectingCountry>
, prov. Mae-Hong-Son,
<location id="7F146026E876AA6C3D28F8FF05C7C556" LSID="urn:lsid:plazi:treatment:F26287EBE876AA6C38CEF9AB0210C5E6:7F146026E876AA6C3D28F8FF05C7C556" box="[1393,1431,1796,1822]" country="Thailand" county="Tan Jet Ton village" municipality="river Pai" name="Pai" pageId="3" pageNumber="44">Pai</location>
,
<collectingMunicipality id="9A10AC87E876AA6C38CEF8D300A9C50A" box="[151,249,1832,1858]" pageId="3" pageNumber="44">river Pai</collectingMunicipality>
upstream
<collectingCounty id="93154E71E876AA6C3928F8D2021DC50A" box="[369,589,1832,1859]" pageId="3" pageNumber="44">Tan Jet Ton village</collectingCounty>
,
<date id="0E75103DE876AA6C3A03F8D30294C50B" box="[602,708,1832,1859]" pageId="3" pageNumber="44" value="2015-02-09">
<collectingDate id="1E31E9D5E876AA6C3A03F8D30294C50B" box="[602,708,1832,1859]" pageId="3" pageNumber="44" value="2015-02-09">9.II.2015</collectingDate>
</date>
, coll.
<collectorName id="D73E532BE876AA6C3B52F8D20323C50A" box="[779,883,1832,1859]" pageId="3" pageNumber="44">N. Kluge</collectorName>
&amp;
<collectorName id="D73E532BE876AA6C3BC2F8D20447C50A" box="[923,1047,1832,1859]" pageId="3" pageNumber="44">L. Sheyko.</collectorName>
</materialsCitation>
<materialsCitation id="CAA33CA0E876AA6C3C79F8D2024DC52F" ID-GBIF-Occurrence="3897738301" collectingDate="2015-02-05" collectingDateMax="2015-02-14" collectingDateMin="2015-02-05" collectorName="N. Kluge &amp; L. Sheyko." country="Thailand" county="Tan Jet Ton village" location="Pai" municipality="river Pai" pageId="3" pageNumber="44" specimenCount="1" specimenCount-male="1" typeStatus="paratype">
<emphasis id="48BFEAEFE876AA6C3C79F8D204F1C50B" bold="true" box="[1056,1185,1833,1859]" pageId="3" pageNumber="44">
<typeStatus id="A570885FE876AA6C3C79F8D204CCC50B" box="[1056,1180,1833,1859]" pageId="3" pageNumber="44" type="paratype">Paratypes</typeStatus>
:
</emphasis>
the same locality and collectors,
<date id="0E75103DE876AA6C394AF8B701F8C52F" box="[275,424,1868,1895]" pageId="3" pageNumber="44" value="2015-02-05" valueMax="2015-02-14" valueMin="2015-02-05">
<collectingDate id="1E31E9D5E876AA6C394AF8B701F8C52F" box="[275,424,1868,1895]" pageId="3" pageNumber="44" value="2015-02-05" valueMax="2015-02-14" valueMin="2015-02-05">514.II.2015</collectingDate>
</date>
: 2 L-S-I
<specimenCount id="6CCDFD74E876AA6C3A53F8B6024DC52F" box="[522,541,1869,1895]" pageId="3" pageNumber="44" type="male"></specimenCount>
</materialsCitation>
, 1 L-S/I
<specimenCount id="6CCDFD74E876AA6C3A26F8B602C3C52F" box="[639,659,1869,1895]" pageId="3" pageNumber="44" type="male"></specimenCount>
, 3 L-S-I
<specimenCount id="6CCDFD74E876AA6C3AA3F8B60358C52F" box="[762,776,1869,1895]" pageId="3" pageNumber="44" type="female"></specimenCount>
,
<specimenCount id="6CCDFD74E876AA6C3B4CF8B703D1C52E" box="[789,897,1868,1894]" pageId="3" pageNumber="44" type="larva">16 larvae</specimenCount>
.
<emphasis id="48BFEAEFE876AA6C3BD5F8B705CCC52F" bold="true" box="[908,1436,1868,1895]" pageId="3" pageNumber="44">Additional material reported by D. Palatov:</emphasis>
<materialsCitation id="CAA33CA0E876AA6C38CEF88A026DC5E6" ID-GBIF-Occurrence="3897738303" collectingDate="2011-02-05" collectorName="D. Palatov" country="India" elevation="700" latitude="30.302452" location="Mandakini river" longLatPrecision="1" longitude="78.97886" municipality="Rudraprayag" pageId="3" pageNumber="44" specimenCount="1" specimenCount-larva="1" stateProvince="Uttarakhand">
<collectingCountry id="02DC766DE876AA6C38CEF88A00B9C5C3" box="[151,233,1905,1931]" name="India" pageId="3" pageNumber="44">INDIA</collectingCountry>
, state
<collectingRegion id="B80FF81FE876AA6C3969F88B01EAC5C2" box="[304,442,1904,1930]" country="India" name="Uttarakhand" pageId="3" pageNumber="44">Uttarakhand</collectingRegion>
,
<collectingMunicipality id="9A10AC87E876AA6C3991F88B020CC5C2" box="[456,604,1904,1930]" pageId="3" pageNumber="44">Rudraprayag</collectingMunicipality>
,
<location id="7F146026E876AA6C3A3EF88B034DC5C2" LSID="urn:lsid:plazi:treatment:F26287EBE876AA6C38CEF9AB0210C5E6:7F146026E876AA6C3A3EF88B034DC5C2" box="[615,797,1904,1930]" country="India" latitude="30.302452" longLatPrecision="1" longitude="78.97886" municipality="Rudraprayag" name="Mandakini river" pageId="3" pageNumber="44" stateProvince="Uttarakhand">Mandakini river</location>
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from mouth,
<geoCoordinate id="1FFF503AE876AA6C3BA1F88904C5C5C5" box="[1016,1173,1906,1933]" degrees="30" direction="north" minutes="18" orientation="latitude" pageId="3" pageNumber="44" precision="1" seconds="8.83" value="30.302452">30°188.83″N</geoCoordinate>
,
<geoCoordinate id="1FFF503AE876AA6C3CF8F8890519C5C5" box="[1185,1353,1906,1933]" degrees="78" direction="east" minutes="58" orientation="longitude" pageId="3" pageNumber="44" precision="1" seconds="43.89" value="78.97886">78°5843.89″E</geoCoordinate>
, h=
<quantity id="BD339B18E876AA6C3D28F88B00FFC5E7" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="3" pageNumber="44" unit="m" value="700.0">
<elevation id="F1E6D1CEE876AA6C3D28F88B00FFC5E7" metricMagnitude="2" metricUnit="m" metricValue="7.0" pageId="3" pageNumber="44" unit="m" value="700.0">700 m</elevation>
</quantity>
,
<date id="0E75103DE876AA6C38E2F86F0173C5E7" box="[187,291,1940,1967]" pageId="3" pageNumber="44" value="2011-02-05">
<collectingDate id="1E31E9D5E876AA6C38E2F86F0173C5E7" box="[187,291,1940,1967]" pageId="3" pageNumber="44" value="2011-02-05">5.II.2011</collectingDate>
</date>
, coll.
<collectorName id="D73E532BE876AA6C393EF86E01B1C5E6" box="[359,481,1940,1967]" pageId="3" pageNumber="44">D. Palatov</collectorName>
:
<specimenCount id="6CCDFD74E876AA6C39B5F86E026DC5E6" box="[492,573,1940,1967]" pageId="3" pageNumber="44" type="larva">1 larva</specimenCount>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>