treatments-xml/data/03/ED/87/03ED87ACFF9FFFBEFF2DFF4F82BF49FD.xml

<document id="DF59A08CB8C129A7FB265ADEA336F47B" ID-CLB-Dataset="7326" ID-DOI="10.11646/zootaxa.4952.2.9" ID-GBIF-Dataset="4c07e3a9-3c3b-436c-852f-3864fe68fc8e" ID-ISSN="1175-5326" ID-Zenodo-Dep="4674237" ID-ZooBank="28006273-1456-47EC-8888-17CAB686C99F" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1617950919278" checkinUser="plazi" docAuthor="Ng, Peter K. L." docDate="2021" docId="03ED87ACFF9FFFBEFF2DFF4F82BF49FD" docLanguage="en" docName="zootaxa.4952.2.9.pdf" docOrigin="Zootaxa 4952 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Ovilyra Ng 2021, n. gen." docType="treatment" docVersion="7" lastPageNumber="371" masterDocId="FFD4FFD4FF9EFFBCFFBAFFD880334850" masterDocTitle="Ovilyra, a new genus of leucosiid crab (Crustacea: Decapoda: Brachyura) from the West Pacific" masterLastPageNumber="380" masterPageNumber="369" pageNumber="370" updateTime="1698936277179" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="6B05F719921FAAAA9073BBA75477B0A3">Ovilyra, a new genus of leucosiid crab (Crustacea: Decapoda: Brachyura) from the West Pacific</mods:title>
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<mods:namePart id="88973687D9B6837821DF15592CE34E7C">Ng, Peter K. L.</mods:namePart>
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<mods:affiliation id="94C9B0AF6D5CB2673EAFE021D01D0DF7">Lee Kong Chian Natural History Museum, National University of Singapore, 2 Conservatory Drive, Singapore 117377, Republic of Singapore. peterng @ nus. edu. sg; https: // orcid. org / 0000 - 0001 - 5946 - 0608</mods:affiliation>
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<treatment id="03ED87ACFF9FFFBEFF2DFF4F82BF49FD" ID-DOI="http://doi.org/10.5281/zenodo.4695028" ID-GBIF-Taxon="181011204" ID-Zenodo-Dep="4695028" LSID="urn:lsid:plazi:treatment:03ED87ACFF9FFFBEFF2DFF4F82BF49FD" httpUri="http://treatment.plazi.org/id/03ED87ACFF9FFFBEFF2DFF4F82BF49FD" lastPageId="2" lastPageNumber="371" pageId="1" pageNumber="370">
<subSubSection id="C35E6531FF9FFFBDFF2DFF4F817948E1" box="[151,330,151,178]" pageId="1" pageNumber="370" type="nomenclature">
<paragraph id="8BFB36BAFF9FFFBDFF2DFF4F817948E1" blockId="1.[151,330,151,178]" box="[151,330,151,178]" pageId="1" pageNumber="370">
<heading id="D0B381D6FF9FFFBDFF2DFF4F817948E1" bold="true" box="[151,330,151,178]" fontSize="11" level="1" pageId="1" pageNumber="370" reason="1">
<emphasis id="B930EAA8FF9FFFBDFF2DFF4F817948E1" bold="true" box="[151,330,151,178]" pageId="1" pageNumber="370">
<taxonomicName id="4C444D39FF9FFFBDFF2DFF4F80C248E2" ID-CoL="8753R" authority="Ng, 2021" authorityName="Ng" authorityYear="2021" box="[151,241,151,178]" class="Malacostraca" family="Leucosiidae" genus="Ovilyra" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="370" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B930EAA8FF9FFFBDFF2DFF4F80C248E2" bold="true" box="[151,241,151,178]" italics="true" pageId="1" pageNumber="370">Ovilyra</emphasis>
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<taxonomicNameLabel id="A20357D3FF9FFFBDFF42FF4F817948E1" box="[248,330,151,177]" pageId="1" pageNumber="370" rank="genus">n. gen.</taxonomicNameLabel>
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<paragraph id="8BFB36BAFF9FFFBDFF2DFF0483FE48A4" blockId="1.[151,973,219,244]" box="[151,973,219,244]" pageId="1" pageNumber="370">
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<typeStatus id="54FF8818FF9FFFBDFF2DFF0480FF48A4" box="[151,204,220,244]" pageId="1" pageNumber="370">Type</typeStatus>
species
</emphasis>
. 
<taxonomicName id="4C444D39FF9FFFBDFE90FF0482EB48A4" ID-CoL="856BT" authority="Targioni-Tozzetti, 1877" authorityName="Targioni-Tozzetti" authorityYear="1877" box="[298,728,220,244]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="370" phylum="Arthropoda" rank="species" species="fuliginosa">
<emphasis id="B930EAA8FF9FFFBDFE90FF0481D248A4" box="[298,481,220,244]" italics="true" pageId="1" pageNumber="370">Philyra fuliginosa</emphasis>
<bibRefCitation id="EFD54B4BFF9FFFBDFE5DFF0482EB48A4" author="Targioni-Tozzetti, A." box="[487,728,220,244]" pageId="1" pageNumber="370" pagination="1 - 12" refId="ref7486" refString="Targioni-Tozzetti, A. (1877) Crostacei Brachyuri e Anomuri. Zoologia del viaggio intorno al Globo della R. Pirocorvetta Magenta durante gli anni 1865 - 1868. Pubblicazioni del R. Istituto di Studi superiori pratici e di Perfezionamento in Firenze, Sezione di Scienze fisiche e naturali, Successori le Monnier, Firenze, 1, i - xxix + 1 - 257 pp., pls. 1 - 12." type="book chapter" year="1877">Targioni-Tozzetti, 1877</bibRefCitation>
</taxonomicName>
, by present designation.
</paragraph>
</subSubSection>
<subSubSection id="C35E6531FF9FFFBDFF2DFEF983C54C60" pageId="1" pageNumber="370" type="diagnosis">
<paragraph id="8BFB36BAFF9FFFBDFF2DFEF983C54C60" blockId="1.[151,1437,289,2008]" pageId="1" pageNumber="370">
<emphasis id="B930EAA8FF9FFFBDFF2DFEF98139496B" bold="true" box="[151,266,289,315]" pageId="1" pageNumber="370">Diagnosis</emphasis>
. Carapace longitudinally ovate, distinctly longer than wide (length to width ratio 1.17–1.26); dorsal surface smooth, covered with minute to small granules and small depressions (
<figureCitation id="137F2A3FFF9FFFBDFBABFE9E84A44930" box="[1041,1175,326,352]" captionStart="FIGURE 1" captionStartId="3.[151,250,1921,1945]" captionTargetBox="[169,1418,181,1897]" captionTargetId="figure-15@3.[169,1418,181,1897]" captionTargetPageId="3" captionText="FIGURE 1. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia.A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, anterior thoracic sternum, pleonal somites 3–6 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674239" httpUri="https://zenodo.org/record/4674239/files/figure.png" pageId="1" pageNumber="370">Figs. 1A, B</figureCitation>
, 
<figureCitation id="137F2A3FFF9FFFBDFB19FE9E84DA4930" box="[1187,1257,326,352]" captionStart="FIGURE 2" captionStartId="4.[151,250,1934,1958]" captionTargetBox="[164,1422,181,1910]" captionTargetId="figure-19@4.[164,1422,181,1910]" captionTargetPageId="4" captionText="FIGURE 2. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore. A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, buccal cavity, anterior thoracic sternum, pleonal somites 3–6 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674241" httpUri="https://zenodo.org/record/4674241/files/figure.png" pageId="1" pageNumber="370">2A, B</figureCitation>
, 
<figureCitation id="137F2A3FFF9FFFBDFB4CFE9E850E4930" box="[1270,1341,326,352]" captionStart="FIGURE 3" captionStartId="5.[151,250,1254,1278]" captionTargetBox="[151,1436,181,1230]" captionTargetId="figure-15@5.[151,1436,181,1230]" captionTargetPageId="5" captionText="FIGURE 3. Ovilyra fuliginosa (Targioni-Tozzetti, 1877). A, male (11.2 × 14.0 mm) (ZRC 2009.0217), Singapore; B, male (13.3 × 16.2 mm) (ZRC 2009.0217), Singapore; C–E, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Muar, Peninsular Malaysia.A–C, overall dorsal view; D, anterior thoracic sternum, pleonal somites 3–6 and telson; E, left third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.4674243" httpUri="https://zenodo.org/record/4674243/files/figure.png" pageId="1" pageNumber="370">3A–C</figureCitation>
); frontal region not produced anteriorly, with shallow median longitudinal depression; frontal margin granulated, with low median triangular tooth (
<figureCitation id="137F2A3FFF9FFFBDFE08FE56822749F8" box="[434,532,398,424]" captionStart="FIGURE 1" captionStartId="3.[151,250,1921,1945]" captionTargetBox="[169,1418,181,1897]" captionTargetId="figure-15@3.[169,1418,181,1897]" captionTargetPageId="3" captionText="FIGURE 1. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia.A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, anterior thoracic sternum, pleonal somites 3–6 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674239" httpUri="https://zenodo.org/record/4674239/files/figure.png" pageId="1" pageNumber="370">Figs. 1B</figureCitation>
, 
<figureCitation id="137F2A3FFF9FFFBDFD98FE56827549F8" box="[546,582,398,424]" captionStart="FIGURE 2" captionStartId="4.[151,250,1934,1958]" captionTargetBox="[164,1422,181,1910]" captionTargetId="figure-19@4.[164,1422,181,1910]" captionTargetPageId="4" captionText="FIGURE 2. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore. A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, buccal cavity, anterior thoracic sternum, pleonal somites 3–6 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674241" httpUri="https://zenodo.org/record/4674241/files/figure.png" pageId="1" pageNumber="370">2B</figureCitation>
, 
<figureCitation id="137F2A3FFF9FFFBDFDE8FE5682A449F8" box="[594,663,398,425]" captionStart="FIGURE 5" captionStartId="7.[151,250,1928,1952]" captionTargetBox="[215,1387,688,1892]" captionTargetId="figure-278@7.[151,1436,669,1905]" captionTargetPageId="7" captionText="FIGURE 5. Ovilyra fuliginosa (Targioni-Tozzetti, 1877).A, D, F, male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia; B, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Malaysia; C, E, G, male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore.A–C, left third maxilliped; D, E, frontal margin; F, G, pleon. Setae on structures not drawn. Scales: A, D = 1.0 mm; B, C, E–G = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4674247" httpUri="https://zenodo.org/record/4674247/files/figure.png" pageId="1" pageNumber="370">5D, E</figureCitation>
, 
<figureCitation id="137F2A3FFF9FFFBDFD1FFE5682DE49F8" box="[677,749,398,424]" captionStart="FIGURE 7" captionStartId="9.[151,250,1754,1778]" captionTargetBox="[151,1436,181,1730]" captionTargetId="figure-15@9.[151,1436,181,1730]" captionTargetPageId="9" captionText="FIGURE 7. Ovilyra fuliginosa (Targioni-Tozzetti, 1877). A–C, female (8.5 × 10.1 mm) (ZRC 2020.0345), Malaysia. D–F, female (9.3 × 11.6 mm) (ZRC 2009.0217), Singapore. A, D, overall dorsal view; B, E, pleon; C, F, vulvae." figureDoi="http://doi.org/10.5281/zenodo.4674251" httpUri="https://zenodo.org/record/4674251/files/figure.png" pageId="1" pageNumber="370">7A, D</figureCitation>
); hepatic facet well defined by rows of granules and cristae, elongate, distal part of lower margin prominently angular, with broad tooth on distal one-third (
<figureCitation id="137F2A3FFF9FFFBDFB78FE6A8574499C" box="[1218,1351,434,460]" captionStart="FIGURE 1" captionStartId="3.[151,250,1921,1945]" captionTargetBox="[169,1418,181,1897]" captionTargetId="figure-15@3.[169,1418,181,1897]" captionTargetPageId="3" captionText="FIGURE 1. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia.A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, anterior thoracic sternum, pleonal somites 3–6 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674239" httpUri="https://zenodo.org/record/4674239/files/figure.png" pageId="1" pageNumber="370">Figs. 1B, D</figureCitation>
, 
<figureCitation id="137F2A3FFF9FFFBDFAE8FE6A85AB499C" box="[1362,1432,434,460]" captionStart="FIGURE 2" captionStartId="4.[151,250,1934,1958]" captionTargetBox="[164,1422,181,1910]" captionTargetId="figure-19@4.[164,1422,181,1910]" captionTargetPageId="4" captionText="FIGURE 2. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore. A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, buccal cavity, anterior thoracic sternum, pleonal somites 3–6 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674241" httpUri="https://zenodo.org/record/4674241/files/figure.png" pageId="1" pageNumber="370">2B, D</figureCitation>
, 
<figureCitation id="137F2A3FFF9FFFBDFF2DFE0E808E49A0" box="[151,189,470,496]" captionStart="FIGURE 4" captionStartId="6.[151,250,1167,1191]" captionTargetBox="[151,1436,417,1143]" captionTargetId="figure-141@6.[151,1436,417,1143]" captionTargetPageId="6" captionText="FIGURE 4. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia. A, buccal cavity, anterior thoracic sternum and pleonal somites 3–6 and telson; B, posterior thoracic sternum and pleonal somites 1–4. Abbreviations: ca = carapace; cx5 = coxa of fourth ambulatory leg; so1–so6 = pleonal somites 1–6; s1–s8 = sternites 1–8; te = telson." figureDoi="http://doi.org/10.5281/zenodo.4674245" httpUri="https://zenodo.org/record/4674245/files/figure.png" pageId="1" pageNumber="370">4A</figureCitation>
); posterolateral margin demarcated from posterior margin by lobiform tooth (
<figureCitation id="137F2A3FFF9FFFBDFBA0FE0E84AE49A0" box="[1050,1181,470,496]" captionStart="FIGURE 1" captionStartId="3.[151,250,1921,1945]" captionTargetBox="[169,1418,181,1897]" captionTargetId="figure-15@3.[169,1418,181,1897]" captionTargetPageId="3" captionText="FIGURE 1. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia.A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, anterior thoracic sternum, pleonal somites 3–6 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674239" httpUri="https://zenodo.org/record/4674239/files/figure.png" pageId="1" pageNumber="370">Figs. 1A, B</figureCitation>
, 
<figureCitation id="137F2A3FFF9FFFBDFB12FE0E84DF49A0" box="[1192,1260,470,496]" captionStart="FIGURE 2" captionStartId="4.[151,250,1934,1958]" captionTargetBox="[164,1422,181,1910]" captionTargetId="figure-19@4.[164,1422,181,1910]" captionTargetPageId="4" captionText="FIGURE 2. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore. A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, buccal cavity, anterior thoracic sternum, pleonal somites 3–6 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674241" httpUri="https://zenodo.org/record/4674241/files/figure.png" pageId="1" pageNumber="370">2A, B</figureCitation>
, 
<figureCitation id="137F2A3FFF9FFFBDFB4DFE0E850F49A0" box="[1271,1340,470,496]" captionStart="FIGURE 3" captionStartId="5.[151,250,1254,1278]" captionTargetBox="[151,1436,181,1230]" captionTargetId="figure-15@5.[151,1436,181,1230]" captionTargetPageId="5" captionText="FIGURE 3. Ovilyra fuliginosa (Targioni-Tozzetti, 1877). A, male (11.2 × 14.0 mm) (ZRC 2009.0217), Singapore; B, male (13.3 × 16.2 mm) (ZRC 2009.0217), Singapore; C–E, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Muar, Peninsular Malaysia.A–C, overall dorsal view; D, anterior thoracic sternum, pleonal somites 3–6 and telson; E, left third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.4674243" httpUri="https://zenodo.org/record/4674243/files/figure.png" pageId="1" pageNumber="370">3A–C</figureCitation>
, 
<figureCitation id="137F2A3FFF9FFFBDFAFDFE0E85BD49A0" box="[1351,1422,470,496]" captionStart="FIGURE 7" captionStartId="9.[151,250,1754,1778]" captionTargetBox="[151,1436,181,1730]" captionTargetId="figure-15@9.[151,1436,181,1730]" captionTargetPageId="9" captionText="FIGURE 7. Ovilyra fuliginosa (Targioni-Tozzetti, 1877). A–C, female (8.5 × 10.1 mm) (ZRC 2020.0345), Malaysia. D–F, female (9.3 × 11.6 mm) (ZRC 2009.0217), Singapore. A, D, overall dorsal view; B, E, pleon; C, F, vulvae." figureDoi="http://doi.org/10.5281/zenodo.4674251" httpUri="https://zenodo.org/record/4674251/files/figure.png" pageId="1" pageNumber="370">7A, D</figureCitation>
); third maxilliped exopod wider than basal part of endopod (
<figureCitation id="137F2A3FFF9FFFBDFC8FFE2283A44A44" box="[821,919,506,532]" captionStart="FIGURE 1" captionStartId="3.[151,250,1921,1945]" captionTargetBox="[169,1418,181,1897]" captionTargetId="figure-15@3.[169,1418,181,1897]" captionTargetPageId="3" captionText="FIGURE 1. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia.A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, anterior thoracic sternum, pleonal somites 3–6 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674239" httpUri="https://zenodo.org/record/4674239/files/figure.png" pageId="1" pageNumber="370">Figs. 1C</figureCitation>
, 
<figureCitation id="137F2A3FFF9FFFBDFC1FFE2283FB4A44" box="[933,968,506,532]" captionStart="FIGURE 2" captionStartId="4.[151,250,1934,1958]" captionTargetBox="[164,1422,181,1910]" captionTargetId="figure-19@4.[164,1422,181,1910]" captionTargetPageId="4" captionText="FIGURE 2. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore. A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, buccal cavity, anterior thoracic sternum, pleonal somites 3–6 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674241" httpUri="https://zenodo.org/record/4674241/files/figure.png" pageId="1" pageNumber="370">2C</figureCitation>
, 
<figureCitation id="137F2A3FFF9FFFBDFC6FFE2283C54A44" box="[981,1014,506,532]" captionStart="FIGURE 3" captionStartId="5.[151,250,1254,1278]" captionTargetBox="[151,1436,181,1230]" captionTargetId="figure-15@5.[151,1436,181,1230]" captionTargetPageId="5" captionText="FIGURE 3. Ovilyra fuliginosa (Targioni-Tozzetti, 1877). A, male (11.2 × 14.0 mm) (ZRC 2009.0217), Singapore; B, male (13.3 × 16.2 mm) (ZRC 2009.0217), Singapore; C–E, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Muar, Peninsular Malaysia.A–C, overall dorsal view; D, anterior thoracic sternum, pleonal somites 3–6 and telson; E, left third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.4674243" httpUri="https://zenodo.org/record/4674243/files/figure.png" pageId="1" pageNumber="370">3E</figureCitation>
, 
<figureCitation id="137F2A3FFF9FFFBDFBB9FE2284784A45" box="[1027,1099,506,533]" captionStart="FIGURE 5" captionStartId="7.[151,250,1928,1952]" captionTargetBox="[215,1387,688,1892]" captionTargetId="figure-278@7.[151,1436,669,1905]" captionTargetPageId="7" captionText="FIGURE 5. Ovilyra fuliginosa (Targioni-Tozzetti, 1877).A, D, F, male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia; B, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Malaysia; C, E, G, male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore.A–C, left third maxilliped; D, E, frontal margin; F, G, pleon. Setae on structures not drawn. Scales: A, D = 1.0 mm; B, C, E–G = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4674247" httpUri="https://zenodo.org/record/4674247/files/figure.png" pageId="1" pageNumber="370">5A–C</figureCitation>
); endopod with basis-ischium longer than acutely triangular merus (
<figureCitation id="137F2A3FFF9FFFBDFDFBFDC682904A68" box="[577,675,542,568]" captionStart="FIGURE 1" captionStartId="3.[151,250,1921,1945]" captionTargetBox="[169,1418,181,1897]" captionTargetId="figure-15@3.[169,1418,181,1897]" captionTargetPageId="3" captionText="FIGURE 1. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia.A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, anterior thoracic sternum, pleonal somites 3–6 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674239" httpUri="https://zenodo.org/record/4674239/files/figure.png" pageId="1" pageNumber="370">Figs. 1C</figureCitation>
, 
<figureCitation id="137F2A3FFF9FFFBDFD0BFDC682E74A68" box="[689,724,542,568]" captionStart="FIGURE 2" captionStartId="4.[151,250,1934,1958]" captionTargetBox="[164,1422,181,1910]" captionTargetId="figure-19@4.[164,1422,181,1910]" captionTargetPageId="4" captionText="FIGURE 2. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore. A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, buccal cavity, anterior thoracic sternum, pleonal somites 3–6 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674241" httpUri="https://zenodo.org/record/4674241/files/figure.png" pageId="1" pageNumber="370">2C</figureCitation>
, 
<figureCitation id="137F2A3FFF9FFFBDFD5BFDC683314A68" box="[737,770,542,568]" captionStart="FIGURE 3" captionStartId="5.[151,250,1254,1278]" captionTargetBox="[151,1436,181,1230]" captionTargetId="figure-15@5.[151,1436,181,1230]" captionTargetPageId="5" captionText="FIGURE 3. Ovilyra fuliginosa (Targioni-Tozzetti, 1877). A, male (11.2 × 14.0 mm) (ZRC 2009.0217), Singapore; B, male (13.3 × 16.2 mm) (ZRC 2009.0217), Singapore; C–E, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Muar, Peninsular Malaysia.A–C, overall dorsal view; D, anterior thoracic sternum, pleonal somites 3–6 and telson; E, left third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.4674243" httpUri="https://zenodo.org/record/4674243/files/figure.png" pageId="1" pageNumber="370">3E</figureCitation>
, 
<figureCitation id="137F2A3FFF9FFFBDFCB5FDC683654A69" box="[783,854,542,569]" captionStart="FIGURE 5" captionStartId="7.[151,250,1928,1952]" captionTargetBox="[215,1387,688,1892]" captionTargetId="figure-278@7.[151,1436,669,1905]" captionTargetPageId="7" captionText="FIGURE 5. Ovilyra fuliginosa (Targioni-Tozzetti, 1877).A, D, F, male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia; B, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Malaysia; C, E, G, male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore.A–C, left third maxilliped; D, E, frontal margin; F, G, pleon. Setae on structures not drawn. Scales: A, D = 1.0 mm; B, C, E–G = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4674247" httpUri="https://zenodo.org/record/4674247/files/figure.png" pageId="1" pageNumber="370">5A–C</figureCitation>
); cheliped merus subcylindrical, symmetrical along entire length, chela stout, pollex bent at angle of 45–60º along longitudinal axis, cutting edge of pollex with broad, large subproximal tooth lined with denticles, that of dactylus with subproximal concavity lined with denticles (
<figureCitation id="137F2A3FFF9FFFBDFADEFDBE808F4AF4" captionStart="FIGURE 1" captionStartId="3.[151,250,1921,1945]" captionTargetBox="[169,1418,181,1897]" captionTargetId="figure-15@3.[169,1418,181,1897]" captionTargetPageId="3" captionText="FIGURE 1. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia.A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, anterior thoracic sternum, pleonal somites 3–6 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674239" httpUri="https://zenodo.org/record/4674239/files/figure.png" pageId="1" pageNumber="370">Figs. 1A</figureCitation>
, 
<figureCitation id="137F2A3FFF9FFFBDFF73FD5280DE4AF4" box="[201,237,650,676]" captionStart="FIGURE 2" captionStartId="4.[151,250,1934,1958]" captionTargetBox="[164,1422,181,1910]" captionTargetId="figure-19@4.[164,1422,181,1910]" captionTargetPageId="4" captionText="FIGURE 2. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore. A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, buccal cavity, anterior thoracic sternum, pleonal somites 3–6 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674241" httpUri="https://zenodo.org/record/4674241/files/figure.png" pageId="1" pageNumber="370">2A</figureCitation>
, 
<figureCitation id="137F2A3FFF9FFFBDFF41FD5281714AF4" box="[251,322,650,676]" captionStart="FIGURE 3" captionStartId="5.[151,250,1254,1278]" captionTargetBox="[151,1436,181,1230]" captionTargetId="figure-15@5.[151,1436,181,1230]" captionTargetPageId="5" captionText="FIGURE 3. Ovilyra fuliginosa (Targioni-Tozzetti, 1877). A, male (11.2 × 14.0 mm) (ZRC 2009.0217), Singapore; B, male (13.3 × 16.2 mm) (ZRC 2009.0217), Singapore; C–E, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Muar, Peninsular Malaysia.A–C, overall dorsal view; D, anterior thoracic sternum, pleonal somites 3–6 and telson; E, left third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.4674243" httpUri="https://zenodo.org/record/4674243/files/figure.png" pageId="1" pageNumber="370">3A–C</figureCitation>
); P2–P5 slender, short, merus unarmed (
<figureCitation id="137F2A3FFF9FFFBDFCB5FD5283A14AF4" box="[783,914,650,676]" captionStart="FIGURE 1" captionStartId="3.[151,250,1921,1945]" captionTargetBox="[169,1418,181,1897]" captionTargetId="figure-15@3.[169,1418,181,1897]" captionTargetPageId="3" captionText="FIGURE 1. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia.A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, anterior thoracic sternum, pleonal somites 3–6 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674239" httpUri="https://zenodo.org/record/4674239/files/figure.png" pageId="1" pageNumber="370">Figs. 1A, F</figureCitation>
, 
<figureCitation id="137F2A3FFF9FFFBDFC25FD5283D04AF4" box="[927,995,650,676]" captionStart="FIGURE 2" captionStartId="4.[151,250,1934,1958]" captionTargetBox="[164,1422,181,1910]" captionTargetId="figure-19@4.[164,1422,181,1910]" captionTargetPageId="4" captionText="FIGURE 2. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore. A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, buccal cavity, anterior thoracic sternum, pleonal somites 3–6 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674241" httpUri="https://zenodo.org/record/4674241/files/figure.png" pageId="1" pageNumber="370">2A, F</figureCitation>
); male thoracic sternite 8 visible when pleon closed, between margins of somites 2 and 3 (
<figureCitation id="137F2A3FFF9FFFBDFD71FD7683114A98" box="[715,802,686,712]" captionStart="FIGURE 4" captionStartId="6.[151,250,1167,1191]" captionTargetBox="[151,1436,417,1143]" captionTargetId="figure-141@6.[151,1436,417,1143]" captionTargetPageId="6" captionText="FIGURE 4. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia. A, buccal cavity, anterior thoracic sternum and pleonal somites 3–6 and telson; B, posterior thoracic sternum and pleonal somites 1–4. Abbreviations: ca = carapace; cx5 = coxa of fourth ambulatory leg; so1–so6 = pleonal somites 1–6; s1–s8 = sternites 1–8; te = telson." figureDoi="http://doi.org/10.5281/zenodo.4674245" httpUri="https://zenodo.org/record/4674245/files/figure.png" pageId="1" pageNumber="370">Fig. 4B</figureCitation>
); sternopleonal cavity deep, reaching to mid-distance between fused thoracic sternites 1–3 (
<figureCitation id="137F2A3FFF9FFFBDFD9BFD0A82B34ABC" box="[545,640,722,748]" captionStart="FIGURE 1" captionStartId="3.[151,250,1921,1945]" captionTargetBox="[169,1418,181,1897]" captionTargetId="figure-15@3.[169,1418,181,1897]" captionTargetPageId="3" captionText="FIGURE 1. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia.A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, anterior thoracic sternum, pleonal somites 3–6 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674239" httpUri="https://zenodo.org/record/4674239/files/figure.png" pageId="1" pageNumber="370">Figs. 1E</figureCitation>
, 
<figureCitation id="137F2A3FFF9FFFBDFD36FD0A829E4ABC" box="[652,685,722,748]" captionStart="FIGURE 2" captionStartId="4.[151,250,1934,1958]" captionTargetBox="[164,1422,181,1910]" captionTargetId="figure-19@4.[164,1422,181,1910]" captionTargetPageId="4" captionText="FIGURE 2. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore. A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, buccal cavity, anterior thoracic sternum, pleonal somites 3–6 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674241" httpUri="https://zenodo.org/record/4674241/files/figure.png" pageId="1" pageNumber="370">2E</figureCitation>
, 
<figureCitation id="137F2A3FFF9FFFBDFD03FD0A82EC4ABC" box="[697,735,722,748]" captionStart="FIGURE 4" captionStartId="6.[151,250,1167,1191]" captionTargetBox="[151,1436,417,1143]" captionTargetId="figure-141@6.[151,1436,417,1143]" captionTargetPageId="6" captionText="FIGURE 4. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia. A, buccal cavity, anterior thoracic sternum and pleonal somites 3–6 and telson; B, posterior thoracic sternum and pleonal somites 1–4. Abbreviations: ca = carapace; cx5 = coxa of fourth ambulatory leg; so1–so6 = pleonal somites 1–6; s1–s8 = sternites 1–8; te = telson." figureDoi="http://doi.org/10.5281/zenodo.4674245" httpUri="https://zenodo.org/record/4674245/files/figure.png" pageId="1" pageNumber="370">4A</figureCitation>
); male pleon narrow, with somites 1 and 2 free, somite 1 longitudinally narrow, somite 2 yoke-like, reaching coxae of fourth ambulatory leg (
<figureCitation id="137F2A3FFF9FFFBDFBBBFD2E846C4B40" box="[1025,1119,758,784]" captionStart="FIGURE 1" captionStartId="3.[151,250,1921,1945]" captionTargetBox="[169,1418,181,1897]" captionTargetId="figure-15@3.[169,1418,181,1897]" captionTargetPageId="3" captionText="FIGURE 1. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia.A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, anterior thoracic sternum, pleonal somites 3–6 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674239" httpUri="https://zenodo.org/record/4674239/files/figure.png" pageId="1" pageNumber="370">Figs. 1E</figureCitation>
, 
<figureCitation id="137F2A3FFF9FFFBDFBD1FD2E84BF4B40" box="[1131,1164,758,784]" captionStart="FIGURE 2" captionStartId="4.[151,250,1934,1958]" captionTargetBox="[164,1422,181,1910]" captionTargetId="figure-19@4.[164,1422,181,1910]" captionTargetPageId="4" captionText="FIGURE 2. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore. A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, buccal cavity, anterior thoracic sternum, pleonal somites 3–6 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674241" httpUri="https://zenodo.org/record/4674241/files/figure.png" pageId="1" pageNumber="370">2E</figureCitation>
, 
<figureCitation id="137F2A3FFF9FFFBDFB2DFD2E848A4B40" box="[1175,1209,758,784]" captionStart="FIGURE 4" captionStartId="6.[151,250,1167,1191]" captionTargetBox="[151,1436,417,1143]" captionTargetId="figure-141@6.[151,1436,417,1143]" captionTargetPageId="6" captionText="FIGURE 4. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia. A, buccal cavity, anterior thoracic sternum and pleonal somites 3–6 and telson; B, posterior thoracic sternum and pleonal somites 1–4. Abbreviations: ca = carapace; cx5 = coxa of fourth ambulatory leg; so1–so6 = pleonal somites 1–6; s1–s8 = sternites 1–8; te = telson." figureDoi="http://doi.org/10.5281/zenodo.4674245" httpUri="https://zenodo.org/record/4674245/files/figure.png" pageId="1" pageNumber="370">4B</figureCitation>
, 
<figureCitation id="137F2A3FFF9FFFBDFB7EFD2E85304B40" box="[1220,1283,758,785]" captionStart="FIGURE 5" captionStartId="7.[151,250,1928,1952]" captionTargetBox="[215,1387,688,1892]" captionTargetId="figure-278@7.[151,1436,669,1905]" captionTargetPageId="7" captionText="FIGURE 5. Ovilyra fuliginosa (Targioni-Tozzetti, 1877).A, D, F, male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia; B, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Malaysia; C, E, G, male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore.A–C, left third maxilliped; D, E, frontal margin; F, G, pleon. Setae on structures not drawn. Scales: A, D = 1.0 mm; B, C, E–G = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4674247" httpUri="https://zenodo.org/record/4674247/files/figure.png" pageId="1" pageNumber="370">5F, G</figureCitation>
), somites 3–6 fused, forming elongate trapezoidal plate, surface smooth, unarmed (
<figureCitation id="137F2A3FFF9FFFBDFC1AFCC284364B64" box="[928,1029,794,820]" captionStart="FIGURE 3" captionStartId="5.[151,250,1254,1278]" captionTargetBox="[151,1436,181,1230]" captionTargetId="figure-15@5.[151,1436,181,1230]" captionTargetPageId="5" captionText="FIGURE 3. Ovilyra fuliginosa (Targioni-Tozzetti, 1877). A, male (11.2 × 14.0 mm) (ZRC 2009.0217), Singapore; B, male (13.3 × 16.2 mm) (ZRC 2009.0217), Singapore; C–E, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Muar, Peninsular Malaysia.A–C, overall dorsal view; D, anterior thoracic sternum, pleonal somites 3–6 and telson; E, left third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.4674243" httpUri="https://zenodo.org/record/4674243/files/figure.png" pageId="1" pageNumber="370">Figs. 3D</figureCitation>
, 
<figureCitation id="137F2A3FFF9FFFBDFBABFCC284074B64" box="[1041,1076,794,820]" captionStart="FIGURE 4" captionStartId="6.[151,250,1167,1191]" captionTargetBox="[151,1436,417,1143]" captionTargetId="figure-141@6.[151,1436,417,1143]" captionTargetPageId="6" captionText="FIGURE 4. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia. A, buccal cavity, anterior thoracic sternum and pleonal somites 3–6 and telson; B, posterior thoracic sternum and pleonal somites 1–4. Abbreviations: ca = carapace; cx5 = coxa of fourth ambulatory leg; so1–so6 = pleonal somites 1–6; s1–s8 = sternites 1–8; te = telson." figureDoi="http://doi.org/10.5281/zenodo.4674245" httpUri="https://zenodo.org/record/4674245/files/figure.png" pageId="1" pageNumber="370">4B</figureCitation>
, 
<figureCitation id="137F2A3FFF9FFFBDFBFBFCC284B14B64" box="[1089,1154,794,821]" captionStart="FIGURE 5" captionStartId="7.[151,250,1928,1952]" captionTargetBox="[215,1387,688,1892]" captionTargetId="figure-278@7.[151,1436,669,1905]" captionTargetPageId="7" captionText="FIGURE 5. Ovilyra fuliginosa (Targioni-Tozzetti, 1877).A, D, F, male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia; B, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Malaysia; C, E, G, male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore.A–C, left third maxilliped; D, E, frontal margin; F, G, pleon. Setae on structures not drawn. Scales: A, D = 1.0 mm; B, C, E–G = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4674247" httpUri="https://zenodo.org/record/4674247/files/figure.png" pageId="1" pageNumber="370">5F, G</figureCitation>
), somite 6 longitudinally subrectangular, free, surface unarmed (
<figureCitation id="137F2A3FFF9FFFBDFDF5FCE6829C4B08" box="[591,687,830,856]" captionStart="FIGURE 1" captionStartId="3.[151,250,1921,1945]" captionTargetBox="[169,1418,181,1897]" captionTargetId="figure-15@3.[169,1418,181,1897]" captionTargetPageId="3" captionText="FIGURE 1. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia.A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, anterior thoracic sternum, pleonal somites 3–6 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674239" httpUri="https://zenodo.org/record/4674239/files/figure.png" pageId="1" pageNumber="370">Figs. 1E</figureCitation>
, 
<figureCitation id="137F2A3FFF9FFFBDFD07FCE682ED4B08" box="[701,734,830,856]" captionStart="FIGURE 2" captionStartId="4.[151,250,1934,1958]" captionTargetBox="[164,1422,181,1910]" captionTargetId="figure-19@4.[164,1422,181,1910]" captionTargetPageId="4" captionText="FIGURE 2. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore. A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, buccal cavity, anterior thoracic sternum, pleonal somites 3–6 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674241" httpUri="https://zenodo.org/record/4674241/files/figure.png" pageId="1" pageNumber="370">2E</figureCitation>
, 
<figureCitation id="137F2A3FFF9FFFBDFD50FCE6833C4B08" box="[746,783,830,856]" captionStart="FIGURE 3" captionStartId="5.[151,250,1254,1278]" captionTargetBox="[151,1436,181,1230]" captionTargetId="figure-15@5.[151,1436,181,1230]" captionTargetPageId="5" captionText="FIGURE 3. Ovilyra fuliginosa (Targioni-Tozzetti, 1877). A, male (11.2 × 14.0 mm) (ZRC 2009.0217), Singapore; B, male (13.3 × 16.2 mm) (ZRC 2009.0217), Singapore; C–E, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Muar, Peninsular Malaysia.A–C, overall dorsal view; D, anterior thoracic sternum, pleonal somites 3–6 and telson; E, left third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.4674243" httpUri="https://zenodo.org/record/4674243/files/figure.png" pageId="1" pageNumber="370">3D</figureCitation>
, 
<figureCitation id="137F2A3FFF9FFFBDFCA6FCE683734B08" box="[796,832,830,856]" captionStart="FIGURE 4" captionStartId="6.[151,250,1167,1191]" captionTargetBox="[151,1436,417,1143]" captionTargetId="figure-141@6.[151,1436,417,1143]" captionTargetPageId="6" captionText="FIGURE 4. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia. A, buccal cavity, anterior thoracic sternum and pleonal somites 3–6 and telson; B, posterior thoracic sternum and pleonal somites 1–4. Abbreviations: ca = carapace; cx5 = coxa of fourth ambulatory leg; so1–so6 = pleonal somites 1–6; s1–s8 = sternites 1–8; te = telson." figureDoi="http://doi.org/10.5281/zenodo.4674245" httpUri="https://zenodo.org/record/4674245/files/figure.png" pageId="1" pageNumber="370">4A</figureCitation>
, 
<figureCitation id="137F2A3FFF9FFFBDFCF7FCE683BD4B08" box="[845,910,830,857]" captionStart="FIGURE 5" captionStartId="7.[151,250,1928,1952]" captionTargetBox="[215,1387,688,1892]" captionTargetId="figure-278@7.[151,1436,669,1905]" captionTargetPageId="7" captionText="FIGURE 5. Ovilyra fuliginosa (Targioni-Tozzetti, 1877).A, D, F, male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia; B, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Malaysia; C, E, G, male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore.A–C, left third maxilliped; D, E, frontal margin; F, G, pleon. Setae on structures not drawn. Scales: A, D = 1.0 mm; B, C, E–G = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4674247" httpUri="https://zenodo.org/record/4674247/files/figure.png" pageId="1" pageNumber="370">5F, G</figureCitation>
), telson triangular, longer than wide (
<figureCitation id="137F2A3FFF9FFFBDFA8CFCE685A44B08" box="[1334,1431,830,856]" captionStart="FIGURE 1" captionStartId="3.[151,250,1921,1945]" captionTargetBox="[169,1418,181,1897]" captionTargetId="figure-15@3.[169,1418,181,1897]" captionTargetPageId="3" captionText="FIGURE 1. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia.A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, anterior thoracic sternum, pleonal somites 3–6 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674239" httpUri="https://zenodo.org/record/4674239/files/figure.png" pageId="1" pageNumber="370">Figs. 1E</figureCitation>
, 
<figureCitation id="137F2A3FFF9FFFBDFF2DFCBA808A4B2C" box="[151,185,866,892]" captionStart="FIGURE 2" captionStartId="4.[151,250,1934,1958]" captionTargetBox="[164,1422,181,1910]" captionTargetId="figure-19@4.[164,1422,181,1910]" captionTargetPageId="4" captionText="FIGURE 2. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore. A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, buccal cavity, anterior thoracic sternum, pleonal somites 3–6 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674241" httpUri="https://zenodo.org/record/4674241/files/figure.png" pageId="1" pageNumber="370">2E</figureCitation>
, 
<figureCitation id="137F2A3FFF9FFFBDFF79FCBA80D44B2C" box="[195,231,866,892]" captionStart="FIGURE 3" captionStartId="5.[151,250,1254,1278]" captionTargetBox="[151,1436,181,1230]" captionTargetId="figure-15@5.[151,1436,181,1230]" captionTargetPageId="5" captionText="FIGURE 3. Ovilyra fuliginosa (Targioni-Tozzetti, 1877). A, male (11.2 × 14.0 mm) (ZRC 2009.0217), Singapore; B, male (13.3 × 16.2 mm) (ZRC 2009.0217), Singapore; C–E, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Muar, Peninsular Malaysia.A–C, overall dorsal view; D, anterior thoracic sternum, pleonal somites 3–6 and telson; E, left third maxilliped." figureDoi="http://doi.org/10.5281/zenodo.4674243" httpUri="https://zenodo.org/record/4674243/files/figure.png" pageId="1" pageNumber="370">3D</figureCitation>
, 
<figureCitation id="137F2A3FFF9FFFBDFF48FCBA81244B2C" box="[242,279,866,892]" captionStart="FIGURE 4" captionStartId="6.[151,250,1167,1191]" captionTargetBox="[151,1436,417,1143]" captionTargetId="figure-141@6.[151,1436,417,1143]" captionTargetPageId="6" captionText="FIGURE 4. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia. A, buccal cavity, anterior thoracic sternum and pleonal somites 3–6 and telson; B, posterior thoracic sternum and pleonal somites 1–4. Abbreviations: ca = carapace; cx5 = coxa of fourth ambulatory leg; so1–so6 = pleonal somites 1–6; s1–s8 = sternites 1–8; te = telson." figureDoi="http://doi.org/10.5281/zenodo.4674245" httpUri="https://zenodo.org/record/4674245/files/figure.png" pageId="1" pageNumber="370">4A</figureCitation>
, 
<figureCitation id="137F2A3FFF9FFFBDFE9BFCBA81524B2C" box="[289,353,866,893]" captionStart="FIGURE 5" captionStartId="7.[151,250,1928,1952]" captionTargetBox="[215,1387,688,1892]" captionTargetId="figure-278@7.[151,1436,669,1905]" captionTargetPageId="7" captionText="FIGURE 5. Ovilyra fuliginosa (Targioni-Tozzetti, 1877).A, D, F, male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia; B, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Malaysia; C, E, G, male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore.A–C, left third maxilliped; D, E, frontal margin; F, G, pleon. Setae on structures not drawn. Scales: A, D = 1.0 mm; B, C, E–G = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4674247" httpUri="https://zenodo.org/record/4674247/files/figure.png" pageId="1" pageNumber="370">5F, G</figureCitation>
); G1 elongate, slender, distal quarter sharply bent backwards 120–150º from the longitudinal axis; with curved, tapering subdistal process, outer margin lined with 1 or 2 rows of 5–8 elongate spines, distal projection elongate, curved, hook-shape, reaches to, overlaps or overreaches shaft, opening distal (
<figureCitation id="137F2A3FFF9FFFBDFBD5FC7285554B94" box="[1135,1382,938,964]" captionStart="FIGURE 6" captionStartId="8.[151,250,1906,1930]" captionTargetBox="[193,1352,191,1868]" captionTargetId="figure-19@8.[178,1409,181,1882]" captionTargetPageId="8" captionText="FIGURE 6. Ovilyra fuliginosa (Targioni-Tozzetti, 1877).A–D, male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia; E–H, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Malaysia; I–L, male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore.A, E, I, left G1 (ventral view); B, F, J, distal part of left G1 (ventral view); C, G, K, distal part of left G1 (dorsal view); D, H, L, left G2. Setae on structures not drawn. Scales: A, D = 1.0 mm; B, C, E–L, = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4674249" httpUri="https://zenodo.org/record/4674249/files/figure.png" pageId="1" pageNumber="370">Fig. 6A–C, E–G, I–K</figureCitation>
); G2 short, ca. one-third length of straight part of G1, apex subspatuliform (
<figureCitation id="137F2A3FFF9FFFBDFC09FC1684634BB8" box="[947,1104,974,1000]" captionStart="FIGURE 6" captionStartId="8.[151,250,1906,1930]" captionTargetBox="[193,1352,191,1868]" captionTargetId="figure-19@8.[178,1409,181,1882]" captionTargetPageId="8" captionText="FIGURE 6. Ovilyra fuliginosa (Targioni-Tozzetti, 1877).A–D, male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia; E–H, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Malaysia; I–L, male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore.A, E, I, left G1 (ventral view); B, F, J, distal part of left G1 (ventral view); C, G, K, distal part of left G1 (dorsal view); D, H, L, left G2. Setae on structures not drawn. Scales: A, D = 1.0 mm; B, C, E–L, = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4674249" httpUri="https://zenodo.org/record/4674249/files/figure.png" pageId="1" pageNumber="370">Fig. 6D, H, L</figureCitation>
); female pleon longitudinally ovate, somites 1 and 2 free, somites 3–6 completely fused to form domed plate (
<figureCitation id="137F2A3FFF9FFFBDFBA8FC2A84BB4C5C" box="[1042,1160,1010,1036]" captionStart="FIGURE 7" captionStartId="9.[151,250,1754,1778]" captionTargetBox="[151,1436,181,1730]" captionTargetId="figure-15@9.[151,1436,181,1730]" captionTargetPageId="9" captionText="FIGURE 7. Ovilyra fuliginosa (Targioni-Tozzetti, 1877). A–C, female (8.5 × 10.1 mm) (ZRC 2020.0345), Malaysia. D–F, female (9.3 × 11.6 mm) (ZRC 2009.0217), Singapore. A, D, overall dorsal view; B, E, pleon; C, F, vulvae." figureDoi="http://doi.org/10.5281/zenodo.4674251" httpUri="https://zenodo.org/record/4674251/files/figure.png" pageId="1" pageNumber="370">Fig. 7B, E</figureCitation>
); vulvae large, obliquely ovate, on anterior part of sternite 6, without sternal vulvar cover (
<figureCitation id="137F2A3FFF9FFFBDFCC9FBCE83D94C60" box="[883,1002,1046,1072]" captionStart="FIGURE 7" captionStartId="9.[151,250,1754,1778]" captionTargetBox="[151,1436,181,1730]" captionTargetId="figure-15@9.[151,1436,181,1730]" captionTargetPageId="9" captionText="FIGURE 7. Ovilyra fuliginosa (Targioni-Tozzetti, 1877). A–C, female (8.5 × 10.1 mm) (ZRC 2020.0345), Malaysia. D–F, female (9.3 × 11.6 mm) (ZRC 2009.0217), Singapore. A, D, overall dorsal view; B, E, pleon; C, F, vulvae." figureDoi="http://doi.org/10.5281/zenodo.4674251" httpUri="https://zenodo.org/record/4674251/files/figure.png" pageId="1" pageNumber="370">Fig. 7C, F</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="C35E6531FF9FFFBDFF7DFBE282844C28" pageId="1" pageNumber="370" type="etymology">
<paragraph id="8BFB36BAFF9FFFBDFF7DFBE282844C28" blockId="1.[151,1437,289,2008]" pageId="1" pageNumber="370">
<emphasis id="B930EAA8FF9FFFBDFF7DFBE2817B4C04" bold="true" box="[199,328,1082,1108]" pageId="1" pageNumber="370">Etymology</emphasis>
. The genus name is derived from the Latin “ovalis” for oval, in arbitrary combination with the suffix of the genus name 
<taxonomicName id="4C444D39FF9FFFBDFE2BFB8681D64C28" authorityName="Leach" authorityYear="1817" box="[401,485,1118,1144]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="370" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAA8FF9FFFBDFE2BFB8681D64C28" box="[401,485,1118,1144]" italics="true" pageId="1" pageNumber="370">Philyra</emphasis>
</taxonomicName>
. Gender feminine.
</paragraph>
</subSubSection>
<subSubSection id="C35E6531FF9FFFBEFF7DFB5A82BF49FD" lastPageId="2" lastPageNumber="371" pageId="1" pageNumber="370" type="discussion">
<paragraph id="8BFB36BAFF9FFFBDFF7DFB5A813D4FC0" blockId="1.[151,1437,289,2008]" pageId="1" pageNumber="370">
<emphasis id="B930EAA8FF9FFFBDFF7DFB5A81074CCC" bold="true" box="[199,308,1154,1180]" pageId="1" pageNumber="370">Remarks</emphasis>
. In 
<bibRefCitation id="EFD54B4BFF9FFFBDFED8FB5A82334CCC" author="Galil, B. S." box="[354,512,1154,1180]" pageId="1" pageNumber="370" pagination="279 - 320" refId="ref6530" refString="Galil, B. S. (2009) An examination of the genus Philyra Leach, 1817 (Crustacea, Decapoda, Leucosiidae) with descriptions of seven new genera and six new species. Zoosystema, 31 (2), 279 - 320. https: // doi. org / 10.5252 / z 2009 n 2 a 4" type="journal article" year="2009">Galil’s (2009)</bibRefCitation>
revision of 
<taxonomicName id="4C444D39FF9FFFBDFD33FB5A83404CCC" authority="Leach, 1817" authorityName="Leach" authorityYear="1817" box="[649,883,1154,1180]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="370" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAA8FF9FFFBDFD33FB5A82EE4CCC" box="[649,733,1154,1180]" italics="true" pageId="1" pageNumber="370">Philyra</emphasis>
<bibRefCitation id="EFD54B4BFF9FFFBDFD5FFB5A83404CCC" author="Leach, W. E." box="[741,883,1154,1180]" pageId="1" pageNumber="370" refId="ref6700" refString="Leach, W. E. (1817) The Zoological Miscellany; being descriptions of new or interesting animals … illustrated with coloured figures engraved from original drawings by R. P. Nodder &amp; c. Vol. 3: i-vii, 1 - 152, pls. 121 - 135, 135 B- 149. R. P. Nodder, London." type="book" year="1817">Leach, 1817</bibRefCitation>
</taxonomicName>
, the articulation of the first two male pleonal somites is a major generic character, separating 
<taxonomicName id="4C444D39FF9FFFBDFD1FFB7E82CA4C90" authorityName="Leach" authorityYear="1817" box="[677,761,1190,1216]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="370" phylum="Arthropoda" rank="genus" sensu="stricto">
<emphasis id="B930EAA8FF9FFFBDFD1FFB7E82CA4C90" box="[677,761,1190,1216]" italics="true" pageId="1" pageNumber="370">Philyra</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A20357D3FF9FFFBDFCBBFB7E83734C90" box="[769,832,1190,1216]" pageId="1" pageNumber="370" sensu="stricto">s. str.</taxonomicNameLabel>
, 
<taxonomicName id="4C444D39FF9FFFBDFCF4FB7E84774C90" authority="Galil, 2009" authorityName="Galil" authorityYear="2009" box="[846,1092,1190,1216]" class="Malacostraca" family="Leucosiidae" genus="Afrophila" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="370" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAA8FF9FFFBDFCF4FB7E838B4C90" box="[846,952,1190,1216]" italics="true" pageId="1" pageNumber="370">Afrophila</emphasis>
<bibRefCitation id="EFD54B4BFF9FFFBDFC7BFB7E84774C90" author="Galil, B. S." box="[961,1092,1190,1216]" pageId="1" pageNumber="370" pagination="279 - 320" refId="ref6530" refString="Galil, B. S. (2009) An examination of the genus Philyra Leach, 1817 (Crustacea, Decapoda, Leucosiidae) with descriptions of seven new genera and six new species. Zoosystema, 31 (2), 279 - 320. https: // doi. org / 10.5252 / z 2009 n 2 a 4" type="journal article" year="2009">Galil, 2009</bibRefCitation>
</taxonomicName>
, 
<taxonomicName id="4C444D39FF9FFFBDFBEBFB7E85544C90" authority="Galil, 2009" authorityName="Galil" authorityYear="2009" box="[1105,1383,1190,1216]" class="Malacostraca" family="Leucosiidae" genus="Atlantolocia" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="370" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAA8FF9FFFBDFBEBFB7E84E84C90" box="[1105,1243,1190,1216]" italics="true" pageId="1" pageNumber="370">Atlantolocia</emphasis>
<bibRefCitation id="EFD54B4BFF9FFFBDFB5EFB7E85544C90" author="Galil, B. S." box="[1252,1383,1190,1216]" pageId="1" pageNumber="370" pagination="279 - 320" refId="ref6530" refString="Galil, B. S. (2009) An examination of the genus Philyra Leach, 1817 (Crustacea, Decapoda, Leucosiidae) with descriptions of seven new genera and six new species. Zoosystema, 31 (2), 279 - 320. https: // doi. org / 10.5252 / z 2009 n 2 a 4" type="journal article" year="2009">Galil, 2009</bibRefCitation>
</taxonomicName>
, and 
<taxonomicName id="4C444D39FF9FFFBDFF2DFB12814A4CB4" authority="Galil, 2009" authorityName="Galil" authorityYear="2009" box="[151,377,1226,1252]" class="Malacostraca" family="Leucosiidae" genus="Ryphila" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="370" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAA8FF9FFFBDFF2DFB1280DD4CB4" box="[151,238,1226,1252]" italics="true" pageId="1" pageNumber="370">Ryphila</emphasis>
<bibRefCitation id="EFD54B4BFF9FFFBDFF4CFB12814A4CB4" author="Galil, B. S." box="[246,377,1226,1252]" pageId="1" pageNumber="370" pagination="279 - 320" refId="ref6530" refString="Galil, B. S. (2009) An examination of the genus Philyra Leach, 1817 (Crustacea, Decapoda, Leucosiidae) with descriptions of seven new genera and six new species. Zoosystema, 31 (2), 279 - 320. https: // doi. org / 10.5252 / z 2009 n 2 a 4" type="journal article" year="2009">Galil, 2009</bibRefCitation>
</taxonomicName>
, from other genera. With regards to this character, 
<taxonomicName id="4C444D39FF9FFFBDFC05FB1285AF4CB4" authority="Targioni-Tozzetti, 1877" authorityName="Targioni-Tozzetti" authorityYear="1877" box="[959,1436,1226,1252]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="370" phylum="Arthropoda" rank="species" species="fuliginosa">
<emphasis id="B930EAA8FF9FFFBDFC05FB1284BF4CB4" box="[959,1164,1226,1252]" italics="true" pageId="1" pageNumber="370">Philyra fuliginosa</emphasis>
<bibRefCitation id="EFD54B4BFF9FFFBDFB2EFB1285AF4CB4" author="Targioni-Tozzetti, A." box="[1172,1436,1226,1252]" pageId="1" pageNumber="370" pagination="1 - 12" refId="ref7486" refString="Targioni-Tozzetti, A. (1877) Crostacei Brachyuri e Anomuri. Zoologia del viaggio intorno al Globo della R. Pirocorvetta Magenta durante gli anni 1865 - 1868. Pubblicazioni del R. Istituto di Studi superiori pratici e di Perfezionamento in Firenze, Sezione di Scienze fisiche e naturali, Successori le Monnier, Firenze, 1, i - xxix + 1 - 257 pp., pls. 1 - 12." type="book chapter" year="1877">Targioni-Tozzetti, 1877</bibRefCitation>
</taxonomicName>
(and its junior synonym, 
<taxonomicName id="4C444D39FF9FFFBDFE0CFB3782D04D58" authority="Rathbun, 1909" authorityName="Rathbun" authorityYear="1909" box="[438,739,1262,1288]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="370" phylum="Arthropoda" rank="species" species="olivacea">
<emphasis id="B930EAA8FF9FFFBDFE0CFB3782014D58" box="[438,562,1262,1288]" italics="true" pageId="1" pageNumber="370">P. olivacea</emphasis>
<bibRefCitation id="EFD54B4BFF9FFFBDFD80FB3682D04D58" author="Rathbun, M. J." box="[570,739,1262,1288]" pageId="1" pageNumber="370" pagination="107 - 114" refId="ref7110" refString="Rathbun, M. J. (1909) New crabs from the Gulf of Siam. Proceedings of the Biological Society of Washington, 22, 107 - 114." type="journal article" year="1909">Rathbun, 1909</bibRefCitation>
</taxonomicName>
, see below), clearly belongs with the species in this group of genera. 
<taxonomicName id="4C444D39FF9FFFBDFF4AFACA818F4D7C" authorityName="Targioni-Tozzetti" authorityYear="1877" box="[240,444,1298,1324]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="370" phylum="Arthropoda" rank="species" species="fuliginosa">
<emphasis id="B930EAA8FF9FFFBDFF4AFACA818F4D7C" box="[240,444,1298,1324]" italics="true" pageId="1" pageNumber="370">Philyra fuliginosa</emphasis>
</taxonomicName>
can, however, be immediately distinguished by its carapace being distinctly longer than wide (length to width ratio 1.17–126); the others being all wider than long or at most subequal. The lobiform tooth present at the junction between the posterolateral and posterior carapace margins (
<figureCitation id="137F2A3FFF9FFFBDFBFBFA8284974D24" box="[1089,1188,1370,1396]" captionStart="FIGURE 1" captionStartId="3.[151,250,1921,1945]" captionTargetBox="[169,1418,181,1897]" captionTargetId="figure-15@3.[169,1418,181,1897]" captionTargetPageId="3" captionText="FIGURE 1. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia.A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, anterior thoracic sternum, pleonal somites 3–6 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674239" httpUri="https://zenodo.org/record/4674239/files/figure.png" pageId="1" pageNumber="370">Figs. 1B</figureCitation>
, 
<figureCitation id="137F2A3FFF9FFFBDFB08FA8284E74D24" box="[1202,1236,1370,1396]" captionStart="FIGURE 2" captionStartId="4.[151,250,1934,1958]" captionTargetBox="[164,1422,181,1910]" captionTargetId="figure-19@4.[164,1422,181,1910]" captionTargetPageId="4" captionText="FIGURE 2. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore. A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, buccal cavity, anterior thoracic sternum, pleonal somites 3–6 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674241" httpUri="https://zenodo.org/record/4674241/files/figure.png" pageId="1" pageNumber="370">2B</figureCitation>
) is diagnostic for 
<taxonomicName id="4C444D39FF9FFFBDFF2DFAA781174DC8" authorityName="Targioni-Tozzetti" authorityYear="1877" box="[151,292,1406,1432]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="370" phylum="Arthropoda" rank="species" species="fuliginosa">
<emphasis id="B930EAA8FF9FFFBDFF2DFAA781174DC8" box="[151,292,1406,1432]" italics="true" pageId="1" pageNumber="370">P. fuliginosa</emphasis>
</taxonomicName>
, a feature absent in the other genera (
<bibRefCitation id="EFD54B4BFF9FFFBDFD6DFAA683664DC8" author="Galil, B. S." box="[727,853,1406,1432]" pageId="1" pageNumber="370" pagination="279 - 320" refId="ref6530" refString="Galil, B. S. (2009) An examination of the genus Philyra Leach, 1817 (Crustacea, Decapoda, Leucosiidae) with descriptions of seven new genera and six new species. Zoosystema, 31 (2), 279 - 320. https: // doi. org / 10.5252 / z 2009 n 2 a 4" type="journal article" year="2009">Galil 2009</bibRefCitation>
: figs. 1A, 4A, 5A, 22A; 
<bibRefCitation id="EFD54B4BFF9FFFBDFBC8FAA6850B4DC9" author="Galil, B. S. &amp; Ng, P. K. L." box="[1138,1336,1406,1433]" pageId="1" pageNumber="370" pagination="451 - 486" refId="ref6592" refString="Galil, B. S. &amp; Ng, P. K. L. (2015) Leucosiid crabs from Papua New Guinea, with descriptions of eight new species (Crustacea: Decapoda: Brachyura). Zootaxa, 4027 (4), 451 - 486. https: // doi. org / 10.11646 / zootaxa. 4027.4.1" type="journal article" year="2015">Galil &amp; Ng 2015</bibRefCitation>
: fig. 9A, B). 
<taxonomicName id="4C444D39FF9FFFBDFF78FA7A811F4DEC" authorityName="Galil" authorityYear="2009" box="[194,300,1442,1468]" class="Malacostraca" family="Leucosiidae" genus="Afrophila" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="370" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAA8FF9FFFBDFF78FA7A811F4DEC" box="[194,300,1442,1468]" italics="true" pageId="1" pageNumber="370">Afrophila</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C444D39FF9FFFBDFED9FA7A81894DEC" authorityName="Galil" authorityYear="2009" box="[355,442,1442,1468]" class="Malacostraca" family="Leucosiidae" genus="Ryphila" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="370" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAA8FF9FFFBDFED9FA7A81894DEC" box="[355,442,1442,1468]" italics="true" pageId="1" pageNumber="370">Ryphila</emphasis>
</taxonomicName>
have visible hepatic facets (
<bibRefCitation id="EFD54B4BFF9FFFBDFD42FA7A83464DEC" author="Galil, B. S." box="[760,885,1442,1468]" pageId="1" pageNumber="370" pagination="279 - 320" refId="ref6530" refString="Galil, B. S. (2009) An examination of the genus Philyra Leach, 1817 (Crustacea, Decapoda, Leucosiidae) with descriptions of seven new genera and six new species. Zoosystema, 31 (2), 279 - 320. https: // doi. org / 10.5252 / z 2009 n 2 a 4" type="journal article" year="2009">Galil 2009</bibRefCitation>
: figs. 4A, 22A; 
<bibRefCitation id="EFD54B4BFF9FFFBDFB93FA7A84DF4DED" author="Galil, B. S. &amp; Ng, P. K. L." box="[1065,1260,1442,1469]" pageId="1" pageNumber="370" pagination="451 - 486" refId="ref6592" refString="Galil, B. S. &amp; Ng, P. K. L. (2015) Leucosiid crabs from Papua New Guinea, with descriptions of eight new species (Crustacea: Decapoda: Brachyura). Zootaxa, 4027 (4), 451 - 486. https: // doi. org / 10.11646 / zootaxa. 4027.4.1" type="journal article" year="2015">Galil &amp; Ng 2015</bibRefCitation>
: fig. 9A, B) but these are short compared to those in 
<taxonomicName id="4C444D39FF9FFFBDFD8AFA1F828F4DB0" authorityName="Targioni-Tozzetti" authorityYear="1877" box="[560,700,1478,1504]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="370" phylum="Arthropoda" rank="species" species="fuliginosa">
<emphasis id="B930EAA8FF9FFFBDFD8AFA1F828F4DB0" box="[560,700,1478,1504]" italics="true" pageId="1" pageNumber="370">P. fuliginosa</emphasis>
</taxonomicName>
(
<figureCitation id="137F2A3FFF9FFFBDFD71FA1E83634DB0" box="[715,848,1478,1504]" captionStart="FIGURE 1" captionStartId="3.[151,250,1921,1945]" captionTargetBox="[169,1418,181,1897]" captionTargetId="figure-15@3.[169,1418,181,1897]" captionTargetPageId="3" captionText="FIGURE 1. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia.A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, anterior thoracic sternum, pleonal somites 3–6 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674239" httpUri="https://zenodo.org/record/4674239/files/figure.png" pageId="1" pageNumber="370">Figs. 1B, D</figureCitation>
, 
<figureCitation id="137F2A3FFF9FFFBDFCE1FA1E83924DB0" box="[859,929,1478,1504]" captionStart="FIGURE 2" captionStartId="4.[151,250,1934,1958]" captionTargetBox="[164,1422,181,1910]" captionTargetId="figure-19@4.[164,1422,181,1910]" captionTargetPageId="4" captionText="FIGURE 2. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore. A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, buccal cavity, anterior thoracic sternum, pleonal somites 3–6 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674241" httpUri="https://zenodo.org/record/4674241/files/figure.png" pageId="1" pageNumber="370">2B, D</figureCitation>
), and the ventral margins do not have a distal submedian lobe; 
<taxonomicName id="4C444D39FF9FFFBDFEEDFA3281984E54" authorityName="Leach" authorityYear="1817" box="[343,427,1514,1540]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="370" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAA8FF9FFFBDFEEDFA3281984E54" box="[343,427,1514,1540]" italics="true" pageId="1" pageNumber="370">Philyra</emphasis>
</taxonomicName>
and 
<taxonomicName id="4C444D39FF9FFFBDFE5BFA3282584E54" authorityName="Galil" authorityYear="2009" box="[481,619,1514,1540]" class="Malacostraca" family="Leucosiidae" genus="Atlantolocia" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="370" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAA8FF9FFFBDFE5BFA3282584E54" box="[481,619,1514,1540]" italics="true" pageId="1" pageNumber="370">Atlantolocia</emphasis>
</taxonomicName>
have no clear hepatic facets (
<bibRefCitation id="EFD54B4BFF9FFFBDFC0DFA3284074E54" author="Galil, B. S." box="[951,1076,1514,1540]" pageId="1" pageNumber="370" pagination="279 - 320" refId="ref6530" refString="Galil, B. S. (2009) An examination of the genus Philyra Leach, 1817 (Crustacea, Decapoda, Leucosiidae) with descriptions of seven new genera and six new species. Zoosystema, 31 (2), 279 - 320. https: // doi. org / 10.5252 / z 2009 n 2 a 4" type="journal article" year="2009">Galil 2009</bibRefCitation>
: figs. 1A, 5A). The cutting edge of the fingers of the chelae are distinctively structured in 
<taxonomicName id="4C444D39FF9FFFBDFCB6F9D783A44E78" authorityName="Targioni-Tozzetti" authorityYear="1877" box="[780,919,1550,1576]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="370" phylum="Arthropoda" rank="species" species="fuliginosa">
<emphasis id="B930EAA8FF9FFFBDFCB6F9D783A44E78" box="[780,919,1550,1576]" italics="true" pageId="1" pageNumber="370">P. fuliginosa</emphasis>
</taxonomicName>
, with a subproximal lobe present on the pollex which fits into a concavity on the opposing cutting edge of the dactylus (
<figureCitation id="137F2A3FFF9FFFBDFC71F9EA841C4E1C" box="[971,1071,1586,1612]" captionStart="FIGURE 1" captionStartId="3.[151,250,1921,1945]" captionTargetBox="[169,1418,181,1897]" captionTargetId="figure-15@3.[169,1418,181,1897]" captionTargetPageId="3" captionText="FIGURE 1. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia.A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, anterior thoracic sternum, pleonal somites 3–6 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674239" httpUri="https://zenodo.org/record/4674239/files/figure.png" pageId="1" pageNumber="370">Figs. 1A</figureCitation>
, 
<figureCitation id="137F2A3FFF9FFFBDFB81F9EA84524E1C" box="[1083,1121,1586,1612]" captionStart="FIGURE 2" captionStartId="4.[151,250,1934,1958]" captionTargetBox="[164,1422,181,1910]" captionTargetId="figure-19@4.[164,1422,181,1910]" captionTargetPageId="4" captionText="FIGURE 2. Ovilyra fuliginosa (Targioni-Tozzetti, 1877), male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore. A, overall dorsal view; B, dorsal view of carapace; C, left third maxilliped; D, frontal view of cephalothorax; E, buccal cavity, anterior thoracic sternum, pleonal somites 3–6 and telson; F, left fourth ambulatory leg." figureDoi="http://doi.org/10.5281/zenodo.4674241" httpUri="https://zenodo.org/record/4674241/files/figure.png" pageId="1" pageNumber="370">2A</figureCitation>
). Such a structure is present in 
<taxonomicName id="4C444D39FF9FFFBDFF0FF98E813A4E20" authorityName="Leach" authorityYear="1817" box="[181,265,1622,1648]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="370" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAA8FF9FFFBDFF0FF98E813A4E20" box="[181,265,1622,1648]" italics="true" pageId="1" pageNumber="370">Philyra</emphasis>
</taxonomicName>
(
<bibRefCitation id="EFD54B4BFF9FFFBDFEA3F98E81A44E20" author="Galil, B. S." box="[281,407,1622,1648]" pageId="1" pageNumber="370" pagination="279 - 320" refId="ref6530" refString="Galil, B. S. (2009) An examination of the genus Philyra Leach, 1817 (Crustacea, Decapoda, Leucosiidae) with descriptions of seven new genera and six new species. Zoosystema, 31 (2), 279 - 320. https: // doi. org / 10.5252 / z 2009 n 2 a 4" type="journal article" year="2009">Galil 2009</bibRefCitation>
: fig. 1A), but absent or indistinct in the other three genera (
<bibRefCitation id="EFD54B4BFF9FFFBDFB84F98E848F4E20" author="Galil, B. S." box="[1086,1212,1622,1648]" pageId="1" pageNumber="370" pagination="279 - 320" refId="ref6530" refString="Galil, B. S. (2009) An examination of the genus Philyra Leach, 1817 (Crustacea, Decapoda, Leucosiidae) with descriptions of seven new genera and six new species. Zoosystema, 31 (2), 279 - 320. https: // doi. org / 10.5252 / z 2009 n 2 a 4" type="journal article" year="2009">Galil 2009</bibRefCitation>
: figs. 4A, 5A, 22A; 
<bibRefCitation id="EFD54B4BFF9FFFBDFF2DF9A2816A4EC5" author="Galil, B. S. &amp; Ng, P. K. L." box="[151,345,1658,1685]" pageId="1" pageNumber="370" pagination="451 - 486" refId="ref6592" refString="Galil, B. S. &amp; Ng, P. K. L. (2015) Leucosiid crabs from Papua New Guinea, with descriptions of eight new species (Crustacea: Decapoda: Brachyura). Zootaxa, 4027 (4), 451 - 486. https: // doi. org / 10.11646 / zootaxa. 4027.4.1" type="journal article" year="2015">Galil &amp; Ng 2015</bibRefCitation>
: fig. 10A). The narrow and slender male pleon of 
<taxonomicName id="4C444D39FF9FFFBDFC37F9A3842B4EC4" authorityName="Targioni-Tozzetti" authorityYear="1877" box="[909,1048,1658,1684]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="370" phylum="Arthropoda" rank="species" species="fuliginosa">
<emphasis id="B930EAA8FF9FFFBDFC37F9A3842B4EC4" box="[909,1048,1658,1684]" italics="true" pageId="1" pageNumber="370">P. fuliginosa</emphasis>
</taxonomicName>
(
<figureCitation id="137F2A3FFF9FFFBDFB92F9A284AF4EC4" box="[1064,1180,1658,1685]" captionStart="FIGURE 5" captionStartId="7.[151,250,1928,1952]" captionTargetBox="[215,1387,688,1892]" captionTargetId="figure-278@7.[151,1436,669,1905]" captionTargetPageId="7" captionText="FIGURE 5. Ovilyra fuliginosa (Targioni-Tozzetti, 1877).A, D, F, male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia; B, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Malaysia; C, E, G, male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore.A–C, left third maxilliped; D, E, frontal margin; F, G, pleon. Setae on structures not drawn. Scales: A, D = 1.0 mm; B, C, E–G = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4674247" httpUri="https://zenodo.org/record/4674247/files/figure.png" pageId="1" pageNumber="370">Fig. 5F, G</figureCitation>
) is very different from the proportionately more triangular structures of 
<taxonomicName id="4C444D39FF9FFFBDFD7CF94683294EE8" authorityName="Leach" authorityYear="1817" box="[710,794,1694,1720]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="370" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAA8FF9FFFBDFD7CF94683294EE8" box="[710,794,1694,1720]" italics="true" pageId="1" pageNumber="370">Philyra</emphasis>
</taxonomicName>
, 
<taxonomicName id="4C444D39FF9FFFBDFC90F94683A74EE8" authorityName="Galil" authorityYear="2009" box="[810,916,1694,1720]" class="Malacostraca" family="Leucosiidae" genus="Afrophila" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="370" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAA8FF9FFFBDFC90F94683A74EE8" box="[810,916,1694,1720]" italics="true" pageId="1" pageNumber="370">Afrophila</emphasis>
</taxonomicName>
or 
<taxonomicName id="4C444D39FF9FFFBDFC07F94684274EE8" authorityName="Galil" authorityYear="2009" box="[957,1044,1694,1720]" class="Malacostraca" family="Leucosiidae" genus="Ryphila" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="370" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAA8FF9FFFBDFC07F94684274EE8" box="[957,1044,1694,1720]" italics="true" pageId="1" pageNumber="370">Ryphila</emphasis>
</taxonomicName>
where somite 3 is proportionately wider (
<bibRefCitation id="EFD54B4BFF9FFFBDFF5DF91A81554E8C" author="Galil, B. S." box="[231,358,1730,1756]" pageId="1" pageNumber="370" pagination="279 - 320" refId="ref6530" refString="Galil, B. S. (2009) An examination of the genus Philyra Leach, 1817 (Crustacea, Decapoda, Leucosiidae) with descriptions of seven new genera and six new species. Zoosystema, 31 (2), 279 - 320. https: // doi. org / 10.5252 / z 2009 n 2 a 4" type="journal article" year="2009">Galil 2009</bibRefCitation>
: figs. 1B, 4B, 22B), and in 
<taxonomicName id="4C444D39FF9FFFBDFD1FF91A831C4E8C" authorityName="Galil" authorityYear="2009" box="[677,815,1730,1756]" class="Malacostraca" family="Leucosiidae" genus="Atlantolocia" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="370" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAA8FF9FFFBDFD1FF91A831C4E8C" box="[677,815,1730,1756]" italics="true" pageId="1" pageNumber="370">Atlantolocia</emphasis>
</taxonomicName>
, the pleon is wider and subrectangular in shape (
<bibRefCitation id="EFD54B4BFF9FFFBDFAD9F91A80E04F50" author="Galil, B. S." pageId="1" pageNumber="370" pagination="279 - 320" refId="ref6530" refString="Galil, B. S. (2009) An examination of the genus Philyra Leach, 1817 (Crustacea, Decapoda, Leucosiidae) with descriptions of seven new genera and six new species. Zoosystema, 31 (2), 279 - 320. https: // doi. org / 10.5252 / z 2009 n 2 a 4" type="journal article" year="2009">Galil 2009</bibRefCitation>
: fig. 5B). The strongly bent distal quarter of the G1 with the presence of a long subdistal process is also a character unique to 
<taxonomicName id="4C444D39FF9FFFBDFEFDF8D381E04F74" authorityName="Targioni-Tozzetti" authorityYear="1877" box="[327,467,1802,1828]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="370" phylum="Arthropoda" rank="species" species="fuliginosa">
<emphasis id="B930EAA8FF9FFFBDFEFDF8D381E04F74" box="[327,467,1802,1828]" italics="true" pageId="1" pageNumber="370">P. fuliginosa</emphasis>
</taxonomicName>
(
<figureCitation id="137F2A3FFF9FFFBDFE59F8D282474F74" box="[483,628,1802,1828]" captionStart="FIGURE 6" captionStartId="8.[151,250,1906,1930]" captionTargetBox="[193,1352,191,1868]" captionTargetId="figure-19@8.[178,1409,181,1882]" captionTargetPageId="8" captionText="FIGURE 6. Ovilyra fuliginosa (Targioni-Tozzetti, 1877).A–D, male (10.4 × 12.2 mm) (ZRC 2020.0345), Malaysia; E–H, male (6.9 × 8.7 mm) (ZRC 1970.8.8.7), Malaysia; I–L, male (12.3 × 14.8 mm) (ZRC 2009.0217), Singapore.A, E, I, left G1 (ventral view); B, F, J, distal part of left G1 (ventral view); C, G, K, distal part of left G1 (dorsal view); D, H, L, left G2. Setae on structures not drawn. Scales: A, D = 1.0 mm; B, C, E–L, = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.4674249" httpUri="https://zenodo.org/record/4674249/files/figure.png" pageId="1" pageNumber="370">Fig. 6A, E, I</figureCitation>
) and is not shared by these genera; all of which have straight or almost straight G1 structures (e.g., 
<bibRefCitation id="EFD54B4BFF9FFFBDFE62F8F682664F18" author="Galil, B. S." box="[472,597,1838,1864]" pageId="1" pageNumber="370" pagination="279 - 320" refId="ref6530" refString="Galil, B. S. (2009) An examination of the genus Philyra Leach, 1817 (Crustacea, Decapoda, Leucosiidae) with descriptions of seven new genera and six new species. Zoosystema, 31 (2), 279 - 320. https: // doi. org / 10.5252 / z 2009 n 2 a 4" type="journal article" year="2009">Galil 2009</bibRefCitation>
: figs. 2A–E, 21). In fact, none of the genera which have been previously associated with 
<taxonomicName id="4C444D39FF9FFFBDFEF0F88A81AD4F3C" authorityName="Leach" authorityYear="1817" box="[330,414,1874,1900]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="370" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAA8FF9FFFBDFEF0F88A81AD4F3C" box="[330,414,1874,1900]" italics="true" pageId="1" pageNumber="370">Philyra</emphasis>
</taxonomicName>
have such a strongly bent G1. As such, we here establish a new genus, 
<taxonomicName id="4C444D39FF9FFFBDFB07F88A85204F3C" authorityName="Ng" authorityYear="2021" box="[1213,1299,1874,1900]" class="Malacostraca" family="Leucosiidae" genus="Ovilyra" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="370" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAA8FF9FFFBDFB07F88A85204F3C" box="[1213,1299,1874,1900]" italics="true" pageId="1" pageNumber="370">Ovilyra</emphasis>
</taxonomicName>
, for 
<taxonomicName id="4C444D39FF9FFFBDFAF2F88A81344FC0" authorityName="Targioni-Tozzetti" authorityYear="1877" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="370" phylum="Arthropoda" rank="species" species="fuliginosa">
<emphasis id="B930EAA8FF9FFFBDFAF2F88A81344FC0" italics="true" pageId="1" pageNumber="370">Philyra fuliginosa</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8BFB36BAFF9FFFBEFF7DF842830849D9" blockId="1.[151,1437,289,2008]" lastBlockId="2.[151,1437,151,429]" lastPageId="2" lastPageNumber="371" pageId="1" pageNumber="370">
In describing 
<taxonomicName id="4C444D39FF9FFFBDFEDEF84282224FE4" authorityName="Kemp" authorityYear="1915" box="[356,529,1946,1972]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="370" phylum="Arthropoda" rank="species" species="alcocki">
<emphasis id="B930EAA8FF9FFFBDFEDEF84282224FE4" box="[356,529,1946,1972]" italics="true" pageId="1" pageNumber="370">Philyra alcocki</emphasis>
</taxonomicName>
from Lake Chilka in eastern 
<collectingCountry id="F353762AFF9FFFBDFCD8F84283AD4FE4" box="[866,926,1946,1972]" name="India" pageId="1" pageNumber="370">India</collectingCountry>
, 
<bibRefCitation id="EFD54B4BFF9FFFBDFC11F84284454FE4" author="Kemp, S." box="[939,1142,1946,1973]" pageId="1" pageNumber="370" pagination="199 - 325" refId="ref6654" refString="Kemp, S. (1915) Crustacea Decapoda. Fauna of the Chilka Lake No. 3. Memoirs of the Indian Museum, 5 (3), 199 - 325, figs. 1 - 37, pls. 12, 13." type="journal article" year="1915">Kemp (1915: 214</bibRefCitation>
, 215) commented that his species was closest to 
<taxonomicName id="4C444D39FF9FFFBDFE2EF867823C4F88" authorityName="Rathbun" authorityYear="1909" box="[404,527,1982,2008]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="370" phylum="Arthropoda" rank="species" species="olivacea">
<emphasis id="B930EAA8FF9FFFBDFE2EF867823C4F88" box="[404,527,1982,2008]" italics="true" pageId="1" pageNumber="370">P. olivacea</emphasis>
</taxonomicName>
and to a lesser degree to 
<taxonomicName id="4C444D39FF9FFFBDFC95F86783884F88" authorityName="Targioni-Tozzetti" authorityYear="1877" box="[815,955,1982,2008]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="1" pageNumber="370" phylum="Arthropoda" rank="species" species="fuliginosa">
<emphasis id="B930EAA8FF9FFFBDFC95F86783884F88" box="[815,955,1982,2008]" italics="true" pageId="1" pageNumber="370">P. fuliginosa</emphasis>
</taxonomicName>
. Although the carapace shape and features are superficially similar, 
<taxonomicName id="4C444D39FF9CFFBEFE17FF4F822A48E1" authorityName="Kemp" authorityYear="1915" box="[429,537,151,177]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="alcocki">
<emphasis id="B930EAA8FF9CFFBEFE17FF4F822A48E1" box="[429,537,151,177]" italics="true" pageId="2" pageNumber="371">P. alcocki</emphasis>
</taxonomicName>
is probably not congeneric with 
<taxonomicName id="4C444D39FF9CFFBEFC32FF4F842748E1" authorityName="Targioni-Tozzetti" authorityYear="1877" box="[904,1044,151,177]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="fuliginosa">
<emphasis id="B930EAA8FF9CFFBEFC32FF4F842748E1" box="[904,1044,151,177]" italics="true" pageId="2" pageNumber="371">P. fuliginosa</emphasis>
</taxonomicName>
as its carapace is only slightly longer than wide, being almost round, the male thoracic sternum is proportionately much wider, male pleonal somites 1–5 are fused (although sutures between somites 1–3 are still visible) with somite 3 very wide, giving the male pleon a distinctly triangular shape (
<bibRefCitation id="EFD54B4BFF9CFFBEFE62FEDB82AB494D" author="Kemp, S." box="[472,664,258,285]" pageId="2" pageNumber="371" pagination="199 - 325" refId="ref6654" refString="Kemp, S. (1915) Crustacea Decapoda. Fauna of the Chilka Lake No. 3. Memoirs of the Indian Museum, 5 (3), 199 - 325, figs. 1 - 37, pls. 12, 13." type="journal article" year="1915">Kemp 1915: 214</bibRefCitation>
, fig. 2, pl. 12 fig. 2). In 
<taxonomicName id="4C444D39FF9CFFBEFC26FEDB83C1494D" authorityName="Ng" authorityYear="2021" box="[924,1010,259,285]" class="Malacostraca" family="Leucosiidae" genus="Ovilyra" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="371" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAA8FF9CFFBEFC26FEDB83C1494D" box="[924,1010,259,285]" italics="true" pageId="2" pageNumber="371">Ovilyra</emphasis>
</taxonomicName>
species, the carapace is longitudinally ovate and much longer than wide, the male thoracic sternum proportionately narrow and male pleonal somite 3 is only slightly wider than the other somites, with the male pleon not obviously triangular in shape. “ 
<taxonomicName id="4C444D39FF9CFFBEFB5EFE93850B4935" authorityName="Leach" authorityYear="1817" box="[1252,1336,331,357]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="371" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAA8FF9CFFBEFB5EFE93850B4935" box="[1252,1336,331,357]" italics="true" pageId="2" pageNumber="371">Philyra</emphasis>
</taxonomicName>
” 
<taxonomicName id="4C444D39FF9CFFBEFAF1FE9385AF4935" authorityName="Kemp" authorityYear="1915" box="[1355,1436,331,357]" class="Malacostraca" family="Leucosiidae" genus="Philyra" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="371" phylum="Arthropoda" rank="species" species="alcocki">
<emphasis id="B930EAA8FF9CFFBEFAF1FE9385AF4935" box="[1355,1436,331,357]" italics="true" pageId="2" pageNumber="371">alcocki</emphasis>
</taxonomicName>
will need to be re-examined to ascertain its generic position.
</paragraph>
<paragraph id="8BFB36BAFF9CFFBEFF7DFE4B82BF49FD" blockId="2.[151,1437,151,429]" box="[199,652,403,429]" pageId="2" pageNumber="371">
<taxonomicName id="4C444D39FF9CFFBEFF7DFE4B812E49FD" authorityName="Ng" authorityYear="2021" box="[199,285,403,429]" class="Malacostraca" family="Leucosiidae" genus="Ovilyra" kingdom="Animalia" order="Decapoda" pageId="2" pageNumber="371" phylum="Arthropoda" rank="genus">
<emphasis id="B930EAA8FF9CFFBEFF7DFE4B812E49FD" box="[199,285,403,429]" italics="true" pageId="2" pageNumber="371">Ovilyra</emphasis>
</taxonomicName>
is monotypic for the time being.
</paragraph>
</subSubSection>
</treatment>
</document>