215 lines
25 KiB
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215 lines
25 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zitteliana.97.115682" ID-Pensoft-Pub="2747-8106-97-53" ID-Pensoft-UUID="4A3318772B185D35B1E7EC9578EF4A47" ID-ZooBank="933EC356F21C45AF9CFA563E64D27953" ModsDocID="2747-8106-97-53" checkinTime="1702462880200" checkinUser="pensoft" docAuthor="Neubauer, Thomas A. & Wesselingh, Frank P." docDate="2023" docId="640D826023515A5AAC5C2C9A3E02F552" docLanguage="en" docName="Zitteliana 97: 53-88" docOrigin="Zitteliana 97" docPubDate="2023-12-12" docSource="http://dx.doi.org/10.3897/zitteliana.97.115682" docTitle="Theodoxus percarinatus Neubauer & Wesselingh 2023, comb. nov." docType="treatment" docVersion="1" id="4A3318772B185D35B1E7EC9578EF4A47" lastPageNumber="53" masterDocId="4A3318772B185D35B1E7EC9578EF4A47" masterDocTitle="The Early Pleistocene freshwater mollusks of the Denizli Basin (Turkey): a new long-lived lake fauna at the crossroads of Pontocaspian and Aegean-Anatolian realms" masterLastPageNumber="88" masterPageNumber="53" pageNumber="53" updateTime="1702462880200" updateUser="pensoft">
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<mods:titleInfo id="A378A72881CB909DC57D5CB49527E017">
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<mods:title id="3676E6315D531D9FE918B6D2054DA9A6">The Early Pleistocene freshwater mollusks of the Denizli Basin (Turkey): a new long-lived lake fauna at the crossroads of Pontocaspian and Aegean-Anatolian realms</mods:title>
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<mods:name id="E706446F5AEC648D37EE7B2B5380B8A8" type="personal">
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<mods:roleTerm id="85A684167DA914AC855E1D924C24C86C">Author</mods:roleTerm>
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<mods:namePart id="C0D28F2F61B601D31D93F3F3E75D98D4">Neubauer, Thomas A.</mods:namePart>
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<mods:nameIdentifier id="554F436662B1F5BD33808BAAD9372109" type="ORCID">https://orcid.org/0000-0002-1398-9941</mods:nameIdentifier>
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<mods:affiliation id="E8FEEAE692CFFBB0390E065EA7E13CEA">SNSB - Bavarian State Collection for Palaeontology and Geology, Richard-Wagner-Strasse 10, 80333 Munich, Germany & Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, Netherlands</mods:affiliation>
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<mods:nameIdentifier id="21A67E7108FF9424BE744355643E07B0" type="email">neubauer@snsb.de</mods:nameIdentifier>
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<mods:roleTerm id="8C5BF4A387E707B5852371938A012D13">Author</mods:roleTerm>
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<mods:namePart id="6686680404ED67BE4B0B6F168426564F">Wesselingh, Frank P.</mods:namePart>
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<mods:nameIdentifier id="2423DF55F910AE914176A6517D25481D" type="ORCID">https://orcid.org/0000-0003-3655-0701</mods:nameIdentifier>
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<mods:affiliation id="B59DD922F7126FC7328A6E503C953CCB">Naturalis Biodiversity Center, Darwinweg 2, 2333 CR Leiden, Netherlands & Department of Earth Sciences, Utrecht University, Princetonlaan 8 a, 3584 CB Utrecht, Netherlands</mods:affiliation>
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<mods:typeOfResource id="7BCC992BF45EE07539CBB98802D63014">text</mods:typeOfResource>
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<mods:title id="17E3D365755B4C2139E31F3ACFA0B4F8">Zitteliana</mods:title>
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<mods:part id="490A6FB62724999B041A30DCBE646F20">
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<mods:date id="AAED5EDD8E838076058CC5A96C7331B8">2023</mods:date>
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<mods:number id="E48F71DCE7C642129498EFC38309E559">2023-12-12</mods:number>
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<mods:detail id="0722AE225C32B30D6E7485857885E13C" type="volume">
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<mods:number id="B737991EA7C14574961425A7BAFAC339">97</mods:number>
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<mods:start id="2304C6865952B6B167B66CE5D62CCF97">53</mods:start>
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<mods:end id="A01E4BF9496CC2C1A1ADC2EF00F8BC74">88</mods:end>
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<mods:url id="391846D1114F380031DBD4E0E22E185C">http://dx.doi.org/10.3897/zitteliana.97.115682</mods:url>
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<mods:classification id="7CD7259224186FEC83DA5C7E7103CBBD">journal article</mods:classification>
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<mods:identifier id="04C076C56251FCB5A3E78E147FA95E97" type="DOI">http://dx.doi.org/10.3897/zitteliana.97.115682</mods:identifier>
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<treatment id="640D826023515A5AAC5C2C9A3E02F552" LSID="urn:lsid:plazi:treatment:640D826023515A5AAC5C2C9A3E02F552" httpUri="http://treatment.plazi.org/id/640D826023515A5AAC5C2C9A3E02F552" lastPageNumber="53" pageId="0" pageNumber="53">
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<subSubSection id="EFA2974117EB24DB04C535085FCCD912" pageId="0" pageNumber="53" type="nomenclature">
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<paragraph id="F48F4761F3F96A027412ECA692C021D1" pageId="0" pageNumber="53">
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<taxonomicName id="09BD27921496B162B0F5BA84775E1A3B" ID-CoL="56C83" LSID="640D8260-2351-5A5A-AC5C-2C9A3E02F552" authority="(Oppenheim, 1919)" authorityName="Neubauer & Wesselingh" authorityYear="2023" baseAuthorityName="Oppenheim" baseAuthorityYear="1919" class="Gastropoda" family="Neritidae" genus="Theodoxus" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Theodoxus percarinatus" order="Cycloneritida" pageId="0" pageNumber="53" phylum="Mollusca" rank="species" species="percarinatus" status="comb. nov.">Theodoxus percarinatus (Oppenheim, 1919)</taxonomicName>
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<taxonomicNameLabel id="4C30C9F805480EDBC14498FE00B47AD5" pageId="0" pageNumber="53">comb. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection id="8C038B8055A9332F5837B96C98D09ACB" pageId="0" pageNumber="53" type="description">
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<paragraph id="8F188DF42D008B5939D5EA1A48837CE4" pageId="0" pageNumber="53">
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<figureCitation id="AB6E256B45EEA87EB3BE3D0AE4B831EB" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Neritidae of the Kolankaya I fauna. A-D. Theodoxus percarinatus (Oppenheim, 1919), RGM 1310797, sample 2. E-H. T. percarinatus, SNSB-BSPG 2023 XII 2, sample 2. I-L. T. percarinatus, Neotype, RGM 962607, sample 2. M-P. T. percarinatus, SNSB-BSPG 2023 XII 1, sample 1. Q-T. T. cf. percarinatus, RGM 962606, sample 2. Scale bars: 1 mm." figureDoi="10.3897/zitteliana.97.115682.figure3" httpUri="https://binary.pensoft.net/fig/950740" pageId="0" pageNumber="53">Fig. 3A-T</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection id="BFF28BD719FBAFC4B13213ECB903CB14" pageId="0" pageNumber="53" type="reference_group">
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<paragraph id="240BB36673F182C5F8F65B099A9C6FA3" pageId="0" pageNumber="53">
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<taxonomicName id="3C39D718E2742DB1F7940476DD8DB230" authorityName="Neubauer & Wesselingh" authorityYear="2023" class="Gastropoda" family="Neritidae" genus="Neritina" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Neritina percarinata" order="Cycloneritida" pageId="0" pageNumber="53" phylum="Mollusca" rank="species" species="percarinata" status="comb. nov.">*1919 Neritina percarinata</taxonomicName>
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sp. nov. - Oppenheim: 128-129, pl. 9, fig. 7.
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</paragraph>
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</subSubSection>
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<subSubSection id="F475E31E3492B2ECA0046DD389549F29" pageId="0" pageNumber="53" type="materials_examined">
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<paragraph id="C10104E780A0F38AB39502F00417FF31" pageId="0" pageNumber="53">Material.</paragraph>
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<paragraph id="FAAE3F3E655F8833B9BC80F31C15673A" pageId="0" pageNumber="53">
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Sample 1: c.
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<specimenCount id="473AD3B48AB9294B8841837A5201FBAF" type="generic">500 specimens</specimenCount>
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(RGM 1310838) and
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<specimenCount id="069618799F5C10546B78D7C13E543721" type="generic">1 specimen</specimenCount>
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(SNSB-BSPG
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<date id="9D9CF0ACBDCEEAA2F812F6589AF1F76C" value="2023-12-01">2023 XII 1</date>
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); sample 2:
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<specimenCount id="BC21AC06297316AC9E2EA9E947B2F4D5" type="generic">1 specimen</specimenCount>
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tentatively attributed to the species (RGM 962606),
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<specimenCount id="0C944C0C68D642C3E702438EDDA8AA7B" type="generic">1 specimen</specimenCount>
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(RGM 962607),
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<specimenCount id="1A300881B24F72B38631B416D21D0EA3" type="generic">1 specimen</specimenCount>
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(RGM 1310797),
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<specimenCount id="096E4C0F408FE5FA50DC788399FF7AE5" type="generic">123 specimens</specimenCount>
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(RGM 1310839), 2 fragments and fragmented opercula (RGM 1310840),
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<specimenCount id="F1869A856B1C636461DFA017C7A84681" type="generic">1 specimen</specimenCount>
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(SNSB-BSPG
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<date id="41AFE1CA7ADC075742E21CD9E41D3CE8" value="2023-12-02">2023 XII 2</date>
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); sample 3:
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<specimenCount id="439C80133A9E8F5DFEC183D483208DD7" type="generic">53 specimens</specimenCount>
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(RGM 1310841).
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</paragraph>
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</subSubSection>
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<subSubSection id="7CB79CB939AB0CB1B20E09AA6D5ECED0" pageId="0" pageNumber="53" type="neotype">
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<paragraph id="54F7592A929A6C8688949C9618F6824A" pageId="0" pageNumber="53">Neotype.</paragraph>
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<paragraph id="6633A3A8FA3A9C81703622DFB76C5EAE" pageId="0" pageNumber="53">
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<normalizedToken id="FB3F9CE1406EE4AF9F00B9B18089CCED" originalValue="Oppenheim’s">Oppenheim's</normalizedToken>
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type material could not be found despite considerable efforts and is presumably lost (see Material and Methods section). To ensure nomenclatural stability and link the name to a type specimen, we herewith designate a neotype. The original type locality includes two localities, Giralan near Pamukkale and Bozalan near Buldan ("Abstieg nach Giralan.-Bosalan bei Bulladan.") in the Denizli Basin (Fig.
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<figureCitation id="F8FF42E1FD3CC0712EDA24B51173703C" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Geographical and geological overview of the study area in SW Turkey (modified from Alcicek et al. 2015)." figureDoi="10.3897/zitteliana.97.115682.figure1" httpUri="https://binary.pensoft.net/fig/950738" pageId="0" pageNumber="53">1</figureCitation>
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). No material from those localities is available to us, but considering the geographic vicinity of the newly collected material as well as the distinct morphological features, there can be no doubt that our material represents the same species as the one described by
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<bibRefCitation id="4CAE5AFA3179E99FE826CA7B34FF086C" author="Oppenheim, P" journalOrPublisher="Zeitschrift der Deutschen Geologischen Gesellschaft" pageId="0" pageNumber="53" pagination="1 - 210" publicationUrl="https://www.biodiversitylibrary.org/page/43778447" refId="B102" refString="Oppenheim, P, 1919. Das Neogen in Kleinasien. Zeitschrift der Deutschen Geologischen Gesellschaft 70: 1 - 210, https://www.biodiversitylibrary.org/page/43778447" title="Das Neogen in Kleinasien." url="https://www.biodiversitylibrary.org/page/43778447" volume="70" year="1919">Oppenheim (1919)</bibRefCitation>
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. We select as neotype the specimen illustrated on Fig.
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<figureCitation id="4AD36C9B6A7BCCECD917E88304B27657" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Neritidae of the Kolankaya I fauna. A-D. Theodoxus percarinatus (Oppenheim, 1919), RGM 1310797, sample 2. E-H. T. percarinatus, SNSB-BSPG 2023 XII 2, sample 2. I-L. T. percarinatus, Neotype, RGM 962607, sample 2. M-P. T. percarinatus, SNSB-BSPG 2023 XII 1, sample 1. Q-T. T. cf. percarinatus, RGM 962606, sample 2. Scale bars: 1 mm." figureDoi="10.3897/zitteliana.97.115682.figure3" httpUri="https://binary.pensoft.net/fig/950740" pageId="0" pageNumber="53">3I-L</figureCitation>
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, which comes from sample 2 (RGM 962607).
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</paragraph>
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<caption id="2762FECB5126DCDD81848E2A4EA237A7" doi="10.3897/zitteliana.97.115682.figure3" httpUri="https://binary.pensoft.net/fig/950740" pageId="0" pageNumber="53" start="Figure 3" startId="F3">
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<paragraph id="793844F40C6E09B4C3FD38428E9C2ACD" pageId="0" pageNumber="53">
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<emphasis id="9DFC64E65E2965B548C3CFAD0016C804" bold="true" pageId="0" pageNumber="53">Figure 3.</emphasis>
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<taxonomicName id="C7C6FE693DE0F476C59FA908C93C6582" class="Gastropoda" family="Neritidae" higherTaxonomySource="GBIF" kingdom="Animalia" lsidName="" order="Cycloneritida" pageId="0" pageNumber="53" phylum="Mollusca" rank="family">Neritidae</taxonomicName>
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of the Kolankaya I fauna.
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<emphasis id="5A6D547F136067817AC8237B1507D92F" bold="true" pageId="0" pageNumber="53">A-D.</emphasis>
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<taxonomicName id="6F1211F70B9DCF89EEF8E5273DC7A7B2" authorityName="Neubauer & Wesselingh" authorityYear="2023" baseAuthorityName="Oppenheim" baseAuthorityYear="1919" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" lsidName="Theodoxus percarinatus" order="Cycloneritida" pageId="0" pageNumber="53" phylum="Mollusca" rank="species" species="percarinatus">
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<emphasis id="1A59B994107D4D927A230D4261DF206B" italics="true" pageId="0" pageNumber="53">Theodoxus percarinatus</emphasis>
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</taxonomicName>
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(Oppenheim, 1919), RGM 1310797, sample 2.
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<emphasis id="A069044B307872E62DDF976EBC28E9C5" bold="true" pageId="0" pageNumber="53">E-H</emphasis>
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.
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<taxonomicName id="4A3C02037BDBCC747F9BEC35D4BC2612" lsidName="T. percarinatus" pageId="0" pageNumber="53" rank="species" species="percarinatus">
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<emphasis id="B75612E5B619EDB78824E46CB711E9C2" italics="true" pageId="0" pageNumber="53">T. percarinatus</emphasis>
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</taxonomicName>
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, SNSB-BSPG 2023 XII 2, sample 2.
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<emphasis id="016C6BE2E6B991E6C0D020DF776CDDC5" bold="true" pageId="0" pageNumber="53">I-L.</emphasis>
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<taxonomicName id="EDAA566E84281F514C495CF5AF70D21C" lsidName="T. percarinatus" pageId="0" pageNumber="53" rank="species" species="percarinatus">
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<emphasis id="D1030911FF8C3450126610EEFE20D136" italics="true" pageId="0" pageNumber="53">T. percarinatus</emphasis>
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</taxonomicName>
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, Neotype, RGM 962607, sample 2.
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<emphasis id="0FA95EE314968D9699A18801CFA01F62" bold="true" pageId="0" pageNumber="53">M-P.</emphasis>
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<taxonomicName id="1A02A14185A3AAD5FA7AB543B5EFA578" lsidName="T. percarinatus" pageId="0" pageNumber="53" rank="species" species="percarinatus">
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<emphasis id="24C30F488CC9DD8CC909D9C6657A6B94" italics="true" pageId="0" pageNumber="53">T. percarinatus</emphasis>
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</taxonomicName>
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, SNSB-BSPG 2023 XII 1, sample 1.
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<emphasis id="2AB976F0176EEB36D2D4783DC9580EE3" bold="true" pageId="0" pageNumber="53">Q-T.</emphasis>
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<taxonomicName id="813103A3BC1606D4F2E3D455D401C12B" lsidName="T. percarinatus" pageId="0" pageNumber="53" rank="species" species="percarinatus">T. cf. percarinatus</taxonomicName>
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, RGM 962606, sample 2. Scale bars: 1 mm.
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</paragraph>
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</caption>
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</subSubSection>
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<subSubSection id="F071944059BE0D6BFE0F2A765095073A" pageId="0" pageNumber="53" type="description">
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<paragraph id="365707141A2457E51290A6CFAF48C3A5" pageId="0" pageNumber="53">Description.</paragraph>
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<paragraph id="77154097262A09EBF674F179B3626A27" pageId="0" pageNumber="53">Globular neritid with short, flattened spire and up to 3.3 whorls. Characteristic keel is observed in part of the material. In well preserved, keeled specimens narrow, distinct keel develops early in ontogeny. In first whorl it aligns with suture, but as shell growth becomes slightly more abapically oriented, keel emerges more clearly and separates whorl top and whorl flank. Expression and extent of keel varies, it may appear and disappear within 0.2 whorls or range until last whorl; however, no specimen is found with keel covering entire shell. Strength of keel very variable, ranging from robust and well delimited to almost obsolete. In much of the material the keel is worn. Aperture broadly semicircular with broad, smooth callus that is weakly thickened especially in its center (see lateral views). Coloration variable, including fully black, white with dark stripes, irregular mix of dark and white, white blotches arranged in spiral bands or entirely white.</paragraph>
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</subSubSection>
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<subSubSection id="9735A8F1FDAB380CFE38A398CC88B5EF" pageId="0" pageNumber="53" type="description">
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<paragraph id="B0AD947B74904528268BAFB17C6624BB" pageId="0" pageNumber="53">Dimensions.</paragraph>
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<paragraph id="11955317BFD780C1D310089AB9B57702" pageId="0" pageNumber="53">
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6.64
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<normalizedToken id="7A290B4955F9E20ACBC829B9D6AAF5B8" originalValue="×">x</normalizedToken>
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7.76 mm (neotype, RGM 962607; Fig.
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<figureCitation id="F47B9091B475096D4324CE70FC803B96" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Geographical and geological overview of the study area in SW Turkey (modified from Alcicek et al. 2015)." figureDoi="10.3897/zitteliana.97.115682.figure1" httpUri="https://binary.pensoft.net/fig/950738" pageId="0" pageNumber="53">1I-L</figureCitation>
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), 10.66
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<normalizedToken id="57F42B404B1D05F3AF55C79D839C6A91" originalValue="×">x</normalizedToken>
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10.51 mm (RGM 1310797; Fig.
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<figureCitation id="2C661F0F10B0EFAB3E6359B66CF48742" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Geographical and geological overview of the study area in SW Turkey (modified from Alcicek et al. 2015)." figureDoi="10.3897/zitteliana.97.115682.figure1" httpUri="https://binary.pensoft.net/fig/950738" pageId="0" pageNumber="53">1A-D</figureCitation>
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), 6.72
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<normalizedToken id="AE3CA27ADB1D7368421027E3DFA46AD0" originalValue="×">x</normalizedToken>
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7.24 mm (SNSB-BSPG 2023 XII 1; Fig.
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<figureCitation id="67F394E753E9FFD437D69125CD45C71E" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Geographical and geological overview of the study area in SW Turkey (modified from Alcicek et al. 2015)." figureDoi="10.3897/zitteliana.97.115682.figure1" httpUri="https://binary.pensoft.net/fig/950738" pageId="0" pageNumber="53">1M-P</figureCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection id="000F3F6D7BF00C024C1752CC7E70F2AF" pageId="0" pageNumber="53" type="remarks">
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<paragraph id="803FCC8AFA5609060BDF5D836EE940F9" pageId="0" pageNumber="53">Remarks.</paragraph>
|
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<paragraph id="377CB5275768D3E2F9AE13CDD97BEA1B" pageId="0" pageNumber="53">
|
||
The species has not been treated in the literature since the original description. It was neither included in the Fossilium Catalogus by Wenz (1930), perhaps because he considered this form not to be a freshwater taxon, nor was it listed by
|
||
<bibRefCitation id="2A12E2D743B5FF58FE9BA5B479E16D30" author="Taner, G" journalOrPublisher="Bulletin of Mineral Research and Exploration Institute of Turkey (MTA)" pageId="0" pageNumber="53" pagination="89 - 126" refId="B148" refString="Taner, G, 1974a. Denizli boelgesi Neojeninin paleontolojik ve stratigrafik etuedue. Bulletin of Mineral Research and Exploration Institute of Turkey (MTA) 82: 89 - 126" title="Denizli boelgesi Neojeninin paleontolojik ve stratigrafik etuedue." volume="82" year="1974 a">Taner (1974a</bibRefCitation>
|
||
,
|
||
<bibRefCitation id="AE0243294C539B79DF6CAB4343ED4877" author="Taner, G" journalOrPublisher="Bulletin of Mineral Research and Exploration Institute of Turkey (MTA)" pageId="0" pageNumber="53" pagination="45 - 66" refId="B150" refString="Taner, G, 1975. Denizli boelgesi Neojeninin paleontolojik ve stratigrafik etuedue. Bulletin of Mineral Research and Exploration Institute of Turkey (MTA) 85: 45 - 66" title="Denizli boelgesi Neojeninin paleontolojik ve stratigrafik etuedue." volume="85" year="1975">1975</bibRefCitation>
|
||
). The combination with the genus
|
||
<taxonomicName id="870DB93120A05F209C81A872B0B25550" authorityName="Montfort" authorityYear="1810" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" lsidName="Theodoxus" order="Cycloneritida" pageId="0" pageNumber="53" phylum="Mollusca" rank="genus">
|
||
<emphasis id="D4D9FB9C01418352E88BFD8BB9380FFA" italics="true" pageId="0" pageNumber="53">Theodoxus</emphasis>
|
||
</taxonomicName>
|
||
appears for the first time in
|
||
<bibRefCitation id="DCB2EEF7A5310956F3549DE99287556A" DOI="https://doi.org/10.1016/j.palaeo.2010.03.002" author="Alcicek, H" journalOrPublisher="Palaeogeography, Palaeoclimatology, Palaeoecology" pageId="0" pageNumber="53" pagination="297 - 318" refId="B2" refString="Alcicek, H, 2010. Stratigraphic correlation of the Neogene basins in southwestern Anatolia: Regional palaeogeographical, palaeoclimatic and tectonic implications. Palaeogeography, Palaeoclimatology, Palaeoecology 291: 297 - 318, DOI: https://doi.org/10.1016/j.palaeo.2010.03.002" title="Stratigraphic correlation of the Neogene basins in southwestern Anatolia: Regional palaeogeographical, palaeoclimatic and tectonic implications." url="https://doi.org/10.1016/j.palaeo.2010.03.002" volume="291" year="2010">
|
||
<normalizedToken id="0116596A972011B268A4BEF82B5D274C" originalValue="Alçiçek">Alcicek</normalizedToken>
|
||
(2010)
|
||
</bibRefCitation>
|
||
, but without discussion. Here, we formalize the classification and re-describe the species.
|
||
</paragraph>
|
||
<paragraph id="1AC9C9DEBA5AF13ECFF80810E0F86E06" pageId="0" pageNumber="53">
|
||
The diagnostic character of this species is its name-giving thin keel at the transition between whorl top and flank, which distinguishes the species from most others. A similarly keeled species is
|
||
<taxonomicName id="7973B4BF19231166698817B9672CDB71" baseAuthorityName="T.Fuchs" baseAuthorityYear="1877" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" lsidName="Theodoxus carinatus" order="Cycloneritida" pageId="0" pageNumber="53" phylum="Mollusca" rank="species" species="carinatus">
|
||
<emphasis id="6204088AEE096DA1C2DBBDA1A3F8BA7F" italics="true" pageId="0" pageNumber="53">Theodoxus carinatus</emphasis>
|
||
</taxonomicName>
|
||
(Fuchs, 1877) from the Lower Pleistocene (Calabrian) of Livonates (mainland Greece). It differs in the flattened whorl flanks, the weakly raised spire, the broader keel and the color pattern consisting of wavy axial lines.
|
||
</paragraph>
|
||
<paragraph id="65FD98482B1D5B186254442973009BCF" pageId="0" pageNumber="53">
|
||
In addition to the majority of keeled specimens of
|
||
<taxonomicName id="56ABF1934023A2B89E3F024961255208" authorityName="Neubauer & Wesselingh" authorityYear="2023" baseAuthorityName="Oppenheim" baseAuthorityYear="1919" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" lsidName="Theodoxus percarinatus" order="Cycloneritida" pageId="0" pageNumber="53" phylum="Mollusca" rank="species" species="percarinatus">
|
||
<emphasis id="8E7D0B412FE80247F4F51C0EC9B04F19" italics="true" pageId="0" pageNumber="53">Theodoxus percarinatus</emphasis>
|
||
</taxonomicName>
|
||
, shells occur that lack the keel entirely and have a slightly more elevated spire (Fig.
|
||
<figureCitation id="B2250682EABB6D7487DCD1A43E35DE69" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Neritidae of the Kolankaya I fauna. A-D. Theodoxus percarinatus (Oppenheim, 1919), RGM 1310797, sample 2. E-H. T. percarinatus, SNSB-BSPG 2023 XII 2, sample 2. I-L. T. percarinatus, Neotype, RGM 962607, sample 2. M-P. T. percarinatus, SNSB-BSPG 2023 XII 1, sample 1. Q-T. T. cf. percarinatus, RGM 962606, sample 2. Scale bars: 1 mm." figureDoi="10.3897/zitteliana.97.115682.figure3" httpUri="https://binary.pensoft.net/fig/950740" pageId="0" pageNumber="53">3Q-T</figureCitation>
|
||
). In terms of size, general shape, and color patterns these specimens fall well within the range of
|
||
<taxonomicName id="EA2B23CD714AD6C6C862F4480544C17A" lsidName="T. percarinatus" pageId="0" pageNumber="53" rank="species" species="percarinatus">
|
||
<emphasis id="A9546D3CB6D253B4048CA19109A8AC8A" italics="true" pageId="0" pageNumber="53">T. percarinatus</emphasis>
|
||
</taxonomicName>
|
||
. Many
|
||
<taxonomicName id="BE8FDAC1D1665E29EF164EA593E6909A" authorityName="Montfort" authorityYear="1810" class="Gastropoda" family="Neritidae" genus="Theodoxus" kingdom="Animalia" lsidName="Theodoxus" order="Cycloneritida" pageId="0" pageNumber="53" phylum="Mollusca" rank="genus">
|
||
<emphasis id="BC2E4F52210A2D4991F0D106575D036A" italics="true" pageId="0" pageNumber="53">Theodoxus</emphasis>
|
||
</taxonomicName>
|
||
species are known to have highly variable shells (
|
||
<bibRefCitation id="B1FE2CF8C9E2A809E378102681652349" DOI="https://doi.org/10.3897/zse.96.48312" author="Sands, AF" journalOrPublisher="Zoosystematics and Evolution" pageId="0" pageNumber="53" pagination="25 - 66" refId="B125" refString="Sands, AF, Gloeer, P, Guerlek, ME, Albrecht, C, Neubauer, TA, 2020. A revision of the extant species of Theodoxus (Gastropoda, Neritidae) in Asia, with the description of three new species. Zoosystematics and Evolution 96: 25 - 66, DOI: https://doi.org/10.3897/zse.96.48312" title="A revision of the extant species of Theodoxus (Gastropoda, Neritidae) in Asia, with the description of three new species." url="https://doi.org/10.3897/zse.96.48312" volume="96" year="2020">Sands et al. 2020</bibRefCitation>
|
||
), which complicates a taxonomic decision about the status of these specimens. Apparent intermediate specimens occur in our material, but the worn nature of much of the material, the small numbers of specimens, and the general variability of the genus preclude confirmation whether it concerns untypically unkeeled representatives of
|
||
<taxonomicName id="82C035831BE2ADF10E5B224B18C017DA" lsidName="T. percarinatus" pageId="0" pageNumber="53" rank="species" species="percarinatus">
|
||
<emphasis id="677289F044B2988AC74C02F94BA632FA" italics="true" pageId="0" pageNumber="53">T. percarinatus</emphasis>
|
||
</taxonomicName>
|
||
or constitute a separate, rare, and potentially new species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="2A7986B76379F4B641D9E06F8B06315E" pageId="0" pageNumber="53" type="distribution">
|
||
<paragraph id="8D8AEA893BEE36FD309AE2F9BC0513CD" pageId="0" pageNumber="53">Distribution.</paragraph>
|
||
<paragraph id="0B753662984C497D6DC4C032F66DB0E5" pageId="0" pageNumber="53">Only known from the Denizli Basin.</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |