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<document id="705E981901D9FEBB01B194F4B1F207B2" ID-CLB-Dataset="42402" ID-DOI="10.11646/zootaxa.3872.5.9" ID-GBIF-Dataset="099868dc-c782-452c-aaa8-d3375dcbb364" ID-ISSN="1175-5326" ID-Zenodo-Dep="251251" ID-ZooBank="41BDCF15-0657-4CC8-82DC-37D04C5DDE38" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1460933014911" checkinUser="plazi" docAuthor="Jin, Mengjie, Weir, Tom, Ślipiński, Adam &amp; Pang, Hong" docDate="2014" docId="214087F6FF8C0832FF262E48CB1007EE" docLanguage="en" docName="zt03872p600.pdf" docOrigin="Zootaxa 3872 (5)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Anomala Samouelle" docType="treatment" docVersion="9" lastPageNumber="593" masterDocId="DD79FF8EFF8D0830FFB12D20C9180666" masterDocTitle="Description of a new species of Anomala Samouelle (Coleoptera: Scarabaeidae: Rutelinae) from Northern Territory, Australia" masterLastPageNumber="600" masterPageNumber="591" pageNumber="592" updateTime="1698582436333" updateUser="ExternalLinkService">
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<mods:title id="84F06B5D8F16700B3891BC14EBCA1BF2">Description of a new species of Anomala Samouelle (Coleoptera: Scarabaeidae: Rutelinae) from Northern Territory, Australia</mods:title>
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<mods:namePart id="D5AD45CCA642AE402EBBA40BABE9D54F">Jin, Mengjie</mods:namePart>
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<treatment id="214087F6FF8C0832FF262E48CB1007EE" ID-DOI="http://doi.org/10.5281/zenodo.6141590" ID-GBIF-Taxon="119590227" ID-Zenodo-Dep="6141590" LSID="urn:lsid:plazi:treatment:214087F6FF8C0832FF262E48CB1007EE" httpUri="http://treatment.plazi.org/id/214087F6FF8C0832FF262E48CB1007EE" lastPageId="2" lastPageNumber="593" pageId="1" pageNumber="592">
<subSubSection id="E1F3656BFF8C0831FF262E48C89505E5" box="[151,397,872,899]" pageId="1" pageNumber="592" type="nomenclature">
<paragraph id="A95636E0FF8C0831FF262E48C89505E5" blockId="1.[151,397,872,899]" box="[151,397,872,899]" pageId="1" pageNumber="592">
<heading id="F21E818CFF8C0831FF262E48C89505E5" bold="true" box="[151,397,872,899]" fontSize="11" level="1" pageId="1" pageNumber="592" reason="1">
<taxonomicName id="6EE94D63FF8C0831FF262E48C89505E5" authority="Samouelle" authorityName="Samouelle" box="[151,397,872,899]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="592" phylum="Arthropoda" rank="genus">
<emphasis id="9B9DEAF2FF8C0831FF262E48C89505E5" bold="true" box="[151,397,872,899]" pageId="1" pageNumber="592">
<emphasis id="9B9DEAF2FF8C0831FF262E48C81E05E4" bold="true" box="[151,262,872,898]" italics="true" pageId="1" pageNumber="592">Anomala</emphasis>
Samouelle
</emphasis>
</taxonomicName>
</heading>
</paragraph>
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<subSubSection id="E1F3656BFF8C0831FF262E89CB1405B8" pageId="1" pageNumber="592" type="reference_group">
<paragraph id="A95636E0FF8C0831FF262E89CB1405B8" blockId="1.[151,984,937,990]" pageId="1" pageNumber="592">
<treatmentCitationGroup id="89F911CEFF8C0831FF262E89CB1405B8" pageId="1" pageNumber="592">
<taxonomicName id="6EE94D63FF8C0831FF262E89C8C105D9" authority="Samouelle, 1819: 191" authorityName="Samouelle" authorityPageNumber="191" authorityYear="1819" box="[151,473,937,959]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="592" phylum="Arthropoda" rank="genus">
<emphasis id="9B9DEAF2FF8C0831FF262E89C9EB05D9" box="[151,243,937,959]" italics="true" pageId="1" pageNumber="592">Anomala</emphasis>
<treatmentCitation id="284810F1FF8C0831FF492E89C8C105D9" author="Samouelle" box="[248,473,937,959]" page="191" pageId="1" pageNumber="592" year="1819">
<bibRefCitation id="CD784B11FF8C0831FF492E89C8BE05D9" author="Samouelle" box="[248,422,937,959]" pageId="1" pageNumber="592" refString="Samouelle, G. (1819) The Entomologist's Useful Compendium: or an Introduction to the Knowledge of British Insects. Thomas Boys, London, 496 pp., 12 plates. [United Kingdom] http: // dx. doi. org / 10.5962 / bhl. title. 5527" type="book" year="1819">Samouelle, 1819</bibRefCitation>
: 191
</treatmentCitation>
</taxonomicName>
.
<typeStatus id="76528842FF8C0831FE532E8ACB0C05D9" box="[482,532,938,959]" pageId="1" pageNumber="592">Type</typeStatus>
species
<taxonomicName id="6EE94D63FF8C0831FDDA2E89CACC05D9" authority="Fabricius, 1775" authorityName="Fabricius" authorityYear="1775" box="[619,980,937,959]" class="Insecta" family="Melolonthidae" genus="Melolontha" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="592" phylum="Arthropoda" rank="species" species="frischii">
<emphasis id="9B9DEAF2FF8C0831FDDA2E89CA3605D9" box="[619,814,937,959]" italics="true" pageId="1" pageNumber="592">Melolontha frischii</emphasis>
Fabricius, 1775
</taxonomicName>
. Full synonymy see
<treatmentCitation id="284810F1FF8C0831FEED2EE9CB1005B8" author="Zorn" box="[348,520,968,990]" page="256" pageId="1" pageNumber="592" year="2006">
<bibRefCitation id="CD784B11FF8C0831FEED2EE9C8CD05B8" author="Zorn" box="[348,469,968,990]" pageId="1" pageNumber="592" refString="Zorn, C. (2006 a) Rutelinae. In: Lobl, I. &amp; Smetana, A. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 3. Apollo Books, Stenstrup, pp. 248 - 277. [Denmark]" type="book" year="2006" yearSuffix="a">Zorn 2006a</bibRefCitation>
: 256
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.
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</paragraph>
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<subSubSection id="E1F3656BFF8C0832FF262924CA000726" lastPageId="2" lastPageNumber="593" pageId="1" pageNumber="592" type="diagnosis">
<paragraph id="A95636E0FF8C0831FF262924C80502ED" blockId="1.[151,1437,1028,2027]" pageId="1" pageNumber="592">
<emphasis id="9B9DEAF2FF8C0831FF262924C809027A" bold="true" box="[151,273,1028,1053]" pageId="1" pageNumber="592">Diagnosis.</emphasis>
<taxonomicName id="6EE94D63FF8C0831FEA82924C866027A" box="[281,382,1028,1052]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="592" phylum="Arthropoda" rank="genus">
<emphasis id="9B9DEAF2FF8C0831FEA82924C866027A" box="[281,382,1028,1052]" italics="true" pageId="1" pageNumber="592">Anomala</emphasis>
</taxonomicName>
can easily be separated from remaining genera of Australian
<taxonomicName id="6EE94D63FF8C0831FB8F2924CDB1027B" box="[1086,1193,1028,1053]" class="Insecta" family="Scarabaeidae" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="592" phylum="Arthropoda" rank="subFamily" subFamily="Rutelinae">Rutelinae</taxonomicName>
by having the tergite and sternite of the 7th abdominal segment completely fused into a ring without visible sutures (
<figureCitation id="31D22A65FF8C0831FB63290ACC3B0225" box="[1234,1315,1066,1091]" captionStart="FIGURES 2 A J" captionStartId="4.[151,264,1852,1875]" captionTargetBox="[178,1392,219,1788]" captionTargetId="figure@4.[160,1426,219,1799]" captionTargetPageId="4" captionText="FIGURES 2 A J. Anomala katherine new species, male: A, Prothorax, dorsal; B, prothorax ventral with both coxae removed; C, metendosternite, ventral; D, pterothorax, ventral with both coxae removed; E, pterothorax, dorsal with wings removed; F, proleg; G, mesoleg; H, metaleg; I, abdomen, ventral; J, abdomen, dorsal." httpUri="https://zenodo.org/record/251253/files/figure.png" pageId="1" pageNumber="592">Fig. 2J</figureCitation>
), and by a combination of 9 antennomeres (
<figureCitation id="31D22A65FF8C0831FDBA296ECB7B0201" box="[523,611,1102,1127]" captionStart="FIGURES 1 A M" captionStartId="3.[151,264,1948,1971]" captionTargetBox="[162,1431,369,1909]" captionTargetId="figure@3.[151,1436,359,1927]" captionTargetPageId="3" captionText="FIGURES 1 A M. Anomala spp. A L, A. katherine new species: A, Head, dorsal; B, head, ventral; C, head, lateral; D, left mandible, dorsal; E, right mandible, dorsal; F, left mandible, inner; G, right mandible, inner; H, antenna; I, maxilla, ventral; J, labrum; K, labium, ventral, L, hind wing. M, A. antiqua (Gyllenhal): maxilla, ventral." httpUri="https://zenodo.org/record/251252/files/figure.png" pageId="1" pageNumber="592">Fig. 1H</figureCitation>
) and the base of the labrum horizontal and separated by membrane from the clypeus.
</paragraph>
<paragraph id="A95636E0FF8C0831FF7629B6C89103A9" blockId="1.[151,1437,1028,2027]" pageId="1" pageNumber="592">
<emphasis id="9B9DEAF2FF8C0831FF7629B6C8FF02C9" bold="true" box="[199,487,1174,1199]" pageId="1" pageNumber="592">Diagnostic combination.</emphasis>
Length 6.018.0 mm. Body elongate oval, convex; dorsum glabrous, green or brown often with iridescent shine. Head with completely visible frontoclypeal suture (
<figureCitation id="31D22A65FF8C0831FBE2299ACDAC02B5" box="[1107,1204,1210,1235]" captionStart="FIGURES 1 A M" captionStartId="3.[151,264,1948,1971]" captionTargetBox="[162,1431,369,1909]" captionTargetId="figure@3.[151,1436,359,1927]" captionTargetPageId="3" captionText="FIGURES 1 A M. Anomala spp. A L, A. katherine new species: A, Head, dorsal; B, head, ventral; C, head, lateral; D, left mandible, dorsal; E, right mandible, dorsal; F, left mandible, inner; G, right mandible, inner; H, antenna; I, maxilla, ventral; J, labrum; K, labium, ventral, L, hind wing. M, A. antiqua (Gyllenhal): maxilla, ventral." httpUri="https://zenodo.org/record/251252/files/figure.png" pageId="1" pageNumber="592">Fig. 1A</figureCitation>
). Labrum (
<figureCitation id="31D22A65FF8C0831FA8E299ACC8B02B5" box="[1343,1427,1210,1235]" captionStart="FIGURES 1 A M" captionStartId="3.[151,264,1948,1971]" captionTargetBox="[162,1431,369,1909]" captionTargetId="figure@3.[151,1436,359,1927]" captionTargetPageId="3" captionText="FIGURES 1 A M. Anomala spp. A L, A. katherine new species: A, Head, dorsal; B, head, ventral; C, head, lateral; D, left mandible, dorsal; E, right mandible, dorsal; F, left mandible, inner; G, right mandible, inner; H, antenna; I, maxilla, ventral; J, labrum; K, labium, ventral, L, hind wing. M, A. antiqua (Gyllenhal): maxilla, ventral." httpUri="https://zenodo.org/record/251252/files/figure.png" pageId="1" pageNumber="592">Fig. 1J</figureCitation>
) separated by exposed membrane from clypeus, horizontal at base and not extending far beyond clypeus (
<figureCitation id="31D22A65FF8C0831FA8429FECC940291" box="[1333,1420,1246,1271]" captionStart="FIGURES 1 A M" captionStartId="3.[151,264,1948,1971]" captionTargetBox="[162,1431,369,1909]" captionTargetId="figure@3.[151,1436,359,1927]" captionTargetPageId="3" captionText="FIGURES 1 A M. Anomala spp. A L, A. katherine new species: A, Head, dorsal; B, head, ventral; C, head, lateral; D, left mandible, dorsal; E, right mandible, dorsal; F, left mandible, inner; G, right mandible, inner; H, antenna; I, maxilla, ventral; J, labrum; K, labium, ventral, L, hind wing. M, A. antiqua (Gyllenhal): maxilla, ventral." httpUri="https://zenodo.org/record/251252/files/figure.png" pageId="1" pageNumber="592">Fig. 1C</figureCitation>
); anterior edge emarginate medially with dense bristles laterally. Antenna with 9 antennomeres (
<figureCitation id="31D22A65FF8C0831FB082822CC09037D" box="[1209,1297,1282,1307]" captionStart="FIGURES 1 A M" captionStartId="3.[151,264,1948,1971]" captionTargetBox="[162,1431,369,1909]" captionTargetId="figure@3.[151,1436,359,1927]" captionTargetPageId="3" captionText="FIGURES 1 A M. Anomala spp. A L, A. katherine new species: A, Head, dorsal; B, head, ventral; C, head, lateral; D, left mandible, dorsal; E, right mandible, dorsal; F, left mandible, inner; G, right mandible, inner; H, antenna; I, maxilla, ventral; J, labrum; K, labium, ventral, L, hind wing. M, A. antiqua (Gyllenhal): maxilla, ventral." httpUri="https://zenodo.org/record/251252/files/figure.png" pageId="1" pageNumber="592">Fig. 1H</figureCitation>
). Mandibles multidentate apically, asymmetrical (
<figureCitation id="31D22A65FF8C0831FD892806CBAF0359" box="[568,695,1318,1343]" captionStart="FIGURES 1 A M" captionStartId="3.[151,264,1948,1971]" captionTargetBox="[162,1431,369,1909]" captionTargetId="figure@3.[151,1436,359,1927]" captionTargetPageId="3" captionText="FIGURES 1 A M. Anomala spp. A L, A. katherine new species: A, Head, dorsal; B, head, ventral; C, head, lateral; D, left mandible, dorsal; E, right mandible, dorsal; F, left mandible, inner; G, right mandible, inner; H, antenna; I, maxilla, ventral; J, labrum; K, labium, ventral, L, hind wing. M, A. antiqua (Gyllenhal): maxilla, ventral." httpUri="https://zenodo.org/record/251252/files/figure.png" pageId="1" pageNumber="592">Figs 1AG</figureCitation>
) with right mandible having larger and more complex mola (
<figureCitation id="31D22A65FF8C0831FADE2806C9AF0305" captionStart="FIGURES 1 A M" captionStartId="3.[151,264,1948,1971]" captionTargetBox="[162,1431,369,1909]" captionTargetId="figure@3.[151,1436,359,1927]" captionTargetPageId="3" captionText="FIGURES 1 A M. Anomala spp. A L, A. katherine new species: A, Head, dorsal; B, head, ventral; C, head, lateral; D, left mandible, dorsal; E, right mandible, dorsal; F, left mandible, inner; G, right mandible, inner; H, antenna; I, maxilla, ventral; J, labrum; K, labium, ventral, L, hind wing. M, A. antiqua (Gyllenhal): maxilla, ventral." httpUri="https://zenodo.org/record/251252/files/figure.png" pageId="1" pageNumber="592">Fig. 1F</figureCitation>
); prostheca membranous with dense fringe of marginal setae. Maxilla (
<figureCitation id="31D22A65FF8C0831FC5C286ACDB20305" box="[1005,1194,1354,1379]" captionStart="FIGURES 1 A M" captionStartId="3.[151,264,1948,1971]" captionTargetBox="[162,1431,369,1909]" captionTargetId="figure@3.[151,1436,359,1927]" captionTargetPageId="3" captionText="FIGURES 1 A M. Anomala spp. A L, A. katherine new species: A, Head, dorsal; B, head, ventral; C, head, lateral; D, left mandible, dorsal; E, right mandible, dorsal; F, left mandible, inner; G, right mandible, inner; H, antenna; I, maxilla, ventral; J, labrum; K, labium, ventral, L, hind wing. M, A. antiqua (Gyllenhal): maxilla, ventral." httpUri="https://zenodo.org/record/251252/files/figure.png" pageId="1" pageNumber="592">
Figs 1A, 1M,
<date id="DD571020FF8C0831FB22286BCDB20305" box="[1171,1194,1355,1379]" pageId="1" pageNumber="592">1I</date>
</figureCitation>
) with lacinia absent; galea well developed bearing several heavily sclerotised teeth apically; terminal maxillary palpomere without sensory palp organ. Labium (
<figureCitation id="31D22A65FF8C0831FE5128B2CB7203CD" box="[480,618,1426,1451]" captionStart="FIGURES 1 A M" captionStartId="3.[151,264,1948,1971]" captionTargetBox="[162,1431,369,1909]" captionTargetId="figure@3.[151,1436,359,1927]" captionTargetPageId="3" captionText="FIGURES 1 A M. Anomala spp. A L, A. katherine new species: A, Head, dorsal; B, head, ventral; C, head, lateral; D, left mandible, dorsal; E, right mandible, dorsal; F, left mandible, inner; G, right mandible, inner; H, antenna; I, maxilla, ventral; J, labrum; K, labium, ventral, L, hind wing. M, A. antiqua (Gyllenhal): maxilla, ventral." httpUri="https://zenodo.org/record/251252/files/figure.png" pageId="1" pageNumber="592">Figs 1B, 1K</figureCitation>
) heavily sclerotised, ligula weakly emarginate medially, without median projection or process.
</paragraph>
<paragraph id="A95636E0FF8C0831FF7628FACB4E00AD" blockId="1.[151,1437,1028,2027]" pageId="1" pageNumber="592">
Prothorax transverse; prosternum (
<figureCitation id="31D22A65FF8C0831FDE328FACBB30395" box="[594,683,1498,1523]" captionStart="FIGURES 2 A J" captionStartId="4.[151,264,1852,1875]" captionTargetBox="[178,1392,219,1788]" captionTargetId="figure@4.[160,1426,219,1799]" captionTargetPageId="4" captionText="FIGURES 2 A J. Anomala katherine new species, male: A, Prothorax, dorsal; B, prothorax ventral with both coxae removed; C, metendosternite, ventral; D, pterothorax, ventral with both coxae removed; E, pterothorax, dorsal with wings removed; F, proleg; G, mesoleg; H, metaleg; I, abdomen, ventral; J, abdomen, dorsal." httpUri="https://zenodo.org/record/251253/files/figure.png" pageId="1" pageNumber="592">Fig. 2B</figureCitation>
) short in front of coxae with narrow prosternal process; procoxal cavities open externally and internally. Scutellar shield well developed (
<figureCitation id="31D22A65FF8C0831FBB528DECD7B0071" box="[1028,1123,1534,1559]" captionStart="FIGURES 2 A J" captionStartId="4.[151,264,1852,1875]" captionTargetBox="[178,1392,219,1788]" captionTargetId="figure@4.[160,1426,219,1799]" captionTargetPageId="4" captionText="FIGURES 2 A J. Anomala katherine new species, male: A, Prothorax, dorsal; B, prothorax ventral with both coxae removed; C, metendosternite, ventral; D, pterothorax, ventral with both coxae removed; E, pterothorax, dorsal with wings removed; F, proleg; G, mesoleg; H, metaleg; I, abdomen, ventral; J, abdomen, dorsal." httpUri="https://zenodo.org/record/251253/files/figure.png" pageId="1" pageNumber="592">Fig. 2E</figureCitation>
). Mesoventrite flat, broad anteriorly, separated from small mesanepisternum by complete sutures (
<figureCitation id="31D22A65FF8C0831FC6D2B02CD22005D" box="[988,1082,1570,1595]" captionStart="FIGURES 2 A J" captionStartId="4.[151,264,1852,1875]" captionTargetBox="[178,1392,219,1788]" captionTargetId="figure@4.[160,1426,219,1799]" captionTargetPageId="4" captionText="FIGURES 2 A J. Anomala katherine new species, male: A, Prothorax, dorsal; B, prothorax ventral with both coxae removed; C, metendosternite, ventral; D, pterothorax, ventral with both coxae removed; E, pterothorax, dorsal with wings removed; F, proleg; G, mesoleg; H, metaleg; I, abdomen, ventral; J, abdomen, dorsal." httpUri="https://zenodo.org/record/251253/files/figure.png" pageId="1" pageNumber="592">Fig. 2D</figureCitation>
). Mesoventral process narrow, extending beyond middle of mesocoxal cavity. Mesocoxal cavity obliquely transverse; laterally closed by broad mesepimeron. Mesotrochantin narrow, exposed. Metaventrite with complete discrimen. Elytra 1.3 times as long as wide and 2.5 times as long as pronotum, with fine strial punctures; epipleura glabrous, obsolete near apices; apical half with membranous border (
<figureCitation id="31D22A65FF8C0831FE432B92CB5200AD" box="[498,586,1714,1739]" captionStart="FIGURES 4 A K" captionStartId="6.[151,264,1757,1780]" captionTargetBox="[189,1406,202,1733]" captionTargetId="figure@6.[179,1407,193,1737]" captionTargetPageId="6" captionText="FIGURES 4 A K. Anomala spp. A C, Posterior part of pronotum and scutellum; D, left elytron showing striae and intervals; E, apical portion of elytra showing membranous border; F, pygidium; G, lateral portion of pronotum showing colour pattern; H, male protarsus; I, female protarsus; J, female ovipositor and internal track; K, spiculum gastrale. A, A. antiqua (Gyllenhal); B, F, G, A. aeneotincta Fairmaire; C E, H K, A. katherine new species." httpUri="https://zenodo.org/record/251255/files/figure.png" pageId="1" pageNumber="592">Fig. 4E</figureCitation>
).
</paragraph>
<paragraph id="A95636E0FF8C0831FF762BF6CAA4013C" blockId="1.[151,1437,1028,2027]" pageId="1" pageNumber="592">
<collectingCountry id="D1FE7670FF8C0831FF762BF6C9E70089" box="[199,255,1750,1775]" name="India" pageId="1" pageNumber="592">Hind</collectingCountry>
wing (
<figureCitation id="31D22A65FF8C0831FEE02BF6C8BE0089" box="[337,422,1750,1775]" captionStart="FIGURES 1 A M" captionStartId="3.[151,264,1948,1971]" captionTargetBox="[162,1431,369,1909]" captionTargetId="figure@3.[151,1436,359,1927]" captionTargetPageId="3" captionText="FIGURES 1 A M. Anomala spp. A L, A. katherine new species: A, Head, dorsal; B, head, ventral; C, head, lateral; D, left mandible, dorsal; E, right mandible, dorsal; F, left mandible, inner; G, right mandible, inner; H, antenna; I, maxilla, ventral; J, labrum; K, labium, ventral, L, hind wing. M, A. antiqua (Gyllenhal): maxilla, ventral." httpUri="https://zenodo.org/record/251252/files/figure.png" pageId="1" pageNumber="592">Fig. 1L</figureCitation>
) about 2.6 times as long as wide; radial bending zone relatively short, setose; R-M loop quite strong; apical field about 0.4 times total wing length, with two
<collectingCountry id="D1FE7670FF8C0831FC142BDBCADF0174" box="[933,967,1787,1810]" name="Poland" pageId="1" pageNumber="592">RP</collectingCountry>
veins, MP1+2 very long and detached to base; medial field with 4 free veins, MP3+4 extremely narrow and detached, AA vein forms a basal cell with Cu; anal lobe well-developed, anal notch absent, jugal veins reduced to one.
</paragraph>
<paragraph id="A95636E0FF8C0831FF762A46CB68018D" blockId="1.[151,1437,1028,2027]" pageId="1" pageNumber="592">
Abdomen with 7 visible segments (IIVIII, ventrites 17) (
<figureCitation id="31D22A65FF8C0831FCDB2A46CAC00119" box="[874,984,1894,1919]" captionStart="FIGURES 2 A J" captionStartId="4.[151,264,1852,1875]" captionTargetBox="[178,1392,219,1788]" captionTargetId="figure@4.[160,1426,219,1799]" captionTargetPageId="4" captionText="FIGURES 2 A J. Anomala katherine new species, male: A, Prothorax, dorsal; B, prothorax ventral with both coxae removed; C, metendosternite, ventral; D, pterothorax, ventral with both coxae removed; E, pterothorax, dorsal with wings removed; F, proleg; G, mesoleg; H, metaleg; I, abdomen, ventral; J, abdomen, dorsal." httpUri="https://zenodo.org/record/251253/files/figure.png" pageId="1" pageNumber="592">
Figs
<date id="DD571020FF8C0831FC152A47CAA60119" box="[932,958,1895,1919]" pageId="1" pageNumber="592">2I</date>
, J
</figureCitation>
); segments I and II connate and mostly hidden under very large and flat metacoxae (
<figureCitation id="31D22A65FF8C0831FD192AAACA1E01C5" box="[680,774,1930,1955]" captionStart="FIGURES 2 A J" captionStartId="4.[151,264,1852,1875]" captionTargetBox="[178,1392,219,1788]" captionTargetId="figure@4.[160,1426,219,1799]" captionTargetPageId="4" captionText="FIGURES 2 A J. Anomala katherine new species, male: A, Prothorax, dorsal; B, prothorax ventral with both coxae removed; C, metendosternite, ventral; D, pterothorax, ventral with both coxae removed; E, pterothorax, dorsal with wings removed; F, proleg; G, mesoleg; H, metaleg; I, abdomen, ventral; J, abdomen, dorsal." httpUri="https://zenodo.org/record/251253/files/figure.png" pageId="1" pageNumber="592">Fig. 2H</figureCitation>
); intercoxal process indistinct. Abdomen with 7 pairs of spiracles, terminal spiracle exposed and placed on evenly sclerotised and fused sternite and tergite VII. Pygidium convex in profile, apex narrowly rounded.
</paragraph>
<paragraph id="A95636E0FF8F0832FF762DB7CA000726" blockId="2.[151,1437,151,392]" pageId="2" pageNumber="593">
Male genitalia trilobate, symmetrical; parameres short, simple, weakly bilobed; phallobase with laterobasal apodemes; penis simple, with slender struts; endophallus with exposed setal patch. Spiculum gastrale short and “Y” shaped (
<figureCitation id="31D22A65FF8F0832FE892DFFC883069E" box="[312,411,223,248]" captionStart="FIGURES 4 A K" captionStartId="6.[151,264,1757,1780]" captionTargetBox="[189,1406,202,1733]" captionTargetId="figure@6.[179,1407,193,1737]" captionTargetPageId="6" captionText="FIGURES 4 A K. Anomala spp. A C, Posterior part of pronotum and scutellum; D, left elytron showing striae and intervals; E, apical portion of elytra showing membranous border; F, pygidium; G, lateral portion of pronotum showing colour pattern; H, male protarsus; I, female protarsus; J, female ovipositor and internal track; K, spiculum gastrale. A, A. antiqua (Gyllenhal); B, F, G, A. aeneotincta Fairmaire; C E, H K, A. katherine new species." httpUri="https://zenodo.org/record/251255/files/figure.png" pageId="2" pageNumber="593">Fig. 4K</figureCitation>
), associated plates large, not overlapping at centre. Ovipositor very short (
<figureCitation id="31D22A65FF8F0832FA842DFFCC96069E" box="[1333,1422,223,248]" captionStart="FIGURES 4 A K" captionStartId="6.[151,264,1757,1780]" captionTargetBox="[189,1406,202,1733]" captionTargetId="figure@6.[179,1407,193,1737]" captionTargetPageId="6" captionText="FIGURES 4 A K. Anomala spp. A C, Posterior part of pronotum and scutellum; D, left elytron showing striae and intervals; E, apical portion of elytra showing membranous border; F, pygidium; G, lateral portion of pronotum showing colour pattern; H, male protarsus; I, female protarsus; J, female ovipositor and internal track; K, spiculum gastrale. A, A. antiqua (Gyllenhal); B, F, G, A. aeneotincta Fairmaire; C E, H K, A. katherine new species." httpUri="https://zenodo.org/record/251255/files/figure.png" pageId="2" pageNumber="593">Fig. 4J</figureCitation>
); gonocoxites sclerotised, kidney shaped, without gonostyli; small spermatheca and a pair of large sclerotised vaginal glands open to short vagina and bursa copulatrix.
</paragraph>
</subSubSection>
<subSubSection id="E1F3656BFF8F0832FF762C6CCB1007EE" pageId="2" pageNumber="593" type="discussion">
<paragraph id="A95636E0FF8F0832FF762C6CCB1007EE" blockId="2.[151,1437,151,392]" pageId="2" pageNumber="593">
<emphasis id="9B9DEAF2FF8F0832FF762C6CC8570702" bold="true" box="[199,335,332,357]" pageId="2" pageNumber="593">Comments.</emphasis>
The diagnosis and the diagnostic combination provided above are based on the Australian representatives of
<taxonomicName id="6EE94D63FF8F0832FED52C50C8D107EE" box="[356,457,368,392]" class="Insecta" family="Rutelidae" genus="Anomala" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="593" phylum="Arthropoda" rank="genus">
<emphasis id="9B9DEAF2FF8F0832FED52C50C8D107EE" box="[356,457,368,392]" italics="true" pageId="2" pageNumber="593">Anomala</emphasis>
</taxonomicName>
only.
</paragraph>
</subSubSection>
</treatment>
</document>