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<document id="28948DE413A1DA7DF1F39A910E67BB3D" ID-CLB-Dataset="29406" ID-DOI="10.11646/zootaxa.4462.2.1" ID-GBIF-Dataset="b52189b5-2b57-474a-86b6-40580b016ecc" ID-ISSN="1175-5326" ID-Zenodo-Dep="1441457" ID-ZooBank="FA92FB55-EA2E-4193-946E-F780A16EB3CC" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1538402499097" checkinUser="plazi" docAuthor="Janák, Jiří" docDate="2018" docId="03DD5963FFE4FFCFFF7AFE00FE47EB6F" docLanguage="en" docName="zootaxa.4462.2.1.pdf" docOrigin="Zootaxa 4462 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Acylophorus hova Janák 2018, sp. nov." docType="treatment" docVersion="7" lastPageNumber="170" masterDocId="FFE4211BFFF5FFDCFFEDFFE7FFFCED62" masterDocTitle="Revision of the genus Acylophorus Nordmann, 1837 from Madagascar and Mascarene Islands (Coleoptera: Staphylinidae: Staphylininae: Acylophorina)" masterLastPageNumber="191" masterPageNumber="151" pageNumber="168" updateTime="1698561012784" updateUser="ExternalLinkService">
<mods:mods id="3A196DB844CC86394D6DD02C7B812588" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="FAD9AA2230E5EDE4DCE55E11DADCDDAA">
<mods:title id="A35B37D14E979D38F48B2E6BF82562EE">Revision of the genus Acylophorus Nordmann, 1837 from Madagascar and Mascarene Islands (Coleoptera: Staphylinidae: Staphylininae: Acylophorina)</mods:title>
</mods:titleInfo>
<mods:name id="15F5AB828F7F639E70570EDCAF77824B" type="personal">
<mods:role id="7ED5C0FD7648B3AA8AF3CBF018243BD8">
<mods:roleTerm id="32DCAEDCCFBB0A978438EE9FB87E051C">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="0EA35D440720C79DC551323888AE1D5D">Janák, Jiří</mods:namePart>
</mods:name>
<mods:typeOfResource id="9A6A77F0E3C4275F27C38F1374007928">text</mods:typeOfResource>
<mods:relatedItem id="62C13E2EBA92ABD9A3EF04EFD93462B7" type="host">
<mods:titleInfo id="4930E5B4C3F71592F2D025E943B74CFF">
<mods:title id="0B335F68538D36EA9E01C14F38D37F76">Zootaxa</mods:title>
</mods:titleInfo>
<mods:part id="E1394F84244F56CD9059F3DE7D3D30EA">
<mods:date id="E29F14BCC9458C3378A4511FA8915FF8">2018</mods:date>
<mods:detail id="CEB8CC3397E4781A3739CFED359837EB" type="pubDate">
<mods:number id="F6605A7D0EA307FD1E94A25F7F78192A">2018-08-27</mods:number>
</mods:detail>
<mods:detail id="44B9A400CF7876E5061427A6451A6C73" type="volume">
<mods:number id="D104494FC33A97940D94B48C5D794083">4462</mods:number>
</mods:detail>
<mods:detail id="F6B96E77E0228E79777E05AB409151EE" type="issue">
<mods:number id="AC484E4E5ADF865E2E89CC90EA2D854C">2</mods:number>
</mods:detail>
<mods:extent id="B43A9D2BBF0696FC634EEF5264D00C00" unit="page">
<mods:start id="4F8108A988C508FB2D4EC058FC646429">151</mods:start>
<mods:end id="4B47745C5F235EEF422DAA64B189249A">191</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="9F7CED8FFB390F6BA5328B03078B0DE2">journal article</mods:classification>
<mods:identifier id="56F0B27B1E32A837A84D3C0AB07C29BE" type="CLB-Dataset">29406</mods:identifier>
<mods:identifier id="24CF138BB4ED2736B7369AC2EBE2DA78" type="DOI">10.11646/zootaxa.4462.2.1</mods:identifier>
<mods:identifier id="DEB167996DA0B1A32C64B1D9A5FA9C1A" type="GBIF-Dataset">b52189b5-2b57-474a-86b6-40580b016ecc</mods:identifier>
<mods:identifier id="0DA5C64190D77693DEF78953868CE175" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="EBDCBDB2E49775FB2FCB85FE27BDE758" type="Zenodo-Dep">1441457</mods:identifier>
<mods:identifier id="6FF4C391BDA5199B1B4F30259E6F9088" type="ZooBank">FA92FB55-EA2E-4193-946E-F780A16EB3CC</mods:identifier>
</mods:mods>
<treatment id="03DD5963FFE4FFCFFF7AFE00FE47EB6F" ID-DOI="http://doi.org/10.5281/zenodo.5960301" ID-GBIF-Taxon="148424846" ID-Zenodo-Dep="5960301" LSID="urn:lsid:plazi:treatment:03DD5963FFE4FFCFFF7AFE00FE47EB6F" httpUri="http://treatment.plazi.org/id/03DD5963FFE4FFCFFF7AFE00FE47EB6F" lastPageId="19" lastPageNumber="170" pageId="17" pageNumber="168">
<subSubSection id="C36EBBFEFFE4FFCDFF7AFE00FE9AEF41" pageId="17" pageNumber="168" type="nomenclature">
<paragraph id="8BCBE875FFE4FFCDFF7AFE00FE2BEF60" blockId="17.[151,471,487,547]" box="[151,471,487,514]" pageId="17" pageNumber="168">
<heading id="D0835F19FFE4FFCDFF7AFE00FE2BEF60" bold="true" box="[151,471,487,514]" fontSize="11" level="1" pageId="17" pageNumber="168" reason="1">
<emphasis id="B9003467FFE4FFCDFF7AFE00FE2BEF60" bold="true" box="[151,471,487,514]" pageId="17" pageNumber="168">
<taxonomicName id="4C7493F6FFE4FFCDFF7AFE00FE8EEF63" ID-CoL="9W7M" authority="Janák, 2018" authorityName="Janák" authorityYear="2018" box="[151,370,487,513]" class="Insecta" family="Staphylinidae" genus="Acylophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="168" phylum="Arthropoda" rank="species" species="hova" status="sp. nov.">
<emphasis id="B9003467FFE4FFCDFF7AFE00FE8EEF63" bold="true" box="[151,370,487,513]" italics="true" pageId="17" pageNumber="168">Acylophorus hova</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A233891CFFE4FFCDFE94FE0FFE2BEF60" box="[377,471,488,514]" pageId="17" pageNumber="168" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8BCBE875FFE4FFCDFF7AFDEDFE9AEF41" blockId="17.[151,471,487,547]" box="[151,358,522,547]" pageId="17" pageNumber="168">
(
<figureCitation id="134FF4F0FFE4FFCDFF72FDEDFF06EF41" box="[159,250,522,547]" captionStart="FIGURES 78–86" captionStartId="20.[151,264,1932,1954]" captionTargetBox="[156,1422,209,1857]" captionTargetId="figure@20.[151,1435,200,1892]" captionTargetPageId="20" captionText="FIGURES 78–86. Acylophorus hova sp. nov., 78, 79, 85, 86, holotype; 80–84 paratypes. 78, habitus; 79, head and pronotum; 80, mandibles; 81, maxillary palpus; 82, 85, aedeagus lateral; 83, aedeagus ventro-lateral; 84, 86, aedeagus ventral. 85, 86, transmitted light. 82–86, same scale bars." httpUri="https://zenodo.org/record/1441481/files/figure.png" pageId="17" pageNumber="168">Figs. 78</figureCitation>
–92, 176)
</paragraph>
</subSubSection>
<subSubSection id="C36EBBFEFFE4FFCDFF7AFDB5FEF4EF99" pageId="17" pageNumber="168" type="materials_examined">
<paragraph id="8BCBE875FFE4FFCDFF7AFDB5FB82EF08" blockId="17.[151,1437,594,2023]" box="[151,1150,594,619]" pageId="17" pageNumber="168">
<materialsCitation id="3B1CE228FFE4FFCDFF7AFDB5FB82EF08" ID-GBIF-Occurrence="1913892117" box="[151,1150,594,619]" country="Madagascar" latitude="-46.8" location="Ankarafantsika National Park" pageId="17" pageNumber="168" specimenCount="1" typeStatus="holotype">
<emphasis id="B9003467FFE4FFCDFF7AFDB5FEC9EF09" bold="true" box="[151,309,594,619]" pageId="17" pageNumber="168">
<typeStatus id="54CF56D7FFE4FFCDFF7AFDB5FF2EEF09" box="[151,210,594,619]" pageId="17" pageNumber="168">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F363A8E5FFE4FFCDFED1FDB5FE39EF09" box="[316,453,594,619]" name="Madagascar" pageId="17" pageNumber="168">Madagascar</collectingCountry>
,
<location id="8EABBEAEFFE4FFCDFE3DFDB5FCDCEF09" LSID="urn:lsid:plazi:treatment:03DD5963FFE4FFCFFF7AFE00FE47EB6F:8EABBEAEFFE4FFCDFE3DFDB5FCDCEF09" box="[464,800,594,619]" country="Madagascar" latitude="-46.8" name="Ankarafantsika National Park" pageId="17" pageNumber="168">Ankarafantsika National Park</location>
, 16 18' 10.7&quot;,
<geoCoordinate id="EE408EB2FFE4FFCDFC2DFDB4FBE0EF09" box="[960,1052,594,619]" direction="south" orientation="latitude" pageId="17" pageNumber="168" precision="925" value="-46.8">S 46 48'</geoCoordinate>
47.1&quot; E.
</materialsCitation>
</paragraph>
<paragraph id="8BCBE875FFE4FFCDFF2AFD91FB63EFEC" blockId="17.[151,1437,594,2023]" box="[199,1183,630,655]" pageId="17" pageNumber="168">
<emphasis id="B9003467FFE4FFCDFF2AFD91FE15EFED" bold="true" box="[199,489,630,655]" pageId="17" pageNumber="168">
<typeStatus id="54CF56D7FFE4FFCDFF2AFD91FEFEEFED" box="[199,258,630,655]" pageId="17" pageNumber="168">Type</typeStatus>
material.
<typeStatus id="54CF56D7FFE4FFCDFE91FD91FE15EFED" box="[380,489,630,655]" pageId="17" pageNumber="168" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="9D7223FCFFE4FFCDFE02FD90FDFBEFEC" box="[495,519,631,654]" pageId="17" pageNumber="168" type="male">♂</specimenCount>
:
<collectingCountry id="F363A8E5FFE4FFCDFDFEFD91FD1CEFED" box="[531,736,630,655]" name="Madagascar" pageId="17" pageNumber="168">
<emphasis id="B9003467FFE4FFCDFDFEFD91FD1CEFED" bold="true" box="[531,736,630,655]" pageId="17" pageNumber="168">MADAGASCAR</emphasis>
</collectingCountry>
: “AKF/08/2011
<collectingCountry id="F363A8E5FFE4FFCDFC70FD90FB98EFED" box="[925,1124,631,655]" name="Madagascar" pageId="17" pageNumber="168">MADAGASCAR</collectingCountry>
NW,
</paragraph>
<paragraph id="8BCBE875FFE4FFCDFF2AFD7CFEF4EF99" blockId="17.[151,1437,594,2023]" pageId="17" pageNumber="168">
ANKARAFANTSIKA N.P.;
<date id="FFCACEB5FFE4FFCDFDFCFD7DFD75EFD1" box="[529,649,666,691]" pageId="17" pageNumber="168" value="2011-04-23">23.iv.2011</date>
, S16°18’10.7‘‘E46°48’47.1‘‘;
<quantity id="4C8C4590FFE4FFCDFC0BFD7CFBE6EFD1" box="[998,1050,667,691]" metricMagnitude="1" metricUnit="m" metricValue="8.1" pageId="17" pageNumber="168" unit="m" value="81.0">81m</quantity>
, sifting forest litter, Winkler app. extr., L.S.Rahanitriniaina &amp; R.Raveloson lgt.”, „HOLOTYPUS
<taxonomicName id="4C7493F6FFE4FFCDFC9AFD59FBBBEFB5" authority="Janák, 2018" authorityName="Janák" authorityYear="2018" box="[887,1095,702,727]" class="Insecta" family="Staphylinidae" genus="Acylophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="168" phylum="Arthropoda" rank="species" species="hova" status="sp. nov.">Acylophorus hova</taxonomicName>
<taxonomicNameLabel id="A233891CFFE4FFCDFBBCFD59FB52EFB5" box="[1105,1198,702,727]" pageId="17" pageNumber="168" rank="species">
<emphasis id="B9003467FFE4FFCDFBBCFD59FB52EFB5" bold="true" box="[1105,1198,702,727]" pageId="17" pageNumber="168">sp. nov.</emphasis>
</taxonomicNameLabel>
, J. Janák det. 2017” (MMBC).
</paragraph>
</subSubSection>
<subSubSection id="C36EBBFEFFE4FFCDFF2AFCE1FBD8EA85" pageId="17" pageNumber="168" type="description">
<paragraph id="8BCBE875FFE4FFCDFF2AFCE1FD53E901" blockId="17.[151,1437,594,2023]" pageId="17" pageNumber="168">
<materialsCitation id="3B1CE228FFE4FFCDFF2AFCE1FC57EE7D" ID-GBIF-Occurrence="1913892133" box="[199,939,774,799]" collectionCode="MMBC, JJRC" pageId="17" pageNumber="168" specimenCount="6" specimenCount-male="6" typeStatus="paratype">
<typeStatus id="54CF56D7FFE4FFCDFF2AFCE1FEC3EE7D" box="[199,319,774,799]" pageId="17" pageNumber="168" type="paratype">
<emphasis id="B9003467FFE4FFCDFF2AFCE1FEC3EE7D" bold="true" box="[199,319,774,799]" pageId="17" pageNumber="168">Paratypes</emphasis>
</typeStatus>
:
<specimenCount id="9D7223FCFFE4FFCDFEA0FCE0FE86EE7C" box="[333,378,775,799]" pageId="17" pageNumber="168" type="male">3 ♂</specimenCount>
: same data as holotype (
<specimenCount id="9D7223FCFFE4FFCDFD61FCE0FD46EE7C" box="[652,698,775,799]" pageId="17" pageNumber="168" type="male">2 ♂</specimenCount>
:
<collectionCode id="ED6570B0FFE4FFCDFD2BFCE0FCDCEE7D" box="[710,800,775,799]" country="Czech Republic" httpUri="http://grbio.org/cool/w50r-ge1v" name="Moravske Muzeum [Moravian Museum]" pageId="17" pageNumber="168">MMBC</collectionCode>
,
<specimenCount id="9D7223FCFFE4FFCDFCC7FCE0FCABEE7C" box="[810,855,775,799]" pageId="17" pageNumber="168" type="male">1 ♂</specimenCount>
:
<collectionCode id="ED6570B0FFE4FFCDFC8EFCE0FC5FEE7D" box="[867,931,775,799]" pageId="17" pageNumber="168">JJRC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B1CE228FFE4FFCDFC5AFCE0FB63EE05" ID-GBIF-Occurrence="1913892132" collectingDate="2011-04-23" collectionCode="MMBC, JJRC" collectorName="Winkler &amp; L. S. Rahanitriniaina &amp; R. Raveloson" country="Madagascar" elevation="89" latitude="-16.301416" location="MADAGASCAR" longLatPrecision="1" longitude="46.81897" pageId="17" pageNumber="168" specimenCount="14" specimenCount-female="4" specimenCount-male="10" typeStatus="paratype">
<specimenCount id="9D7223FCFFE4FFCDFC5AFCE0FC1EEE7C" box="[951,994,775,799]" pageId="17" pageNumber="168" type="male">4 ♂</specimenCount>
,
<specimenCount id="9D7223FCFFE4FFCDFC03FCE0FBE6EE7C" box="[1006,1050,775,799]" pageId="17" pageNumber="168" type="female">3 ♀</specimenCount>
: “AKF/05/2011
<collectingCountry id="F363A8E5FFE4FFCDFB38FCE0FA60EE7D" box="[1237,1436,775,799]" name="Madagascar" pageId="17" pageNumber="168">MADAGASCAR</collectingCountry>
NW, ANKARAFANTSIKA N.P.;
<collectingDate id="EF8E375DFFE4FFCDFDF7FCCDFD6EEE21" box="[538,658,810,835]" pageId="17" pageNumber="168" value="2011-04-23">
<date id="FFCACEB5FFE4FFCDFDF7FCCDFD6EEE21" box="[538,658,810,835]" pageId="17" pageNumber="168" value="2011-04-23">23.iv.2011</date>
</collectingDate>
,
<geoCoordinate id="EE408EB2FFE4FFCDFD72FCCDFCC4EE21" box="[671,824,810,835]" direction="south" orientation="latitude" pageId="17" pageNumber="168" precision="1" value="-16.301416">S16°18’05.1‘‘</geoCoordinate>
<geoCoordinate id="EE408EB2FFE4FFCDFCD5FCCDFC22EE21" box="[824,990,810,835]" direction="east" orientation="longitude" pageId="17" pageNumber="168" precision="1" value="46.81897">E46°49’08.3‘‘</geoCoordinate>
;
<elevation id="00590F46FFE4FFCDFC01FCCCFBDDEE21" box="[1004,1057,811,835]" metricMagnitude="1" metricUnit="m" metricValue="8.9" pageId="17" pageNumber="168" unit="m" value="89.0">
<quantity id="4C8C4590FFE4FFCDFC01FCCCFBDDEE21" box="[1004,1057,811,835]" metricMagnitude="1" metricUnit="m" metricValue="8.9" pageId="17" pageNumber="168" unit="m" value="89.0">89m</quantity>
</elevation>
, sifting forest litter,
<collectorName id="26818DA3FFE4FFCDFAE4FCCDFA98EE21" box="[1289,1380,810,835]" pageId="17" pageNumber="168">Winkler</collectorName>
app. extr.,
<collectorName id="26818DA3FFE4FFCDFF31FCA9FE4BEE05" box="[220,439,846,871]" pageId="17" pageNumber="168">L.S.Rahanitriniaina</collectorName>
&amp;
<collectorName id="26818DA3FFE4FFCDFE0AFCA9FD8BEE05" box="[487,631,846,871]" pageId="17" pageNumber="168">R.Raveloson</collectorName>
lgt. ” (
<specimenCount id="9D7223FCFFE4FFCDFD35FCA8FCF7EE04" box="[728,779,847,871]" pageId="17" pageNumber="168" type="male">2 ♂</specimenCount>
,
<specimenCount id="9D7223FCFFE4FFCDFCF0FCA8FCACEE04" box="[797,848,847,871]" pageId="17" pageNumber="168" type="male">2 ♂</specimenCount>
:
<collectionCode id="ED6570B0FFE4FFCDFC8FFCA8FC40EE05" box="[866,956,847,871]" country="Czech Republic" httpUri="http://grbio.org/cool/w50r-ge1v" name="Moravske Muzeum [Moravian Museum]" pageId="17" pageNumber="168">MMBC</collectionCode>
,
<specimenCount id="9D7223FCFFE4FFCDFC23FCA8FBFCEE04" box="[974,1024,847,871]" pageId="17" pageNumber="168" type="male">2 ♂</specimenCount>
,
<specimenCount id="9D7223FCFFE4FFCDFBFFFCA8FBB8EE04" box="[1042,1092,847,871]" pageId="17" pageNumber="168" type="female">1 ♀</specimenCount>
:
<collectionCode id="ED6570B0FFE4FFCDFBBAFCA8FB6BEE05" box="[1111,1175,847,871]" pageId="17" pageNumber="168">JJRC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B1CE228FFE4FFCDFB5DFCA8FC19EECD" ID-GBIF-Occurrence="1913892128" collectingDate="2011-04-23" collectionCode="MMBC" collectorName="Winkler &amp; L. S. Rahanitriniaina &amp; R. Raveloson" country="Madagascar" elevation="79" latitude="-16.302166" location="MADAGASCAR" longLatPrecision="1" longitude="46.81653" pageId="17" pageNumber="168" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D7223FCFFE4FFCDFB5DFCA8FB1EEE04" box="[1200,1250,847,871]" pageId="17" pageNumber="168" type="female">1 ♀</specimenCount>
: “AKF/07/2011
<collectingCountry id="F363A8E5FFE4FFCDFF7AFC94FEA2EEE9" box="[151,350,883,907]" name="Madagascar" pageId="17" pageNumber="168">MADAGASCAR</collectingCountry>
NW, ANKARAFANTSIKA N.P.;
<collectingDate id="EF8E375DFFE4FFCDFD1FFC95FC95EEE9" box="[754,873,882,907]" pageId="17" pageNumber="168" value="2011-04-23">
<date id="FFCACEB5FFE4FFCDFD1FFC95FC95EEE9" box="[754,873,882,907]" pageId="17" pageNumber="168" value="2011-04-23">23.iv.2011</date>
</collectingDate>
,
<geoCoordinate id="EE408EB2FFE4FFCDFC95FC95FBEEEEE9" box="[888,1042,882,907]" direction="south" orientation="latitude" pageId="17" pageNumber="168" precision="1" value="-16.302166">S16°18’07.8‘‘</geoCoordinate>
<geoCoordinate id="EE408EB2FFE4FFCDFBFEFC95FB44EEE9" box="[1043,1208,882,907]" direction="east" orientation="longitude" pageId="17" pageNumber="168" precision="1" value="46.81653">E46°48’59.5‘‘</geoCoordinate>
;
<elevation id="00590F46FFE4FFCDFB2AFC94FB00EEE9" box="[1223,1276,883,907]" metricMagnitude="1" metricUnit="m" metricValue="7.9" pageId="17" pageNumber="168" unit="m" value="79.0">
<quantity id="4C8C4590FFE4FFCDFB2AFC94FB00EEE9" box="[1223,1276,883,907]" metricMagnitude="1" metricUnit="m" metricValue="7.9" pageId="17" pageNumber="168" unit="m" value="79.0">79m</quantity>
</elevation>
, sifting forest litter,
<collectorName id="26818DA3FFE4FFCDFF35FC71FECFEECD" box="[216,307,918,943]" pageId="17" pageNumber="168">Winkler</collectorName>
app. extr.,
<collectorName id="26818DA3FFE4FFCDFE40FC71FD74EECD" box="[429,648,918,943]" pageId="17" pageNumber="168">L.S.Rahanitriniaina</collectorName>
&amp;
<collectorName id="26818DA3FFE4FFCDFD46FC71FCC7EECD" box="[683,827,918,943]" pageId="17" pageNumber="168">R.Raveloson</collectorName>
lgt.” (
<collectionCode id="ED6570B0FFE4FFCDFC6DFC70FC21EECD" box="[896,989,919,943]" country="Czech Republic" httpUri="http://grbio.org/cool/w50r-ge1v" name="Moravske Muzeum [Moravian Museum]" pageId="17" pageNumber="168">MMBC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B1CE228FFE4FFCDFC1DFC70FC9CEE95" ID-GBIF-Occurrence="1913892141" collectingDate="2011-04-23" collectionCode="MMBC" collectorName="Winkler &amp; L. S. Rahanitriniaina &amp; R. Raveloson" country="Madagascar" elevation="87" latitude="-16.30275" location="MADAGASCAR" longLatPrecision="1" longitude="46.810917" pageId="17" pageNumber="168" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D7223FCFFE4FFCDFC1DFC70FBE0EECC" box="[1008,1052,919,943]" pageId="17" pageNumber="168" type="female">1 ♀</specimenCount>
: “AKF/09/2011
<collectingCountry id="F363A8E5FFE4FFCDFB38FC70FA60EECD" box="[1237,1436,919,943]" name="Madagascar" pageId="17" pageNumber="168">MADAGASCAR</collectingCountry>
NW, ANKARAFANTSIKA N.P.;
<collectingDate id="EF8E375DFFE4FFCDFDF7FC5DFD6EEEB1" box="[538,658,954,979]" pageId="17" pageNumber="168" value="2011-04-23">
<date id="FFCACEB5FFE4FFCDFDF7FC5DFD6EEEB1" box="[538,658,954,979]" pageId="17" pageNumber="168" value="2011-04-23">23.iv.2011</date>
</collectingDate>
,
<geoCoordinate id="EE408EB2FFE4FFCDFD72FC5DFCC6EEB1" box="[671,826,954,979]" direction="south" orientation="latitude" pageId="17" pageNumber="168" precision="1" value="-16.30275">S16°18’09.9‘‘</geoCoordinate>
<geoCoordinate id="EE408EB2FFE4FFCDFCD7FC5DFC22EEB1" box="[826,990,954,979]" direction="east" orientation="longitude" pageId="17" pageNumber="168" precision="1" value="46.810917">E46°48’39.3‘‘</geoCoordinate>
;
<elevation id="00590F46FFE4FFCDFC01FC5CFBDCEEB1" box="[1004,1056,955,979]" metricMagnitude="1" metricUnit="m" metricValue="8.7" pageId="17" pageNumber="168" unit="m" value="87.0">
<quantity id="4C8C4590FFE4FFCDFC01FC5CFBDCEEB1" box="[1004,1056,955,979]" metricMagnitude="1" metricUnit="m" metricValue="8.7" pageId="17" pageNumber="168" unit="m" value="87.0">87m</quantity>
</elevation>
, sifting forest litter,
<collectorName id="26818DA3FFE4FFCDFAE4FC5DFA98EEB1" box="[1289,1380,954,979]" pageId="17" pageNumber="168">Winkler</collectorName>
app. extr.,
<collectorName id="26818DA3FFE4FFCDFF08FC39FE3DEE95" box="[229,449,990,1015]" pageId="17" pageNumber="168">L.S.Rahanitriniaina</collectorName>
&amp;
<collectorName id="26818DA3FFE4FFCDFDE9FC39FD68EE95" box="[516,660,990,1015]" pageId="17" pageNumber="168">R.Raveloson</collectorName>
lgt.” (
<collectionCode id="ED6570B0FFE4FFCDFD17FC38FCABEE95" box="[762,855,991,1015]" country="Czech Republic" httpUri="http://grbio.org/cool/w50r-ge1v" name="Moravske Muzeum [Moravian Museum]" pageId="17" pageNumber="168">MMBC</collectionCode>
)
</materialsCitation>
;
<materialsCitation id="3B1CE228FFE4FFCDFC96FC38FD57E901" ID-GBIF-Occurrence="1913892129" collectingDate="2011-04-24" collectionCode="MMBC" collectorName="Winkler &amp; L. S. Rahanitriniaina &amp; Raveloson &amp; All" country="Madagascar" determinerName="J. Janak" elevation="83" latitude="-16.3025" location="MADAGASCAR" longLatPrecision="1" longitude="46.815445" pageId="17" pageNumber="168" specimenCount="1" specimenCount-female="1" typeStatus="paratype">
<specimenCount id="9D7223FCFFE4FFCDFC96FC38FC4AEE94" box="[891,950,991,1015]" pageId="17" pageNumber="168" type="female">1 ♀</specimenCount>
: “AKF/11/2011
<collectingCountry id="F363A8E5FFE4FFCDFB62FC38FAAAEE95" box="[1167,1366,991,1015]" name="Madagascar" pageId="17" pageNumber="168">MADAGASCAR</collectingCountry>
NW ANKARAFANTSIKA N.P.;
<collectingDate id="EF8E375DFFE4FFCDFE33FBE5FDA9E979" box="[478,597,1026,1051]" pageId="17" pageNumber="168" value="2011-04-24">
<date id="FFCACEB5FFE4FFCDFE33FBE5FDA9E979" box="[478,597,1026,1051]" pageId="17" pageNumber="168" value="2011-04-24">24.iv.2011</date>
</collectingDate>
,
<geoCoordinate id="EE408EB2FFE4FFCDFD8FFBE5FD00E979" box="[610,764,1026,1051]" direction="south" orientation="latitude" pageId="17" pageNumber="168" precision="1" value="-16.3025">S16°18’09.0‘‘</geoCoordinate>
<geoCoordinate id="EE408EB2FFE4FFCDFD11FBE5FC5DE979" box="[764,929,1026,1051]" direction="east" orientation="longitude" pageId="17" pageNumber="168" precision="1" value="46.815445">E46°48’55.6‘‘</geoCoordinate>
;
<elevation id="00590F46FFE4FFCDFC43FBE4FC1EE979" box="[942,994,1027,1051]" metricMagnitude="1" metricUnit="m" metricValue="8.3" pageId="17" pageNumber="168" unit="m" value="83.0">
<quantity id="4C8C4590FFE4FFCDFC43FBE4FC1EE979" box="[942,994,1027,1051]" metricMagnitude="1" metricUnit="m" metricValue="8.3" pageId="17" pageNumber="168" unit="m" value="83.0">83m</quantity>
</elevation>
, sifting forest litter,
<collectorName id="26818DA3FFE4FFCDFB27FBE5FAD9E979" box="[1226,1317,1026,1051]" pageId="17" pageNumber="168">Winkler</collectorName>
app. extr.,
<collectorName id="26818DA3FFE4FFCDFF7AFBC1FE8EE95D" box="[151,370,1062,1087]" pageId="17" pageNumber="168">L.S.Rahanitriniaina</collectorName>
&amp; R.
<collectorName id="26818DA3FFE4FFCDFE2BFBC1FDBCE95D" box="[454,576,1062,1087]" pageId="17" pageNumber="168">Raveloson</collectorName>
lgt.” (
<collectionCode id="ED6570B0FFE4FFCDFD4CFBC0FD02E95D" box="[673,766,1063,1087]" country="Czech Republic" httpUri="http://grbio.org/cool/w50r-ge1v" name="Moravske Muzeum [Moravian Museum]" pageId="17" pageNumber="168">MMBC</collectionCode>
).
<collectorName id="26818DA3FFE4FFCDFCF3FBC1FCBEE95D" box="[798,834,1062,1087]" pageId="17" pageNumber="168">All</collectorName>
<typeStatus id="54CF56D7FFE4FFCDFCBBFBC0FC3DE95D" box="[854,961,1063,1087]" pageId="17" pageNumber="168" type="paratype">paratypes</typeStatus>
with additional label: „
<typeStatus id="54CF56D7FFE4FFCDFB11FBC0FA60E95D" box="[1276,1436,1063,1087]" pageId="17" pageNumber="168" type="paratype">PARATYPUS</typeStatus>
<taxonomicName id="4C7493F6FFE4FFCDFF7AFBADFE98E901" authority="Janák, 2018" authorityName="Janák" authorityYear="2018" box="[151,356,1098,1123]" class="Insecta" family="Staphylinidae" genus="Acylophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="168" phylum="Arthropoda" rank="species" species="hova" status="sp. nov.">Acylophorus hova</taxonomicName>
<taxonomicNameLabel id="A233891CFFE4FFCDFE81FBADFE39E901" box="[364,453,1098,1123]" pageId="17" pageNumber="168" rank="species">
<emphasis id="B9003467FFE4FFCDFE81FBADFE39E901" bold="true" box="[364,453,1098,1123]" pageId="17" pageNumber="168">sp. nov.</emphasis>
</taxonomicNameLabel>
,
<determinerName id="7E1704B1FFE4FFCDFE3EFBACFDD0E901" box="[467,556,1098,1123]" pageId="17" pageNumber="168">J. Janák</determinerName>
det. 2017”
</materialsCitation>
.
</paragraph>
<paragraph id="8BCBE875FFE4FFCDFF2AFB89FA7DE991" blockId="17.[151,1437,594,2023]" pageId="17" pageNumber="168">
<emphasis id="B9003467FFE4FFCDFF2AFB89FEA5E9E5" bold="true" box="[199,345,1134,1159]" pageId="17" pageNumber="168">Description.</emphasis>
Body length
<quantity id="4C8C4590FFE4FFCDFE1EFB89FD80E9E4" box="[499,636,1134,1159]" metricMagnitude="-3" metricUnit="m" metricValue="7.1" metricValueMax="7.7" metricValueMin="6.5" pageId="17" pageNumber="168" unit="mm" value="7.1" valueMax="7.7" valueMin="6.5">6.5–7.7 mm</quantity>
(M=7.0 mm, HT=
<quantity id="4C8C4590FFE4FFCDFCBCFB88FC50E9E5" box="[849,940,1135,1159]" metricMagnitude="-3" metricUnit="m" metricValue="7.3" pageId="17" pageNumber="168" unit="mm" value="7.3">7.3 mm</quantity>
), forebody length
<quantity id="4C8C4590FFE4FFCDFB91FB88FAF8E9E4" box="[1148,1284,1135,1159]" metricMagnitude="-3" metricUnit="m" metricValue="2.5999999999999996" metricValueMax="2.8" metricValueMin="2.4" pageId="17" pageNumber="168" unit="mm" value="2.5999999999999996" valueMax="2.8" valueMin="2.4">2.4–2.8 mm</quantity>
(M=
<quantity id="4C8C4590FFE4FFCDFAD0FB89FA64E9E4" box="[1341,1432,1134,1159]" metricMagnitude="-3" metricUnit="m" metricValue="2.6" pageId="17" pageNumber="168" unit="mm" value="2.6">2.6 mm</quantity>
, HT=
<quantity id="4C8C4590FFE4FFCDFF21FB74FEDBE9C9" box="[204,295,1171,1195]" metricMagnitude="-3" metricUnit="m" metricValue="2.7" pageId="17" pageNumber="168" unit="mm" value="2.7">2.7 mm</quantity>
). Body dark brown (
<figureCitation id="134FF4F0FFE4FFCDFDFFFB75FD9BE9C9" box="[530,615,1170,1195]" captionStart="FIGURES 78–86" captionStartId="20.[151,264,1932,1954]" captionTargetBox="[156,1422,209,1857]" captionTargetId="figure@20.[151,1435,200,1892]" captionTargetPageId="20" captionText="FIGURES 78–86. Acylophorus hova sp. nov., 78, 79, 85, 86, holotype; 80–84 paratypes. 78, habitus; 79, head and pronotum; 80, mandibles; 81, maxillary palpus; 82, 85, aedeagus lateral; 83, aedeagus ventro-lateral; 84, 86, aedeagus ventral. 85, 86, transmitted light. 82–86, same scale bars." httpUri="https://zenodo.org/record/1441481/files/figure.png" pageId="17" pageNumber="168">Fig. 78</figureCitation>
), pronotum reddish-brown with reddish margins, apical margin of elytra reddish-brown, abdominal tergites brown, markedly bluish iridescent; antennae brown with first segment almost completely and sometimes last 1–2 segments partly reddish; legs reddish, tarsi and maxillary palpi light reddish.
</paragraph>
<paragraph id="8BCBE875FFE4FFCDFF2AFB19FEF2EB55" blockId="17.[151,1437,594,2023]" pageId="17" pageNumber="168">
Head suboval (
<figureCitation id="134FF4F0FFE4FFCDFE9BFB19FE30E875" box="[374,460,1278,1303]" captionStart="FIGURES 78–86" captionStartId="20.[151,264,1932,1954]" captionTargetBox="[156,1422,209,1857]" captionTargetId="figure@20.[151,1435,200,1892]" captionTargetPageId="20" captionText="FIGURES 78–86. Acylophorus hova sp. nov., 78, 79, 85, 86, holotype; 80–84 paratypes. 78, habitus; 79, head and pronotum; 80, mandibles; 81, maxillary palpus; 82, 85, aedeagus lateral; 83, aedeagus ventro-lateral; 84, 86, aedeagus ventral. 85, 86, transmitted light. 82–86, same scale bars." httpUri="https://zenodo.org/record/1441481/files/figure.png" pageId="17" pageNumber="168">Fig. 79</figureCitation>
), about as long as wide (L/W=0.94–1.05, M=0.98, HT=0.99), widened behind eyes, with two pairs of interocular setae. Temples markedly shorter than eyes (LT/LE=0.60–0.74, M=0.68, HT=0.60). Dorsal side of head shining, without micropunctures. Only sparse pale grey pubescence behind eyes. Four postocular setae visible on each side, additional seta on hind margin of eye absent. Mandibles with large teeth (
<figureCitation id="134FF4F0FFE4FFCDFA82FA8DFF4BE8C5" captionStart="FIGURES 78–86" captionStartId="20.[151,264,1932,1954]" captionTargetBox="[156,1422,209,1857]" captionTargetId="figure@20.[151,1435,200,1892]" captionTargetPageId="20" captionText="FIGURES 78–86. Acylophorus hova sp. nov., 78, 79, 85, 86, holotype; 80–84 paratypes. 78, habitus; 79, head and pronotum; 80, mandibles; 81, maxillary palpus; 82, 85, aedeagus lateral; 83, aedeagus ventro-lateral; 84, 86, aedeagus ventral. 85, 86, transmitted light. 82–86, same scale bars." httpUri="https://zenodo.org/record/1441481/files/figure.png" pageId="17" pageNumber="168">Fig. 80</figureCitation>
). Maxillary palpi with terminal segment wide and quite short, pubescent, asymmetric, as wide as or only slightly wider and longer than glabrous penultimate segment which is triangular (
<figureCitation id="134FF4F0FFE4FFCDFC25FA55FBE6E8A9" box="[968,1050,1458,1483]" captionStart="FIGURES 78–86" captionStartId="20.[151,264,1932,1954]" captionTargetBox="[156,1422,209,1857]" captionTargetId="figure@20.[151,1435,200,1892]" captionTargetPageId="20" captionText="FIGURES 78–86. Acylophorus hova sp. nov., 78, 79, 85, 86, holotype; 80–84 paratypes. 78, habitus; 79, head and pronotum; 80, mandibles; 81, maxillary palpus; 82, 85, aedeagus lateral; 83, aedeagus ventro-lateral; 84, 86, aedeagus ventral. 85, 86, transmitted light. 82–86, same scale bars." httpUri="https://zenodo.org/record/1441481/files/figure.png" pageId="17" pageNumber="168">Fig. 81</figureCitation>
). First segment of antenna slightly longer than next five; segments 1 to 4 elongate, segment 5 longer than wide (L/W=1.04–1.37, M=1.20, HT=1.38), segments 6 as long as wide, segments 7–9 transverse, segment 10 markedly transverse (R=0.73–0.83, M=0.77, HT=0.83).
</paragraph>
<paragraph id="8BCBE875FFE4FFCDFF2AF9A4FC5EEA51" blockId="17.[151,1437,594,2023]" pageId="17" pageNumber="168">
Pronotum (
<figureCitation id="134FF4F0FFE4FFCDFEAAF9A5FE66EB39" box="[327,410,1602,1627]" captionStart="FIGURES 78–86" captionStartId="20.[151,264,1932,1954]" captionTargetBox="[156,1422,209,1857]" captionTargetId="figure@20.[151,1435,200,1892]" captionTargetPageId="20" captionText="FIGURES 78–86. Acylophorus hova sp. nov., 78, 79, 85, 86, holotype; 80–84 paratypes. 78, habitus; 79, head and pronotum; 80, mandibles; 81, maxillary palpus; 82, 85, aedeagus lateral; 83, aedeagus ventro-lateral; 84, 86, aedeagus ventral. 85, 86, transmitted light. 82–86, same scale bars." httpUri="https://zenodo.org/record/1441481/files/figure.png" pageId="17" pageNumber="168">Fig. 79</figureCitation>
) slightly transverse (W/L=1.12–1.22, M=1.17, HT=1.17), markedly widened behind with sides slightly rounded, widest in basal half, shining, without micropunctures. One pair of dorsal setae and one pair of lateral setae. Marginal setae short. Elytra markedly transverse (R=1.42–1.74, M=1.53, HT=1.74) with short, pale, but not shining pubescence arising from dense aspirate punctures. Apical fringe of bristles slightly longer than pubescence on other parts of the elytra. Abdominal tergites with short, moderately dense pubescence, interstices in transverse direction on middle part of tergite IV about two times as large as diameter of punctures. Meso- and metatarsi with short empodial setae, shorter than claws (
<figureCitation id="134FF4F0FFE4FFCDFCE0F8FDFC69EA51" box="[781,917,1818,1843]" captionStart="FIGURES 87–90" captionStartId="21.[151,264,1952,1974]" captionTargetBox="[156,1424,258,1831]" captionTargetId="figure@21.[151,1435,250,1855]" captionTargetPageId="21" captionText="FIGURES 87–90. Acylophorus hova sp. nov., paratypes. 87, metatarsus lateral; 88, metatarsus dorsal; 89, male tergites 9, 10; 90, male sternite 9; 91, female tergites 9, 10; 92, valves. 87, 88, same scale bars." httpUri="https://zenodo.org/record/1441483/files/figure.png" pageId="17" pageNumber="168">Figs. 87, 88</figureCitation>
).
</paragraph>
<paragraph id="8BCBE875FFE4FFCDFF2AF8D9FCB3EAA1" blockId="17.[151,1437,594,2023]" pageId="17" pageNumber="168">
Male (n=5). Sternite 9 long and narrow (
<figureCitation id="134FF4F0FFE4FFCDFD7EF8D9FD0EEA35" box="[659,754,1854,1879]" captionStart="FIGURES 87–90" captionStartId="21.[151,264,1952,1974]" captionTargetBox="[156,1424,258,1831]" captionTargetId="figure@21.[151,1435,250,1855]" captionTargetPageId="21" captionText="FIGURES 87–90. Acylophorus hova sp. nov., paratypes. 87, metatarsus lateral; 88, metatarsus dorsal; 89, male tergites 9, 10; 90, male sternite 9; 91, female tergites 9, 10; 92, valves. 87, 88, same scale bars." httpUri="https://zenodo.org/record/1441483/files/figure.png" pageId="17" pageNumber="168">Figs. 90</figureCitation>
), apex of tergite 10 narrowly rounded, with very long apical setae (
<figureCitation id="134FF4F0FFE4FFCDFF0FF885FEC5EA19" box="[226,313,1890,1915]" captionStart="FIGURES 87–90" captionStartId="21.[151,264,1952,1974]" captionTargetBox="[156,1424,258,1831]" captionTargetId="figure@21.[151,1435,250,1855]" captionTargetPageId="21" captionText="FIGURES 87–90. Acylophorus hova sp. nov., paratypes. 87, metatarsus lateral; 88, metatarsus dorsal; 89, male tergites 9, 10; 90, male sternite 9; 91, female tergites 9, 10; 92, valves. 87, 88, same scale bars." httpUri="https://zenodo.org/record/1441483/files/figure.png" pageId="17" pageNumber="168">Fig. 89</figureCitation>
). Aedeagus (
<figureCitation id="134FF4F0FFE4FFCDFE3EF885FDA2EA19" box="[467,606,1890,1915]" captionStart="FIGURES 78–86" captionStartId="20.[151,264,1932,1954]" captionTargetBox="[156,1422,209,1857]" captionTargetId="figure@20.[151,1435,200,1892]" captionTargetPageId="20" captionText="FIGURES 78–86. Acylophorus hova sp. nov., 78, 79, 85, 86, holotype; 80–84 paratypes. 78, habitus; 79, head and pronotum; 80, mandibles; 81, maxillary palpus; 82, 85, aedeagus lateral; 83, aedeagus ventro-lateral; 84, 86, aedeagus ventral. 85, 86, transmitted light. 82–86, same scale bars." httpUri="https://zenodo.org/record/1441481/files/figure.png" pageId="17" pageNumber="168">Figs. 82–86</figureCitation>
)
<quantity id="4C8C4590FFE4FFCDFD9DF885FCEBEA18" box="[624,791,1890,1915]" metricMagnitude="-4" metricUnit="m" metricValue="8.049999999999999" metricValueMax="8.5" metricValueMin="7.6" pageId="17" pageNumber="168" unit="mm" value="0.8049999999999999" valueMax="0.85" valueMin="0.76">0.76–0.85 mm</quantity>
long (M=
<quantity id="4C8C4590FFE4FFCDFC62F884FC06EA18" box="[911,1018,1891,1915]" metricMagnitude="-4" metricUnit="m" metricValue="7.9" pageId="17" pageNumber="168" unit="mm" value="0.79">0.79 mm</quantity>
, HT=
<quantity id="4C8C4590FFE4FFCDFBD2F884FB57EA19" box="[1087,1195,1891,1915]" metricMagnitude="-4" metricUnit="m" metricValue="7.8" pageId="17" pageNumber="168" unit="mm" value="0.78">0.78 mm</quantity>
). Parameres bilobed, lobes markedly separated, sensory pegs confusedly arranged in posterior quarter (
<figureCitation id="134FF4F0FFE4FFCDFBAFF861FB64EAFD" box="[1090,1176,1926,1951]" captionStart="FIGURES 78–86" captionStartId="20.[151,264,1932,1954]" captionTargetBox="[156,1422,209,1857]" captionTargetId="figure@20.[151,1435,200,1892]" captionTargetPageId="20" captionText="FIGURES 78–86. Acylophorus hova sp. nov., 78, 79, 85, 86, holotype; 80–84 paratypes. 78, habitus; 79, head and pronotum; 80, mandibles; 81, maxillary palpus; 82, 85, aedeagus lateral; 83, aedeagus ventro-lateral; 84, 86, aedeagus ventral. 85, 86, transmitted light. 82–86, same scale bars." httpUri="https://zenodo.org/record/1441481/files/figure.png" pageId="17" pageNumber="168">Fig. 86</figureCitation>
). Median lobe slightly overreaching paramere, apex narrowly rounded (
<figureCitation id="134FF4F0FFE4FFCDFD54F84DFCBEEAA1" box="[697,834,1962,1987]" captionStart="FIGURES 78–86" captionStartId="20.[151,264,1932,1954]" captionTargetBox="[156,1422,209,1857]" captionTargetId="figure@20.[151,1435,200,1892]" captionTargetPageId="20" captionText="FIGURES 78–86. Acylophorus hova sp. nov., 78, 79, 85, 86, holotype; 80–84 paratypes. 78, habitus; 79, head and pronotum; 80, mandibles; 81, maxillary palpus; 82, 85, aedeagus lateral; 83, aedeagus ventro-lateral; 84, 86, aedeagus ventral. 85, 86, transmitted light. 82–86, same scale bars." httpUri="https://zenodo.org/record/1441481/files/figure.png" pageId="17" pageNumber="168">Figs. 84, 86</figureCitation>
).
</paragraph>
<paragraph id="8BCBE875FFE4FFCDFF2AF829FBD8EA85" blockId="17.[151,1437,594,2023]" box="[199,1060,1998,2023]" pageId="17" pageNumber="168">Female. Posterior margin of tergite 10 pointed (Fig. 91), valves as in Fig. 92.</paragraph>
</subSubSection>
<caption id="DF0BB8FDFFE7FFCEFF7AF894FAF2EAC5" httpUri="https://zenodo.org/record/1441477/files/figure.png" pageId="18" pageNumber="169" startId="18.[151,264,1907,1929]" targetBox="[163,1428,201,1881]" targetPageId="18">
<paragraph id="8BCBE875FFE7FFCEFF7AF894FAF2EAC5" blockId="18.[151,1436,1907,1959]" pageId="18" pageNumber="169">
<emphasis id="B9003467FFE7FFCEFF7AF894FEA5EAEB" bold="true" box="[151,345,1907,1929]" pageId="18" pageNumber="169">FIGURES 67–73.</emphasis>
<taxonomicName id="4C7493F6FFE7FFCEFE89F894FD94EAEB" authority="Janák, 2018" authorityName="Janák" authorityYear="2018" box="[356,616,1907,1929]" class="Insecta" family="Staphylinidae" genus="Acylophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="18" pageNumber="169" phylum="Arthropoda" rank="species" species="madecassus" status="sp. nov.">
<emphasis id="B9003467FFE7FFCEFE89F894FD94EAEB" box="[356,616,1907,1929]" italics="true" pageId="18" pageNumber="169">Acylophorus madecassus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A233891CFFE7FFCEFD9EF894FD35EAEB" box="[627,713,1907,1929]" pageId="18" pageNumber="169" rank="species">
<emphasis id="B9003467FFE7FFCEFD9EF894FD35EAEB" bold="true" box="[627,713,1907,1929]" pageId="18" pageNumber="169">sp. nov.</emphasis>
</taxonomicNameLabel>
, holotype. 67, habitus; 68, head and pronotum; 69, mandibles; 70, maxillary palpus; 71, aedeagus lateral; 72, aedeagus ventro-lateral; 73, aedeagus ventral. 71–73, same scale bars.
</paragraph>
</caption>
<caption id="DF0BB8FDFFE6FFCFFF7AFBCBFCD5E903" httpUri="https://zenodo.org/record/1441479/files/figure.png" pageId="19" pageNumber="170" startId="19.[151,264,1068,1090]" targetBox="[165,1408,201,1026]" targetPageId="19">
<paragraph id="8BCBE875FFE6FFCFFF7AFBCBFCD5E903" blockId="19.[151,1436,1068,1121]" pageId="19" pageNumber="170">
<emphasis id="B9003467FFE6FFCFFF7AFBCBFEA8E920" bold="true" box="[151,340,1068,1090]" pageId="19" pageNumber="170">FIGURES 74–77.</emphasis>
<taxonomicName id="4C7493F6FFE6FFCFFEB7FBCBFDA6E920" authority="Janák, 2018" authorityName="Janák" authorityYear="2018" box="[346,602,1068,1090]" class="Insecta" family="Staphylinidae" genus="Acylophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="170" phylum="Arthropoda" rank="species" species="madecassus" status="sp. nov.">
<emphasis id="B9003467FFE6FFCFFEB7FBCBFDA6E920" box="[346,602,1068,1090]" italics="true" pageId="19" pageNumber="170">Acylophorus madecassus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A233891CFFE6FFCFFD8CFBCBFD4DE920" box="[609,689,1068,1090]" pageId="19" pageNumber="170" rank="species">
<emphasis id="B9003467FFE6FFCFFD8CFBCBFD4DE920" bold="true" box="[609,689,1068,1090]" pageId="19" pageNumber="170">sp. nov.</emphasis>
</taxonomicNameLabel>
, holotype. 74, aedeagus lateral; 75, aedeagus ventral; 76, male tergites 9, 10; 77, male sternite 9. 74, 75, transmitted light, same scale bars.
</paragraph>
</caption>
<subSubSection id="C36EBBFEFFE6FFCFFF2AFB6AFB0BE98C" pageId="19" pageNumber="170" type="diagnosis">
<paragraph id="8BCBE875FFE6FFCFFF2AFB6AFB0BE98C" blockId="19.[151,1436,1165,1550]" pageId="19" pageNumber="170">
<emphasis id="B9003467FFE6FFCFFF2AFB6AFE31E9C4" bold="true" box="[199,461,1165,1190]" pageId="19" pageNumber="170">Differential diagnosis.</emphasis>
<taxonomicName id="4C7493F6FFE6FFCFFE3EFB6AFD61E9C7" authority="Janák, 2018" authorityName="Janák" authorityYear="2018" box="[467,669,1165,1189]" class="Insecta" family="Staphylinidae" genus="Acylophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="170" phylum="Arthropoda" rank="species" species="hova" status="sp. nov.">
<emphasis id="B9003467FFE6FFCFFE3EFB6AFD61E9C7" box="[467,669,1165,1189]" italics="true" pageId="19" pageNumber="170">Acylophorus hova</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A233891CFFE6FFCFFD49FB6AFD02E9C4" box="[676,766,1165,1190]" pageId="19" pageNumber="170" rank="species">
<emphasis id="B9003467FFE6FFCFFD49FB6AFD02E9C4" bold="true" box="[676,766,1165,1190]" pageId="19" pageNumber="170">sp. nov.</emphasis>
</taxonomicNameLabel>
is very similar to
<taxonomicName id="4C7493F6FFE6FFCFFC2AFB69FBAEE9C7" authority="Janák, 2018" authorityName="Janák" authorityYear="2018" box="[967,1106,1166,1189]" class="Insecta" family="Staphylinidae" genus="Acylophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="170" phylum="Arthropoda" rank="species" species="montanus" status="sp. nov.">
<emphasis id="B9003467FFE6FFCFFC2AFB69FBAEE9C7" box="[967,1106,1166,1189]" italics="true" pageId="19" pageNumber="170">A. montanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A233891CFFE6FFCFFBB4FB6AFB4EE9C4" box="[1113,1202,1165,1190]" pageId="19" pageNumber="170" rank="species">
<emphasis id="B9003467FFE6FFCFFBB4FB6AFB4EE9C4" bold="true" box="[1113,1202,1165,1190]" pageId="19" pageNumber="170">sp. nov.</emphasis>
</taxonomicNameLabel>
, but differs by wider pronotum, which is more widened behind, at most partly lighter last antennal segments, densely punctate abdomen, narrowely rounded apex of median lobe in males and pointed apical margin of tergite
<quantity id="4C8C4590FFE6FFCFFBB4FB31FB6EE98C" box="[1113,1170,1237,1262]" metricMagnitude="-1" metricUnit="m" metricValue="2.54" pageId="19" pageNumber="170" unit="in" value="10.0">10 in</quantity>
females.
</paragraph>
</subSubSection>
<subSubSection id="C36EBBFEFFE6FFCFFF2AFB1EFAF3E838" pageId="19" pageNumber="170" type="distribution">
<paragraph id="8BCBE875FFE6FFCFFF2AFB1EFEFBE854" blockId="19.[151,1436,1165,1550]" pageId="19" pageNumber="170">
<emphasis id="B9003467FFE6FFCFFF2AFB1EFEA5E870" bold="true" box="[199,345,1273,1298]" pageId="19" pageNumber="170">Distribution</emphasis>
. The new species is known only from Ankarafantsika in northern part of
<collectingRegion id="49B02697FFE6FFCFFB55FB1EFAF7E870" box="[1208,1291,1273,1298]" country="Sri Lanka" name="Central" pageId="19" pageNumber="170">Central</collectingRegion>
<collectingCountry id="F363A8E5FFE6FFCFFAF8FB1EFA60E870" box="[1301,1436,1273,1298]" name="Madagascar" pageId="19" pageNumber="170">Madagascar</collectingCountry>
(
<figureCitation id="134FF4F0FFE6FFCFFF72FAFAFF06E854" box="[159,250,1309,1334]" captionStart="FIGURES 172–177" captionStartId="38.[151,264,1927,1949]" captionTargetBox="[265,1321,625,1895]" captionTargetId="figure@38.[265,1321,625,1909]" captionTargetPageId="38" captionText="FIGURES 172–177. Distribution maps. 172, Acylophorus tristis Jarrige; 173, A. derougemonti sp. nov.; 174, A. muscorum Jarrige; 175, A. delphinus Fauvel (D), A. madecassus sp. nov. (Ŧ); 176, A. minutus Jarrige (D), A. hova sp. nov. (Ɨ), A. montanus sp. nov. (Δ); 177, A. sogai Jarrige." httpUri="https://zenodo.org/record/1441507/files/figure.png" pageId="19" pageNumber="170">Fig.176</figureCitation>
).
</paragraph>
<paragraph id="8BCBE875FFE6FFCFFF2AFAA6FAF3E838" blockId="19.[151,1436,1165,1550]" box="[199,1295,1345,1370]" pageId="19" pageNumber="170">
<emphasis id="B9003467FFE6FFCFFF2AFAA6FEBFE838" bold="true" box="[199,323,1345,1370]" pageId="19" pageNumber="170">Bionomics</emphasis>
. All specimens of
<taxonomicName id="4C7493F6FFE6FFCFFDFBFAA6FD1CE83B" authority="Janák, 2018" authorityName="Janák" authorityYear="2018" box="[534,736,1345,1369]" class="Insecta" family="Staphylinidae" genus="Acylophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="170" phylum="Arthropoda" rank="species" species="hova" status="sp. nov.">
<emphasis id="B9003467FFE6FFCFFDFBFAA6FD1CE83B" box="[534,736,1345,1369]" italics="true" pageId="19" pageNumber="170">Acylophorus hova</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A233891CFFE6FFCFFD05FAA6FCBDE838" box="[744,833,1345,1370]" pageId="19" pageNumber="170" rank="species">
<emphasis id="B9003467FFE6FFCFFD05FAA6FCBDE838" bold="true" box="[744,833,1345,1370]" pageId="19" pageNumber="170">sp. nov.</emphasis>
</taxonomicNameLabel>
were collected by siftings of forest litter.
</paragraph>
</subSubSection>
<subSubSection id="C36EBBFEFFE6FFCFFF2AFA82FE47EB6F" pageId="19" pageNumber="170" type="discussion">
<paragraph id="8BCBE875FFE6FFCFFF2AFA82FE47EB6F" blockId="19.[151,1436,1165,1550]" pageId="19" pageNumber="170">
<emphasis id="B9003467FFE6FFCFFF2AFA82FEB0E81C" bold="true" box="[199,332,1381,1406]" pageId="19" pageNumber="170">Discussion.</emphasis>
The new species shares very similar shape of aedeagus with
<taxonomicName id="4C7493F6FFE6FFCFFBC1FA82FACEE81F" authority="Janák, 2018" authorityName="Janák" authorityYear="2018" box="[1068,1330,1381,1405]" class="Insecta" family="Staphylinidae" genus="Acylophorus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="19" pageNumber="170" phylum="Arthropoda" rank="species" species="montanus" status="sp. nov.">
<emphasis id="B9003467FFE6FFCFFBC1FA82FACEE81F" box="[1068,1330,1381,1405]" italics="true" pageId="19" pageNumber="170">Acylophorus montanus</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A233891CFFE6FFCFFAD3FA82FA60E81C" box="[1342,1436,1381,1406]" pageId="19" pageNumber="170" rank="species">
<emphasis id="B9003467FFE6FFCFFAD3FA82FA60E81C" bold="true" box="[1342,1436,1381,1406]" pageId="19" pageNumber="170">sp. nov.</emphasis>
</taxonomicNameLabel>
Originally I supposed that all the differences between the two species fell within an intraspecific variability of one species. However after examination of females, where significant difference between the shape of apical margin of tergite 10 was found, it became apparent that these are two different species. Later I found a few additional differencies in both sexes.
</paragraph>
</subSubSection>
</treatment>
</document>
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