273 lines
34 KiB
XML
273 lines
34 KiB
XML
<document id="6DEAABE7CBF64D491FDAE1BC202459EF" ID-CLB-Dataset="307152" ID-DOI="10.11646/zootaxa.5562.1.10" ID-GBIF-Dataset="43c9f6fc-9bf7-4741-b8b2-653bdc9253da" ID-ISSN="1175-5326" ID-Zenodo-Dep="14595649" ID-ZooBank="63D02C0B-C7D3-4959-BB5F-34772BCF72F6" IM.bibliography_approvedBy="felipe" IM.illustrations_approvedBy="felipe" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" IM.treatments_approvedBy="felipe" checkinTime="1735931758043" checkinUser="plazi" docAuthor="Cai, Chenyang, Zhao, Qian & Azar, Dany" docDate="2024" docId="03EAC54AFE10FF9AC682FCF4FBA8FDF3" docLanguage="en" docName="zootaxa.5562.1.10.pdf" docOrigin="Zootaxa 5562 (1)" docSource="https://doi.org/10.11646/zootaxa.5562.1.10" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Elodophthalmus maksoudae Cai, Zhao & Azar, 2024, sp. nov." docType="treatment" docVersion="4" lastPageNumber="70" masterDocId="FFD3BD32FE12FF9FC615FFAAFF9BFFEA" masterDocTitle="A new species of Elodophthalmus from Lower Cretaceous Lebanese amber unravels the systematic placement of Elodophthalmidae and reconciling phylogenetic conflicts within Tenebrionoidea" masterLastPageNumber="75" masterPageNumber="65" pageNumber="67" updateTime="1735934705980" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
|
||
<mods:mods id="720237853387514D79845583A35087A7" xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo id="482CA2DB3960BAC7CBDAA63A3F390793">
|
||
<mods:title id="C17886D3914527394351EF4DDACAC1EF">A new species of Elodophthalmus from Lower Cretaceous Lebanese amber unravels the systematic placement of Elodophthalmidae and reconciling phylogenetic conflicts within Tenebrionoidea</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name id="B073FC64CDC5D3352E3D1957BC628292" type="personal">
|
||
<mods:role id="334A7B3A0A2DACD1433E4D84F3AF3212">
|
||
<mods:roleTerm id="EA5105200A031604C584D66FF7948E56">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="E60C0E6A199E494E9362595BB9B7E09A">Cai, Chenyang</mods:namePart>
|
||
<mods:affiliation id="8C4192EC298A843FA3A105F943A7340E">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="6FAF6E4354CB34BAB6803620CBA4FDA4" type="personal">
|
||
<mods:role id="1A6D5D8A4031670E0761FA241184FAAC">
|
||
<mods:roleTerm id="172007AD874B9C3C9C7939B3BE41494B">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="197008F9B9B3010C6A206465732962FE">Zhao, Qian</mods:namePart>
|
||
<mods:affiliation id="027A45AB41C0B7BCDF7D286A2FBAAEA6">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China</mods:affiliation>
|
||
</mods:name>
|
||
<mods:name id="A229B680EBCB87789ABB48F7B79ED4D5" type="personal">
|
||
<mods:role id="82AE79A3ABA9CDDCF301796CAC61DE5C">
|
||
<mods:roleTerm id="BF84316FABF60034BECB154E20C46A46">Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart id="E5B7A1DAA957633F3D79A88385D1F5CE">Azar, Dany</mods:namePart>
|
||
<mods:affiliation id="1F540C9419D9E19A531D7E85F40F45F9">State Key Laboratory of Palaeobiology and Stratigraphy, Nanjing Institute of Geology and Palaeontology, Chinese Academy of Sciences, Nanjing 210008, China & Faculty of Science II, Natural Sciences Department, Lebanese University, Fanar - El-Matn, PO Box 90656 Jdeideh, Lebanon * Corresponding authors</mods:affiliation>
|
||
</mods:name>
|
||
<mods:typeOfResource id="784D031CA702122B55DBDA5EFD797FD3">text</mods:typeOfResource>
|
||
<mods:relatedItem id="DF2445610F4C912484780CB7671E6191" type="host">
|
||
<mods:titleInfo id="A023CAA69A0BC779DDDB584663865C04">
|
||
<mods:title id="7E1641CDE7081DE8799E7855293D0123">Zootaxa</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part id="37A02D37095361E0B1F8A84C0B3659A2">
|
||
<mods:date id="3CC33DDE8AF9D3EF67973E658AC738C9">2024</mods:date>
|
||
<mods:detail id="9424B85C2DC7F08B684046EA358D9311" type="pubDate">
|
||
<mods:number id="741E7DF4FB328550893B10C2836C313D">2024-12-31</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="DFC9CAEE7C060E0A56E64712536FD6DE" type="volume">
|
||
<mods:number id="AC5BDABCBE8BA63053040744185C1893">5562</mods:number>
|
||
</mods:detail>
|
||
<mods:detail id="AFB269C6282B14979B8D5DB7401AAB63" type="issue">
|
||
<mods:number id="7CDB7BDEEDDA988D7C22B6E4E359EC04">1</mods:number>
|
||
</mods:detail>
|
||
<mods:extent id="FFD7E5A174BD28A8DC651DC9550CB9F9" unit="page">
|
||
<mods:start id="7E19D65ECADAC4DFB5D693D2BBA787C8">65</mods:start>
|
||
<mods:end id="FD54073BDDC856DDBC0069A67840AEC0">75</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:location id="D6A543B113AF5C8B2A0A437DC1E5AA0A">
|
||
<mods:url id="4840B69F710F424038F65EF785E2D9FB">https://doi.org/10.11646/zootaxa.5562.1.10</mods:url>
|
||
</mods:location>
|
||
<mods:classification id="2C11C6838410E32D1A35C38D4FC6D5ED">journal article</mods:classification>
|
||
<mods:identifier id="960752A5E46A1FD572434210B6522002" type="CLB-Dataset">307152</mods:identifier>
|
||
<mods:identifier id="00F6D2343514DBB7377E8ADD8CB143A8" type="DOI">10.11646/zootaxa.5562.1.10</mods:identifier>
|
||
<mods:identifier id="DD492DAE3B49FC8189702DB4E19B04C0" type="GBIF-Dataset">43c9f6fc-9bf7-4741-b8b2-653bdc9253da</mods:identifier>
|
||
<mods:identifier id="C8EF2A0BCCA5365E8D280957A8464AB0" type="ISSN">1175-5326</mods:identifier>
|
||
<mods:identifier id="7439E48A0C548A479B7C8C59C146DFD1" type="Zenodo-Dep">14595649</mods:identifier>
|
||
<mods:identifier id="3B9C2D16AD985C6116372BD3FBC1F2FE" type="ZooBank">63D02C0B-C7D3-4959-BB5F-34772BCF72F6</mods:identifier>
|
||
</mods:mods>
|
||
<treatment id="03EAC54AFE10FF9AC682FCF4FBA8FDF3" LSID="urn:lsid:plazi:treatment:03EAC54AFE10FF9AC682FCF4FBA8FDF3" httpUri="http://treatment.plazi.org/id/03EAC54AFE10FF9AC682FCF4FBA8FDF3" lastPageId="5" lastPageNumber="70" pageId="2" pageNumber="67" scope_family="Elodophthalmidae" scope_order="Coleoptera" scope_superFamily="Tenebrionoidea">
|
||
<subSubSection id="C35927D7FE10FF9DC682FCF4FDCEFC93" box="[151,597,862,889]" pageId="2" pageNumber="67" type="nomenclature">
|
||
<paragraph id="8BFC745CFE10FF9DC682FCF4FDCEFC93" blockId="2.[151,597,862,925]" box="[151,597,862,889]" pageId="2" pageNumber="67">
|
||
<heading id="D0B4C330FE10FF9DC682FCF4FDCEFC93" bold="true" box="[151,597,862,889]" fontSize="11" level="1" pageId="2" pageNumber="67" reason="1">
|
||
<emphasis id="B937A84EFE10FF9DC682FCF4FDCEFC93" bold="true" box="[151,597,862,889]" pageId="2" pageNumber="67">
|
||
<taxonomicName id="4C430FDFFE10FF9DC682FCF4FE6BFC93" authorityName="Cai & Zhao & Azar" authorityYear="2024" box="[151,496,862,889]" class="Insecta" family="Elodophthalmidae" genus="Elodophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="67" phylum="Arthropoda" rank="species" species="maksoudae" status="sp. nov.">
|
||
<emphasis id="B937A84EFE10FF9DC682FCF4FE6BFC93" bold="true" box="[151,496,862,889]" italics="true" pageId="2" pageNumber="67">Elodophthalmus maksoudae</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A2041535FE10FF9DC7E2FCF5FDCEFC93" box="[503,597,863,889]" pageId="2" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35927D7FE10FF9DC682FC29FE90FC77" box="[151,267,899,925]" pageId="2" pageNumber="67" type="description">
|
||
<paragraph id="8BFC745CFE10FF9DC682FC29FE90FC77" blockId="2.[151,597,862,925]" box="[151,267,899,925]" pageId="2" pageNumber="67">
|
||
(
|
||
<figureCitation id="137868D9FE10FF9DC68AFC29FE99FC77" box="[159,258,899,925]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="3.[151,250,1011,1035]" captionStartId-1="3.[151,250,1971,1995]" captionStartId-2="4.[151,250,1889,1913]" captionTargetBox-0="[168,1417,188,984]" captionTargetBox-1="[456,1131,1093,1945]" captionTargetBox-2="[230,1357,179,1864]" captionTargetId-0="figure-21@3.[158,1430,179,984]" captionTargetId-1="figure-57@3.[456,1131,1093,1945]" captionTargetId-2="figure-29@4.[230,1357,179,1864]" captionTargetPageId-0="3" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 1. General habitus of Elodophthalmus maksoudae sp. nov., holotype specimen 1821, D. Azar coll., under incident light. A, Dorsal view. B, Ventral view. Scale bars: 200 μm." captionText-1="FIGURE 2. General habitus of Elodophthalmus maksoudae sp. nov., holotype specimen 1821, D. Azar coll., under confocal microscopy. Ventral view. Scale bars: 200 μm." captionText-2="FIGURE 3. General habitus of Elodophthalmus maksoudae sp. nov., holotype specimen 1821, D. Azar coll., under confocal microscopy. A, Ventral view of body. B, Ventral view of head. C, Left antenna. D, Meso- and metatarsi. Abbreviations: a, antennomere; ey, compound eye; la, labrum; mp, maxillary palpomere; mst, mesotarsomere; mtt, metatarsimere. Scale bars: 200 μm in A and C; 100 μm in B and D." figureDoi-0="http://doi.org/10.5281/zenodo.14595651" figureDoi-1="http://doi.org/10.5281/zenodo.14595653" figureDoi-2="http://doi.org/10.5281/zenodo.14595657" httpUri-0="https://zenodo.org/record/14595651/files/figure.png" httpUri-1="https://zenodo.org/record/14595653/files/figure.png" httpUri-2="https://zenodo.org/record/14595657/files/figure.png" pageId="2" pageNumber="67">Figs 1–3</figureCitation>
|
||
)
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35927D7FE10FF9DC682FC60FE45FBE3" pageId="2" pageNumber="67" type="materials_examined">
|
||
<paragraph id="8BFC745CFE10FF9DC682FC60FE45FBE3" blockId="2.[151,1437,970,1933]" pageId="2" pageNumber="67">
|
||
<emphasis id="B937A84EFE10FF9DC682FC60FE92FC0E" bold="true" box="[151,265,970,996]" pageId="2" pageNumber="67">Material.</emphasis>
|
||
<materialsCitation id="3B2B7E01FE10FF9DC700FC61FE45FBE3" collectorName="D. Azar" country="Lebanon" location="Central" pageId="2" pageNumber="67" specimenCount="1" typeStatus="holotype">
|
||
<typeStatus id="54F8CAFEFE10FF9DC700FC61FE18FC0F" box="[277,387,971,997]" pageId="2" pageNumber="67" type="holotype">Holotype</typeStatus>
|
||
, 1821,
|
||
<collectorName id="26B6118AFE10FF9DC7CAFC61FDA5FC0F" box="[479,574,971,997]" pageId="2" pageNumber="67">D. Azar</collectorName>
|
||
collection, Hammana/Mdeyrij outcrop, Caza Baabda, Mouhafazet Jabal Loubnan (
|
||
<location id="8E9C2287FE10FF9DC719FC45FEF9FBE3" LSID="urn:lsid:plazi:treatment:03EAC54AFE10FF9AC682FCF4FBA8FDF3:8E9C2287FE10FF9DC719FC45FEF9FBE3" box="[268,354,1007,1033]" country="Lebanon" name="Central" pageId="2" pageNumber="67">Central</location>
|
||
<collectingCountry id="F35434CCFE10FF9DC77CFC45FE49FBE3" box="[361,466,1007,1033]" name="Lebanon" pageId="2" pageNumber="67">Lebanon</collectingCountry>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35927D7FE10FF9DC6D2FBB9FC29FBBB" pageId="2" pageNumber="67" type="etymology">
|
||
<paragraph id="8BFC745CFE10FF9DC6D2FBB9FC29FBBB" blockId="2.[151,1437,970,1933]" pageId="2" pageNumber="67">
|
||
<emphasis id="B937A84EFE10FF9DC6D2FBB9FED5FBC7" bold="true" box="[199,334,1043,1069]" pageId="2" pageNumber="67">Etymology.</emphasis>
|
||
The specific name is dedicated to Professor Sibelle Maksoud, who has made great contribution of the study of Lebanese amber and palaeontology in
|
||
<collectingCountry id="F35434CCFE10FF9DC4C7FB9DFCAFFBBB" box="[722,820,1079,1105]" name="Lebanon" pageId="2" pageNumber="67">Lebanon</collectingCountry>
|
||
in general.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35927D7FE10FF9DC6D2FBF0FE69FA53" pageId="2" pageNumber="67" type="diagnosis">
|
||
<paragraph id="8BFC745CFE10FF9DC6D2FBF0FD90FA9B" blockId="2.[151,1437,970,1933]" pageId="2" pageNumber="67">
|
||
<emphasis id="B937A84EFE10FF9DC6D2FBF0FED9FB9E" bold="true" box="[199,322,1114,1140]" pageId="2" pageNumber="67">Diagnosis.</emphasis>
|
||
<taxonomicName id="4C430FDFFE10FF9DC752FBF1FD18FB9F" authorityName="Cai & Zhao & Azar" authorityYear="2024" box="[327,643,1115,1141]" class="Insecta" family="Elodophthalmidae" genus="Elodophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="67" phylum="Arthropoda" rank="species" species="maksoudae" status="sp. nov.">
|
||
<emphasis id="B937A84EFE10FF9DC752FBF1FD18FB9F" box="[327,643,1115,1141]" italics="true" pageId="2" pageNumber="67">Elodophthalmus maksoudae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B937A84EFE10FF9DC49CFBF1FD79FB9F" bold="true" box="[649,738,1115,1141]" pageId="2" pageNumber="67">
|
||
<taxonomicNameLabel id="A2041535FE10FF9DC49CFBF1FD79FB9F" box="[649,738,1115,1141]" pageId="2" pageNumber="67" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
differs from the other two
|
||
<taxonomicName id="4C430FDFFE10FF9DC21EFBF1FA82FB9F" box="[1035,1305,1115,1141]" pageId="2" pageNumber="67">
|
||
<emphasis id="B937A84EFE10FF9DC21EFBF1FB58FB9F" box="[1035,1219,1115,1141]" italics="true" pageId="2" pageNumber="67">Elodophthalmus</emphasis>
|
||
species
|
||
</taxonomicName>
|
||
(
|
||
<taxonomicName id="4C430FDFFE10FF9DC33CFBF1FEF7FB73" authority="Kirejtshuk & Azar" authorityName="Kirejtshuk & Azar" authorityYear="2008" class="Insecta" family="Elodophthalmidae" genus="Elodophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="67" phylum="Arthropoda" rank="species" species="gracilis">
|
||
<emphasis id="B937A84EFE10FF9DC33CFBF1FA06FB9F" box="[1321,1437,1115,1141]" italics="true" pageId="2" pageNumber="67">E. gracilis</emphasis>
|
||
Kirejtshuk & Azar
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName id="4C430FDFFE10FF9DC7BFFBD5FDCBFB73" authorityName="Kirejtshuk & Azar" authorityYear="2008" box="[426,592,1151,1177]" class="Insecta" family="Elodophthalmidae" genus="Elodophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="67" phylum="Arthropoda" rank="species" species="harmonicus">
|
||
<emphasis id="B937A84EFE10FF9DC7BFFBD5FE20FB72" box="[426,443,1151,1176]" italics="true" pageId="2" pageNumber="67">E</emphasis>
|
||
.
|
||
<emphasis id="B937A84EFE10FF9DC7D9FBD5FDCBFB73" box="[460,592,1151,1177]" italics="true" pageId="2" pageNumber="67">harmonicus</emphasis>
|
||
</taxonomicName>
|
||
) primarily by the minimum dorsal distance between the compound eyes and the antennal morphology. The compound eyes of
|
||
<taxonomicName id="4C430FDFFE10FF9DC4EBFB09FC07FB57" authorityName="Cai & Zhao & Azar" authorityYear="2024" box="[766,924,1187,1213]" class="Insecta" family="Elodophthalmidae" genus="Elodophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="67" phylum="Arthropoda" rank="species" species="maksoudae">
|
||
<emphasis id="B937A84EFE10FF9DC4EBFB09FC07FB57" box="[766,924,1187,1213]" italics="true" pageId="2" pageNumber="67">E. maksoudae</emphasis>
|
||
</taxonomicName>
|
||
are more closely spaced, (Ocular Index [OI] = 17, widely spaced in
|
||
<taxonomicName id="4C430FDFFE10FF9DC7BEFB6DFDCAFB0B" authorityName="Kirejtshuk & Azar" authorityYear="2008" box="[427,593,1223,1249]" class="Insecta" family="Elodophthalmidae" genus="Elodophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="67" phylum="Arthropoda" rank="species" species="harmonicus">
|
||
<emphasis id="B937A84EFE10FF9DC7BEFB6DFDCAFB0B" box="[427,593,1223,1249]" italics="true" pageId="2" pageNumber="67">E. harmonicus</emphasis>
|
||
</taxonomicName>
|
||
(OI = 41);
|
||
<bibRefCitation id="EFD209ADFE10FF9DC4C8FB6DFB86FB0B" author="Campbell, J. M. & Marshall, J. D." box="[733,1053,1223,1249]" pageId="2" pageNumber="67" refId="ref5419" refString="Campbell, J. M. & Marshall, J. D. (1964) The ocular index and its application to the taxonomy of the Alleculidae (Coleoptera). The Coleopterists Bulletin, 18 (2), 42. https: // doi. org / 10.5962 / p. 372397" type="journal volume" year="1964">Campbell & Marshall 1964</bibRefCitation>
|
||
). More importantly, the antennae of
|
||
<taxonomicName id="4C430FDFFE10FF9DC6ACFB41FEC1FAEF" authorityName="Cai & Zhao & Azar" authorityYear="2024" box="[185,346,1259,1285]" class="Insecta" family="Elodophthalmidae" genus="Elodophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="67" phylum="Arthropoda" rank="species" species="maksoudae">
|
||
<emphasis id="B937A84EFE10FF9DC6ACFB41FF51FAEE" box="[185,202,1259,1284]" italics="true" pageId="2" pageNumber="67">E</emphasis>
|
||
.
|
||
<emphasis id="B937A84EFE10FF9DC6C9FB41FEC1FAEF" box="[220,346,1259,1285]" italics="true" pageId="2" pageNumber="67">maksoudae</emphasis>
|
||
</taxonomicName>
|
||
are much longer than those of the other species. The antennomeres 3–5 are strongly reduced and transverse (antennomeres 3–6 of
|
||
<taxonomicName id="4C430FDFFE10FF9DC454FAA5FD2CFAC3" authorityName="Kirejtshuk & Azar" authorityYear="2008" box="[577,695,1295,1321]" class="Insecta" family="Elodophthalmidae" genus="Elodophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="67" phylum="Arthropoda" rank="species" species="gracilis">
|
||
<emphasis id="B937A84EFE10FF9DC454FAA5FD2CFAC3" box="[577,695,1295,1321]" italics="true" pageId="2" pageNumber="67">E. gracilis</emphasis>
|
||
</taxonomicName>
|
||
are reduced, but not transverse, whereas antennomeres 3–5 of
|
||
<taxonomicName id="4C430FDFFE10FF9DC391FAA5FE80FAA7" authorityName="Kirejtshuk & Azar" authorityYear="2008" class="Insecta" family="Elodophthalmidae" genus="Elodophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="67" phylum="Arthropoda" rank="species" species="harmonicus">
|
||
<emphasis id="B937A84EFE10FF9DC391FAA5FE80FAA7" italics="true" pageId="2" pageNumber="67">E. harmonicus</emphasis>
|
||
</taxonomicName>
|
||
are reduced, but not transverse). The antennomeres 6–11 of
|
||
<taxonomicName id="4C430FDFFE10FF9DC5DEFA99FBF2FAA7" authorityName="Cai & Zhao & Azar" authorityYear="2024" box="[971,1129,1331,1357]" class="Insecta" family="Elodophthalmidae" genus="Elodophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="67" phylum="Arthropoda" rank="species" species="maksoudae">
|
||
<emphasis id="B937A84EFE10FF9DC5DEFA99FBF2FAA7" box="[971,1129,1331,1357]" italics="true" pageId="2" pageNumber="67">E. maksoudae</emphasis>
|
||
</taxonomicName>
|
||
are elongate (less elongate and triangular in
|
||
<taxonomicName id="4C430FDFFE10FF9DC742FAFDFE61FA9B" authorityName="Kirejtshuk & Azar" authorityYear="2008" box="[343,506,1367,1393]" class="Insecta" family="Elodophthalmidae" genus="Elodophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="67" phylum="Arthropoda" rank="species" species="harmonicus">
|
||
<emphasis id="B937A84EFE10FF9DC742FAFDFE61FA9B" box="[343,506,1367,1393]" italics="true" pageId="2" pageNumber="67">E. harmonicus</emphasis>
|
||
</taxonomicName>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BFC745CFE10FF9DC6D2FAD0FE69FA53" blockId="2.[151,1437,970,1933]" pageId="2" pageNumber="67">
|
||
<emphasis id="B937A84EFE10FF9DC6D2FAD0FE5FFA7E" bold="true" box="[199,452,1402,1429]" pageId="2" pageNumber="67">Locality and horizon.</emphasis>
|
||
Hammana/Mdeyrij, Caza Baabda, Mouhafazet Jabal Loubnan (Central
|
||
<collectingCountry id="F35434CCFE10FF9DC2CEFAD1FAD9FA7F" box="[1243,1346,1403,1429]" name="Lebanon" pageId="2" pageNumber="67">Lebanon</collectingCountry>
|
||
); Lower Cretaceous (lower Barremian).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C35927D7FE10FF9AC6D2FA68FBA8FDF3" lastPageId="5" lastPageNumber="70" pageId="2" pageNumber="67" type="description">
|
||
<paragraph id="8BFC745CFE10FF9DC6D2FA68FF6FF9EB" blockId="2.[151,1437,970,1933]" pageId="2" pageNumber="67">
|
||
<emphasis id="B937A84EFE10FF9DC6D2FA68FEC7FA36" bold="true" box="[199,348,1474,1500]" pageId="2" pageNumber="67">Description.</emphasis>
|
||
Body
|
||
<emphasis id="B937A84EFE10FF9DC7BEFA69FE56FA36" box="[427,461,1475,1500]" italics="true" pageId="2" pageNumber="67">ca.</emphasis>
|
||
<quantity id="4CBBD9B9FE10FF9DC7C0FA69FDA7FA37" box="[469,572,1475,1501]" metricMagnitude="-3" metricUnit="m" metricValue="1.14" pageId="2" pageNumber="67" unit="mm" value="1.14">1.14 mm</quantity>
|
||
long, moderately convex. Upper surfaces clothed with fine, suberect hairs (
|
||
<figureCitation id="137868D9FE10FF9DC6B5FA4DFF78F9EB" box="[160,227,1511,1537]" captionStart="FIGURE 1" captionStartId="3.[151,250,1011,1035]" captionTargetBox="[168,1417,188,984]" captionTargetId="figure-21@3.[158,1430,179,984]" captionTargetPageId="3" captionText="FIGURE 1. General habitus of Elodophthalmus maksoudae sp. nov., holotype specimen 1821, D. Azar coll., under incident light. A, Dorsal view. B, Ventral view. Scale bars: 200 μm." figureDoi="http://doi.org/10.5281/zenodo.14595651" httpUri="https://zenodo.org/record/14595651/files/figure.png" pageId="2" pageNumber="67">Fig. 1</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="8BFC745CFE10FF9DC6D2F9A1FBB5F867" blockId="2.[151,1437,970,1933]" pageId="2" pageNumber="67">
|
||
Head
|
||
<quantity id="4CBBD9B9FE10FF9DC71BF9A1FEE3F9CF" box="[270,376,1547,1573]" metricMagnitude="-4" metricUnit="m" metricValue="4.0" pageId="2" pageNumber="67" unit="mm" value="0.4">0.40 mm</quantity>
|
||
wide, strongly declined (
|
||
<figureCitation id="137868D9FE10FF9DC4BEF9A1FC9CF9CF" box="[683,775,1547,1573]" captionStart="FIGURE 2" captionStartId="3.[151,250,1971,1995]" captionTargetBox="[456,1131,1093,1945]" captionTargetId="figure-57@3.[456,1131,1093,1945]" captionTargetPageId="3" captionText="FIGURE 2. General habitus of Elodophthalmus maksoudae sp. nov., holotype specimen 1821, D. Azar coll., under confocal microscopy. Ventral view. Scale bars: 200 μm." figureDoi="http://doi.org/10.5281/zenodo.14595653" httpUri="https://zenodo.org/record/14595653/files/figure.png" pageId="2" pageNumber="67">Fig. 2A</figureCitation>
|
||
); abruptly constricted posteriorly to form narrow neck. Transverse occipital ridge developed. Eyes (
|
||
<figureCitation id="137868D9FE10FF9DC4ADF985FC83F9A3" box="[696,792,1583,1609]" captionStart="FIGURE 2" captionStartId="3.[151,250,1971,1995]" captionTargetBox="[456,1131,1093,1945]" captionTargetId="figure-57@3.[456,1131,1093,1945]" captionTargetPageId="3" captionText="FIGURE 2. General habitus of Elodophthalmus maksoudae sp. nov., holotype specimen 1821, D. Azar coll., under confocal microscopy. Ventral view. Scale bars: 200 μm." figureDoi="http://doi.org/10.5281/zenodo.14595653" httpUri="https://zenodo.org/record/14595653/files/figure.png" pageId="2" pageNumber="67">Fig. 2B</figureCitation>
|
||
) strongly protuberant and coarsely facetted, slightly emarginate, with interfacetal setae; close to each other, with OI = 17. Antennal insertions (
|
||
<figureCitation id="137868D9FE10FF9DC28FF9F9FB69F987" box="[1178,1266,1619,1645]" captionStart="FIGURE 2" captionStartId="3.[151,250,1971,1995]" captionTargetBox="[456,1131,1093,1945]" captionTargetId="figure-57@3.[456,1131,1093,1945]" captionTargetPageId="3" captionText="FIGURE 2. General habitus of Elodophthalmus maksoudae sp. nov., holotype specimen 1821, D. Azar coll., under confocal microscopy. Ventral view. Scale bars: 200 μm." figureDoi="http://doi.org/10.5281/zenodo.14595653" httpUri="https://zenodo.org/record/14595653/files/figure.png" pageId="2" pageNumber="67">Fig. 2C</figureCitation>
|
||
) exposed from above, located at inner edges of eyes. Subantennal grooves absent. Frontoclypeal suture absent. Labrum (
|
||
<figureCitation id="137868D9FE10FF9DC328F9DDFA0FF97B" box="[1341,1428,1655,1681]" captionStart="FIGURE 2" captionStartId="3.[151,250,1971,1995]" captionTargetBox="[456,1131,1093,1945]" captionTargetId="figure-57@3.[456,1131,1093,1945]" captionTargetPageId="3" captionText="FIGURE 2. General habitus of Elodophthalmus maksoudae sp. nov., holotype specimen 1821, D. Azar coll., under confocal microscopy. Ventral view. Scale bars: 200 μm." figureDoi="http://doi.org/10.5281/zenodo.14595653" httpUri="https://zenodo.org/record/14595653/files/figure.png" pageId="2" pageNumber="67">Fig. 2B</figureCitation>
|
||
) free, strongly transverse, rounded apically. Antennae (
|
||
<figureCitation id="137868D9FE10FF9DC516F931FCC0F95F" box="[771,859,1691,1717]" captionStart="FIGURE 2" captionStartId="3.[151,250,1971,1995]" captionTargetBox="[456,1131,1093,1945]" captionTargetId="figure-57@3.[456,1131,1093,1945]" captionTargetPageId="3" captionText="FIGURE 2. General habitus of Elodophthalmus maksoudae sp. nov., holotype specimen 1821, D. Azar coll., under confocal microscopy. Ventral view. Scale bars: 200 μm." figureDoi="http://doi.org/10.5281/zenodo.14595653" httpUri="https://zenodo.org/record/14595653/files/figure.png" pageId="2" pageNumber="67">Fig. 2C</figureCitation>
|
||
)
|
||
<emphasis id="B937A84EFE10FF9DC57EF931FC16F95E" box="[875,909,1691,1716]" italics="true" pageId="2" pageNumber="67">ca.</emphasis>
|
||
<quantity id="4CBBD9B9FE10FF9DC581F931FC61F95F" box="[916,1018,1691,1717]" metricMagnitude="-4" metricUnit="m" metricValue="9.2" pageId="2" pageNumber="67" unit="mm" value="0.92">0.92 mm</quantity>
|
||
long, 11-segmented, densely setose, markedly elongate; antennomere 1 subquadrate; antennomere 2 slightly longer than antennomere 1; antennomeres 3–5 strongly reduced in length, slightly narrower than antennomere 2; antennomeres 4 and 5 strongly transverse; antennomeres 6–11 slender, antennomeres 6 as long as antennomeres 1–5 combined, gradually widened apically; antennomere 7 slightly shorter than antennomere 6; antennomeres 8–10 each similar to shape and size of antennomere 7; antennomere 11 slightly longer than antennomere 10. Mandible curved. Maxillary lobes short, apically pubescent; apical maxillary palpomeres slightly expanded and truncate.
|
||
</paragraph>
|
||
<caption id="DF3C24D4FE11FF9CC682FC59FD71FBC5" ID-DOI="http://doi.org/10.5281/zenodo.14595651" ID-Zenodo-Dep="14595651" httpUri="https://zenodo.org/record/14595651/files/figure.png" pageId="3" pageNumber="68" startId="3.[151,250,1011,1035]" targetBox="[168,1417,188,984]" targetPageId="3" targetType="figure">
|
||
<paragraph id="8BFC745CFE11FF9CC682FC59FD71FBC5" blockId="3.[151,1437,1011,1071]" pageId="3" pageNumber="68">
|
||
<emphasis id="B937A84EFE11FF9CC682FC59FE8EFBE1" bold="true" box="[151,277,1011,1035]" pageId="3" pageNumber="68">FIGURE 1.</emphasis>
|
||
General habitus of
|
||
<taxonomicName id="4C430FDFFE11FF9CC7F6FC59FC98FBE1" authorityName="Cai & Zhao & Azar" authorityYear="2024" box="[483,771,1011,1035]" class="Insecta" family="Elodophthalmidae" genus="Elodophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="species" species="maksoudae" status="sp. nov.">
|
||
<emphasis id="B937A84EFE11FF9CC7F6FC59FC98FBE1" box="[483,771,1011,1035]" italics="true" pageId="3" pageNumber="68">Elodophthalmus maksoudae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B937A84EFE11FF9CC51EFC59FCC5FBE1" bold="true" box="[779,862,1011,1035]" pageId="3" pageNumber="68">
|
||
<taxonomicNameLabel id="A2041535FE11FF9CC51EFC59FCC5FBE1" box="[779,862,1011,1035]" pageId="3" pageNumber="68" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype specimen 1821, D. Azar coll., under incident light.
|
||
<emphasis id="B937A84EFE11FF9CC6C7FBBDFF7FFBC5" bold="true" box="[210,228,1047,1071]" pageId="3" pageNumber="68">A</emphasis>
|
||
, Dorsal view.
|
||
<emphasis id="B937A84EFE11FF9CC762FBBDFE13FBC5" bold="true" box="[375,392,1047,1071]" pageId="3" pageNumber="68">B</emphasis>
|
||
, Ventral view. Scale bars: 200 μm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF3C24D4FE11FF9CC682F819FDF6F81A" ID-DOI="http://doi.org/10.5281/zenodo.14595653" ID-Zenodo-Dep="14595653" httpUri="https://zenodo.org/record/14595653/files/figure.png" pageId="3" pageNumber="68" startId="3.[151,250,1971,1995]" targetBox="[456,1131,1093,1945]" targetPageId="3" targetType="figure">
|
||
<paragraph id="8BFC745CFE11FF9CC682F819FDF6F81A" blockId="3.[151,1436,1971,2032]" pageId="3" pageNumber="68">
|
||
<emphasis id="B937A84EFE11FF9CC682F819FE8FF821" bold="true" box="[151,276,1971,1995]" pageId="3" pageNumber="68">FIGURE 2.</emphasis>
|
||
General habitus of
|
||
<taxonomicName id="4C430FDFFE11FF9CC7F7F819FC9AF821" authorityName="Cai & Zhao & Azar" authorityYear="2024" box="[482,769,1971,1995]" class="Insecta" family="Elodophthalmidae" genus="Elodophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="68" phylum="Arthropoda" rank="species" species="maksoudae" status="sp. nov.">
|
||
<emphasis id="B937A84EFE11FF9CC7F7F819FC9AF821" box="[482,769,1971,1995]" italics="true" pageId="3" pageNumber="68">Elodophthalmus maksoudae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B937A84EFE11FF9CC51CF819FCC0F821" bold="true" box="[777,859,1971,1995]" pageId="3" pageNumber="68">
|
||
<taxonomicNameLabel id="A2041535FE11FF9CC51CF819FCC0F821" box="[777,859,1971,1995]" pageId="3" pageNumber="68" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype specimen 1821, D. Azar coll., under confocal microscopy. Ventral view. Scale bars: 200 μm.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF3C24D4FE16FF9BC682F8CBFDA1F80F" ID-DOI="http://doi.org/10.5281/zenodo.14595657" ID-Zenodo-Dep="14595657" httpUri="https://zenodo.org/record/14595657/files/figure.png" pageId="4" pageNumber="69" startId="4.[151,250,1889,1913]" targetBox="[230,1357,179,1864]" targetPageId="4" targetType="figure">
|
||
<paragraph id="8BFC745CFE16FF9BC682F8CBFDA1F80F" blockId="4.[151,1437,1889,2022]" pageId="4" pageNumber="69">
|
||
<emphasis id="B937A84EFE16FF9BC682F8CBFE8FF890" bold="true" box="[151,276,1889,1914]" pageId="4" pageNumber="69">FIGURE 3.</emphasis>
|
||
General habitus of
|
||
<taxonomicName id="4C430FDFFE16FF9BC7F7F8C8FC9AF890" authorityName="Cai & Zhao & Azar" authorityYear="2024" box="[482,769,1890,1914]" class="Insecta" family="Elodophthalmidae" genus="Elodophthalmus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="69" phylum="Arthropoda" rank="species" species="maksoudae" status="sp. nov.">
|
||
<emphasis id="B937A84EFE16FF9BC7F7F8C8FC9AF890" box="[482,769,1890,1914]" italics="true" pageId="4" pageNumber="69">Elodophthalmus maksoudae</emphasis>
|
||
</taxonomicName>
|
||
<emphasis id="B937A84EFE16FF9BC51CF8C8FCC0F890" bold="true" box="[777,859,1890,1914]" pageId="4" pageNumber="69">
|
||
<taxonomicNameLabel id="A2041535FE16FF9BC51CF8C8FCC0F890" box="[777,859,1890,1914]" pageId="4" pageNumber="69" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</emphasis>
|
||
, holotype specimen 1821, D. Azar coll., under confocal microscopy.
|
||
<emphasis id="B937A84EFE16FF9BC70EF82FFEB6F877" bold="true" box="[283,301,1925,1949]" pageId="4" pageNumber="69">A</emphasis>
|
||
, Ventral view of body.
|
||
<emphasis id="B937A84EFE16FF9BC43DF82CFDA2F874" bold="true" box="[552,569,1926,1950]" pageId="4" pageNumber="69">B</emphasis>
|
||
, Ventral view of head.
|
||
<emphasis id="B937A84EFE16FF9BC526F82FFCDEF877" bold="true" box="[819,837,1925,1949]" pageId="4" pageNumber="69">C</emphasis>
|
||
, Left antenna.
|
||
<emphasis id="B937A84EFE16FF9BC5F3F82CFC63F874" bold="true" box="[998,1016,1926,1950]" pageId="4" pageNumber="69">D</emphasis>
|
||
, Meso- and metatarsi. Abbreviations: a, antennomere; ey, compound eye; la, labrum; mp, maxillary palpomere; mst, mesotarsomere; mtt, metatarsimere. Scale bars: 200 μm in
|
||
<emphasis id="B937A84EFE16FF9BC71DF867FE81F80F" bold="true" box="[264,282,1997,2021]" pageId="4" pageNumber="69">A</emphasis>
|
||
and
|
||
<emphasis id="B937A84EFE16FF9BC758F867FEC4F80F" bold="true" box="[333,351,1997,2021]" pageId="4" pageNumber="69">C</emphasis>
|
||
; 100 μm in
|
||
<emphasis id="B937A84EFE16FF9BC7CBF864FE74F80C" bold="true" box="[478,495,1998,2022]" pageId="4" pageNumber="69">B</emphasis>
|
||
and
|
||
<emphasis id="B937A84EFE16FF9BC434F864FDA8F80C" bold="true" box="[545,563,1998,2022]" pageId="4" pageNumber="69">D</emphasis>
|
||
.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BFC745CFE17FF9AC6D2FF3DFC72FE3B" blockId="5.[151,1437,150,537]" pageId="5" pageNumber="70">
|
||
Pronotum transverse,
|
||
<quantity id="4CBBD9B9FE17FF9AC7AAFF3CFDBFFF5B" box="[447,548,150,177]" metricMagnitude="-4" metricUnit="m" metricValue="5.2" pageId="5" pageNumber="70" unit="mm" value="0.52">0.52 mm</quantity>
|
||
wide; sides slightly curved; base slightly narrower than elytral bases; anterior angles not produced; posterior angles broadly rounded; posterior edge simple; disc without impressions. Scutellar shield not abruptly elevated anteriorly; posteriorly rounded. Elytron
|
||
<quantity id="4CBBD9B9FE17FF9AC5A3FF75FB85FF13" box="[950,1054,223,249]" metricMagnitude="-4" metricUnit="m" metricValue="8.9" pageId="5" pageNumber="70" unit="mm" value="0.89">0.89 mm</quantity>
|
||
long along suture,
|
||
<emphasis id="B937A84EFE17FF9AC310FF75FABCFF12" box="[1285,1319,223,248]" italics="true" pageId="5" pageNumber="70">ca.</emphasis>
|
||
3.8 times as long as its width; without scutellary striole; epipleura incomplete. Metaventrite moderately convex; discrimen absent; postcoxal lines absent. Metacoxae contiguous, extending laterally to meet elytra; metacoxal plates present, small. Hind wing well developed. Trochanterofemoral joint slightly oblique with base of femur separate from coxa. Mesotibia not strongly widened; outer edge simple; outer apical angle simple. Tibial spurs developed, simple. Tarsi (
|
||
<figureCitation id="137868D9FE17FF9AC68AFE39FF6DFE47" box="[159,246,403,429]" captionStart="FIGURE 2" captionStartId="3.[151,250,1971,1995]" captionTargetBox="[456,1131,1093,1945]" captionTargetId="figure-57@3.[456,1131,1093,1945]" captionTargetPageId="3" captionText="FIGURE 2. General habitus of Elodophthalmus maksoudae sp. nov., holotype specimen 1821, D. Azar coll., under confocal microscopy. Ventral view. Scale bars: 200 μm." figureDoi="http://doi.org/10.5281/zenodo.14595653" httpUri="https://zenodo.org/record/14595653/files/figure.png" pageId="5" pageNumber="70">Fig. 2D</figureCitation>
|
||
) 5-5-4, simple; pro- and mesotarsomeres 1–4 successively shortened; pro- and mesotarsomeres 5 elongate; metatarsomeres 1 longer than other metatarsomeres. Pretarsal claws simple.
|
||
</paragraph>
|
||
<paragraph id="8BFC745CFE17FF9AC6D2FE71FBA8FDF3" blockId="5.[151,1437,150,537]" pageId="5" pageNumber="70">
|
||
Abdomen (
|
||
<figureCitation id="137868D9FE17FF9AC750FE71FE07FE1F" box="[325,412,475,501]" captionStart="FIGURE 2" captionStartId="3.[151,250,1971,1995]" captionTargetBox="[456,1131,1093,1945]" captionTargetId="figure-57@3.[456,1131,1093,1945]" captionTargetPageId="3" captionText="FIGURE 2. General habitus of Elodophthalmus maksoudae sp. nov., holotype specimen 1821, D. Azar coll., under confocal microscopy. Ventral view. Scale bars: 200 μm." figureDoi="http://doi.org/10.5281/zenodo.14595653" httpUri="https://zenodo.org/record/14595653/files/figure.png" pageId="5" pageNumber="70">Fig. 2A</figureCitation>
|
||
) with five free ventrites, first two of which not connate; ventrite 1 not much longer than 2; suture between ventrites 1 and 2 distinct; ventrite 3 slightly shorter than ventrite 2.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |