treatments-xml/data/21/6F/4E/216F4E11FF887302FD3AFD6719E4FEF2.xml

<document ID-DOI="http://doi.org/10.5281/zenodo.3671810" ID-GBIF-Dataset="f033f07a-e903-4e39-81d4-2e9a157ac80d" ID-GBIF-Taxon="162136341" ID-Zenodo-Dep="3671810" checkinTime="1581620919828" checkinUser="plazi" docAuthor="Lücking, Robert, Moncada, Bibiana, Habibe, María Cristina Martínez-, Salgado- Negret, Beatriz E., Celis, Marcela, Zamora, Oscar Rojas-, Rodríguez- M., Gina M., Brokamp, Grischa &amp; Borsch, Thomas" docDate="2019" docId="216F4E11FF887302FD3AFD6719E4FEF2" docLanguage="en" docName="cal-41-01-194.pdf" docOrigin="Caldasia 41 (1)" docStyle="DocumentStyle{}" docTitle="Graphis mokanarum Lucking, Moncada &amp; M. C. Martinez 2019" docType="treatment" docVersion="5" lastPageId="14" lastPageNumber="208" masterDocId="DD563669FF85730CFFBAFFD71A4BFFF8" masterDocTitle="Lichen diversity in colombian caribbean dry forest remnants" masterLastPageNumber="214" masterPageNumber="94" pageId="13" pageNumber="207" updateTime="1636508073100" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Lichen diversity in colombian caribbean dry forest remnants</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Lücking, Robert</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Moncada, Bibiana</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Habibe, María Cristina Martínez-</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Salgado- Negret, Beatriz E.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Celis, Marcela</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Zamora, Oscar Rojas-</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Rodríguez- M., Gina M.</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Brokamp, Grischa</mods:namePart>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Borsch, Thomas</mods:namePart>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>Caldasia</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2019</mods:date>
<mods:detail type="pubDate">
<mods:number>2019-06-30</mods:number>
</mods:detail>
<mods:detail type="volume">
<mods:number>41</mods:number>
</mods:detail>
<mods:detail type="issue">
<mods:number>1</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>94</mods:start>
<mods:end>214</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">https://dx.doi.org/10.15446/caldasia.v41n1.71060</mods:identifier>
<mods:identifier type="GBIF-Dataset">f033f07a-e903-4e39-81d4-2e9a157ac80d</mods:identifier>
<mods:identifier type="Zenodo-Dep">3666769</mods:identifier>
</mods:mods>
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3671810" ID-GBIF-Taxon="162136341" ID-Zenodo-Dep="3671810" LSID="urn:lsid:plazi:treatment:216F4E11FF887302FD3AFD6719E4FEF2" httpUri="http://treatment.plazi.org/id/216F4E11FF887302FD3AFD6719E4FEF2" lastPageId="14" lastPageNumber="208" pageId="13" pageNumber="207">
<subSubSection pageId="13" pageNumber="207" type="nomenclature">
<paragraph blockId="13.[640,1111,688,776]" pageId="13" pageNumber="207">
<emphasis bold="true" pageId="13" pageNumber="207">
<heading bold="true" box="[640,1111,688,713]" centered="true" fontSize="10" level="4" pageId="13" pageNumber="207" reason="0">
<taxonomicName authority="Lucking, Moncada" authorityName="Lucking, Moncada &amp; M. C. Martinez" authorityYear="2019" box="[640,1111,688,713]" class="Gastropoda" family="Cimidae" genus="Graphis" kingdom="Animalia" order="Ostropales" pageId="13" pageNumber="207" phylum="Mollusca" rank="species" species="mokanarum" status="sp. nov.">
<emphasis bold="true" box="[640,877,688,712]" italics="true" pageId="13" pageNumber="207">Graphis mokanarum</emphasis>
Lücking, Moncada
</taxonomicName>
</heading>
&amp; M.C. Martínez sp. nov.
</emphasis>
</paragraph>
<paragraph blockId="13.[640,1111,688,776]" pageId="13" pageNumber="207">
Mycobank 
<accessionNumber pageId="13" pageNumber="207">
<collectionCode box="[1070,1111,721,744]" country="Portugal" name="Universidade de Lisboa, Museu Bocage" pageId="13" pageNumber="207">MB</collectionCode>
828116
</accessionNumber>
</paragraph>
</subSubSection>
<subSubSection box="[726,824,752,776]" pageId="13" pageNumber="207" type="description">
<paragraph blockId="13.[640,1111,688,776]" box="[726,824,752,776]" pageId="13" pageNumber="207">
(
<figureCitation box="[733,811,752,776]" captionStart="Figure 3" captionStartId="12.[136,204,1591,1613]" captionTargetBox="[139,1109,843,1570]" captionTargetId="figure@12.[139,1109,843,1570]" captionTargetPageId="12" captionText="Figure 3. Species of lichens new to science. a. Fissurina linoana (42977b, holotype); b. Graphis lurizana (42922, holotype); c. Graphis mokanarum (42930, holotype); d. Phaeographis galeanoae (42981, holotype). Thalli with ascomata. Scale = 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3666775" httpUri="https://zenodo.org/record/3666775/files/figure.png" pageId="13" pageNumber="207">Fig. 3c</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="207" type="diagnosis">
<paragraph blockId="13.[640,1112,816,1064]" pageId="13" pageNumber="207">
<emphasis bold="true" box="[640,757,816,840]" pageId="13" pageNumber="207">Diagnosis.</emphasis>
Differing from 
<taxonomicName class="Gastropoda" family="Cimidae" genus="Graphis" kingdom="Animalia" order="Ostropales" pageId="13" pageNumber="207" phylum="Mollusca" rank="species" species="microsperma">
<emphasis italics="true" pageId="13" pageNumber="207">Graphis microsperma</emphasis>
</taxonomicName>
in the larger ascospores and the stictic acid chemistry.
</paragraph>
</subSubSection>
<subSubSection pageId="13" pageNumber="207" type="materials_examined">
<paragraph blockId="13.[640,1112,816,1064]" pageId="13" pageNumber="207">
<emphasis bold="true" box="[889,952,881,905]" pageId="13" pageNumber="207">Type.</emphasis>
<materialsCitation ID-GBIF-Occurrence="2565375026" collectingDate="2015-12-01" collectionCode="DMI, R, UNO" collectorName="Lucking, B" country="Colombia" county="Usiacuri" elevation="150" latitude="10.739445" location="Luriza" longLatPrecision="21" longitude="-75.02334" municipality="Distrito Regional de Manejo Integrado" pageId="13" pageNumber="207" specimenCount="1" stateProvince="Atlantico" typeStatus="holotype">
<collectingCountry box="[962,1108,880,904]" name="Colombia" pageId="13" pageNumber="207">COLOMBIA</collectingCountry>
. 
<emphasis bold="true" box="[640,754,912,936]" pageId="13" pageNumber="207">
<collectingRegion box="[640,749,912,936]" country="Colombia" name="Atlantico" pageId="13" pageNumber="207">Atlántico</collectingRegion>
:
</emphasis>
<collectingCounty box="[768,861,912,936]" pageId="13" pageNumber="207">Usiacurí</collectingCounty>
, 
<collectingMunicipality pageId="13" pageNumber="207">Distrito Regional de Manejo Integrado</collectingMunicipality>
(
<collectionCode box="[847,902,944,968]" pageId="13" pageNumber="207">DMI</collectionCode>
) 
<location LSID="urn:lsid:plazi:treatment:216F4E11FF887302FD3AFD6719E4FEF2:AC19A9DCFF887301FC2FFC671995FC30" box="[917,990,944,968]" country="Colombia" county="Usiacuri" latitude="10.739445" longLatPrecision="21" longitude="-75.02334" municipality="Distrito Regional de Manejo Integrado" name="Luriza" pageId="13" pageNumber="207" stateProvince="Atlantico">Luriza</location>
; 
<geoCoordinate degrees="10" direction="north" minutes="44" orientation="latitude" pageId="13" pageNumber="207" precision="15" seconds="22" value="10.739445">10º44’22” N</geoCoordinate>
, 
<geoCoordinate box="[676,821,976,1000]" degrees="75" direction="west" minutes="01" orientation="longitude" pageId="13" pageNumber="207" precision="15" seconds="24" value="-75.02334">75º01’24” W</geoCoordinate>
, 
<quantity box="[835,963,976,1000]" metricMagnitude="2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="13" pageNumber="207" unit="m" value="150.0" valueMax="200.0" valueMin="100.0">
<elevation box="[835,963,976,1000]" metricMagnitude="2" metricUnit="m" metricValue="1.5" metricValueMax="2.0" metricValueMin="1.0" pageId="13" pageNumber="207" unit="m" value="150.0" valueMax="200.0" valueMin="100.0">100–200 m</elevation>
</quantity>
; 
<date pageId="13" pageNumber="207" value="2015-12-01">
<collectingDate pageId="13" pageNumber="207" value="2015-12-01">1 December 2015</collectingDate>
</date>
; 
<emphasis box="[709,1112,1008,1033]" italics="true" pageId="13" pageNumber="207">
<collectionCode box="[709,728,1009,1032]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="13" pageNumber="207">R</collectionCode>
. 
<collectorName box="[740,860,1008,1032]" pageId="13" pageNumber="207">Lücking, B</collectorName>
. Moncada et al. 42930
</emphasis>
(
<collectionCode box="[647,709,1040,1064]" country="0" httpUri="http://grbio.org/cool/fyxp-7ves" name="University of Nebraska at Omaha" pageId="13" pageNumber="207">UNO</collectionCode>
, 
<typeStatus box="[720,817,1040,1064]" pageId="13" pageNumber="207">holotype</typeStatus>
; B, isotype)
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection lastPageId="14" lastPageNumber="208" pageId="13" pageNumber="207" type="description">
<paragraph blockId="13.[640,1112,1104,1672]" lastBlockId="14.[136,608,176,425]" lastPageId="14" lastPageNumber="208" pageId="13" pageNumber="207">
<emphasis bold="true" box="[640,779,1104,1128]" pageId="13" pageNumber="207">Description.</emphasis>
Thallus corticolous, epiperidermal, up to 
<quantity box="[928,1016,1136,1160]" metricMagnitude="-1" metricUnit="m" metricValue="1.0" pageId="13" pageNumber="207" unit="cm" value="10.0">10 cm</quantity>
diam., continuous, light greenish grey, uneven to minutely verruculose, opaque; prothallus not observed. Thallus in section 
<quantity box="[938,1046,1232,1257]" metricMagnitude="-2" metricUnit="m" metricValue="4.5" metricValueMax="5.0" metricValueMin="4.0" pageId="13" pageNumber="207" unit="mm" value="45.0" valueMax="50.0" valueMin="40.0">40–50 µm</quantity>
thick, with distinct. prosoplectenchymatous cortex and distinct photobiont layer, blurry through incrustation with numerous small crystals. Ascomata lirellate, irregularly branched, erumpent, 
<quantity box="[769,871,1392,1416]" metricMagnitude="-3" metricUnit="m" metricValue="3.0" metricValueMax="5.0" metricValueMin="1.0" pageId="13" pageNumber="207" unit="mm" value="3.0" valueMax="5.0" valueMin="1.0">1–5 mm</quantity>
long, 
<quantity box="[968,1111,1392,1416]" metricMagnitude="-4" metricUnit="m" metricValue="3.5" metricValueMax="4.0" metricValueMin="3.0" pageId="13" pageNumber="207" unit="mm" value="0.35" valueMax="0.4" valueMin="0.3">0.3–0.4 mm</quantity>
broad, 
<quantity box="[719,865,1424,1448]" metricMagnitude="-4" metricUnit="m" metricValue="1.75" metricValueMax="2.0" metricValueMin="1.5" pageId="13" pageNumber="207" unit="mm" value="0.175" valueMax="0.2" valueMin="0.15">0.15–0.2 mm</quantity>
high; disc concealed; labia entire, apically exposed, whitish, with thick, lateral to almost complete, uneven to verruculose thalline margin. Excipulum uncarbonized, 
<quantity box="[808,928,1552,1577]" metricMagnitude="-2" metricUnit="m" metricValue="4.0" metricValueMax="5.0" metricValueMin="3.0" pageId="13" pageNumber="207" unit="mm" value="40.0" valueMax="50.0" valueMin="30.0">30–50 µm</quantity>
broad, orangebrown; hypothecium 
<quantity box="[904,1032,1584,1609]" metricMagnitude="-2" metricUnit="m" metricValue="1.25" metricValueMax="1.5" metricValueMin="1.0" pageId="13" pageNumber="207" unit="mm" value="12.5" valueMax="15.0" valueMin="10.0">10–15 µm</quantity>
high, hyaline to yellowish. Hymenium 
<quantity box="[990,1111,1616,1641]" metricMagnitude="-2" metricUnit="m" metricValue="9.0" metricValueMax="10.0" metricValueMin="8.0" pageId="13" pageNumber="207" unit="mm" value="90.0" valueMax="100.0" valueMin="80.0">80–100 µm</quantity>
high, clear; paraphyses unbranched, apically smooth. Asci 80–90 
<emphasis box="[369,384,177,200]" italics="true" pageId="14" pageNumber="208">×</emphasis>
<quantity box="[395,511,176,201]" metricMagnitude="-2" metricUnit="m" metricValue="2.25" metricValueMax="2.5" metricValueMin="2.0" pageId="14" pageNumber="208" unit="mm" value="22.5" valueMax="25.0" valueMin="20.0">20–25 µm</quantity>
, oblong. Ascospores 8 per ascus, muriform, with (5–) 7 transverse septa and 1–3 longitudinal septa per segment, 25–30 
<emphasis box="[366,381,273,296]" italics="true" pageId="14" pageNumber="208">×</emphasis>
<quantity box="[392,508,273,297]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.2" metricValueMin="1.0" pageId="14" pageNumber="208" unit="mm" value="11.0" valueMax="12.0" valueMin="10.0">10–12 µm</quantity>
, oblongellipsoid, with thickened septa and rounded lumina, hyaline, I+ violet-blue. Secondary chemistry: stictic acid, thallus in section with K+ persistently yellow efflux.
</paragraph>
</subSubSection>
<subSubSection pageId="14" pageNumber="208" type="etymology">
<paragraph blockId="14.[136,608,464,833]" pageId="14" pageNumber="208">
<emphasis bold="true" box="[136,265,464,488]" pageId="14" pageNumber="208">Etymology.</emphasis>
This new species is dedicated to the Mokaná, the only indigenous community in 
<collectingRegion box="[325,424,527,551]" country="Colombia" name="Atlantico" pageId="14" pageNumber="208">Atlántico</collectingRegion>
(
<bibRefCitation author="Borda C." box="[458,600,527,551]" journalOrPublisher="Eur. Rev. Latin Am. Caribb. Stud." pageId="14" pageNumber="208" pagination="39 - 57" part="86" refId="ref9525" refString="Borda C. 2009. Diversidad etnica y la reconstruccion de identidades: el grupo Mokana en el departamento del Atlantico, Colombia. Eur. Rev. Latin Am. Caribb. Stud. 86: 39 - 57." title="Diversidad etnica y la reconstruccion de identidades: el grupo Mokana en el departamento del Atlantico, Colombia" type="journal article" year="2009">Borda 2009</bibRefCitation>
, 
<bibRefCitation author="Montoya AB &amp; Siegler AH" box="[136,417,558,582]" journalOrPublisher="Barranquilla, Colombia: Ediciones UniNorte" pageId="14" pageNumber="208" refId="ref11818" refString="Montoya AB, Siegler AH. 2010. Cultura y tradicion oral en el Caribe colombiano: propuesta pedagogica para incorporar la investigacion: recoleccion de la tradicion oral Mokana en el Departamento del Atlantico. Barranquilla, Colombia: Ediciones UniNorte." title="Cultura y tradicion oral en el Caribe colombiano: propuesta pedagogica para incorporar la investigacion: recoleccion de la tradicion oral Mokana en el Departamento del Atlantico" type="book" year="2010">Montoya and Siegler 2010</bibRefCitation>
, 
<bibRefCitation author="Montoya AB &amp; Siegler AH" box="[429,481,558,582]" journalOrPublisher="Memorias" pageId="14" pageNumber="208" pagination="9" part="14" refId="ref11861" refString="Montoya AB, Siegler AH. 2011. La historia de los Mokana. Un capitulo de la historia en la region Caribe Colombiana. Memorias 14: 9." title="La historia de los Mokana. Un capitulo de la historia en la region Caribe Colombiana" type="journal article" year="2011">2011</bibRefCitation>
). The name is said to mean “ 
<emphasis box="[325,566,589,613]" italics="true" pageId="14" pageNumber="208">those without feathers</emphasis>
” in the Mokaná language, but this interpretation is disputed (
<bibRefCitation author="Montoya AB &amp; Siegler AH" box="[283,592,652,676]" journalOrPublisher="Barranquilla, Colombia: Ediciones UniNorte" pageId="14" pageNumber="208" refId="ref11818" refString="Montoya AB, Siegler AH. 2010. Cultura y tradicion oral en el Caribe colombiano: propuesta pedagogica para incorporar la investigacion: recoleccion de la tradicion oral Mokana en el Departamento del Atlantico. Barranquilla, Colombia: Ediciones UniNorte." title="Cultura y tradicion oral en el Caribe colombiano: propuesta pedagogica para incorporar la investigacion: recoleccion de la tradicion oral Mokana en el Departamento del Atlantico" type="book" year="2010">Montoya and Siegler 2010</bibRefCitation>
). The Mokaná language has become extinct, but the community, which was considered near-extinct about three decades ago, is recovering its cultural traditions in a process of re-indigenization (
<bibRefCitation author="Borda C." box="[363,487,809,833]" journalOrPublisher="Eur. Rev. Latin Am. Caribb. Stud." pageId="14" pageNumber="208" pagination="39 - 57" part="86" refId="ref9525" refString="Borda C. 2009. Diversidad etnica y la reconstruccion de identidades: el grupo Mokana en el departamento del Atlantico, Colombia. Eur. Rev. Latin Am. Caribb. Stud. 86: 39 - 57." title="Diversidad etnica y la reconstruccion de identidades: el grupo Mokana en el departamento del Atlantico, Colombia" type="journal article" year="2009">Borda 2009</bibRefCitation>
).
</paragraph>
</subSubSection>
<subSubSection pageId="14" pageNumber="208" type="discussion">
<paragraph blockId="14.[136,608,871,1553]" pageId="14" pageNumber="208">
<emphasis bold="true" box="[136,247,871,895]" pageId="14" pageNumber="208">Remarks.</emphasis>
Because of its uncarbonized excipulum, this species corresponds to the concept of the genus 
<taxonomicName authorityName="Trevisan" authorityYear="1853" box="[382,524,934,958]" class="Lecanoromycetes" family="Graphidaceae" genus="Hemithecium" kingdom="Fungi" order="Ostropales" pageId="14" pageNumber="208" phylum="Ascomycota" rank="genus">
<emphasis box="[382,524,934,958]" italics="true" pageId="14" pageNumber="208">Hemithecium</emphasis>
</taxonomicName>
, which has now been subsumed under the genera 
<taxonomicName box="[136,223,997,1021]" class="Gastropoda" family="Cimidae" genus="Graphis" kingdom="Animalia" order="Ostropales" pageId="14" pageNumber="208" phylum="Mollusca" rank="genus">
<emphasis box="[136,223,997,1021]" italics="true" pageId="14" pageNumber="208">Graphis</emphasis>
</taxonomicName>
and 
<taxonomicName authority="(Lucking and Kalb 2018)" baseAuthorityName="Lucking and Kalb" baseAuthorityYear="2018" class="Lecanoromycetes" family="Graphidaceae" genus="Allographa" kingdom="Fungi" order="Ostropales" pageId="14" pageNumber="208" phylum="Ascomycota" rank="genus">
<emphasis box="[306,427,997,1021]" italics="true" pageId="14" pageNumber="208">Allographa</emphasis>
(
<bibRefCitation author="Lucking R &amp; Kalb K." journalOrPublisher="Herzogia" pageId="14" pageNumber="208" pagination="535 - 561" part="31" refId="ref10824" refString="Lucking R, Kalb K. 2018. Formal instatement of Allographa (Graphidaceae): how to deal with a hyperdiverse genus complex with cryptic differentiation and paucity of molecular data. Herzogia 31 (1): 535 - 561. doi: 10.13158 / heia. 31. 1.2018. 535." title="Formal instatement of Allographa (Graphidaceae): how to deal with a hyperdiverse genus complex with cryptic differentiation and paucity of molecular data" type="journal article" year="2018">Lücking and Kalb 2018</bibRefCitation>
)
</taxonomicName>
. Among these two genera there are only two species with uncarbonized excipulum, small, muriform ascospores, and secondary substances: 
<taxonomicName authority="(Chitale, Makhija &amp; B. O. Sharma) Lucking &amp; Kalb" authorityName="Lucking &amp; Kalb" baseAuthorityName="Chitale, Makhija &amp; B. O. Sharma" class="Gastropoda" family="Cimidae" genus="Graphis" kingdom="Animalia" order="Ostropales" pageId="14" pageNumber="208" phylum="Mollusca" rank="species" species="microsperma">
<emphasis box="[432,608,1122,1146]" italics="true" pageId="14" pageNumber="208">G. microsperma</emphasis>
(Chitale, Makhija &amp; B. O. Sharma) Lücking &amp; Kalb
</taxonomicName>
from 
<collectingCountry box="[300,355,1185,1209]" name="India" pageId="14" pageNumber="208">India</collectingCountry>
differs in the smaller ascospores (15–20 
<emphasis box="[359,374,1216,1239]" italics="true" pageId="14" pageNumber="208">×</emphasis>
<quantity box="[390,496,1216,1240]" metricMagnitude="-2" metricUnit="m" metricValue="1.1" metricValueMax="1.4" metricValueMin="0.8" pageId="14" pageNumber="208" unit="mm" value="11.0" valueMax="14.0" valueMin="8.0">8–14 µm</quantity>
) and the formation of norstictic instead of stictic acid, whereas 
<taxonomicName authority="(Redinger) Lucking &amp; Kalb" authorityName="Lucking &amp; Kalb" baseAuthorityName="Redinger" class="Lecanoromycetes" family="Graphidaceae" genus="Allographa" kingdom="Fungi" order="Ostropales" pageId="14" pageNumber="208" phylum="Ascomycota" rank="species" species="dispersa">
<emphasis box="[242,473,1279,1303]" italics="true" pageId="14" pageNumber="208">Allographa dispersa</emphasis>
(Redinger) Lücking &amp; Kalb
</taxonomicName>
agrees in ascospore size, but also differs in the norstictic acid chemistry and the very short, prominent, white lirellae contrasting with the brownish thallus. The new species is morphologically similar to the widespread 
<taxonomicName authority="Fee" authorityName="Fee" box="[270,525,1467,1491]" class="Gastropoda" family="Cimidae" genus="Graphis" kingdom="Animalia" order="Ostropales" pageId="14" pageNumber="208" phylum="Mollusca" rank="species" species="implicata">
<emphasis box="[270,472,1467,1491]" italics="true" pageId="14" pageNumber="208">Graphis implicata</emphasis>
Fée
</taxonomicName>
, which differs in the transversely septate ascospores and lack of secondary substances.
</paragraph>
</subSubSection>
<subSubSection pageId="14" pageNumber="208" type="materials_examined">
<paragraph blockId="14.[136,608,1592,1679]" box="[136,608,1592,1616]" pageId="14" pageNumber="208">
<heading bold="true" box="[136,608,1592,1616]" centered="true" fontSize="10" level="4" pageId="14" pageNumber="208" reason="0">
<emphasis bold="true" box="[136,608,1592,1616]" pageId="14" pageNumber="208">Additional specimen examined.</emphasis>
</heading>
</paragraph>
<paragraph blockId="14.[136,608,1592,1679]" pageId="14" pageNumber="208">
<materialsCitation ID-GBIF-Occurrence="2565375025" collectionCode="RFP" collectorName="El Palomar" country="Colombia" location="Reserva Forestal Protectora" municipality="Piojo" pageId="14" pageNumber="208" specimenCount="1" stateProvince="Atlantico">
<collectingCountry box="[136,282,1623,1647]" name="Colombia" pageId="14" pageNumber="208">COLOMBIA</collectingCountry>
. 
<emphasis bold="true" box="[305,419,1623,1647]" pageId="14" pageNumber="208">
<collectingRegion box="[305,414,1623,1647]" country="Colombia" name="Atlantico" pageId="14" pageNumber="208">Atlántico</collectingRegion>
:
</emphasis>
<collectingMunicipality box="[439,498,1623,1647]" pageId="14" pageNumber="208">Piojó</collectingMunicipality>
, 
<location LSID="urn:lsid:plazi:treatment:216F4E11FF887302FD3AFD6719E4FEF2:AC19A9DCFF8B7302FDB3F98F1B28F976" country="Colombia" municipality="Piojo" name="Reserva Forestal Protectora" pageId="14" pageNumber="208" stateProvince="Atlantico">Reserva Forestal Protectora</location>
(
<collectionCode box="[384,434,1655,1679]" pageId="14" pageNumber="208">RFP</collectionCode>
) 
<collectorName box="[465,603,1655,1679]" pageId="14" pageNumber="208">El Palomar</collectorName>
</materialsCitation>
;
</paragraph>
<paragraph blockId="14.[640,1112,176,266]" pageId="14" pageNumber="208">
<materialsCitation ID-GBIF-Occurrence="2565375024" collectingDate="2016-03-18" collectionCode="R" elevation="400" latitude="10.763056" longLatPrecision="21" longitude="-75.14889" pageId="14" pageNumber="208" specimenCount="1">
<geoCoordinate box="[640,779,176,200]" degrees="10" direction="north" minutes="45" orientation="latitude" pageId="14" pageNumber="208" precision="15" seconds="47" value="10.763056">10º45’47” N</geoCoordinate>
, 
<geoCoordinate box="[792,934,176,200]" degrees="75" direction="west" minutes="08" orientation="longitude" pageId="14" pageNumber="208" precision="15" seconds="56" value="-75.14889">75º08’56” W</geoCoordinate>
, 
<quantity box="[946,1072,176,200]" metricMagnitude="2" metricUnit="m" metricValue="4.0" metricValueMax="6.0" metricValueMin="2.0" pageId="14" pageNumber="208" unit="m" value="400.0" valueMax="600.0" valueMin="200.0">
<elevation box="[946,1072,176,200]" metricMagnitude="2" metricUnit="m" metricValue="4.0" metricValueMax="6.0" metricValueMin="2.0" pageId="14" pageNumber="208" unit="m" value="400.0" valueMax="600.0" valueMin="200.0">200–600 m</elevation>
</quantity>
; 
<date pageId="14" pageNumber="208" value="2016-03-18">
<collectingDate pageId="14" pageNumber="208" value="2016-03-18">18 March 2016</collectingDate>
</date>
, 
<emphasis italics="true" pageId="14" pageNumber="208">
<collectionCode box="[785,804,210,233]" country="Chile" name="Departamento de Geologia, Universidad de Chile" pageId="14" pageNumber="208">R</collectionCode>
. 
<materialsCitation ID-GBIF-Occurrence="2565375029" collectionCode="UNO" collectorName="Lucking, B" pageId="14" pageNumber="208" specimenCount="1" typeStatus="paratype">
<collectorName box="[816,936,209,233]" pageId="14" pageNumber="208">Lücking, B</collectorName>
. Moncada et al. 42975 (B, 
<collectionCode box="[753,813,242,266]" country="0" httpUri="http://grbio.org/cool/fyxp-7ves" name="University of Nebraska at Omaha" pageId="14" pageNumber="208">UNO</collectionCode>
, 
<typeStatus box="[824,931,242,266]" pageId="14" pageNumber="208">paratypes</typeStatus>
)
</materialsCitation>
</emphasis>
</materialsCitation>
.
</paragraph>
</subSubSection>
</treatment>
</document>