245 lines
33 KiB
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245 lines
33 KiB
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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1119.85400" ID-GBIF-Dataset="418bd521-e4b9-4ee6-981f-a6f471eab287" ID-Pensoft-Pub="1313-2970-1119-1" ID-Pensoft-UUID="F158A7C5261D52B69288A62F3C777CAF" ID-ZooBank="A3BE91C6B79344E1A886A803BF104D8B" ModsDocID="1313-2970-1119-1" checkinTime="1661532673136" checkinUser="pensoft" docAuthor="Jirapatrasilp, Parin, Sutcharit, Chirasak & Panha, Somsak" docDate="2022" docId="210059B2AF7151B8821080CDE3E8D9AD" docLanguage="en" docName="ZooKeys 1119: 1-115" docOrigin="ZooKeys 1119" docPubDate="2022-08-25" docSource="http://dx.doi.org/10.3897/zookeys.1119.85400" docTitle="Pupinella illustris Jirapatrasilp & Sutcharit & Panha 2022, comb. nov." docType="treatment" docVersion="4" id="F158A7C5261D52B69288A62F3C777CAF" lastPageNumber="1" masterDocId="F158A7C5261D52B69288A62F3C777CAF" masterDocTitle="Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia" masterLastPageNumber="115" masterPageNumber="1" pageNumber="1" updateTime="1661537468408" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Annotated checklist of the operculated land snails from Thailand (Mollusca, Gastropoda, Caenogastropoda): the family Pupinidae, with descriptions of several new species and subspecies, and notes on classification of Pupina Vignard, 1829 and Pupinella Gray, 1850 from mainland Southeast Asia</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Jirapatrasilp, Parin</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0002-5591-6724</mods:nameIdentifier>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Sutcharit, Chirasak</mods:namePart>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand</mods:affiliation>
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<mods:nameIdentifier type="email">jirasak4@yahoo.com</mods:nameIdentifier>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Panha, Somsak</mods:namePart>
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<mods:affiliation>Animal Systematics Research Unit, Department of Biology, Faculty of Science, Chulalongkorn University, Bangkok 10330, Thailand & Academy of Science, The Royal Society of Thailand, Bangkok 10300, Thailand</mods:affiliation>
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<mods:nameIdentifier type="email">somsak.pan@chula.ac.th</mods:nameIdentifier>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-08-25</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1119</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>115</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1119.85400</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1119.85400</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1119-1</mods:identifier>
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<mods:identifier type="ZooBank">A3BE91C6B79344E1A886A803BF104D8B</mods:identifier>
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<mods:identifier type="Pensoft-UUID">F158A7C5261D52B69288A62F3C777CAF</mods:identifier>
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<treatment ID-GBIF-Taxon="199839070" LSID="urn:lsid:plazi:treatment:210059B2AF7151B8821080CDE3E8D9AD" httpUri="http://treatment.plazi.org/id/210059B2AF7151B8821080CDE3E8D9AD" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="210059B2-AF71-51B8-8210-80CDE3E8D9AD" authority="(Mabille, 1887)" authorityName="Jirapatrasilp & Sutcharit & Panha" authorityYear="2022" baseAuthorityName="Mabille" baseAuthorityYear="1887" class="Gastropoda" family="Pupinidae" genus="Pupinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pupinella illustris" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="illustris" status="comb. nov.">Pupinella illustris (Mabille, 1887)</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">comb. nov.</taxonomicNameLabel>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Shells of Pupinella species from mainland Southeast Asia. A-G Pupinella mansuyi: A syntype of Eupupina mansuyi MNHN-IM- 2000 - 30756 from Deux-Ponts B syntype of Eupupina mansuyi MNHN-IM- 2000 - 36067 from Deux-Ponts C syntype of Eupupina mansuyi MNHN-IM- 2000 - 36068 from Quang-Huyen D syntype of Eupupina mansuyi RBINS MT 970 / 1 from Quang-Huyen E holotype of Pupinella frednaggsi NHMUK 20170285 F specimen CUMZ 12148 from Pha Chu Mount, Nan, and G specimen CUMZ 12149 from Pha Tub Cave, Nan H Pupinella sonlaensis, paratype ZRC. MOL. 9377 I-L Pupinella illustris I, J syntypes of Pupina illustris MNHN-IM- 2000 - 35842 from Tonkin K lectotype of Pupina tonkiniana MNHN-IM- 2000 - 35838 from Lang-Son et That-Khe, and L paralectotype of Pupina tonkiniana SMF 109932 / 10 from Tonkin: That-khe. Photo: A. Lardeur, P. Maestrati, MNHN (A-C, I-K), F. Trus, RBINS (D), S. K. Tan, ZRC (H)." figureDoi="10.3897/zookeys.1119.85400.figure11" httpUri="https://binary.pensoft.net/fig/735372" pageId="0" pageNumber="1">Figs 11I-L</figureCitation>
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. A-C Pupinella mansuyi: A syntype of Eupupina mansuyi RBINS MT 970 / 1 from Quang-Huyen B specimen CUMZ 12148 from Pha Chu Mount, Nan, and C specimen CUMZ 12149 from Pha Tub Cave, Nan. D, E Pupinella illustris D syntype of Pupina illustris MNHN-IM- 2000 - 35842 from Tonkin and E paralectotype of Pupina tonkiniana SMF 109932 / 10 from Tonkin: That-khe F Pupinella sonlaensis, paratype ZRC. MOL. 9377. Photo: F. Trus, RBINS (A), P. Maestrati, MNHN (D), S. K. Tan, ZRC (F)." figureDoi="10.3897/zookeys.1119.85400.figure12" httpUri="https://binary.pensoft.net/fig/735373" pageId="0" pageNumber="1">, 12D, E</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="reference_group">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Jirapatrasilp & Sutcharit & Panha" authorityYear="2022" baseAuthorityName="Jirapatrasilp & Sutcharit & Panha" baseAuthorityYear="2022" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina illustris" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="illustris" status="comb. nov.">Pupina illustris</taxonomicName>
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Mabille, 1887: 136, 137. Type locality: Tonkin.
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<bibRefCitation DOI="https://doi.org/10.5962/bhl.title.14809" author="Fischer, P" journalOrPublisher="Imprimerie Dejussieu Pere et Fils, Autun" pageId="0" pageNumber="1" refId="B41" refString="Fischer, P, 1891. Catalogue et distribution geographique des Mollusques terrestres, fluviatilies & marins ďune partie de ľIndo-Chine (Siam, Laos, Cambodge, Cochinchine, Annam, Tonkin). Imprimerie Dejussieu Pere et Fils, Autun, DOI: https://doi.org/10.5962/bhl.title.14809" title="Catalogue et distribution geographique des Mollusques terrestres, fluviatilies & marins ďune partie de ľIndo-Chine (Siam, Laos, Cambodge, Cochinchine, Annam, Tonkin)." url="https://doi.org/10.5962/bhl.title.14809" year="1891">Fischer 1891</bibRefCitation>
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: 107.
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<bibRefCitation author="Fischer, H" editor="Leroux, E" journalOrPublisher="E. Leroux, Paris" pageId="0" pageNumber="1" pagination="390 - 450" refId="B45" refString="Fischer, H, Dautzenberg, P, 1904. Catalogue des mollusques terrestres et fluviatiles de ľIndo-Chine orientale cites jusqu' a ce jour. In: Leroux, E, Ed., Mission Pavie Indo-Chine 1879-1895, Etudes diverses III Recherches sur l'histoire naturelle de l'Indo-Chine orientale. E. Leroux, Paris: 390 - 450" title="Catalogue des mollusques terrestres et fluviatiles de ľIndo-Chine orientale cites jusqu' a ce jour." volumeTitle="Mission Pavie Indo-Chine 1879 - 1895, Etudes diverses III Recherches sur l'histoire naturelle de l'Indo-Chine orientale." year="1904">Fischer and Dautzenberg 1904</bibRefCitation>
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: 431.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Jirapatrasilp & Sutcharit & Panha" authorityYear="2022" baseAuthorityName="Jirapatrasilp & Sutcharit & Panha" baseAuthorityYear="2022" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina tonkiniana" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tonkiniana" status="comb. nov.">Pupina tonkiniana</taxonomicName>
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Bavay & Dautzenberg, 1899: 54, 55, pl. 3, fig. 6, 6a (as
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<taxonomicName authorityName="Jirapatrasilp & Sutcharit & Panha" authorityYear="2022" baseAuthorityName="Jirapatrasilp & Sutcharit & Panha" baseAuthorityYear="2022" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina tonkiana" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tonkiana" status="comb. nov.">Pupina tonkiana</taxonomicName>
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in the original description). Type locality: Entre Lang-Son [Lang Son Province, Vietnam] et
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<normalizedToken originalValue="That-Khé">That-Khe</normalizedToken>
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[That Ke, Lang Son Province, Vietnam]. Syn. nov.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Jirapatrasilp & Sutcharit & Panha" authorityYear="2022" baseAuthorityName="Jirapatrasilp & Sutcharit & Panha" baseAuthorityYear="2022" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina (Tylotoechus) illustris" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="illustris" status="comb. nov." subGenus="Tylotoechus">Pupina (Tylotoechus) illustris</taxonomicName>
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-
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<bibRefCitation author="Kobelt, W" journalOrPublisher="Friedlaender und Sohn, Berlin" pageId="0" pageNumber="1" refId="B76" refString="Kobelt, W, 1902. Cyclophoridae. Das Tierreich. R. Friedlaender und Sohn, Berlin" title="Cyclophoridae. Das Tierreich. R." year="1902">Kobelt 1902</bibRefCitation>
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: 314, 315.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Jirapatrasilp & Sutcharit & Panha" authorityYear="2022" baseAuthorityName="Jirapatrasilp & Sutcharit & Panha" baseAuthorityYear="2022" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina (Tylotoechus) tonkiniana" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tonkiniana" status="comb. nov." subGenus="Tylotoechus">Pupina (Tylotoechus) tonkiniana</taxonomicName>
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-
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<bibRefCitation author="Kobelt, W" journalOrPublisher="Friedlaender und Sohn, Berlin" pageId="0" pageNumber="1" refId="B76" refString="Kobelt, W, 1902. Cyclophoridae. Das Tierreich. R. Friedlaender und Sohn, Berlin" title="Cyclophoridae. Das Tierreich. R." year="1902">Kobelt 1902</bibRefCitation>
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: 323, 324.
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<bibRefCitation author="Zilch, A" journalOrPublisher="Archiv fuer Molluskenkunde" pageId="0" pageNumber="1" pagination="41 - 56" refId="B198" refString="Zilch, A, 1957. Die Typen und Typoide des Natur-Museums Senckenberg, 19: Mollusca, Cyclophoridae, Pupininae. Archiv fuer Molluskenkunde 86: 41 - 56" title="Die Typen und Typoide des Natur-Museums Senckenberg, 19: Mollusca, Cyclophoridae, Pupininae." volume="86" year="1957">Zilch 1957</bibRefCitation>
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: 48.
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</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Jirapatrasilp & Sutcharit & Panha" authorityYear="2022" baseAuthorityName="Jirapatrasilp & Sutcharit & Panha" baseAuthorityYear="2022" class="Gastropoda" family="Pupinidae" genus="Pupina" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Pupina tonkiniana" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tonkiniana" status="comb. nov.">Pupina tonkiniana</taxonomicName>
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-
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<bibRefCitation author="Fischer, H" editor="Leroux, E" journalOrPublisher="E. Leroux, Paris" pageId="0" pageNumber="1" pagination="390 - 450" refId="B45" refString="Fischer, H, Dautzenberg, P, 1904. Catalogue des mollusques terrestres et fluviatiles de ľIndo-Chine orientale cites jusqu' a ce jour. In: Leroux, E, Ed., Mission Pavie Indo-Chine 1879-1895, Etudes diverses III Recherches sur l'histoire naturelle de l'Indo-Chine orientale. E. Leroux, Paris: 390 - 450" title="Catalogue des mollusques terrestres et fluviatiles de ľIndo-Chine orientale cites jusqu' a ce jour." volumeTitle="Mission Pavie Indo-Chine 1879 - 1895, Etudes diverses III Recherches sur l'histoire naturelle de l'Indo-Chine orientale." year="1904">Fischer and Dautzenberg 1904</bibRefCitation>
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: 432.
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<bibRefCitation author="Fischer-Piette, E" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" pagination="149 - 180" refId="B47" refString="Fischer-Piette, E, 1950. Liste des types decrits dans le Journal de Conchyliologie et conserves dans la collection de ce journal (avec planches) (suite). Journal de Conchyliologie 90: 149 - 180" title="Liste des types decrits dans le Journal de Conchyliologie et conserves dans la collection de ce journal (avec planches) (suite)." volume="90" year="1950">Fischer-Piette 1950</bibRefCitation>
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: 167.
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<bibRefCitation author="Do, DS" journalOrPublisher="Ruthenica" pageId="0" pageNumber="1" pagination="117 - 132" refId="B34" refString="Do, DS, Nguyen, THT, Do, VN, 2015. A checklist and classification of terrestrial prosobranch snails from Son La, north-western Vietnam. Ruthenica 25: 117 - 132" title="A checklist and classification of terrestrial prosobranch snails from Son La, north-western Vietnam." volume="25" year="2015">Do et al. 2015</bibRefCitation>
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: 126, fig. 6b, Son La Province, Vietnam.
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<bibRefCitation author="Raheem, DC" journalOrPublisher="The Natural History Museum, London, UK, the Royal Belgian Institute of Natural Sciences, Brussels, Belgium & the Zoological Society of London, UK" pageId="0" pageNumber="1" refId="B137" refString="Raheem, DC, Backeljau, T, Pearce-Kelly, P, Taylor, H, Fenn, J, Sutcharit, C, Panha, S, von, Oheimb KCM, von, Oheimb PV, Ikebe, C, Pall-Gergely, B, Gargominy, O, Hao, LV, Sang, PV, Tu, DV, Phong, DT, Naggs, M, Ablett, J, Dodds, JM, Wade, CM, Naggs, F, 2017. An Illustrated Guide to the Land Snails and Slugs of Vietnam. The Natural History Museum, London, UK, the Royal Belgian Institute of Natural Sciences, Brussels, Belgium & the Zoological Society of London, UK" title="An Illustrated Guide to the Land Snails and Slugs of Vietnam." year="2017">Raheem et al. 2017</bibRefCitation>
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: 5 (plate figure).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type material">
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<paragraph pageId="0" pageNumber="1">Type material examined.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Syntypes</emphasis>
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of
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<taxonomicName authorityName="Jirapatrasilp & Sutcharit & Panha" authorityYear="2022" baseAuthorityName="Jirapatrasilp & Sutcharit & Panha" baseAuthorityYear="2022" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina illustris" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="illustris">
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<emphasis italics="true" pageId="0" pageNumber="1">Pupina illustris</emphasis>
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</taxonomicName>
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MNHN-IM-2000-35842 (9 shells; Figs
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Shells of Pupinella species from mainland Southeast Asia. A-G Pupinella mansuyi: A syntype of Eupupina mansuyi MNHN-IM- 2000 - 30756 from Deux-Ponts B syntype of Eupupina mansuyi MNHN-IM- 2000 - 36067 from Deux-Ponts C syntype of Eupupina mansuyi MNHN-IM- 2000 - 36068 from Quang-Huyen D syntype of Eupupina mansuyi RBINS MT 970 / 1 from Quang-Huyen E holotype of Pupinella frednaggsi NHMUK 20170285 F specimen CUMZ 12148 from Pha Chu Mount, Nan, and G specimen CUMZ 12149 from Pha Tub Cave, Nan H Pupinella sonlaensis, paratype ZRC. MOL. 9377 I-L Pupinella illustris I, J syntypes of Pupina illustris MNHN-IM- 2000 - 35842 from Tonkin K lectotype of Pupina tonkiniana MNHN-IM- 2000 - 35838 from Lang-Son et That-Khe, and L paralectotype of Pupina tonkiniana SMF 109932 / 10 from Tonkin: That-khe. Photo: A. Lardeur, P. Maestrati, MNHN (A-C, I-K), F. Trus, RBINS (D), S. K. Tan, ZRC (H)." figureDoi="10.3897/zookeys.1119.85400.figure11" httpUri="https://binary.pensoft.net/fig/735372" pageId="0" pageNumber="1">11I, J</figureCitation>
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,
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. A-C Pupinella mansuyi: A syntype of Eupupina mansuyi RBINS MT 970 / 1 from Quang-Huyen B specimen CUMZ 12148 from Pha Chu Mount, Nan, and C specimen CUMZ 12149 from Pha Tub Cave, Nan. D, E Pupinella illustris D syntype of Pupina illustris MNHN-IM- 2000 - 35842 from Tonkin and E paralectotype of Pupina tonkiniana SMF 109932 / 10 from Tonkin: That-khe F Pupinella sonlaensis, paratype ZRC. MOL. 9377. Photo: F. Trus, RBINS (A), P. Maestrati, MNHN (D), S. K. Tan, ZRC (F)." figureDoi="10.3897/zookeys.1119.85400.figure12" httpUri="https://binary.pensoft.net/fig/735373" pageId="0" pageNumber="1">12D</figureCitation>
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) from Tonkin.
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Lectotype</emphasis>
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of
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<taxonomicName authorityName="Jirapatrasilp & Sutcharit & Panha" authorityYear="2022" baseAuthorityName="Jirapatrasilp & Sutcharit & Panha" baseAuthorityYear="2022" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina tonkiniana" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tonkiniana">
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<emphasis italics="true" pageId="0" pageNumber="1">Pupina tonkiniana</emphasis>
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</taxonomicName>
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MNHN-IM-2000-35838 (Fig.
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Shells of Pupinella species from mainland Southeast Asia. A-G Pupinella mansuyi: A syntype of Eupupina mansuyi MNHN-IM- 2000 - 30756 from Deux-Ponts B syntype of Eupupina mansuyi MNHN-IM- 2000 - 36067 from Deux-Ponts C syntype of Eupupina mansuyi MNHN-IM- 2000 - 36068 from Quang-Huyen D syntype of Eupupina mansuyi RBINS MT 970 / 1 from Quang-Huyen E holotype of Pupinella frednaggsi NHMUK 20170285 F specimen CUMZ 12148 from Pha Chu Mount, Nan, and G specimen CUMZ 12149 from Pha Tub Cave, Nan H Pupinella sonlaensis, paratype ZRC. MOL. 9377 I-L Pupinella illustris I, J syntypes of Pupina illustris MNHN-IM- 2000 - 35842 from Tonkin K lectotype of Pupina tonkiniana MNHN-IM- 2000 - 35838 from Lang-Son et That-Khe, and L paralectotype of Pupina tonkiniana SMF 109932 / 10 from Tonkin: That-khe. Photo: A. Lardeur, P. Maestrati, MNHN (A-C, I-K), F. Trus, RBINS (D), S. K. Tan, ZRC (H)." figureDoi="10.3897/zookeys.1119.85400.figure11" httpUri="https://binary.pensoft.net/fig/735372" pageId="0" pageNumber="1">11K</figureCitation>
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) from Lang-Son et
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<normalizedToken originalValue="That-Khé">That-Khe</normalizedToken>
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. Paralectotypes of
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<taxonomicName authorityName="Jirapatrasilp & Sutcharit & Panha" authorityYear="2022" baseAuthorityName="Jirapatrasilp & Sutcharit & Panha" baseAuthorityYear="2022" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina tonkiniana" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tonkiniana">
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<emphasis italics="true" pageId="0" pageNumber="1">Pupina tonkiniana</emphasis>
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</taxonomicName>
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SMF 109932/10 (10 shells; Figs
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Shells of Pupinella species from mainland Southeast Asia. A-G Pupinella mansuyi: A syntype of Eupupina mansuyi MNHN-IM- 2000 - 30756 from Deux-Ponts B syntype of Eupupina mansuyi MNHN-IM- 2000 - 36067 from Deux-Ponts C syntype of Eupupina mansuyi MNHN-IM- 2000 - 36068 from Quang-Huyen D syntype of Eupupina mansuyi RBINS MT 970 / 1 from Quang-Huyen E holotype of Pupinella frednaggsi NHMUK 20170285 F specimen CUMZ 12148 from Pha Chu Mount, Nan, and G specimen CUMZ 12149 from Pha Tub Cave, Nan H Pupinella sonlaensis, paratype ZRC. MOL. 9377 I-L Pupinella illustris I, J syntypes of Pupina illustris MNHN-IM- 2000 - 35842 from Tonkin K lectotype of Pupina tonkiniana MNHN-IM- 2000 - 35838 from Lang-Son et That-Khe, and L paralectotype of Pupina tonkiniana SMF 109932 / 10 from Tonkin: That-khe. Photo: A. Lardeur, P. Maestrati, MNHN (A-C, I-K), F. Trus, RBINS (D), S. K. Tan, ZRC (H)." figureDoi="10.3897/zookeys.1119.85400.figure11" httpUri="https://binary.pensoft.net/fig/735372" pageId="0" pageNumber="1">11L</figureCitation>
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,
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. A-C Pupinella mansuyi: A syntype of Eupupina mansuyi RBINS MT 970 / 1 from Quang-Huyen B specimen CUMZ 12148 from Pha Chu Mount, Nan, and C specimen CUMZ 12149 from Pha Tub Cave, Nan. D, E Pupinella illustris D syntype of Pupina illustris MNHN-IM- 2000 - 35842 from Tonkin and E paralectotype of Pupina tonkiniana SMF 109932 / 10 from Tonkin: That-khe F Pupinella sonlaensis, paratype ZRC. MOL. 9377. Photo: F. Trus, RBINS (A), P. Maestrati, MNHN (D), S. K. Tan, ZRC (F)." figureDoi="10.3897/zookeys.1119.85400.figure12" httpUri="https://binary.pensoft.net/fig/735373" pageId="0" pageNumber="1">12E</figureCitation>
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) from Tonkin:
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<normalizedToken originalValue="That-khé">That-khe</normalizedToken>
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. Paralectotypes of
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<taxonomicName authorityName="Jirapatrasilp & Sutcharit & Panha" authorityYear="2022" baseAuthorityName="Jirapatrasilp & Sutcharit & Panha" baseAuthorityYear="2022" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina tonkiniana" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tonkiniana">
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<emphasis italics="true" pageId="0" pageNumber="1">Pupina tonkiniana</emphasis>
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</taxonomicName>
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RBINS MT976/2 (14 shells) from Lang Son et
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<normalizedToken originalValue="That-khé">That-khe</normalizedToken>
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.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">Shell elongate fusiform; last whorl ca. 55-60% of shell height. Apertural lip expanded and slightly reflected; apertural lip when observed from lateral view almost straight. Parietal callus absent. Parietal tooth pointily sharp, located next to wide posterior canal. Anterior canal funnel-like. Umbilicus closed.</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="differential diagnosis">
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<paragraph pageId="0" pageNumber="1">Differential diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName authorityName="Jirapatrasilp & Sutcharit & Panha" authorityYear="2022" baseAuthorityName="Mabille" baseAuthorityYear="1887" class="Gastropoda" family="Pupinidae" genus="Pupinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pupinella illustris" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="illustris">
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<emphasis italics="true" pageId="0" pageNumber="1">Pupinella illustris</emphasis>
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</taxonomicName>
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can be distinguished from all other species in mainland Southeast Asia by an elongate fusiform shell shape, an absence of parietal callus and a pointily sharp parietal tooth located next to a wide posterior canal.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="distribution">
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<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Northern Vietnam (
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<bibRefCitation author="Do, DS" journalOrPublisher="Ruthenica" pageId="0" pageNumber="1" pagination="117 - 132" refId="B34" refString="Do, DS, Nguyen, THT, Do, VN, 2015. A checklist and classification of terrestrial prosobranch snails from Son La, north-western Vietnam. Ruthenica 25: 117 - 132" title="A checklist and classification of terrestrial prosobranch snails from Son La, north-western Vietnam." volume="25" year="2015">Do et al. 2015</bibRefCitation>
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;
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<bibRefCitation author="Raheem, DC" journalOrPublisher="The Natural History Museum, London, UK, the Royal Belgian Institute of Natural Sciences, Brussels, Belgium & the Zoological Society of London, UK" pageId="0" pageNumber="1" refId="B137" refString="Raheem, DC, Backeljau, T, Pearce-Kelly, P, Taylor, H, Fenn, J, Sutcharit, C, Panha, S, von, Oheimb KCM, von, Oheimb PV, Ikebe, C, Pall-Gergely, B, Gargominy, O, Hao, LV, Sang, PV, Tu, DV, Phong, DT, Naggs, M, Ablett, J, Dodds, JM, Wade, CM, Naggs, F, 2017. An Illustrated Guide to the Land Snails and Slugs of Vietnam. The Natural History Museum, London, UK, the Royal Belgian Institute of Natural Sciences, Brussels, Belgium & the Zoological Society of London, UK" title="An Illustrated Guide to the Land Snails and Slugs of Vietnam." year="2017">Raheem et al. 2017</bibRefCitation>
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).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="remarks">
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<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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This taxon is allocated to the genus
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<taxonomicName authorityName="Gray" authorityYear="1850" class="Gastropoda" family="Pupinidae" genus="Pupinella" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pupinella" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
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<emphasis italics="true" pageId="0" pageNumber="1">Pupinella</emphasis>
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</taxonomicName>
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due to the presence of a funnel-like anterior canal, which is the diagnostic character of this genus. In the original description of
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<taxonomicName authorityName="Jirapatrasilp & Sutcharit & Panha" authorityYear="2022" baseAuthorityName="Jirapatrasilp & Sutcharit & Panha" baseAuthorityYear="2022" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina tonkiniana" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tonkiniana">
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<emphasis italics="true" pageId="0" pageNumber="1">Pupina tonkiniana</emphasis>
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</taxonomicName>
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|
, two ways of spelling were shown: the spelling '
|
|
<taxonomicName authorityName="Jirapatrasilp & Sutcharit & Panha" authorityYear="2022" baseAuthorityName="Jirapatrasilp & Sutcharit & Panha" baseAuthorityYear="2022" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina tonkiana" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tonkiana">
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<emphasis italics="true" pageId="0" pageNumber="1">Pupina tonkiana</emphasis>
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</taxonomicName>
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' in the description, and '
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<taxonomicName authorityName="Jirapatrasilp & Sutcharit & Panha" authorityYear="2022" baseAuthorityName="Jirapatrasilp & Sutcharit & Panha" baseAuthorityYear="2022" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina tonkiniana" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tonkiniana">
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<emphasis italics="true" pageId="0" pageNumber="1">Pupina tonkiniana</emphasis>
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</taxonomicName>
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|
' in the plate caption. Later,
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<bibRefCitation author="Kobelt, W" journalOrPublisher="Friedlaender und Sohn, Berlin" pageId="0" pageNumber="1" refId="B76" refString="Kobelt, W, 1902. Cyclophoridae. Das Tierreich. R. Friedlaender und Sohn, Berlin" title="Cyclophoridae. Das Tierreich. R." year="1902">Kobelt (1902)</bibRefCitation>
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acted as the First Reviser (
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|
<bibRefCitation author="Hwang, C-C" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" refId="B70" refString="ICZN (1999) International Code of Zoological Nomenclature, 4th Edn.International Trust for Zoological Nomenclature, London, 306 pp." year="2014">ICZN 1999</bibRefCitation>
|
|
: Art. 24.2.3) in selecting '
|
|
<taxonomicName authorityName="Jirapatrasilp & Sutcharit & Panha" authorityYear="2022" baseAuthorityName="Jirapatrasilp & Sutcharit & Panha" baseAuthorityYear="2022" class="Gastropoda" family="Pupinidae" genus="Pupina" kingdom="Animalia" lsidName="Pupina tonkiniana" order="Architaenioglossa" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="tonkiniana">
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|
<emphasis italics="true" pageId="0" pageNumber="1">Pupina tonkiniana</emphasis>
|
|
</taxonomicName>
|
|
' as the correct original spelling. As the original description did not explicitly state that the description of
|
|
<taxonomicName class="Gastropoda" family="Pupinidae" kingdom="Animalia" lsidName="P. tonkiniana" order="Architaenioglossa" pageId="0" pageNumber="1" rank="species" species="tonkiniana">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. tonkiniana</emphasis>
|
|
</taxonomicName>
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|
was based on a single specimen (nor could this be inferred), the designation of a holotype by
|
|
<bibRefCitation author="Fischer-Piette, E" journalOrPublisher="Journal de Conchyliologie" pageId="0" pageNumber="1" pagination="149 - 180" refId="B47" refString="Fischer-Piette, E, 1950. Liste des types decrits dans le Journal de Conchyliologie et conserves dans la collection de ce journal (avec planches) (suite). Journal de Conchyliologie 90: 149 - 180" title="Liste des types decrits dans le Journal de Conchyliologie et conserves dans la collection de ce journal (avec planches) (suite)." volume="90" year="1950">Fischer-Piette (1950)</bibRefCitation>
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|
in fact constitutes a lectotype designation (
|
|
<bibRefCitation author="Hwang, C-C" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" refId="B70" refString="ICZN (1999) International Code of Zoological Nomenclature, 4th Edn.International Trust for Zoological Nomenclature, London, 306 pp." year="2014">ICZN 1999</bibRefCitation>
|
|
: Art. 74.6).
|
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</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
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|
Upon examining the type specimens of both
|
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<taxonomicName class="Gastropoda" family="Pupinidae" kingdom="Animalia" lsidName="P. illustris" order="Architaenioglossa" pageId="0" pageNumber="1" rank="species" species="illustris">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. illustris</emphasis>
|
|
</taxonomicName>
|
|
and
|
|
<taxonomicName class="Gastropoda" family="Pupinidae" kingdom="Animalia" lsidName="P. tonkiniana" order="Architaenioglossa" pageId="0" pageNumber="1" rank="species" species="tonkiniana">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. tonkiniana</emphasis>
|
|
</taxonomicName>
|
|
, the type series of
|
|
<taxonomicName class="Gastropoda" family="Pupinidae" kingdom="Animalia" lsidName="P. tonkiniana" order="Architaenioglossa" pageId="0" pageNumber="1" rank="species" species="tonkiniana">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. tonkiniana</emphasis>
|
|
</taxonomicName>
|
|
agree well with all the syntypes of
|
|
<taxonomicName class="Gastropoda" family="Pupinidae" kingdom="Animalia" lsidName="P. illustris" order="Architaenioglossa" pageId="0" pageNumber="1" rank="species" species="illustris">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. illustris</emphasis>
|
|
</taxonomicName>
|
|
in having an elongate fusiform shell shape, an expanded and slightly reflected apertural lip without a parietal callus, and a sharp, tooth-like, parietal tooth located next to a wide posterior canal. Thus,
|
|
<taxonomicName class="Gastropoda" family="Pupinidae" kingdom="Animalia" lsidName="P. tonkiniana" order="Architaenioglossa" pageId="0" pageNumber="1" rank="species" species="tonkiniana">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. tonkiniana</emphasis>
|
|
</taxonomicName>
|
|
is regarded herein as a junior subjective synonym of
|
|
<taxonomicName class="Gastropoda" family="Pupinidae" kingdom="Animalia" lsidName="P. illustris" order="Architaenioglossa" pageId="0" pageNumber="1" rank="species" species="illustris">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. illustris</emphasis>
|
|
</taxonomicName>
|
|
.
|
|
</paragraph>
|
|
</subSubSection>
|
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</treatment>
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</document> |