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<document id="E70FEF4B0D4DDE6E17FAC167BE8B8B49" ID-CLB-Dataset="6390" ID-DOI="10.5252/g2014n4a4" ID-GBIF-Dataset="8bc26961-9853-4113-a63d-4cf691aa7100" ID-ISSN="1638-9395" ID-Zenodo-Dep="4538625" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1613150850454" checkinUser="felipe" docAuthor="Vianey-Liaud, Monique, Comte, Bernard, Marandat, Bernard, Peigné, Stéphane, Rage, Jean-Claude &amp; Sudre, Jean" docDate="2014" docId="03D6987B4450B10FFF4F15C4FD04E3AC" docLanguage="en" docName="Geodiversitas.36.4.565-622.pdf" docOrigin="Geodiversitas 36 (4)" docSource="http://dx.doi.org/10.5252/g2014n4a4" docStyle="DocumentStyle:F8AA0CB5AB58FB1C1EB3E4696C1B2D36.3:Geodiversitas.2013-2014.journal_article" docStyleId="F8AA0CB5AB58FB1C1EB3E4696C1B2D36" docStyleName="Geodiversitas.2013-2014.journal_article" docStyleVersion="3" docTitle="Heterocricetodon gaimersheimensis Freudenberg 1941" docType="treatment" docVersion="7" lastPageNumber="600" masterDocId="FFEFE0034471B12CFF8F106FFF81E50F" masterDocTitle="A new early Late Oligocene (MP 26) continental vertebrate fauna from Saint-Privat-des-Vieux (Alès Basin, Gard, Southern France)" masterLastPageNumber="622" masterPageNumber="565" pageNumber="598" updateTime="1698929460103" updateUser="ExternalLinkService" zenodo-license-document="CC0-1.0" zenodo-license-figures="CC0-1.0">
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<mods:title id="DB32391272542775AE0C4B5A3DF79417">A new early Late Oligocene (MP 26) continental vertebrate fauna from Saint-Privat-des-Vieux (Alès Basin, Gard, Southern France)</mods:title>
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<treatment id="03D6987B4450B10FFF4F15C4FD04E3AC" ID-DOI="http://doi.org/10.5281/zenodo.4837402" ID-GBIF-Taxon="182405889" ID-Zenodo-Dep="4837402" LSID="urn:lsid:plazi:treatment:03D6987B4450B10FFF4F15C4FD04E3AC" httpUri="http://treatment.plazi.org/id/03D6987B4450B10FFF4F15C4FD04E3AC" lastPageId="35" lastPageNumber="600" pageId="33" pageNumber="598">
<subSubSection id="C3657AE64450B10DFF4F15C4FE62E0EA" pageId="33" pageNumber="598" type="nomenclature">
<paragraph id="8BC0296D4450B10DFF4F15C4FE62E0EA" blockId="33.[192,569,1451,1509]" pageId="33" pageNumber="598">
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<taxonomicName id="4C7F52EE4450B10DFF4F15C4FE62E0EA" authority="Freudenberg, 1941" authorityName="Freudenberg" authorityYear="1941" class="Mammalia" family="Cricetidae" genus="Heterocricetodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="33" pageNumber="598" phylum="Chordata" rank="species" species="gaimersheimensis">
<emphasis id="B90BF57F4450B10DFF4F15C4FDB8E0C9" bold="true" box="[192,569,1451,1478]" italics="true" pageId="33" pageNumber="598">Heterocricetodon gaimersheimensis</emphasis>
<bibRefCitation id="EFEE549C4450B10DFE9B15A4FE62E0EA" author="FREUDENBERG H." box="[276,483,1483,1509]" pageId="33" pageNumber="598" pagination="99 - 164" refId="ref32684" refString="FREUDENBERG H. 1941. - Die oberoligozanen Nager von Gaimesrsheim bei Ingolstadt und ihre Verwandten. Palaeontographica Abteilung A 92: 99 - 164." type="journal article" year="1941">Freudenberg, 1941</bibRefCitation>
</taxonomicName>
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</paragraph>
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<subSubSection id="C3657AE64450B10DFEC21583FE2BE309" box="[333,426,1516,1542]" pageId="33" pageNumber="598" type="description">
<paragraph id="8BC0296D4450B10DFEC21583FE2BE309" blockId="33.[333,426,1516,1542]" box="[333,426,1516,1542]" pageId="33" pageNumber="598">
(
<figureCitation id="134435E84450B10DFED91583FE22E309" box="[342,419,1516,1542]" captionStart="FIG" captionStartId="35.[113,124,1274,1292]" captionTargetBox="[114,1210,216,1233]" captionTargetId="figure-221@35.[661,936,502,779]" captionTargetPageId="35" captionText="FIG. 12. — Heterocricetodon gaimersheimensis Freudenberg,1941, from Saint-Privat-des-Vieux (Gard;Lower Chattian,MP 26): A, right M1 (reversed), SPV170: A1, occlusal view, A2, lingual view; B, left M2, SPV171: B1, occlusal view, B2, lingual view; C, right M3 (re- versed), SPV174: C1, occlusal view, C2, lingual view; D, right m1 (reversed), SPV175: D1, occlusal view, D2, lingual view; E, left m2, SPV178, occlusal view; F, left m3, SPV179: F1, occlusal view, F2, lingual view. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.4538653" httpUri="https://zenodo.org/record/4538653/files/figure.png" pageId="33" pageNumber="598">Fig. 12</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C3657AE64450B10DFFFE1649FC12E42B" pageId="33" pageNumber="598" type="materials_examined">
<paragraph id="8BC0296D4450B10DFFFE1649FE74E358" blockId="33.[113,646,1573,1624]" pageId="33" pageNumber="598">
<materialsCitation id="3B1723304450B10DFFFE1649FE74E358" ID-GBIF-Occurrence="3122035308" collectingDate="1939-11-04" collectionCode="BSPG" pageId="33" pageNumber="598" specimenCount="1" typeStatus="holotype">
<typeStatus id="54C497CF4450B10DFFFE1649FF5CE333" box="[113,221,1574,1597]" pageId="33" pageNumber="598" type="holotype">HOLOTYPE</typeStatus>
. — Fragment of right lower jaw, bearing m1-3, BSP
<date id="FFC10FAD4450B10DFF6A162FFECDE357" box="[229,332,1600,1624]" pageId="33" pageNumber="598" value="1939-11-04">
<collectingDate id="EF85F6454450B10DFF6A162FFECDE357" box="[229,332,1600,1624]" pageId="33" pageNumber="598" value="1939-11-04">1939 XI 4</collectingDate>
</date>
,
<collectionCode id="ED6EB1A84450B10DFED8162FFE17E357" box="[343,406,1600,1624]" country="0" httpUri="http://grbio.org/cool/r9n2-w98n" name="Bayerische Staatssammlung fuer Palaeontologie und Geologie" pageId="33" pageNumber="598">BSPG</collectionCode>
, Munich.
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</paragraph>
<paragraph id="8BC0296D4450B10DFFFE161EFF61E3AC" blockId="33.[113,646,1649,1699]" pageId="33" pageNumber="598">
<collectionCode id="ED6EB1A84450B10DFFFE161EFE8BE388" box="[113,266,1649,1672]" pageId="33" pageNumber="598">TYPE LOCALITY</collectionCode>
. — Gaimersheim (South
<collectingCountry id="F36869FD4450B10DFD96161EFDFDE386" box="[537,636,1649,1673]" name="Germany" pageId="33" pageNumber="598">Germany</collectingCountry>
),
<collectionCode id="ED6EB1A84450B10DFFFC16E3FF19E3AC" box="[115,152,1676,1699]" country="0" name="Mohonk Preserve, Inc." pageId="33" pageNumber="598">MP</collectionCode>
27-28.
</paragraph>
<paragraph id="8BC0296D4450B10DFD2A10B7FC12E42B" blockId="33.[675,1211,215,292]" pageId="33" pageNumber="598">
ORIGINAL DIAGNOSIS. — See
<bibRefCitation id="EFEE549C4450B10DFC3010B8FB37E5E0" author="FREUDENBERG H." box="[959,1206,215,239]" pageId="33" pageNumber="598" pagination="99 - 164" refId="ref32684" refString="FREUDENBERG H. 1941. - Die oberoligozanen Nager von Gaimesrsheim bei Ingolstadt und ihre Verwandten. Palaeontographica Abteilung A 92: 99 - 164." type="journal article" year="1941">Freudenberg (1941: 105)</bibRefCitation>
. Description and variability of teeth of the type-population:
<bibRefCitation id="EFEE549C4450B10DFD2C1163FC0EE42B" author="KRISTKOIZ A." box="[675,911,268,292]" pageId="33" pageNumber="598" pagination="1 - 137" refId="ref33559" refString="KRISTKOIZ A. 1992. - Zahnmorphologische und schadelanatomische Untersuchungen an Nagertieren aus dem Oberoligozan von Gaimersheim (Suddeutschland). Abhandlungen / Bayerische Akademie derWissenschaften, Mathematische-Naturwissenschaftliche Klasse 167: 1 - 137." type="journal article" year="1992">Kristkoiz (1992: 63-72)</bibRefCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="C3657AE64450B10DFD2A1153FC20E187" pageId="33" pageNumber="598" type="description">
<paragraph id="8BC0296D4450B10DFD2A1153FB38E45B" blockId="33.[664,1210,316,419]" box="[677,1209,316,341]" pageId="33" pageNumber="598">MEASUREMENTS (in mm). — Eight isolated teeth: 1 M1</paragraph>
<paragraph id="8BC0296D4450B10DFD171139FC76E4AC" blockId="33.[664,1210,316,419]" pageId="33" pageNumber="598">
(2.54 × 1.66), 3 M2 (2.12 × 1.94, 2.17 × 1.86, 1.94 × 1.79), 1 M3 (1.72 × 1.54),
<quantity id="4C8784884450B10DFC38111EFC62E486" box="[951,995,369,393]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="33" pageNumber="598" unit="m" value="1.0">1 m</quantity>
1 (2.65 × 1.60),
<quantity id="4C8784884450B10DFB0A111EFB2FE486" box="[1157,1198,369,393]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="33" pageNumber="598" unit="m" value="1.0">1 m</quantity>
2 (2.54 × 1.47),
<quantity id="4C8784884450B10DFCBC11E4FCDDE4AC" box="[819,860,395,419]" metricMagnitude="0" metricUnit="m" metricValue="1.0" pageId="33" pageNumber="598" unit="m" value="1.0">1 m</quantity>
3 (2.37 × 1.78).
</paragraph>
<paragraph id="8BC0296D4450B10DFD2A11B9FCB4E4E1" blockId="33.[677,1211,470,590]" box="[677,821,470,496]" pageId="33" pageNumber="598">DESCRIPTION</paragraph>
<paragraph id="8BC0296D4450B10DFD2A119AFBBDE741" blockId="33.[677,1211,470,590]" pageId="33" pageNumber="598">
<emphasis id="B90BF57F4450B10DFD2A119AFD55E700" bold="true" box="[677,724,501,527]" pageId="33" pageNumber="598">M1.</emphasis>
Ŋe anterior arm of the protocone and the mesoloph join the labial edge of the tooth, like on several specimens from Gaimersheim.
</paragraph>
<paragraph id="8BC0296D4450B10DFD2A121BFCC3E603" blockId="33.[677,1210,627,780]" pageId="33" pageNumber="598">
<emphasis id="B90BF57F4450B10DFD2A121BFC8BE782" bold="true" box="[677,778,627,654]" pageId="33" pageNumber="598">M2-M3.</emphasis>
Ŋere is a double connecting of the anterior cusps (complete metalophule I and II) and the mesoloph reaches the labial edge of the teeth, as it happens on some specimens from Gaimersheim.
</paragraph>
<paragraph id="8BC0296D4450B10DFD2A135DFC78E664" blockId="33.[677,1209,818,876]" pageId="33" pageNumber="598">
<emphasis id="B90BF57F4450B10DFD2A135DFD50E643" bold="true" box="[677,721,818,844]" pageId="33" pageNumber="598">m1.</emphasis>
Ŋe two mesolophids are short, like it is often the case in the
<typeStatus id="54C497CF4450B10DFCCA133DFCF2E663" box="[837,883,850,876]" pageId="33" pageNumber="598">type</typeStatus>
population.
</paragraph>
<paragraph id="8BC0296D4450B10DFD2A13FEFCBFE105" blockId="33.[677,1210,913,1034]" pageId="33" pageNumber="598">
<emphasis id="B90BF57F4450B10DFD2A13FEFD51E6A4" bold="true" box="[677,720,913,939]" pageId="33" pageNumber="598">m2.</emphasis>
Ŋe posterior arm of the protoconid is short but exceeds the mesolophid length, extending towards the metaconid, as for some specimens from Gaimersheim.
</paragraph>
<paragraph id="8BC0296D4450B10DFD2A1440FC20E187" blockId="33.[677,1210,1071,1160]" pageId="33" pageNumber="598">
<emphasis id="B90BF57F4450B10DFD2A1440FD51E146" bold="true" box="[677,720,1071,1097]" pageId="33" pageNumber="598">m3.</emphasis>
Ŋe posterior arm of the protoconid is long and joins the labial edge of the tooth, at the base of the metaconid ridge.
</paragraph>
</subSubSection>
<subSubSection id="C3657AE64450B10FFD2A14C1FD04E3AC" lastPageId="35" lastPageNumber="600" pageId="33" pageNumber="598" type="discussion">
<paragraph id="8BC0296D4450B10DFD2A14C1FCA7E1C9" blockId="33.[675,1210,1198,1699]" box="[677,806,1198,1224]" pageId="33" pageNumber="598">DISCUSSION</paragraph>
<paragraph id="8BC0296D4450B10EFD2A14A1FCE0E3AB" blockId="33.[675,1210,1198,1699]" lastBlockId="34.[675,1209,1546,1700]" lastPageId="34" lastPageNumber="599" pageId="33" pageNumber="598">
Four species of relatively large size have been referred to this genus. Among them,
<taxonomicName id="4C7F52EE4450B10DFBC91481FCC0E027" authority="Schaub, 1925" authorityName="Schaub" authorityYear="1925" class="Mammalia" family="Cricetidae" genus="Heterocricetodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="33" pageNumber="598" phylum="Chordata" rank="species" species="stehlini">
<emphasis id="B90BF57F4450B10DFBC91481FB38E008" box="[1094,1209,1261,1287]" italics="true" pageId="33" pageNumber="598">H. stehlini</emphasis>
<bibRefCitation id="EFEE549C4450B10DFD2A1562FCC0E027" author="SCHAUB S." box="[677,833,1293,1320]" pageId="33" pageNumber="598" pagination="1 - 114" refId="ref34764" refString="SCHAUB S. 1925. - Die hamsterartigen Nagetiere des Tertiars und ihre lebenden Verwandten. Abhandlungen der Schweizerischen Palaeontologischen Gesellschaft 45: 1 - 114." type="journal article" year="1925">Schaub, 1925</bibRefCitation>
</taxonomicName>
,
<taxonomicName id="4C7F52EE4450B10DFCC01561FB09E027" authority="(Schaub, 1925)" baseAuthorityName="Schaub" baseAuthorityYear="1925" box="[847,1160,1293,1320]" class="Mammalia" family="Cricetidae" genus="Heterocricetodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="33" pageNumber="598" phylum="Chordata" rank="species" species="schlosseri">
<emphasis id="B90BF57F4450B10DFCC01561FC52E028" box="[847,979,1293,1319]" italics="true" pageId="33" pageNumber="598">H. schlosseri</emphasis>
(
<bibRefCitation id="EFEE549C4450B10DFC6D1562FB01E027" author="SCHAUB S." box="[994,1152,1293,1320]" pageId="33" pageNumber="598" pagination="1 - 114" refId="ref34764" refString="SCHAUB S. 1925. - Die hamsterartigen Nagetiere des Tertiars und ihre lebenden Verwandten. Abhandlungen der Schweizerischen Palaeontologischen Gesellschaft 45: 1 - 114." type="journal article" year="1925">Schaub, 1925</bibRefCitation>
)
</taxonomicName>
and
<taxonomicName id="4C7F52EE4450B10DFD2A1542FBD8E048" authority="Stehlin &amp; Schaub, 1951" authorityName="Stehlin &amp; Schaub" authorityYear="1951" box="[677,1113,1324,1351]" class="Mammalia" family="Cricetidae" genus="Heterocricetodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="33" pageNumber="598" phylum="Chordata" rank="species" species="helbingi">
<emphasis id="B90BF57F4450B10DFD2A1542FCA9E048" box="[677,808,1325,1351]" italics="true" pageId="33" pageNumber="598">H. helbingi</emphasis>
Stehlin &amp; Schaub, 1951
</taxonomicName>
, remain poorly defined.
<bibRefCitation id="EFEE549C4450B10DFCDC1522FB9DE069" author="HUGUENEY M." box="[851,1052,1356,1383]" pageId="33" pageNumber="598" pagination="57 - 72" refId="ref33196" refString="HUGUENEY M. 1980. - La faune de mammiferes de l'Oligocene moyen de Saint-Menoux (Allier). Revue Scientifique du Bourbonnais et du Centre de la France 1 re partie: Rongeurs: 57 - 72." type="book chapter" year="1980">Hugueney (1980)</bibRefCitation>
, in her discussion about the species of
<taxonomicName id="4C7F52EE4450B10DFC491503FBFBE089" authorityName="Schaub" authorityYear="1925" box="[966,1146,1388,1414]" class="Mammalia" family="Cricetidae" genus="Heterocricetodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="33" pageNumber="598" phylum="Chordata" rank="genus">
<emphasis id="B90BF57F4450B10DFC491503FBFBE089" box="[966,1146,1388,1414]" italics="true" pageId="33" pageNumber="598">Heterocricetodon</emphasis>
</taxonomicName>
from Saint-Menoux, has suggested that the variations between the three species
<taxonomicName id="4C7F52EE4450B10DFC4515C4FB99E0CA" authorityName="Schaub" authorityYear="1925" box="[970,1048,1451,1477]" class="Mammalia" family="Cricetidae" genus="Heterocricetodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="33" pageNumber="598" phylum="Chordata" rank="species" species="stehlini">
<emphasis id="B90BF57F4450B10DFC4515C4FB99E0CA" box="[970,1048,1451,1477]" italics="true" pageId="33" pageNumber="598">stehlini</emphasis>
</taxonomicName>
,
<taxonomicName id="4C7F52EE4450B10DFBA615C4FB09E0CA" baseAuthorityName="Schaub" baseAuthorityYear="1925" box="[1065,1160,1451,1477]" class="Mammalia" family="Cricetidae" genus="Heterocricetodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="33" pageNumber="598" phylum="Chordata" rank="species" species="schlosseri">
<emphasis id="B90BF57F4450B10DFBA615C4FB09E0CA" box="[1065,1160,1451,1477]" italics="true" pageId="33" pageNumber="598">schlosseri</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7F52EE4450B10DFD2915A4FD7DE0EA" authorityName="Stehlin &amp; Schaub" authorityYear="1951" box="[678,764,1483,1509]" class="Mammalia" family="Cricetidae" genus="Heterocricetodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="33" pageNumber="598" phylum="Chordata" rank="species" species="helbingi">
<emphasis id="B90BF57F4450B10DFD2915A4FD7DE0EA" box="[678,764,1483,1509]" italics="true" pageId="33" pageNumber="598">helbingi</emphasis>
</taxonomicName>
could enter a unique species, when more material would be known. Concerning
<taxonomicName id="4C7F52EE4450B10DFBD71584FC7FE32A" authority="Freudenberg, 1941" authorityName="Freudenberg" authorityYear="1941" class="Mammalia" family="Cricetidae" genus="Heterocricetodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="33" pageNumber="598" phylum="Chordata" rank="species" species="gaimersheimensis">
<emphasis id="B90BF57F4450B10DFBD71584FCA4E32B" italics="true" pageId="33" pageNumber="598">H. gaimersheimensis</emphasis>
<bibRefCitation id="EFEE549C4450B10DFCA51665FC7FE32A" author="FREUDENBERG H." box="[810,1022,1546,1573]" pageId="33" pageNumber="598" pagination="99 - 164" refId="ref32684" refString="FREUDENBERG H. 1941. - Die oberoligozanen Nager von Gaimesrsheim bei Ingolstadt und ihre Verwandten. Palaeontographica Abteilung A 92: 99 - 164." type="journal article" year="1941">Freudenberg, 1941</bibRefCitation>
</taxonomicName>
,
<bibRefCitation id="EFEE549C4450B10DFB861664FB38E32A" author="ENGESSER B." box="[1033,1209,1547,1573]" pageId="33" pageNumber="598" pagination="943 - 994" refId="ref32460" refString="ENGESSER B. 1987. - New Eomyidae, Dipodidae, and Cricetidae (Rodentia, Mammalia) of the Lower Freshwater Molasse of Switzerland and Savoy. Eclogae Geologicae Helvetiae 80: 943 - 994." type="journal article" year="1987">Engesser (1987)</bibRefCitation>
supposed that “the material from Gaimersheim described by Freudenberg includes two species,
<taxonomicName id="4C7F52EE4450B10DFD2A1605FC99E38C" authorityName="Schaub" authorityYear="1925" box="[677,792,1641,1667]" class="Mammalia" family="Cricetidae" genus="Heterocricetodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="33" pageNumber="598" phylum="Chordata" rank="species" species="stehlini">
<emphasis id="B90BF57F4450B10DFD2A1605FC99E38C" box="[677,792,1641,1667]" italics="true" pageId="33" pageNumber="598">H. stehlini</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7F52EE4450B10DFCDD1605FC4FE38C" authorityName="Stehlin &amp; Schaub" authorityYear="1951" box="[850,974,1641,1667]" class="Mammalia" family="Cricetidae" genus="Heterocricetodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="33" pageNumber="598" phylum="Chordata" rank="species" species="helbingi">
<emphasis id="B90BF57F4450B10DFCDD1605FC4FE38C" box="[850,974,1641,1667]" italics="true" pageId="33" pageNumber="598">H. helbingi</emphasis>
</taxonomicName>
”. Later, an extensive study has been achieved by
<bibRefCitation id="EFEE549C4450B10DFC7F16E6FB37E3AC" author="KRISTKOIZ A." box="[1008,1206,1673,1699]" pageId="33" pageNumber="598" pagination="1 - 137" refId="ref33559" refString="KRISTKOIZ A. 1992. - Zahnmorphologische und schadelanatomische Untersuchungen an Nagertieren aus dem Oberoligozan von Gaimersheim (Suddeutschland). Abhandlungen / Bayerische Akademie derWissenschaften, Mathematische-Naturwissenschaftliche Klasse 167: 1 - 137." type="journal article" year="1992">Kristkoiz (1992)</bibRefCitation>
. He established the presence of only one species although with a great variability in the abundant material from Gaimersheim. Ŋerefore,
<taxonomicName id="4C7F52EE4453B10EFD961624FD06E36B" authorityName="Schaub" authorityYear="1925" box="[537,647,1610,1636]" class="Mammalia" family="Cricetidae" genus="Heterocricetodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="34" pageNumber="599" phylum="Chordata" rank="species" species="stehlini">
<emphasis id="B90BF57F4453B10EFD961624FD06E36B" box="[537,647,1610,1636]" italics="true" pageId="34" pageNumber="599">H. stehlini</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7F52EE4453B10EFF101605FE94E38C" authorityName="Stehlin &amp; Schaub" authorityYear="1951" box="[159,277,1641,1667]" class="Mammalia" family="Cricetidae" genus="Heterocricetodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="34" pageNumber="599" phylum="Chordata" rank="species" species="helbingi">
<emphasis id="B90BF57F4453B10EFF101605FE94E38C" box="[159,277,1641,1667]" italics="true" pageId="34" pageNumber="599">H. helbingi</emphasis>
</taxonomicName>
could be two morphotypes within
<taxonomicName id="4C7F52EE4453B10EFFFF16E5FEDAE3AC" authorityName="Freudenberg" authorityYear="1941" box="[112,347,1673,1699]" class="Mammalia" family="Cricetidae" genus="Heterocricetodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="34" pageNumber="599" phylum="Chordata" rank="species" species="gaimersheimensis">
<emphasis id="B90BF57F4453B10EFFFF16E5FEDAE3AC" box="[112,347,1673,1699]" italics="true" pageId="34" pageNumber="599">H. gaimersheimensis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C7F52EE4453B10EFE1316E5FCF8E32A" authority="Freudenberg, 1941" authorityName="Freudenberg" authorityYear="1941" box="[412,889,1546,1699]" class="Mammalia" family="Cricetidae" genus="Heterocricetodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="34" pageNumber="599" phylum="Chordata" rank="species" species="gaimersheimensis">
<emphasis id="B90BF57F4453B10EFE1316E5FD07E3AC" box="[412,646,1673,1699]" italics="true" pageId="34" pageNumber="599">H. gaimersheimensis</emphasis>
<bibRefCitation id="EFEE549C4453B10EFD2A1665FCF8E32A" author="FREUDENBERG H." box="[677,889,1546,1573]" pageId="34" pageNumber="599" pagination="99 - 164" refId="ref32684" refString="FREUDENBERG H. 1941. - Die oberoligozanen Nager von Gaimesrsheim bei Ingolstadt und ihre Verwandten. Palaeontographica Abteilung A 92: 99 - 164." type="journal article" year="1941">Freudenberg, 1941</bibRefCitation>
</taxonomicName>
would be a junior synonyme of
<taxonomicName id="4C7F52EE4453B10EFD4E1644FC48E34A" authority="Shaub, 1925" authorityName="Schaub" authorityYear="1925" box="[705,969,1578,1605]" class="Mammalia" family="Cricetidae" genus="Heterocricetodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="34" pageNumber="599" phylum="Chordata" rank="species" species="stehlini">
<emphasis id="B90BF57F4453B10EFD4E1644FCB2E34B" box="[705,819,1578,1604]" italics="true" pageId="34" pageNumber="599">H. stehlini</emphasis>
Shaub, 1925
</taxonomicName>
. However, the species
<taxonomicName id="4C7F52EE4453B10EFD2C1624FC94E36B" authorityName="Schaub" authorityYear="1925" box="[675,789,1610,1636]" class="Mammalia" family="Cricetidae" genus="Heterocricetodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="34" pageNumber="599" phylum="Chordata" rank="species" species="stehlini">
<emphasis id="B90BF57F4453B10EFD2C1624FC94E36B" box="[675,789,1610,1636]" italics="true" pageId="34" pageNumber="599">H. stehlini</emphasis>
</taxonomicName>
is based on an isolated lower jaw from an undetermined locality from Quercy (“Bach”, Old collections).
</paragraph>
<caption id="DF0079E54453B10EFFFA150EFEC5E0C2" ID-DOI="http://doi.org/10.5281/zenodo.4538651" ID-Zenodo-Dep="4538651" httpUri="https://zenodo.org/record/4538651/files/figure.png" pageId="34" pageNumber="599" startId="34.[117,128,1377,1395]" targetBox="[138,1206,215,1337]" targetPageId="34">
<paragraph id="8BC0296D4453B10EFFFA150EFEC5E0C2" blockId="34.[117,1211,1376,1485]" pageId="34" pageNumber="599">
FIG. 11.
<emphasis id="B90BF57F4453B10EFF38150EFF4BE07C" box="[183,202,1377,1395]" italics="true" pageId="34" pageNumber="599"></emphasis>
Other cricetids from Saint-Privat-des-Vieux (Gard; Lower Chattian, MP 26):
<emphasis id="B90BF57F4453B10EFCB3150EFCC8E07C" bold="true" box="[828,841,1377,1395]" pageId="34" pageNumber="599">A</emphasis>
,
<taxonomicName id="4C7F52EE4453B10EFCDC150EFB36E07C" authority="Vianey-liaud, 1972" authorityName="Vianey-Liaud" authorityYear="1972" box="[851,1207,1377,1395]" class="Mammalia" family="Cricetidae" genus="Eucricetodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="34" pageNumber="597" phylum="Chordata" rank="species" species="huerzeleri">
<emphasis id="B90BF57F4453B10EFCDC150EFB97E07C" box="[851,1046,1377,1395]" italics="true" pageId="34" pageNumber="599">Eucricetodon huerzeleri</emphasis>
Vianey-liaud, 1972
</taxonomicName>
, left M1-M2 (reversed), SPV165:
<emphasis id="B90BF57F4453B10EFE0E1518FE19E086" bold="true" box="[385,408,1399,1417]" pageId="34" pageNumber="599">A1</emphasis>
, occlusal view,
<emphasis id="B90BF57F4453B10EFD921518FDB4E086" bold="true" box="[541,565,1399,1417]" pageId="34" pageNumber="599">A2</emphasis>
, lingual view;
<emphasis id="B90BF57F4453B10EFD231518FD38E086" bold="true" box="[684,697,1399,1417]" pageId="34" pageNumber="599">B</emphasis>
,
<taxonomicName id="4C7F52EE4453B10EFD4B1518FC98E086" authorityName="Fischer" authorityYear="1817" box="[708,793,1399,1417]" class="Mammalia" family="Cricetidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="34" pageNumber="599" phylum="Chordata" rank="family">Cricetidae</taxonomicName>
indet., fragment of left maxillary with M1, SPV20, occlusal view;
<emphasis id="B90BF57F4453B10EFF6015E1FF7CE0AF" bold="true" box="[239,253,1422,1440]" pageId="34" pageNumber="599">C</emphasis>
, bivariate (Length and Width) diagram of M1 of
<taxonomicName id="4C7F52EE4453B10EFD0615E1FC7AE0AF" authority="(Schlosser, 1884)" baseAuthorityName="Schlosser" baseAuthorityYear="1884" box="[649,1019,1422,1440]" class="Mammalia" family="Cricetidae" genus="Pseudocricetodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="34" pageNumber="599" phylum="Chordata" rank="species" species="incertus">
<emphasis id="B90BF57F4453B10EFD0615E1FCE9E0AF" box="[649,872,1422,1440]" italics="true" pageId="34" pageNumber="599">Pseudocricetodon incertus</emphasis>
(Schlosser, 1884)
</taxonomicName>
from Saint-Privat-des- Vieux. The open circle represents the M1 SPV20. The plots for the M1 of
<taxonomicName id="4C7F52EE4453B10EFD5E15CAFCA7E0B8" baseAuthorityName="Schlosser" baseAuthorityYear="1884" box="[721,806,1445,1463]" class="Mammalia" family="Cricetidae" genus="Pseudocricetodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="34" pageNumber="599" phylum="Chordata" rank="species" species="incertus">
<emphasis id="B90BF57F4453B10EFD5E15CAFCA7E0B8" box="[721,806,1445,1463]" italics="true" pageId="34" pageNumber="599">P. incertus</emphasis>
</taxonomicName>
are surrounded by their calculated equiprobable ellipse. Scale bar: 1 mm.
</paragraph>
</caption>
<caption id="DF0079E54452B10FFFFE1495FC10E05F" ID-DOI="http://doi.org/10.5281/zenodo.4538653" ID-Zenodo-Dep="4538653" httpUri="https://zenodo.org/record/4538653/files/figure.png" pageId="35" pageNumber="600" startId="35.[113,124,1274,1292]" targetBox="[114,1210,216,1233]" targetPageId="35">
<paragraph id="8BC0296D4452B10FFFFE1495FC10E05F" blockId="35.[113,1211,1274,1360]" pageId="35" pageNumber="600">
FIG. 12.
<emphasis id="B90BF57F4452B10FFF3E1495FF45E003" box="[177,196,1274,1292]" italics="true" pageId="35" pageNumber="600"></emphasis>
<taxonomicName id="4C7F52EE4452B10FFF481495FD10E003" authority="Freudenberg, 1941" authorityName="Freudenberg" authorityYear="1941" box="[199,657,1274,1292]" class="Mammalia" family="Cricetidae" genus="Heterocricetodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="35" pageNumber="600" phylum="Chordata" rank="species" species="gaimersheimensis">
<emphasis id="B90BF57F4452B10FFF481495FE6EE003" box="[199,495,1274,1292]" italics="true" pageId="35" pageNumber="600">Heterocricetodon gaimersheimensis</emphasis>
<bibRefCitation id="EFEE549C4452B10FFE7C1495FD10E003" author="FREUDENBERG H." box="[499,657,1274,1292]" pageId="35" pageNumber="600" pagination="99 - 164" refId="ref32684" refString="FREUDENBERG H. 1941. - Die oberoligozanen Nager von Gaimesrsheim bei Ingolstadt und ihre Verwandten. Palaeontographica Abteilung A 92: 99 - 164." type="journal article" year="1941">Freudenberg,1941</bibRefCitation>
</taxonomicName>
, from Saint-Privat-des-Vieux (Gard;Lower Chattian,MP 26):
<emphasis id="B90BF57F4452B10FFBF01495FB0DE003" bold="true" box="[1151,1164,1274,1292]" pageId="35" pageNumber="600">A</emphasis>
, right M1 (reversed), SPV170:
<emphasis id="B90BF57F4452B10FFEB3157FFED3E02D" bold="true" box="[316,338,1296,1314]" pageId="35" pageNumber="600">A1</emphasis>
, occlusal view,
<emphasis id="B90BF57F4452B10FFE58157FFE6EE02D" bold="true" box="[471,495,1296,1314]" pageId="35" pageNumber="600">A2</emphasis>
, lingual view;
<emphasis id="B90BF57F4452B10FFDEA157FFDF3E02D" bold="true" box="[613,626,1296,1314]" pageId="35" pageNumber="600">B</emphasis>
, left M2, SPV171:
<emphasis id="B90BF57F4452B10FFC80157FFCA6E02D" bold="true" box="[783,807,1296,1314]" pageId="35" pageNumber="600">B1</emphasis>
, occlusal view,
<emphasis id="B90BF57F4452B10FFC24157FFC42E02D" bold="true" box="[939,963,1296,1314]" pageId="35" pageNumber="600">B2</emphasis>
, lingual view;
<emphasis id="B90BF57F4452B10FFBB5157FFBC9E02D" bold="true" box="[1082,1096,1296,1314]" pageId="35" pageNumber="600">C</emphasis>
, right M3 (re- versed), SPV174:
<emphasis id="B90BF57F4452B10FFE891548FE9FE036" bold="true" box="[262,286,1319,1337]" pageId="35" pageNumber="600">C1</emphasis>
, occlusal view,
<emphasis id="B90BF57F4452B10FFE2D1548FE38E036" bold="true" box="[418,441,1319,1337]" pageId="35" pageNumber="600">C2</emphasis>
, lingual view;
<emphasis id="B90BF57F4452B10FFDA01548FDBCE036" bold="true" box="[559,573,1319,1337]" pageId="35" pageNumber="600">D</emphasis>
, right m1 (reversed), SPV175:
<emphasis id="B90BF57F4452B10FFCB41548FCD2E036" bold="true" box="[827,851,1319,1337]" pageId="35" pageNumber="600">D1</emphasis>
, occlusal view,
<emphasis id="B90BF57F4452B10FFC581548FC6EE036" bold="true" box="[983,1007,1319,1337]" pageId="35" pageNumber="600">D2</emphasis>
, lingual view;
<emphasis id="B90BF57F4452B10FFBEA1548FBF0E036" bold="true" box="[1125,1137,1319,1337]" pageId="35" pageNumber="600">E</emphasis>
, left m2, SPV178, occlusal view;
<emphasis id="B90BF57F4452B10FFEB61551FEC5E05F" bold="true" box="[313,324,1342,1360]" pageId="35" pageNumber="600">F</emphasis>
, left m3, SPV179:
<emphasis id="B90BF57F4452B10FFE501551FE75E05F" bold="true" box="[479,500,1342,1360]" pageId="35" pageNumber="600">F1</emphasis>
, occlusal view,
<emphasis id="B90BF57F4452B10FFDF81551FD0DE05F" bold="true" box="[631,652,1342,1360]" pageId="35" pageNumber="600">F2</emphasis>
, lingual view. Scale bar: 1 mm.
</paragraph>
</caption>
<paragraph id="8BC0296D4452B10FFF0715C4FD04E3AC" blockId="35.[111,648,1451,1699]" pageId="35" pageNumber="600">
Morphological and dimensional variations of the material of Saint Privat-des-Vieux fall within those of
<taxonomicName id="4C7F52EE4452B10FFFFE1584FEBCE30A" authorityName="Freudenberg" authorityYear="1941" box="[113,317,1515,1541]" class="Mammalia" family="Cricetidae" genus="Heterocricetodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="35" pageNumber="600" phylum="Chordata" rank="species" species="gaimersheimensis">
<emphasis id="B90BF57F4452B10FFFFE1584FEBCE30A" box="[113,317,1515,1541]" italics="true" pageId="35" pageNumber="600">H. gaimersheimensis</emphasis>
</taxonomicName>
, and are not completely similar with the
<typeStatus id="54C497CF4452B10FFF5D1664FF7EE32A" box="[210,255,1547,1573]" pageId="35" pageNumber="600">type</typeStatus>
of
<taxonomicName id="4C7F52EE4452B10FFEAA1664FE11E32B" authorityName="Schaub" authorityYear="1925" box="[293,400,1546,1572]" class="Mammalia" family="Cricetidae" genus="Heterocricetodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="35" pageNumber="600" phylum="Chordata" rank="species" species="stehlini">
<emphasis id="B90BF57F4452B10FFEAA1664FE11E32B" box="[293,400,1546,1572]" italics="true" pageId="35" pageNumber="600">H. stehlini</emphasis>
</taxonomicName>
. Ŋe teeth from Saint- Privat-des-Vieux are thus reported to
<taxonomicName id="4C7F52EE4452B10FFD9E1644FF57E36B" authorityName="Freudenberg" authorityYear="1941" class="Mammalia" family="Cricetidae" genus="Heterocricetodon" higherTaxonomySource="GBIF" kingdom="Animalia" order="Rodentia" pageId="35" pageNumber="600" phylum="Chordata" rank="species" species="gaimersheimensis">
<emphasis id="B90BF57F4452B10FFD9E1644FF57E36B" italics="true" pageId="35" pageNumber="600">H. gaimersheimensis</emphasis>
</taxonomicName>
; because it is the only one species whose variability is known until now. It is the same for a few teeth from Sarèle (Vianey-Liaud 1972) and BoujacA.
</paragraph>
</subSubSection>
</treatment>
</document>