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<document id="979DCD8FBC3776915E9E5470871216BC" ID-DOI="10.5281/zenodo.182666" ID-GBIF-Dataset="d1698b4a-ec6b-4a5f-96cf-956535545e85" ID-ISSN="1175-5326" ID-Zenodo-Dep="182666" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1459872547299" checkinUser="plazi" docAuthor="Ellenrieder, Natalia Von &amp; Garrison, Rosser W." docDate="2008" docId="03ED87D9FF93BE63A9D1A6E6FB30611B" docLanguage="en" docName="zt01805p051.pdf" docOrigin="Zootaxa 1805" docStyle="DocumentStyle:6581673A57F01A3145754A1E615EDFF0.4:Zootaxa.2007-2008.journal_article" docStyleId="6581673A57F01A3145754A1E615EDFF0" docStyleName="Zootaxa.2007-2008.journal_article" docStyleVersion="4" docTitle="Oreiallagma acutum Ris 1918, comb. nov." docType="treatment" docVersion="11" lastPageNumber="45" masterDocId="FFD4FFA1FFB8BE4FA946A043FFC86549" masterDocTitle="Oreiallagma gen. nov. with a redefinition of Cyanallagma Kennedy 1920 and Mesamphiagrion Kennedy 1920, and the description of M. dunklei sp. nov. and M. ecuatoriale sp. nov. from Ecuador (Odonata: Coenagrionidae)" masterLastPageNumber="51" masterPageNumber="1" pageNumber="44" updateTime="1698226780440" updateUser="plazi">
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<mods:title id="9272B944C3F0B2CC2216AE784F61C473">Oreiallagma gen. nov. with a redefinition of Cyanallagma Kennedy 1920 and Mesamphiagrion Kennedy 1920, and the description of M. dunklei sp. nov. and M. ecuatoriale sp. nov. from Ecuador (Odonata: Coenagrionidae)</mods:title>
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<mods:namePart id="BC216E3073C11F152A5B33B293F3D871">Ellenrieder, Natalia Von</mods:namePart>
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<mods:namePart id="64A419FAF0E2B7166A1B1ECFB8D7ACB0">Garrison, Rosser W.</mods:namePart>
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<mods:date id="529D341D9F07F989FE0D29E6CE713F05">2008</mods:date>
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<treatment id="03ED87D9FF93BE63A9D1A6E6FB30611B" ID-DOI="http://doi.org/10.5281/zenodo.5691619" ID-GBIF-Taxon="119369806" ID-Zenodo-Dep="5691619" LSID="urn:lsid:plazi:treatment:03ED87D9FF93BE63A9D1A6E6FB30611B" httpUri="http://treatment.plazi.org/id/03ED87D9FF93BE63A9D1A6E6FB30611B" lastPageId="44" lastPageNumber="45" pageId="43" pageNumber="44">
<subSubSection id="C35E6544FF93BE64A9D1A6E6FE1D63AB" pageId="43" pageNumber="44" type="nomenclature">
<paragraph id="8BFB36CFFF93BE64A9D1A6E6FD6C63F6" blockId="43.[151,676,1701,1762]" box="[151,676,1701,1727]" pageId="43" pageNumber="44">
<heading id="D0B381A3FF93BE64A9D1A6E6FD6C63F6" bold="true" box="[151,676,1701,1727]" fontSize="11" level="1" pageId="43" pageNumber="44" reason="1">
<emphasis id="B930EADDFF93BE64A9D1A6E6FD6C63F6" bold="true" box="[151,676,1701,1727]" pageId="43" pageNumber="44">
<taxonomicName id="4C444D4CFF93BE64A9D1A6E6FDDE63F6" ID-CoL="6SSP5" authority="Ris 1918" authorityName="Ris" authorityYear="1918" box="[151,534,1701,1727]" class="Insecta" family="Coenagrionidae" genus="Oreiallagma" kingdom="Animalia" order="Odonata" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="acutum" status="comb. nov.">
<emphasis id="B930EADDFF93BE64A9D1A6E6FE5863F6" box="[151,400,1701,1727]" italics="true" pageId="43" pageNumber="44">Oreiallagma acutum</emphasis>
(
<bibRefCitation id="EFD54B3EFF93BE64A8E6A6E6FDC663F6" author="Ris" box="[416,526,1701,1727]" pageId="43" pageNumber="51" refString="Ris, F. (1918) Libellen (Odonata) aus der Region der amerikanischen Kordilleren von Costarica bis Catamarca. Archiv fur Naturgeschichte, A (9), 1 - 197." type="journal article" year="1918">Ris 1918</bibRefCitation>
)
</taxonomicName>
<taxonomicNameLabel id="A20357A6FF93BE64AB5BA6E6FD6C63F6" box="[541,676,1701,1727]" pageId="43" pageNumber="44">comb. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BFB36CFFF93BE64A9D1A68BFE1D63AB" blockId="43.[151,676,1701,1762]" box="[151,469,1736,1762]" pageId="43" pageNumber="44">
<figureCitation id="137F2A4AFF93BE64A9D1A68BFEDF63AB" box="[151,279,1736,1762]" captionStart="FIGURES 25 29" captionStartId="20.[151,269,1543,1567]" captionTargetBox="[188,1403,280,1513]" captionTargetId="figure@20.[183,1415,274,1525]" captionTargetPageId="20" captionText="FIGURES 25 29. Posterior lobe of prothorax. (25) Oreiallagma prothoracicum, male paratype, Ecuador, Intaj, (a) dorsal view, (b) medio-dorsal view; (26), O. oreas, male holotype, Colombia, Monte Socorro, dorsal view; (27) O. acutum, male lectotype, Bolivia, Río Zongo, dorsal view; (28) O. quadricolor, Peru, (a b) posterior lobe of prothorax, male, holotype, Santa Ana, (a) dorsal view, (b) antero-lateral view, (c) female, Morro Leguía, dorsal view; (29) O. thelkterion, Venezuela, dorsal view, (a) male, Monte Zerpa, (b) female, Guaramacal." httpUri="https://zenodo.org/record/182676/files/figure.png" pageId="43" pageNumber="44">Figures 27</figureCitation>
; 51; 85; 90; 104
</paragraph>
</subSubSection>
<subSubSection id="C35E6544FF93BE64A9D1A75BFDA962C6" pageId="43" pageNumber="44" type="reference_group">
<paragraph id="8BFB36CFFF93BE64A9D1A75BFEDF6206" blockId="43.[151,1436,1816,2031]" pageId="43" pageNumber="44">
<treatmentCitationGroup id="AB5411E1FF93BE64A9D1A75BFEDF6206" pageId="43" pageNumber="44">
<taxonomicName id="4C444D4CFF93BE64A9D1A75BFDEC6266" ID-CoL="9WZMC" authority="Ris 1918: 120" authorityName="Ris" authorityPageNumber="120" authorityYear="1918" box="[151,548,1816,1839]" class="Insecta" family="Coenagrionidae" genus="Acanthagrion" kingdom="Animalia" order="Odonata" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="acutum">
<emphasis id="B930EADDFF93BE64A9D1A75BFE496266" box="[151,385,1816,1839]" italics="true" pageId="43" pageNumber="44">Acanthagrion acutum</emphasis>
<treatmentCitation id="0AE510DEFF93BE64A8CCA75BFDEC6266" author="Ris" box="[394,548,1816,1839]" page="120" pageId="43" pageNumber="44" year="1918">
<bibRefCitation id="EFD54B3EFF93BE64A8CCA75BFE276266" author="Ris" box="[394,495,1816,1839]" pageId="43" pageNumber="51" refString="Ris, F. (1918) Libellen (Odonata) aus der Region der amerikanischen Kordilleren von Costarica bis Catamarca. Archiv fur Naturgeschichte, A (9), 1 - 197." type="journal article" year="1918">Ris 1918</bibRefCitation>
: 120
</treatmentCitation>
</taxonomicName>
122, figs. 6162 [mislabeled as 59] (description of male, illustration of S10 and wings).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BFB36CFFF93BE64A9D1A71BFCB16226" blockId="43.[151,1436,1816,2031]" box="[151,889,1880,1903]" pageId="43" pageNumber="44">
<treatmentCitationGroup id="AB5411E1FF93BE64A9D1A71BFCB16226" box="[151,889,1880,1903]" pageId="43" pageNumber="44">
<taxonomicName id="4C444D4CFF93BE64A9D1A71BFD8F6226" authority="Kennedy 1920: 87" authorityName="Kennedy" authorityPageNumber="87" authorityYear="1920" box="[151,583,1880,1903]" class="Insecta" family="Coenagrionidae" genus="Cyanallagma" kingdom="Animalia" order="Odonata" pageId="43" pageNumber="44" phylum="Arthropoda" rank="species" species="acutum">
<emphasis id="B930EADDFF93BE64A9D1A71BFEB46226" box="[151,380,1880,1903]" italics="true" pageId="43" pageNumber="44">Cyanallagma acutum</emphasis>
<treatmentCitation id="0AE510DEFF93BE64A8C4A71BFD8F6226" author="Kennedy" box="[386,583,1880,1903]" page="87" pageId="43" pageNumber="44" year="1920">
<bibRefCitation id="EFD54B3EFF93BE64A8C4A71BFDEA6226" author="Kennedy" box="[386,546,1880,1903]" pageId="43" pageNumber="50" refString="Kennedy, C. H. (1920) Forty-two hitherto unrecognized genera and subgenera of Zygoptera. Ohio Journal of Science, 21 (2), 83 - 88." type="journal article" year="1920">Kennedy 1920</bibRefCitation>
: 87
</treatmentCitation>
</taxonomicName>
(inclusion in
<taxonomicName id="4C444D4CFF93BE64AB9DA71BFCA26226" ID-CoL="3XMH" box="[731,874,1880,1903]" class="Insecta" family="Coenagrionidae" genus="Cyanallagma" kingdom="Animalia" order="Odonata" pageId="43" pageNumber="44" phylum="Arthropoda" rank="genus">
<emphasis id="B930EADDFF93BE64AB9DA71BFCA26226" box="[731,874,1880,1903]" italics="true" pageId="43" pageNumber="44">Cyanallagma</emphasis>
</taxonomicName>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BFB36CFFF93BE64A9D1A73BFDA962C6" blockId="43.[151,1436,1816,2031]" box="[151,609,1912,1935]" pageId="43" pageNumber="44">
<bibRefCitation id="EFD54B3EFF93BE64A9D1A73BFE4862C6" author="Davies" box="[151,384,1912,1935]" pageId="43" pageNumber="50" refString="Davies, D. A. L. &amp; Tobin, P. (1984) The dragonflies of the world: a systematic list of the extant species of Odonata. Volume 1. Zygoptera, Anisozygoptera. Societas Internationalis Odonatologica Rapid Communications (Supplement), Utrecht, 3: ix + 127 pp." type="book" year="1984">Davies &amp; Tobin 1984</bibRefCitation>
: 66 (synonymic list).
</paragraph>
</subSubSection>
<subSubSection id="C35E6544FF93BE63A9D1A7DBFAEB6673" lastPageId="44" lastPageNumber="45" pageId="43" pageNumber="44" type="description">
<paragraph id="8BFB36CFFF93BE64A9D1A7DBFE516286" blockId="43.[151,1436,1816,2031]" pageId="43" pageNumber="44">
<bibRefCitation id="EFD54B3EFF93BE64A9D1A7DBFE9362E6" author="De" box="[151,347,1944,1967]" pageId="43" pageNumber="50" refString="De Marmels, J. (1989) Notes on Acanthagrion acutum Ris, Enallagma occultum Ris, and E. ovigerum Calvert (Zygoptera: Coenagrionidae). Odonatologica, 18 (3), 245 - 252." type="journal article" year="1989">De Marmels 1989</bibRefCitation>
: 246247, figs. 17 (notes on male and illustrations of head, posterior lobe of prothorax, S710, genital ligula, and S10).
</paragraph>
<paragraph id="8BFB36CFFF93BE64A9D1A79BFDDC62A6" blockId="43.[151,1436,1816,2031]" box="[151,532,2008,2031]" pageId="43" pageNumber="44">
<bibRefCitation id="EFD54B3EFF93BE64A9D1A79BFEFC62A6" author="Garrison" box="[151,308,2008,2031]" pageId="43" pageNumber="50" refString="Garrison, R. W. (1991) A synonymic list of the New World Odonata. Argia, 3 (2), 1 - 30." type="journal article" year="1991">Garrison 1991</bibRefCitation>
: 11 (synonymic list).
</paragraph>
<paragraph id="8BFB36CFFF94BE63A9D1A0DBFDE165E6" blockId="44.[151,1313,152,271]" box="[151,553,152,175]" pageId="44" pageNumber="45">
<bibRefCitation id="EFD54B3EFF94BE63A9D1A0DBFEE165E6" author="Bridges" box="[151,297,152,175]" pageId="44" pageNumber="49" refString="Bridges, C. A. (1994) Catalogue of the family-group, genus-group, and species-group names of the Odonata of the world (Third Edition). Urbana, xlvi + 905 pp." type="book" year="1994">Bridges 1994</bibRefCitation>
:
<date id="FFFA100FFF94BE63A872A0DBFEA565E6" box="[308,365,152,175]" pageId="44" pageNumber="45">VII.3</date>
(synonymic list).
</paragraph>
<paragraph id="8BFB36CFFF94BE63A9D1A0FBFE3F6446" blockId="44.[151,1313,152,271]" pageId="44" pageNumber="45">
<bibRefCitation id="EFD54B3EFF94BE63A9D1A0FBFE806586" author="Steinmann" box="[151,328,184,207]" pageId="44" pageNumber="51" refString="Steinmann, H. (1997) World catalogue of Odonata. Volume I. Zygoptera. In: Wermuth, H. &amp; Fischer, M. (eds.). Das Tierreich. The Animal Kingdom. Eine Zusammenstellung und Kennzeichnung der rezenten Tierformen, Walter de Gruyter, Berlin, 110, xxi + 500 pp." type="book" year="1997">Steinmann 1997</bibRefCitation>
: 247 (synonymic list as
<taxonomicName id="4C444D4CFF94BE63AB0CA0FBFCD36586" box="[586,795,184,207]" class="Insecta" family="Coenagrionidae" genus="Cyanallagma" kingdom="Animalia" order="Odonata" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="acuta">
<emphasis id="B930EADDFF94BE63AB0CA0FBFCD36586" box="[586,795,184,207]" italics="true" pageId="44" pageNumber="45">Cyanallagma acuta</emphasis>
</taxonomicName>
).
<bibRefCitation id="EFD54B3EFF94BE63A9D1A09BFE9365A6" author="De" box="[151,347,216,239]" pageId="44" pageNumber="50" refString="De Marmels, J. (1997) New and little-known species of Cyanallagma Kennedy, 1920 from the Andes and from Pantepui (Zygoptera: Coenagrionidae). Odonatologica, 26, 135 - 157." type="journal article" year="1997">De Marmels 1997</bibRefCitation>
: 138, fig. 82 (key to northern
<taxonomicName id="4C444D4CFF94BE63ABDFA09BFCE065A6" box="[665,808,216,239]" class="Insecta" family="Coenagrionidae" genus="Cyanallagma" kingdom="Animalia" order="Odonata" pageId="44" pageNumber="45" phylum="Arthropoda" rank="genus">
<emphasis id="B930EADDFF94BE63ABDFA09BFCE065A6" box="[665,808,216,239]" italics="true" pageId="44" pageNumber="45">Cyanallagma</emphasis>
</taxonomicName>
species, map, comparison with
<taxonomicName id="4C444D4CFF94BE63AD38A09BFAD965A6" box="[1150,1297,216,239]" class="Insecta" family="Coenagrionidae" genus="Cyanallagma" kingdom="Animalia" order="Odonata" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="thelkterion">
<emphasis id="B930EADDFF94BE63AD38A09BFAD965A6" box="[1150,1297,216,239]" italics="true" pageId="44" pageNumber="45">C. thelkterion</emphasis>
</taxonomicName>
).
<bibRefCitation id="EFD54B3EFF94BE63A9D1A0BBFEDE6446" author="Tsuda" box="[151,278,248,271]" pageId="44" pageNumber="51" refString="Tsuda, S. (2000) A distributional list of world Odonata 2000. Privately published, Osaka, vi + 430 pp." type="book" year="2000">Tsuda 2000</bibRefCitation>
: 31 (synonymic list).
</paragraph>
<paragraph id="8BFB36CFFF94BE63A9D1A17CFB5A676B" blockId="44.[151,1437,319,1106]" pageId="44" pageNumber="45">
<emphasis id="B930EADDFF94BE63A9D1A17CFF2F6410" bold="true" box="[151,231,319,345]" pageId="44" pageNumber="45">
<typeStatus id="54FF886DFF94BE63A9D1A17CFF2A6410" box="[151,226,319,345]" pageId="44" pageNumber="45">Types</typeStatus>
:
</emphasis>
<bibRefCitation id="EFD54B3EFF94BE63A9B4A103FEB86413" author="Ris" box="[242,368,320,346]" pageId="44" pageNumber="51" refString="Ris, F. (1918) Libellen (Odonata) aus der Region der amerikanischen Kordilleren von Costarica bis Catamarca. Archiv fur Naturgeschichte, A (9), 1 - 197." type="journal article" year="1918">Ris (1918)</bibRefCitation>
described this species based on nine males (three from Río Zongo, six from Coroico) but did not designate a
<typeStatus id="54FF886DFF94BE63A8C6A12BFE2364CB" box="[384,491,360,386]" pageId="44" pageNumber="45" type="holotype">holotype</typeStatus>
. We examined a male
<typeStatus id="54FF886DFF94BE63ABBEA12BFC9B64CB" box="[760,851,360,386]" pageId="44" pageNumber="45" type="syntype">syntype</typeStatus>
labeled &quot;Río Zongo,
<specimenCount id="9D42FD46FF94BE63AD12A12BFB6E64CB" box="[1108,1190,360,386]" pageId="44" pageNumber="45" type="generic">
<quantity id="4CBC9B2AFF94BE63AD12A12BFB6A64CB" box="[1108,1186,360,386]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="44" pageNumber="45" unit="m" value="800.0">800 m</quantity>
.
</specimenCount>
/
<collectingCountry id="F353765FFF94BE63ADFEA12BFAFC64CB" box="[1208,1332,360,386]" name="Bolivia" pageId="44" pageNumber="45">BOLIVIA</collectingCountry>
. [La Paz Dept.] 1913 (2), A.H. Fassl&quot;, which also has a red label &quot;Lecto-[in ink by an unknown hand]/Para-/typoid [printed]&quot; affixed to the envelope.
<bibRefCitation id="EFD54B3EFF94BE63AB6DA1FBFCD9649B" box="[555,785,440,466]" pageId="44" pageNumber="50" refString="De Marmels, J. (1989) Notes on Acanthagrion acutum Ris, Enallagma occultum Ris, and E. ovigerum Calvert (Zygoptera: Coenagrionidae). Odonatologica, 18 (3), 245 - 252." type="journal article">De Marmels (1989)</bibRefCitation>
stated that he examined two
<typeStatus id="54FF886DFF94BE63AD21A1FBFAD8649B" box="[1127,1296,440,466]" pageId="44" pageNumber="45" type="paralectotype">paralectotypes</typeStatus>
from Coroico. In order to clarify application of the name, we designate the specimen we examined as
<typeStatus id="54FF886DFF94BE63AD8FA19CFAF164B0" box="[1225,1337,479,505]" pageId="44" pageNumber="45" type="lectotype">
<emphasis id="B930EADDFF94BE63AD8FA19CFAF164B0" bold="true" box="[1225,1337,479,505]" pageId="44" pageNumber="45">lectotype</emphasis>
</typeStatus>
(Senckenberg Register No. 10196) that served as basis for our
<figureCitation id="137F2A4AFF94BE63AA65A24BFC6A676B" box="[803,930,520,546]" captionStart="FIGURES 25 29" captionStartId="20.[151,269,1543,1567]" captionTargetBox="[188,1403,280,1513]" captionTargetId="figure@20.[183,1415,274,1525]" captionTargetPageId="20" captionText="FIGURES 25 29. Posterior lobe of prothorax. (25) Oreiallagma prothoracicum, male paratype, Ecuador, Intaj, (a) dorsal view, (b) medio-dorsal view; (26), O. oreas, male holotype, Colombia, Monte Socorro, dorsal view; (27) O. acutum, male lectotype, Bolivia, Río Zongo, dorsal view; (28) O. quadricolor, Peru, (a b) posterior lobe of prothorax, male, holotype, Santa Ana, (a) dorsal view, (b) antero-lateral view, (c) female, Morro Leguía, dorsal view; (29) O. thelkterion, Venezuela, dorsal view, (a) male, Monte Zerpa, (b) female, Guaramacal." httpUri="https://zenodo.org/record/182676/files/figure.png" pageId="44" pageNumber="45">Figures 27</figureCitation>
; 51; 85; 90. In FNS.
</paragraph>
<paragraph id="8BFB36CFFF94BE63A980A26CFAEB6673" blockId="44.[151,1437,319,1106]" pageId="44" pageNumber="45">
<emphasis id="B930EADDFF94BE63A980A26CFE6B6700" bold="true" box="[198,419,559,585]" pageId="44" pageNumber="45">Characterization.</emphasis>
Medial lobe of posterior lobe of prothorax slightly constricted at base, subquadrate, with shallow medial concavity on posterior margin (
<figureCitation id="137F2A4AFF94BE63AA41A21BFCA8673B" box="[775,864,600,626]" captionStart="FIGURES 25 29" captionStartId="20.[151,269,1543,1567]" captionTargetBox="[188,1403,280,1513]" captionTargetId="figure@20.[183,1415,274,1525]" captionTargetPageId="20" captionText="FIGURES 25 29. Posterior lobe of prothorax. (25) Oreiallagma prothoracicum, male paratype, Ecuador, Intaj, (a) dorsal view, (b) medio-dorsal view; (26), O. oreas, male holotype, Colombia, Monte Socorro, dorsal view; (27) O. acutum, male lectotype, Bolivia, Río Zongo, dorsal view; (28) O. quadricolor, Peru, (a b) posterior lobe of prothorax, male, holotype, Santa Ana, (a) dorsal view, (b) antero-lateral view, (c) female, Morro Leguía, dorsal view; (29) O. thelkterion, Venezuela, dorsal view, (a) male, Monte Zerpa, (b) female, Guaramacal." httpUri="https://zenodo.org/record/182676/files/figure.png" pageId="44" pageNumber="45">Fig. 27</figureCitation>
; as in
<taxonomicName id="4C444D4CFF94BE63AAE9A21BFBDD6738" box="[943,1045,600,625]" class="Insecta" family="Coenagrionidae" genus="Oreiallagma" kingdom="Animalia" order="Odonata" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="oreas">
<emphasis id="B930EADDFF94BE63AAE9A21BFBDD6738" box="[943,1045,600,625]" italics="true" pageId="44" pageNumber="45">O. oreas</emphasis>
</taxonomicName>
and
<taxonomicName id="4C444D4CFF94BE63AD17A21BFACA6738" box="[1105,1282,600,625]" class="Insecta" family="Coenagrionidae" genus="Oreiallagma" kingdom="Animalia" order="Odonata" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="quadricolor">
<emphasis id="B930EADDFF94BE63AD17A21BFACA6738" box="[1105,1282,600,625]" italics="true" pageId="44" pageNumber="45">O. quadricolor</emphasis>
</taxonomicName>
). Distal segment of genital ligula (
<figureCitation id="137F2A4AFF94BE63A8E8A2C3FDCD67D3" box="[430,517,640,666]" captionStart="FIGURES 49 53" captionStartId="26.[151,269,1805,1829]" captionTargetBox="[166,1412,208,1768]" captionTargetId="figure@26.[151,1436,194,1781]" captionTargetPageId="26" captionText="FIGURES 49 53. Distal segment of male genital ligula, (a) lateral view, (b) ectal view, (c) ental view. (49) Oreiallagma thelkterion, paratype, Venezuela, Monte Zerpa; (50) O. quadricolor, holotype, Peru, Santa Ana; (51) O. acutum, lectotype, Bolivia, Río Zongo; (52) O. oreas, holotype, Colombia, Monte Socorro; (53) O. prothoracicum, paratype, Ecuador, Intaj. i. p.: inner process; l-ap. l.: latero-apical lobe; l-m. l.: latero-medial lobe; te. f.: terminal fold; tr. f.: transverse fold; scl. l-ap. f.: sclerotized latero-apical fold." httpUri="https://zenodo.org/record/182680/files/figure.png" pageId="44" pageNumber="45">Fig. 51</figureCitation>
) approximately as wide at apex as at base with a pair of latero-apical lobes recurved and directed basally, no accessory latero-medial lobes (unique), and two ental membranous transverse folds (as in
<taxonomicName id="4C444D4CFF94BE63A821A293FDDF67A0" box="[359,535,720,745]" class="Insecta" family="Coenagrionidae" genus="Oreiallagma" kingdom="Animalia" order="Odonata" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="quadricolor">
<emphasis id="B930EADDFF94BE63A821A293FDDF67A0" box="[359,535,720,745]" italics="true" pageId="44" pageNumber="45">O. quadricolor</emphasis>
</taxonomicName>
and
<taxonomicName id="4C444D4CFF94BE63AB17A293FD3C67A0" box="[593,756,720,745]" class="Insecta" family="Coenagrionidae" genus="Cyanallagma" kingdom="Animalia" order="Odonata" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="thelkterion">
<emphasis id="B930EADDFF94BE63AB17A293FD3C67A0" box="[593,756,720,745]" italics="true" pageId="44" pageNumber="45">O. thelkterion</emphasis>
</taxonomicName>
). Dorsal process of male cercus approximately as long as ventral process (
<figureCitation id="137F2A4AFF94BE63A81EA2BBFE73665B" box="[344,443,760,786]" captionStart="FIGURES 85 89" captionStartId="31.[151,269,1253,1277]" captionTargetBox="[165,1415,272,1213]" captionTargetId="figure@31.[152,1435,244,1279]" captionTargetPageId="31" captionText="FIGURES 85 89. Male caudal appendages, medio-dorsal view, (86 b) medial view of cercus, (88 b) medio-ventral view, (88 c) dorsal view. (85) Oreiallagma acutum, lectotype, Bolivia, Río Zongo; (86) O. thelkterion, paratype, Venezuela, Monte Zerpa; (87) O. prothoracicum, holotype, Ecuador, Intaj; (88) O. quadricolor, holotype, Peru, Santa Ana; (89) O. oreas, holotype, Colombia, Monte Socorro. d. p.: dorsal process; v. ap. p.: ventro-apical process; v-b. p.: ventro-basal process; scal. cu.: scalariform-like cuticle." httpUri="https://zenodo.org/record/182684/files/figure.png" pageId="44" pageNumber="45">Figs. 85</figureCitation>
; 90; unique). Dorsum of S17 black, of S89 blue, sides of S37 reddish-orange, of S8 with a black stripe along anterior 2/3, S9 entirely blue and S10 entirely black. Female unknown.
</paragraph>
</subSubSection>
<subSubSection id="C35E6544FF94BE63A980A304FCBE614B" pageId="44" pageNumber="45" type="diagnosis">
<paragraph id="8BFB36CFFF94BE63A980A304FCBE614B" blockId="44.[151,1437,319,1106]" pageId="44" pageNumber="45">
<emphasis id="B930EADDFF94BE63A980A304FE8E6628" bold="true" box="[198,326,839,865]" pageId="44" pageNumber="45">Diagnosis.</emphasis>
Species unique by cercus morphology: the long, digit-like dorsal process (
<figureCitation id="137F2A4AFF94BE63AD8FA30BFAE8662B" box="[1225,1312,840,866]" captionStart="FIGURES 85 89" captionStartId="31.[151,269,1253,1277]" captionTargetBox="[165,1415,272,1213]" captionTargetId="figure@31.[152,1435,244,1279]" captionTargetPageId="31" captionText="FIGURES 85 89. Male caudal appendages, medio-dorsal view, (86 b) medial view of cercus, (88 b) medio-ventral view, (88 c) dorsal view. (85) Oreiallagma acutum, lectotype, Bolivia, Río Zongo; (86) O. thelkterion, paratype, Venezuela, Monte Zerpa; (87) O. prothoracicum, holotype, Ecuador, Intaj; (88) O. quadricolor, holotype, Peru, Santa Ana; (89) O. oreas, holotype, Colombia, Monte Socorro. d. p.: dorsal process; v. ap. p.: ventro-apical process; v-b. p.: ventro-basal process; scal. cu.: scalariform-like cuticle." httpUri="https://zenodo.org/record/182684/files/figure.png" pageId="44" pageNumber="45">Fig. 85</figureCitation>
) is shared only with
<taxonomicName id="4C444D4CFF94BE63A856A333FE7D66C0" box="[272,437,880,905]" class="Insecta" family="Coenagrionidae" genus="Cyanallagma" kingdom="Animalia" order="Odonata" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="thelkterion">
<emphasis id="B930EADDFF94BE63A856A333FE7D66C0" box="[272,437,880,905]" italics="true" pageId="44" pageNumber="45">O. thelkterion</emphasis>
</taxonomicName>
(
<figureCitation id="137F2A4AFF94BE63A880A333FDD466C3" box="[454,540,880,906]" captionStart="FIGURES 85 89" captionStartId="31.[151,269,1253,1277]" captionTargetBox="[165,1415,272,1213]" captionTargetId="figure@31.[152,1435,244,1279]" captionTargetPageId="31" captionText="FIGURES 85 89. Male caudal appendages, medio-dorsal view, (86 b) medial view of cercus, (88 b) medio-ventral view, (88 c) dorsal view. (85) Oreiallagma acutum, lectotype, Bolivia, Río Zongo; (86) O. thelkterion, paratype, Venezuela, Monte Zerpa; (87) O. prothoracicum, holotype, Ecuador, Intaj; (88) O. quadricolor, holotype, Peru, Santa Ana; (89) O. oreas, holotype, Colombia, Monte Socorro. d. p.: dorsal process; v. ap. p.: ventro-apical process; v-b. p.: ventro-basal process; scal. cu.: scalariform-like cuticle." httpUri="https://zenodo.org/record/182684/files/figure.png" pageId="44" pageNumber="45">Fig. 86</figureCitation>
) but is more robust and as long as or longer than ventral process (
<figureCitation id="137F2A4AFF94BE63AC75A333FA5F66C3" box="[1331,1431,880,906]" captionStart="FIGURES 85 89" captionStartId="31.[151,269,1253,1277]" captionTargetBox="[165,1415,272,1213]" captionTargetId="figure@31.[152,1435,244,1279]" captionTargetPageId="31" captionText="FIGURES 85 89. Male caudal appendages, medio-dorsal view, (86 b) medial view of cercus, (88 b) medio-ventral view, (88 c) dorsal view. (85) Oreiallagma acutum, lectotype, Bolivia, Río Zongo; (86) O. thelkterion, paratype, Venezuela, Monte Zerpa; (87) O. prothoracicum, holotype, Ecuador, Intaj; (88) O. quadricolor, holotype, Peru, Santa Ana; (89) O. oreas, holotype, Colombia, Monte Socorro. d. p.: dorsal process; v. ap. p.: ventro-apical process; v-b. p.: ventro-basal process; scal. cu.: scalariform-like cuticle." httpUri="https://zenodo.org/record/182684/files/figure.png" pageId="44" pageNumber="45">Figs. 85</figureCitation>
; 90); in
<taxonomicName id="4C444D4CFF94BE63A9ABA3DBFE5866F8" box="[237,400,920,945]" class="Insecta" family="Coenagrionidae" genus="Cyanallagma" kingdom="Animalia" order="Odonata" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="thelkterion">
<emphasis id="B930EADDFF94BE63A9ABA3DBFE5866F8" box="[237,400,920,945]" italics="true" pageId="44" pageNumber="45">O. thelkterion</emphasis>
</taxonomicName>
, dorsal process is more acuminate and shorter than ventral process (
<figureCitation id="137F2A4AFF94BE63ADFEA3DBFAD866FB" box="[1208,1296,920,946]" captionStart="FIGURES 85 89" captionStartId="31.[151,269,1253,1277]" captionTargetBox="[165,1415,272,1213]" captionTargetId="figure@31.[152,1435,244,1279]" captionTargetPageId="31" captionText="FIGURES 85 89. Male caudal appendages, medio-dorsal view, (86 b) medial view of cercus, (88 b) medio-ventral view, (88 c) dorsal view. (85) Oreiallagma acutum, lectotype, Bolivia, Río Zongo; (86) O. thelkterion, paratype, Venezuela, Monte Zerpa; (87) O. prothoracicum, holotype, Ecuador, Intaj; (88) O. quadricolor, holotype, Peru, Santa Ana; (89) O. oreas, holotype, Colombia, Monte Socorro. d. p.: dorsal process; v. ap. p.: ventro-apical process; v-b. p.: ventro-basal process; scal. cu.: scalariform-like cuticle." httpUri="https://zenodo.org/record/182684/files/figure.png" pageId="44" pageNumber="45">Fig. 86</figureCitation>
). In
<taxonomicName id="4C444D4CFF94BE63AC00A3DBFED56690" class="Insecta" family="Coenagrionidae" genus="Oreiallagma" kingdom="Animalia" order="Odonata" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="prothoracicum">
<emphasis id="B930EADDFF94BE63AC00A3DBFED56690" italics="true" pageId="44" pageNumber="45">O. prothoracicum</emphasis>
</taxonomicName>
(
<figureCitation id="137F2A4AFF94BE63A869A383FE5C6693" box="[303,404,960,986]" captionStart="FIGURES 85 89" captionStartId="31.[151,269,1253,1277]" captionTargetBox="[165,1415,272,1213]" captionTargetId="figure@31.[152,1435,244,1279]" captionTargetPageId="31" captionText="FIGURES 85 89. Male caudal appendages, medio-dorsal view, (86 b) medial view of cercus, (88 b) medio-ventral view, (88 c) dorsal view. (85) Oreiallagma acutum, lectotype, Bolivia, Río Zongo; (86) O. thelkterion, paratype, Venezuela, Monte Zerpa; (87) O. prothoracicum, holotype, Ecuador, Intaj; (88) O. quadricolor, holotype, Peru, Santa Ana; (89) O. oreas, holotype, Colombia, Monte Socorro. d. p.: dorsal process; v. ap. p.: ventro-apical process; v-b. p.: ventro-basal process; scal. cu.: scalariform-like cuticle." httpUri="https://zenodo.org/record/182684/files/figure.png" pageId="44" pageNumber="45">Figs. 87</figureCitation>
; 92) and
<taxonomicName id="4C444D4CFF94BE63AB4CA383FD756690" box="[522,701,960,985]" class="Insecta" family="Coenagrionidae" genus="Oreiallagma" kingdom="Animalia" order="Odonata" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="quadricolor">
<emphasis id="B930EADDFF94BE63AB4CA383FD756690" box="[522,701,960,985]" italics="true" pageId="44" pageNumber="45">O. quadricolor</emphasis>
</taxonomicName>
(
<figureCitation id="137F2A4AFF94BE63AB96A383FCFC6693" box="[720,820,960,986]" captionStart="FIGURES 85 89" captionStartId="31.[151,269,1253,1277]" captionTargetBox="[165,1415,272,1213]" captionTargetId="figure@31.[152,1435,244,1279]" captionTargetPageId="31" captionText="FIGURES 85 89. Male caudal appendages, medio-dorsal view, (86 b) medial view of cercus, (88 b) medio-ventral view, (88 c) dorsal view. (85) Oreiallagma acutum, lectotype, Bolivia, Río Zongo; (86) O. thelkterion, paratype, Venezuela, Monte Zerpa; (87) O. prothoracicum, holotype, Ecuador, Intaj; (88) O. quadricolor, holotype, Peru, Santa Ana; (89) O. oreas, holotype, Colombia, Monte Socorro. d. p.: dorsal process; v. ap. p.: ventro-apical process; v-b. p.: ventro-basal process; scal. cu.: scalariform-like cuticle." httpUri="https://zenodo.org/record/182684/files/figure.png" pageId="44" pageNumber="45">Figs. 88</figureCitation>
a; 93), dorsal process is small and acute, and in
<taxonomicName id="4C444D4CFF94BE63AC39A383FF1F6148" class="Insecta" family="Coenagrionidae" genus="Oreiallagma" kingdom="Animalia" order="Odonata" pageId="44" pageNumber="45" phylum="Arthropoda" rank="species" species="oreas">
<emphasis id="B930EADDFF94BE63AC39A383FF1F6148" italics="true" pageId="44" pageNumber="45">O. oreas</emphasis>
</taxonomicName>
it is represented by a blunt setose tubercle (
<figureCitation id="137F2A4AFF94BE63AB9EA3ABFCF2614B" box="[728,826,1000,1026]" captionStart="FIGURES 85 89" captionStartId="31.[151,269,1253,1277]" captionTargetBox="[165,1415,272,1213]" captionTargetId="figure@31.[152,1435,244,1279]" captionTargetPageId="31" captionText="FIGURES 85 89. Male caudal appendages, medio-dorsal view, (86 b) medial view of cercus, (88 b) medio-ventral view, (88 c) dorsal view. (85) Oreiallagma acutum, lectotype, Bolivia, Río Zongo; (86) O. thelkterion, paratype, Venezuela, Monte Zerpa; (87) O. prothoracicum, holotype, Ecuador, Intaj; (88) O. quadricolor, holotype, Peru, Santa Ana; (89) O. oreas, holotype, Colombia, Monte Socorro. d. p.: dorsal process; v. ap. p.: ventro-apical process; v-b. p.: ventro-basal process; scal. cu.: scalariform-like cuticle." httpUri="https://zenodo.org/record/182684/files/figure.png" pageId="44" pageNumber="45">Figs. 89</figureCitation>
; 94).
</paragraph>
</subSubSection>
<subSubSection id="C35E6544FF94BE63A980A44CFC9A6163" box="[198,850,1039,1066]" pageId="44" pageNumber="45" type="biology_ecology">
<paragraph id="8BFB36CFFF94BE63A980A44CFC9A6163" blockId="44.[151,1437,319,1106]" box="[198,850,1039,1066]" pageId="44" pageNumber="45">
<emphasis id="B930EADDFF94BE63A980A44CFEE26160" bold="true" box="[198,298,1039,1065]" pageId="44" pageNumber="45">Biology.</emphasis>
Unknown; see remarks under generic account.
</paragraph>
</subSubSection>
<subSubSection id="C35E6544FF94BE63A980A474FB30611B" box="[198,1272,1079,1106]" pageId="44" pageNumber="45" type="distribution">
<paragraph id="8BFB36CFFF94BE63A980A474FB30611B" blockId="44.[151,1437,319,1106]" box="[198,1272,1079,1106]" pageId="44" pageNumber="45">
<emphasis id="B930EADDFF94BE63A980A474FEAF6118" bold="true" box="[198,359,1079,1105]" pageId="44" pageNumber="45">Distribution.</emphasis>
<collectingCountry id="F353765FFF94BE63A828A47BFE0C611B" box="[366,452,1080,1106]" name="Bolivia" pageId="44" pageNumber="45">Bolivia</collectingCountry>
(La Paz Department), from
<quantity id="4CBC9B2AFF94BE63AA56A47BFC73611B" box="[784,955,1080,1106]" metricMagnitude="3" metricUnit="m" metricValue="1.1" metricValueMax="1.4" metricValueMin="0.8" pageId="44" pageNumber="45" unit="m" value="1100.0" valueMax="1400.0" valueMin="800.0">800 to 1400 m</quantity>
above sea level (
<figureCitation id="137F2A4AFF94BE63ADC2A47BFB23611B" box="[1156,1259,1080,1106]" captionStart="FIGURE 104" captionStartId="38.[151,255,1925,1949]" captionTargetBox="[170,1430,210,1889]" captionTargetId="figure@38.[151,1436,194,1902]" captionTargetPageId="38" captionText="FIGURE 104. Distribution of species of Oreiallagma in South America." httpUri="https://zenodo.org/record/182691/files/figure.png" pageId="44" pageNumber="45">Fig. 104</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>