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<document id="4CD7EF72B0FEE23F301357D8634D62C3" ID-DOI="10.1051/acarologia/20162242" ID-ISSN="2107-7207" ID-Zenodo-Dep="5394920" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630324124403" checkinUser="marcus" docAuthor="Lindquist, E. E. &amp; Moraza, M. L." docDate="2016" docId="03D6A053317D5B50FC328327A5FFF857" docLanguage="en" docName="Acarologia.2016.56.3.293-319.pdf" docOrigin="Acarologia 56 (3)" docSource="http://dx.doi.org/10.1051/acarologia/20162242" docStyle="DocumentStyle:8E0DCBB23C2CB3FD20935B7794B24F85.3:Acarologia.2010-2017.journal_article" docStyleId="8E0DCBB23C2CB3FD20935B7794B24F85" docStyleName="Acarologia.2010-2017.journal_article" docStyleVersion="3" docTitle="Calyptoseius Lindquist &amp; Moraza 2016, new genus" docType="treatment" docVersion="3" lastPageNumber="305" masterDocId="FFEFD82B31795B5DFFE08211A17CFFCB" masterDocTitle="A new genus of mites of the subfamily Platyseiinae associated with Azteca ant galleries in Cecropia trees in Costa Rica (Acari: Mesostigmata: Blattisociidae)" masterLastPageNumber="319" masterPageNumber="293" pageNumber="296" updateTime="1699227521836" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-ND-4.0">
<mods:mods id="297954FD5E179E9CC248CB896E935B77" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="3F2750959415771E68D4E5333BFB1FC1">
<mods:title id="A8793480C358A1BADA29562AACCDC384">A new genus of mites of the subfamily Platyseiinae associated with Azteca ant galleries in Cecropia trees in Costa Rica (Acari: Mesostigmata: Blattisociidae)</mods:title>
</mods:titleInfo>
<mods:name id="0A7555D54F9C51DF6B48C6CCDF989A09" type="personal">
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</mods:role>
<mods:namePart id="D822D4DC304C3FEEE557FAB3396EB5A5">Lindquist, E. E.</mods:namePart>
</mods:name>
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<mods:namePart id="7D492643656D65E444FEF9C49072A111">Moraza, M. L.</mods:namePart>
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<mods:title id="09B4718EFBAC3EF11A3B7FF8E0F600B8">Acarologia</mods:title>
</mods:titleInfo>
<mods:part id="32C9A96F2AEBB7E86C520F3C5205D058">
<mods:date id="FC206CF616820C249CA2FD966EA7D25E">2016</mods:date>
<mods:detail id="D3ABD60A0D864777EA1DCAE0CC2C2AE6" type="pubDate">
<mods:number id="9C8D4EC40BA98E411BE5D9985A03A287">2016-07-01</mods:number>
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<mods:number id="2F28A8F53BDBC42D5E9231C127BACA32">56</mods:number>
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<mods:number id="0BE27D6D2801EFBE7627369537FACCF3">3</mods:number>
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<mods:start id="AD7A6F9800FB8C183AAADCBF87A415D8">293</mods:start>
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<mods:url id="0A4B8F3080A44220994C98368A5B1DE2">http://dx.doi.org/10.1051/acarologia/20162242</mods:url>
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<mods:classification id="C1FE0C43BD064B177C190D63E3337A61">journal article</mods:classification>
<mods:identifier id="129132C7996176BDD0D1ED11990F6516" type="DOI">10.1051/acarologia/20162242</mods:identifier>
<mods:identifier id="82C497E78713C5293CA8E79C8E9F6C4D" type="ISSN">2107-7207</mods:identifier>
<mods:identifier id="AAF2D77D3D0CD232E89491DD5EEDA008" type="Zenodo-Dep">5394920</mods:identifier>
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<treatment id="03D6A053317D5B50FC328327A5FFF857" LSID="urn:lsid:plazi:treatment:03D6A053317D5B50FC328327A5FFF857" httpUri="http://treatment.plazi.org/id/03D6A053317D5B50FC328327A5FFF857" lastPageId="13" lastPageNumber="305" pageId="4" pageNumber="296">
<subSubSection id="C36542CE317D5B59FC328327A581FE84" box="[978,1277,310,335]" pageId="4" pageNumber="296" type="nomenclature">
<paragraph id="8BC01145317D5B59FC328327A581FE84" blockId="4.[978,1277,310,370]" box="[978,1277,310,335]" pageId="4" pageNumber="296">
<heading id="D088A629317D5B59FC328327A581FE84" bold="true" box="[978,1277,310,335]" centered="true" fontSize="10" level="2" pageId="4" pageNumber="296" reason="2">
<emphasis id="B90BCD57317D5B59FC328327A581FE84" bold="true" box="[978,1277,310,335]" pageId="4" pageNumber="296">
<taxonomicName id="4C7F6AC6317D5B59FC328327A511FE85" ID-CoL="9CCM2" authority="Lindquist &amp; Moraza, 2016" authorityName="Lindquist &amp; Moraza" authorityYear="2016" box="[978,1133,310,334]" class="Arachnida" family="Blattisociidae" genus="Calyptoseius" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="296" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B90BCD57317D5B59FC328327A511FE85" bold="true" box="[978,1133,310,334]" italics="true" pageId="4" pageNumber="296">Calyptoseius</emphasis>
</taxonomicName>
,
<taxonomicNameLabel id="A238702C317D5B59FB9A8326A581FE84" box="[1146,1277,311,335]" pageId="4" pageNumber="296" rank="genus">new genus</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C36542CE317D5B50FBFC834BA43DF9ED" lastPageId="13" lastPageNumber="305" pageId="4" pageNumber="296" type="description">
<paragraph id="8BC01145317D5B59FBFC834BA5CEFEB9" blockId="4.[978,1277,310,370]" box="[1052,1202,346,370]" pageId="4" pageNumber="296">
<emphasis id="B90BCD57317D5B59FBFC834BA5CEFEB9" bold="true" box="[1052,1202,346,370]" pageId="4" pageNumber="296">
(
<figureCitation id="13440DC0317D5B59FBC4834BA5D6FEB9" box="[1060,1194,346,370]" captionStart-0="FIGURE" captionStart-1="FIGURE" captionStart-2="FIGURE" captionStart-3="FIGURE" captionStart-4="FIGURE" captionStart-5="FIGURE" captionStart-6="FIGURE" captionStart-7="FIGURE" captionStartId-0="3.[159,171,1799,1818]" captionStartId-1="5.[159,171,1700,1719]" captionStartId-2="6.[159,171,1799,1818]" captionStartId-3="7.[159,171,1615,1634]" captionStartId-4="8.[159,171,1587,1606]" captionStartId-5="9.[295,307,1617,1636]" captionStartId-6="10.[159,171,1799,1818]" captionStartId-7="11.[159,171,1042,1061]" captionTargetBox-0="[268,1314,401,1625]" captionTargetBox-1="[265,1319,477,1635]" captionTargetBox-2="[265,1323,396,1751]" captionTargetBox-3="[270,1319,589,1549]" captionTargetBox-4="[265,1316,580,1499]" captionTargetBox-5="[268,1317,623,1499]" captionTargetBox-6="[268,1314,399,1661]" captionTargetBox-7="[265,1323,302,1010]" captionTargetId-0="figure-6@3.[265,1323,396,1767]" captionTargetId-1="figure-6@5.[265,1323,467,1668]" captionTargetId-2="figure-5@6.[265,1323,396,1767]" captionTargetId-3="figure-6@7.[265,1323,579,1583]" captionTargetId-4="figure-5@8.[265,1323,580,1556]" captionTargetId-5="figure-6@9.[265,1323,605,1585]" captionTargetId-6="figure-5@10.[265,1323,396,1767]" captionTargetId-7="figure-6@11.[265,1323,302,1010]" captionTargetPageId-0="3" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="8" captionTargetPageId-5="9" captionTargetPageId-6="10" captionTargetPageId-7="11" captionText-0="FIGURE 1: Calyptoseius longinoi n. sp., adult female: A – Idiosoma, dorsal view; B – Detail of denoted dorsal setae; C – Gnathotectum; D – Subcapitulum; E – Tritosternum; F – Palp, dorsolateral view; dorsal setae in solid black color; G – Detail of palptarsus." captionText-1="FIGURE 2: Calyptoseius longinoi n. sp., adult female: A – Idiosoma, ventral view; B – Detail of proximal region of peritrematal shield and its fusion with exopodal strip; C – Chelicera, anterolateral view; D – Fixed cheliceral digit, lateral detail of distal region; E – Dorsum of fixed digit, with detail of dorsal cheliceral seta." captionText-2="FIGURE 3: Calyptoseius longinoi n. sp., adult male: A – Idiosoma, ventral view; B – Idiosoma, dorsal view; C – Subcapitulum; D – Chelicera and spermatodactyl, lateral view; E – Detail of cheliceral digits and spermatodactyl." captionText-3="FIGURE 4: Calyptoseius longinoi n. sp., legs I-IV excluding tarsus, adult female: A – Leg I, dorsal view; B – Leg II, dorsal view; C – Leg III, anterolateral view; D – Leg IV, anterolateral view. Dorsal setae in solid black color." captionText-4="FIGURE 5: Calyptoseius longinoi n. sp., adult female: A – Tarsus I, detail of distal seta &quot;s&quot;; B – Tarsus I, dorsal view; C – Tarsus II, dorsal view; D – detail of pretarsus, ventral view; E – Detail of basitarsus III; F – Tarsus IV, anterolateral view and detail of dorsal strap-like seta. Fig. B, solid black circles refer to ventral setae; Figs. C-F, dorsal setae in solid black color." captionText-5="FIGURE 6: Calyptoseius longinoi n. sp. deutonymph: A – Idiosoma, dorsal view; B – Idiosoma, ventral view." captionText-6="FIGURE 7: Calyptoseius longinoi n. sp. protonymph: A – Idiosoma, dorsal view; B – Idiosoma, ventral view; C – Detail of corniculi and salivary styli; D – Detail of apex of subcapitulum." captionText-7="FIGURE 8: Calyptoseius longinoi n. sp.: A, B – Female, spermathecal structures, different views, arrow pointing to diverticular lobe; C-E – Male, spermatodactyl, different views." figureDoi-0="http://doi.org/10.5281/zenodo.5394922" figureDoi-1="http://doi.org/10.5281/zenodo.5394924" figureDoi-2="http://doi.org/10.5281/zenodo.5394926" figureDoi-3="http://doi.org/10.5281/zenodo.5394928" figureDoi-4="http://doi.org/10.5281/zenodo.5394930" figureDoi-5="http://doi.org/10.5281/zenodo.5394933" figureDoi-6="http://doi.org/10.5281/zenodo.5394935" figureDoi-7="http://doi.org/10.5281/zenodo.5394937" httpUri-0="https://zenodo.org/record/5394922/files/figure.png" httpUri-1="https://zenodo.org/record/5394924/files/figure.png" httpUri-2="https://zenodo.org/record/5394926/files/figure.png" httpUri-3="https://zenodo.org/record/5394928/files/figure.png" httpUri-4="https://zenodo.org/record/5394930/files/figure.png" httpUri-5="https://zenodo.org/record/5394933/files/figure.png" httpUri-6="https://zenodo.org/record/5394935/files/figure.png" httpUri-7="https://zenodo.org/record/5394937/files/figure.png" pageId="4" pageNumber="296">Figures 1-8</figureCitation>
)
</emphasis>
</paragraph>
<paragraph id="8BC01145317D5B59FCDA838BA2BEFE1F" blockId="4.[826,1430,409,1948]" pageId="4" pageNumber="296">
<typeStatus id="54C4AFE7317D5B59FCDA838BA20FFE79" box="[826,883,410,434]" pageId="4" pageNumber="296">Type</typeStatus>
species:
<taxonomicName id="4C7F6AC6317D5B59FC178388A595FE7A" authority="Lindquist &amp; Moraza, 2016" authorityName="Lindquist &amp; Moraza" authorityYear="2016" box="[1015,1257,409,433]" class="Arachnida" family="Blattisociidae" genus="Calyptoseius" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="296" phylum="Arthropoda" rank="species" species="longinoi" status="sp. nov.">
<emphasis id="B90BCD57317D5B59FC178388A595FE7A" box="[1015,1257,409,433]" italics="true" pageId="4" pageNumber="296">Calyptoseius longinoi</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A238702C317D5B59FB19838BA4EEFE7A" box="[1273,1426,409,433]" pageId="4" pageNumber="296" rank="species">new species</taxonomicNameLabel>
. Monotypic.
</paragraph>
<paragraph id="8BC01145317D5B59FC8083FFA5B3F98E" blockId="4.[826,1430,409,1948]" pageId="4" pageNumber="296">
Diagnosis — Adults (and deutonymphs) of the new genus are immediately distinguished autapomorphically from those of other platyseiines as well as from other members of the superfamily
<taxonomicName id="4C7F6AC6317D5B59FAA68046A2E6FD59" pageId="4" pageNumber="296" rank="superFamily" superFamily="Phytoseioidea">Phytoseioidea</taxonomicName>
in having the tarsi of legs II to IV each with four elongated, strap- or whip-like setae (
<emphasis id="B90BCD57317D5B59FAC1808CA42CFD7E" box="[1313,1360,669,693]" italics="true" pageId="4" pageNumber="296">ad-2</emphasis>
,
<emphasis id="B90BCD57317D5B59FABD808CA4F2FD7E" box="[1373,1422,669,693]" italics="true" pageId="4" pageNumber="296">pd-2</emphasis>
,
<emphasis id="B90BCD57317D5B59FCDA80D1A21FFD13" box="[826,867,704,728]" italics="true" pageId="4" pageNumber="296">al-2</emphasis>
,
<emphasis id="B90BCD57317D5B59FC9280D1A2E0FD13" box="[882,924,704,728]" italics="true" pageId="4" pageNumber="296">pl-2</emphasis>
); those of other platyseiine genera have at most three setae (
<emphasis id="B90BCD57317D5B59FBEE80F2A541FD30" box="[1038,1085,739,763]" italics="true" pageId="4" pageNumber="296">ad-2</emphasis>
,
<emphasis id="B90BCD57317D5B59FBAE80F2A503FD30" box="[1102,1151,739,763]" italics="true" pageId="4" pageNumber="296">pd-2</emphasis>
,
<emphasis id="B90BCD57317D5B59FB6F80F2A5C6FD30" box="[1167,1210,739,763]" italics="true" pageId="4" pageNumber="296">pl-2</emphasis>
) strap-like on tarsus II and two (
<emphasis id="B90BCD57317D5B59FC148117A55FFCD5" box="[1012,1059,774,798]" italics="true" pageId="4" pageNumber="296">ad-2</emphasis>
,
<emphasis id="B90BCD57317D5B59FBD18117A51EFCD5" box="[1073,1122,774,798]" italics="true" pageId="4" pageNumber="296">pd-2</emphasis>
) strap-like on tarsi III- IV. They are also distinguished apomorphically from other platyseines in having the third pair of sternal setae alone clearly longer than the other sternal setae, the dorsal and lateral setae of most leg segments collectively palmate-ciliated, and the male having a discrete ventral shield between the sternitigenital and anal shields. They are distinguished plesiomorphically from those of other platyseiines in having 12 and 11 setae on the femur of legs I and II, respectively (by retaining
<emphasis id="B90BCD57317D5B59FB6D8674A5C1FBB6" box="[1165,1213,1125,1149]" italics="true" pageId="4" pageNumber="296">pv-2</emphasis>
), and 11 and 9 setae on the genu of legs II and III (by retaining
<emphasis id="B90BCD57317D5B59FA918698A23BFB08" italics="true" pageId="4" pageNumber="296">pv- 1</emphasis>
). Among the genera of
<taxonomicName id="4C7F6AC6317D5B59FB8E86BAA582FB08" box="[1134,1278,1195,1219]" class="Arachnida" family="Ascidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="296" phylum="Arthropoda" rank="subFamily" subFamily="Platyseiinae">Platyseiinae</taxonomicName>
,
<taxonomicName id="4C7F6AC6317D5B59FAEB86BAA4E8FB08" authorityName="Lindquist &amp; Moraza" authorityYear="2016" box="[1291,1428,1195,1219]" class="Arachnida" family="Blattisociidae" genus="Calyptoseius" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="296" phylum="Arthropoda" rank="genus">
<emphasis id="B90BCD57317D5B59FAEB86BAA4E8FB08" box="[1291,1428,1195,1219]" italics="true" pageId="4" pageNumber="296">Calyptoseius</emphasis>
</taxonomicName>
shares with
<taxonomicName id="4C7F6AC6317D5B59FC2886DFA526FB2D" baseAuthorityName="Evans and Baker" baseAuthorityYear="1991" box="[968,1114,1230,1254]" class="Arachnida" family="Ascidae" genus="Cheiroseiulus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="296" phylum="Arthropoda" rank="genus">
<emphasis id="B90BCD57317D5B59FC2886DFA526FB2D" box="[968,1114,1230,1254]" italics="true" pageId="4" pageNumber="296">Cheiroseiulus</emphasis>
</taxonomicName>
the following apomorphic attributes: opisthogaster of female and male with an anal shield; gland pores
<emphasis id="B90BCD57317D5B59FB458705A5B1FAE7" box="[1189,1229,1300,1324]" italics="true" pageId="4" pageNumber="296">gv3</emphasis>
on soft integument adjacent to anal shield; and legs I to IV with paradactyli and medial lobe of pulvillus untapered, blunt-tipped. However, adults of
<taxonomicName id="4C7F6AC6317D5B59FB3D876CA41AFA5E" authorityName="Lindquist &amp; Moraza" authorityYear="2016" box="[1245,1382,1405,1429]" class="Arachnida" family="Blattisociidae" genus="Calyptoseius" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="296" phylum="Arthropoda" rank="genus">
<emphasis id="B90BCD57317D5B59FB3D876CA41AFA5E" box="[1245,1382,1405,1429]" italics="true" pageId="4" pageNumber="296">Calyptoseius</emphasis>
</taxonomicName>
are similar plesiomorphically to those of
<taxonomicName id="4C7F6AC6317D5B59FB1987B1A409FA73" box="[1273,1397,1440,1464]" class="Arachnida" family="Ascidae" genus="Cheiroseius" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="4" pageNumber="296" phylum="Arthropoda" rank="genus">
<emphasis id="B90BCD57317D5B59FB1987B1A409FA73" box="[1273,1397,1440,1464]" italics="true" pageId="4" pageNumber="296">Cheiroseius</emphasis>
</taxonomicName>
in having a holotrichous dorsal shield with 21 to 23 pairs of setae on the podonotal region and 15 to 18 pairs on the opisthonotal region, and in having 10 setae on tibia II (seta
<emphasis id="B90BCD57317D5B59FBD1843CA51DF98E" box="[1073,1121,1581,1605]" italics="true" pageId="4" pageNumber="296">pd-2</emphasis>
present).
</paragraph>
<paragraph id="8BC01145317D5B59FC80844FA28CF9BD" blockId="4.[826,1430,409,1948]" box="[864,1008,1630,1654]" pageId="4" pageNumber="296">Description:</paragraph>
<paragraph id="8BC01145317D5B59FC80849EA4E9F857" blockId="4.[826,1430,409,1948]" pageId="4" pageNumber="296">
Idiosomatic dorsum — Adult female. Dorsal shield entire, without lateral incisions, moderately well sclerotized, surrounding soft integument smoothly striate. Dorsal shield with complement of 38 pairs of setae, including 23 podonotal pairs (
<emphasis id="B90BCD57317D5B59FA97850AA232F89D" italics="true" pageId="4" pageNumber="296">j1- j6</emphasis>
,
<emphasis id="B90BCD57317D5B59FCBC852FA2E4F89D" box="[860,920,1854,1878]" italics="true" pageId="4" pageNumber="296">z1-z6</emphasis>
,
<emphasis id="B90BCD57317D5B59FC45852FA2A1F89D" box="[933,989,1854,1878]" italics="true" pageId="4" pageNumber="296">s1-s6</emphasis>
,
<emphasis id="B90BCD57317D5B59FC0B852FA558F89D" box="[1003,1060,1854,1878]" italics="true" pageId="4" pageNumber="296">r2-r6</emphasis>
) and 15 opisthonotal pairs (
<emphasis id="B90BCD57317D5B59FA96852EA22CF8B2" italics="true" pageId="4" pageNumber="296">J1- J5</emphasis>
,
<emphasis id="B90BCD57317D5B59FC808573A2DBF8B1" box="[864,935,1890,1914]" italics="true" pageId="4" pageNumber="296">Z1-Z5</emphasis>
,
<emphasis id="B90BCD57317D5B59FC578573A28BF8B1" box="[951,1015,1890,1914]" italics="true" pageId="4" pageNumber="296">S1-S5</emphasis>
), sometimes
<emphasis id="B90BCD57317D5B59FB778570A5D3F8B2" box="[1175,1199,1889,1913]" italics="true" pageId="4" pageNumber="296">r6</emphasis>
asymmetrically off shield; setae collectively of similar, smooth, attenu-
</paragraph>
<caption id="DF0041CD317C5B58FF7F84B5A3ECF925" ID-DOI="http://doi.org/10.5281/zenodo.5394924" ID-Zenodo-Dep="5394924" httpUri="https://zenodo.org/record/5394924/files/figure.png" pageId="5" pageNumber="297" startId="5.[159,171,1700,1719]" targetBox="[265,1319,477,1635]" targetPageId="5">
<paragraph id="8BC01145317C5B58FF7F84B5A3ECF925" blockId="5.[159,1429,1699,1774]" pageId="5" pageNumber="297">
FIGURE 2:
<taxonomicName id="4C7F6AC6317C5B58FEE484B2A0C3F97D" authority="Lindquist &amp; Moraza, 2016" authorityName="Lindquist &amp; Moraza" authorityYear="2016" box="[260,447,1699,1718]" class="Arachnida" family="Blattisociidae" genus="Calyptoseius" kingdom="Animalia" order="Mesostigmata" pageId="5" pageNumber="297" phylum="Arthropoda" rank="species" species="longinoi" status="sp. nov.">
<emphasis id="B90BCD57317C5B58FEE484B2A0C3F97D" box="[260,447,1699,1718]" italics="true" pageId="5" pageNumber="297">Calyptoseius longinoi</emphasis>
</taxonomicName>
<emphasis id="B90BCD57317C5B58FE2484B2A084F97D" bold="true" box="[452,504,1699,1718]" pageId="5" pageNumber="297">
<taxonomicNameLabel id="A238702C317C5B58FE2484B2A084F97D" box="[452,504,1699,1718]" pageId="5" pageNumber="297" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, adult female: A – Idiosoma, ventral view; B – Detail of proximal region of peritrematal shield and its fusion with exopodal strip; C – Chelicera, anterolateral view; D – Fixed cheliceral digit, lateral detail of distal region; E – Dorsum of fixed digit, with detail of dorsal cheliceral seta.
</paragraph>
</caption>
<caption id="DF0041CD317F5B5BFF7F8516A54DF8FE" ID-DOI="http://doi.org/10.5281/zenodo.5394926" ID-Zenodo-Dep="5394926" httpUri="https://zenodo.org/record/5394926/files/figure.png" pageId="6" pageNumber="298" startId="6.[159,171,1799,1818]" targetBox="[265,1323,396,1751]" targetPageId="6">
<paragraph id="8BC01145317F5B5BFF7F8516A54DF8FE" blockId="6.[159,1429,1798,1845]" pageId="6" pageNumber="298">
FIGURE 3:
<taxonomicName id="4C7F6AC6317F5B5BFEEC8517A0B5F8D2" authority="Lindquist &amp; Moraza, 2016" authorityName="Lindquist &amp; Moraza" authorityYear="2016" box="[268,457,1798,1817]" class="Arachnida" family="Blattisociidae" genus="Calyptoseius" kingdom="Animalia" order="Mesostigmata" pageId="6" pageNumber="298" phylum="Arthropoda" rank="species" species="longinoi" status="sp. nov.">
<emphasis id="B90BCD57317F5B5BFEEC8517A0B5F8D2" box="[268,457,1798,1817]" italics="true" pageId="6" pageNumber="298">Calyptoseius longinoi</emphasis>
</taxonomicName>
<emphasis id="B90BCD57317F5B5BFE308517A370F8D2" bold="true" box="[464,524,1798,1817]" pageId="6" pageNumber="298">
<taxonomicNameLabel id="A238702C317F5B5BFE308517A370F8D2" box="[464,524,1798,1817]" pageId="6" pageNumber="298" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, adult male: A – Idiosoma, ventral view; B – Idiosoma, dorsal view; C – Subcapitulum; D – Chelicera and spermatodactyl, lateral view; E – Detail of cheliceral digits and spermatodactyl.
</paragraph>
</caption>
<caption id="DF0041CD317E5B5AFF7F845EA2B9F9B5" ID-DOI="http://doi.org/10.5281/zenodo.5394928" ID-Zenodo-Dep="5394928" httpUri="https://zenodo.org/record/5394928/files/figure.png" pageId="7" pageNumber="299" startId="7.[159,171,1615,1634]" targetBox="[270,1319,589,1549]" targetPageId="7">
<paragraph id="8BC01145317E5B5AFF7F845EA2B9F9B5" blockId="7.[159,1429,1615,1662]" pageId="7" pageNumber="299">
FIGURE 4:
<taxonomicName id="4C7F6AC6317E5B5AFEE4845EA0C2F9A9" authority="Lindquist &amp; Moraza, 2016" authorityName="Lindquist &amp; Moraza" authorityYear="2016" box="[260,446,1615,1634]" class="Arachnida" family="Blattisociidae" genus="Calyptoseius" kingdom="Animalia" order="Mesostigmata" pageId="7" pageNumber="299" phylum="Arthropoda" rank="species" species="longinoi" status="sp. nov.">
<emphasis id="B90BCD57317E5B5AFEE4845EA0C2F9A9" box="[260,446,1615,1634]" italics="true" pageId="7" pageNumber="299">Calyptoseius longinoi</emphasis>
</taxonomicName>
<emphasis id="B90BCD57317E5B5AFE23845EA08BF9A9" bold="true" box="[451,503,1615,1634]" pageId="7" pageNumber="299">
<taxonomicNameLabel id="A238702C317E5B5AFE23845EA08BF9A9" box="[451,503,1615,1634]" pageId="7" pageNumber="299" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, legs I-IV excluding tarsus, adult female: A – Leg I, dorsal view; B – Leg II, dorsal view; C – Leg III, anterolateral view; D – Leg IV, anterolateral view. Dorsal setae in solid black color.
</paragraph>
</caption>
<caption id="DF0041CD31715B55FF7F8422A54CF9B6" ID-DOI="http://doi.org/10.5281/zenodo.5394930" ID-Zenodo-Dep="5394930" httpUri="https://zenodo.org/record/5394930/files/figure.png" pageId="8" pageNumber="300" startId="8.[159,171,1587,1606]" targetBox="[265,1316,580,1499]" targetPageId="8">
<paragraph id="8BC0114531715B55FF7F8422A54CF9B6" blockId="8.[159,1429,1587,1662]" pageId="8" pageNumber="300">
FIGURE 5:
<taxonomicName id="4C7F6AC631715B55FEE68422A0BDF98D" authority="Lindquist &amp; Moraza, 2016" authorityName="Lindquist &amp; Moraza" authorityYear="2016" box="[262,449,1587,1606]" class="Arachnida" family="Blattisociidae" genus="Calyptoseius" kingdom="Animalia" order="Mesostigmata" pageId="8" pageNumber="300" phylum="Arthropoda" rank="species" species="longinoi" status="sp. nov.">
<emphasis id="B90BCD5731715B55FEE68422A0BDF98D" box="[262,449,1587,1606]" italics="true" pageId="8" pageNumber="300">Calyptoseius longinoi</emphasis>
</taxonomicName>
<emphasis id="B90BCD5731715B55FE278422A081F98D" bold="true" box="[455,509,1587,1606]" pageId="8" pageNumber="300">
<taxonomicNameLabel id="A238702C31715B55FE278422A081F98D" box="[455,509,1587,1606]" pageId="8" pageNumber="300" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
, adult female: A – Tarsus I, detail of distal seta &quot;s&quot;; B – Tarsus I, dorsal view; C – Tarsus II, dorsal view; D – detail of pretarsus, ventral view; E – Detail of basitarsus III; F – Tarsus IV, anterolateral view and detail of dorsal strap-like seta. Fig. B, solid black circles refer to ventral setae; Figs. C-F, dorsal setae in solid black color.
</paragraph>
</caption>
<caption id="DF0041CD31705B54FEC78440A472F9AF" ID-DOI="http://doi.org/10.5281/zenodo.5394933" ID-Zenodo-Dep="5394933" box="[295,1294,1617,1636]" httpUri="https://zenodo.org/record/5394933/files/figure.png" pageId="9" pageNumber="301" startId="9.[295,307,1617,1636]" targetBox="[268,1317,623,1499]" targetPageId="9">
<paragraph id="8BC0114531705B54FEC78440A472F9AF" blockId="9.[295,1294,1617,1636]" box="[295,1294,1617,1636]" pageId="9" pageNumber="301">
FIGURE 6:
<taxonomicName id="4C7F6AC631705B54FE6C8440A33AF9AF" authority="Lindquist &amp; Moraza, 2016" authorityName="Lindquist &amp; Moraza" authorityYear="2016" box="[396,582,1617,1636]" class="Arachnida" family="Blattisociidae" genus="Calyptoseius" kingdom="Animalia" order="Mesostigmata" pageId="9" pageNumber="301" phylum="Arthropoda" rank="species" species="longinoi" status="sp. nov.">
<emphasis id="B90BCD5731705B54FE6C8440A33AF9AF" box="[396,582,1617,1636]" italics="true" pageId="9" pageNumber="301">Calyptoseius longinoi</emphasis>
</taxonomicName>
<emphasis id="B90BCD5731705B54FDAC8440A3FCF9AF" bold="true" box="[588,640,1617,1636]" pageId="9" pageNumber="301">
<taxonomicNameLabel id="A238702C31705B54FDAC8440A3FCF9AF" box="[588,640,1617,1636]" pageId="9" pageNumber="301" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
deutonymph: A – Idiosoma, dorsal view; B – Idiosoma, ventral view.
</paragraph>
</caption>
<caption id="DF0041CD31735B57FF7F8516A3EEF8FE" ID-DOI="http://doi.org/10.5281/zenodo.5394935" ID-Zenodo-Dep="5394935" httpUri="https://zenodo.org/record/5394935/files/figure.png" pageId="10" pageNumber="302" startId="10.[159,171,1799,1818]" targetBox="[268,1314,399,1661]" targetPageId="10">
<paragraph id="8BC0114531735B57FF7F8516A3EEF8FE" blockId="10.[159,1429,1798,1845]" pageId="10" pageNumber="302">
FIGURE 7:
<taxonomicName id="4C7F6AC631735B57FEE78517A0BEF8D2" authority="Lindquist &amp; Moraza, 2016" authorityName="Lindquist &amp; Moraza" authorityYear="2016" box="[263,450,1798,1817]" class="Arachnida" family="Blattisociidae" genus="Calyptoseius" kingdom="Animalia" order="Mesostigmata" pageId="10" pageNumber="302" phylum="Arthropoda" rank="species" species="longinoi" status="sp. nov.">
<emphasis id="B90BCD5731735B57FEE78517A0BEF8D2" box="[263,450,1798,1817]" italics="true" pageId="10" pageNumber="302">Calyptoseius longinoi</emphasis>
</taxonomicName>
<emphasis id="B90BCD5731735B57FE288517A082F8D2" bold="true" box="[456,510,1798,1817]" pageId="10" pageNumber="302">
<taxonomicNameLabel id="A238702C31735B57FE288517A082F8D2" box="[456,510,1798,1817]" pageId="10" pageNumber="302" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
protonymph: A – Idiosoma, dorsal view; B – Idiosoma, ventral view; C – Detail of corniculi and salivary styli; D – Detail of apex of subcapitulum.
</paragraph>
</caption>
<paragraph id="8BC0114531725B56FB8682EBA4E9FEC6" blockId="11.[1126,1429,250,269]" box="[1126,1429,250,269]" pageId="11" pageNumber="303">
<bibCitation id="5E55192931725B56FB8682EBA4E9FEC6" box="[1126,1429,250,269]" issue="3" journal="Acarologia" page="293" pageId="11" pageNumber="303" score="5.405330260433823" volume="56" year="2016">Acarologia 56(3): 293–319 (2016)</bibCitation>
</paragraph>
<caption id="DF0041CD31725B56FF7F8603A357FB8A" ID-DOI="http://doi.org/10.5281/zenodo.5394937" ID-Zenodo-Dep="5394937" httpUri="https://zenodo.org/record/5394937/files/figure.png" pageId="11" pageNumber="303" startId="11.[159,171,1042,1061]" targetBox="[265,1323,302,1010]" targetPageId="11">
<paragraph id="8BC0114531725B56FF7F8603A357FB8A" blockId="11.[159,1429,1042,1089]" pageId="11" pageNumber="303">
FIGURE 8:
<taxonomicName id="4C7F6AC631725B56FEE48603A0C2FBEE" authority="Lindquist &amp; Moraza, 2016" authorityName="Lindquist &amp; Moraza" authorityYear="2016" box="[260,446,1042,1061]" class="Arachnida" family="Blattisociidae" genus="Calyptoseius" kingdom="Animalia" order="Mesostigmata" pageId="11" pageNumber="303" phylum="Arthropoda" rank="species" species="longinoi" status="sp. nov.">
<emphasis id="B90BCD5731725B56FEE48603A0C2FBEE" box="[260,446,1042,1061]" italics="true" pageId="11" pageNumber="303">Calyptoseius longinoi</emphasis>
</taxonomicName>
<emphasis id="B90BCD5731725B56FE238603A084FBEE" bold="true" box="[451,504,1042,1061]" pageId="11" pageNumber="303">
<taxonomicNameLabel id="A238702C31725B56FE238603A084FBEE" box="[451,504,1042,1061]" pageId="11" pageNumber="303" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
: A, B – Female, spermathecal structures, different views, arrow pointing to diverticular lobe; C-E – Male, spermatodactyl, different views.
</paragraph>
</caption>
<paragraph id="8BC0114531725B56FF7F8693A02EF9A9" blockId="11.[159,762,1154,1634]" pageId="11" pageNumber="303">
ated form and lengths except
<emphasis id="B90BCD5731725B56FDEE8693A35FFB51" box="[526,547,1154,1178]" italics="true" pageId="11" pageNumber="303">j1</emphasis>
shorter, palmateserrate, and
<emphasis id="B90BCD5731725B56FEDA86B7A02DFB76" box="[314,337,1190,1213]" italics="true" pageId="11" pageNumber="303">J5</emphasis>
shorter, slightly spiculate. Dorsal shield with complement of 23 pairs of discernible pore-like structures (nine podonotal, 14 opisthonotal), of which seven pairs (four podonotal, three opisthonotal) superficially appear secretory (gland pores) and 16 pairs (five podonotal, 11 opisthonotal) non-secretory (poroids). Soft surrounding cuticle with six or seven pairs of
<emphasis id="B90BCD5731725B56FE1D878AA373FA79" box="[509,527,1435,1458]" italics="true" pageId="11" pageNumber="303">R</emphasis>
-marginal setae and two pairs of marginal poroids including
<emphasis id="B90BCD5731725B56FD9F87AFA3CFFA1D" box="[639,691,1470,1494]" italics="true" pageId="11" pageNumber="303">idRp</emphasis>
, lacking
<emphasis id="B90BCD5731725B56FF3087F3A18AFA32" box="[208,246,1506,1529]" italics="true" pageId="11" pageNumber="303">UR</emphasis>
-setae. Peritrematal plates broadly united anteriorly with dorsal shield to level of setae
<emphasis id="B90BCD5731725B56FD468415A3C1F9D7" box="[678,701,1540,1564]" italics="true" pageId="11" pageNumber="303">r3</emphasis>
; peritremes well developed, reaching to level between setae
<emphasis id="B90BCD5731725B56FF00845BA184F9A9" box="[224,248,1610,1634]" italics="true" pageId="11" pageNumber="303">s1</emphasis>
and
<emphasis id="B90BCD5731725B56FED2845BA037F9A9" box="[306,331,1610,1634]" italics="true" pageId="11" pageNumber="303">z1</emphasis>
.
</paragraph>
<paragraph id="8BC0114531725B56FF25849EA30CF857" blockId="11.[159,762,1679,1948]" pageId="11" pageNumber="303">
Adult male — Dorsal shield with form and sclerotization as in female except more broadly united to peritrematal shields at level of setae
<emphasis id="B90BCD5731725B56FD6D84C4A3D8F926" box="[653,676,1749,1773]" italics="true" pageId="11" pageNumber="303">r4</emphasis>
. Dorsal shield with complement and form of setae similar to that of female, except with usually two more pairs of marginal setae including
<emphasis id="B90BCD5731725B56FDC9852FA33DF89D" box="[553,577,1854,1878]" italics="true" pageId="11" pageNumber="303">r6</emphasis>
and sometimes
<emphasis id="B90BCD5731725B56FF7F8573A1C2F8B1" box="[159,190,1890,1914]" italics="true" pageId="11" pageNumber="303">R1</emphasis>
. Lateral soft cuticle with five or six pairs of
<emphasis id="B90BCD5731725B56FD3F8573A38DF8B2" box="[735,753,1890,1913]" italics="true" pageId="11" pageNumber="303">R</emphasis>
- marginal setae. Peritremes as in female.
</paragraph>
<paragraph id="8BC0114531725B56FC808693A2C4F94E" blockId="11.[826,1429,1154,1669]" pageId="11" pageNumber="303">
Deutonymph — Dorsal shield with short lateral incisions, similarly sclerotized as in adults. Dorsal shield with complement of 30 pairs of setae, including 15 podonotal and 15 opisthonotal pairs, of similar length and shape as in adult; 14 pairs of dorsal setae on surrounding soft integument (
<emphasis id="B90BCD5731725B56FAE48723A461FA81" box="[1284,1309,1330,1354]" italics="true" pageId="11" pageNumber="303">z1</emphasis>
,
<emphasis id="B90BCD5731725B56FACA8723A43EFA81" box="[1322,1346,1330,1354]" italics="true" pageId="11" pageNumber="303">s1</emphasis>
,
<emphasis id="B90BCD5731725B56FAAF8723A41BFA81" box="[1359,1383,1330,1354]" italics="true" pageId="11" pageNumber="303">s2</emphasis>
,
<emphasis id="B90BCD5731725B56FA948723A22EFAA7" italics="true" pageId="11" pageNumber="303">r2- r6</emphasis>
,
<emphasis id="B90BCD5731725B56FC808745A2DAFAA7" box="[864,934,1364,1388]" italics="true" pageId="11" pageNumber="303">R1-R6</emphasis>
). Dorsal complement of discernible porelike structures as in adult. Peritrematal shields free, limited to a narrow strip with two widened areas, one at anteriormost dorsal region bearing
<emphasis id="B90BCD5731725B56FAD387AFA426FA1D" box="[1331,1370,1470,1494]" italics="true" pageId="11" pageNumber="303">gp1</emphasis>
, one at medial area bearing
<emphasis id="B90BCD5731725B56FBA087F0A515FA32" box="[1088,1129,1505,1529]" italics="true" pageId="11" pageNumber="303">gp2</emphasis>
,
<emphasis id="B90BCD5731725B56FB9487F0A5E1FA32" box="[1140,1181,1505,1529]" italics="true" pageId="11" pageNumber="303">ip2;</emphasis>
a short poststigmatic extension with
<emphasis id="B90BCD5731725B56FC118415A561F9D7" box="[1009,1053,1540,1564]" italics="true" pageId="11" pageNumber="303">ips1</emphasis>
; gland pore
<emphasis id="B90BCD5731725B56FB568415A5A2F9D7" box="[1206,1246,1540,1564]" italics="true" pageId="11" pageNumber="303">gp3</emphasis>
and poroid
<emphasis id="B90BCD5731725B56FA938415A4E9F9D7" box="[1395,1429,1540,1564]" italics="true" pageId="11" pageNumber="303">ip3</emphasis>
on a tiny platelet on soft cuticle behind stigma; peritremes well developed, narrow, extending to level of setae
<emphasis id="B90BCD5731725B56FC79847CA2CDF94E" box="[921,945,1645,1669]" italics="true" pageId="11" pageNumber="303">s1</emphasis>
.
</paragraph>
<paragraph id="8BC0114531725B51FC8084A3A3FDFD8F" blockId="11.[826,1429,1714,1949]" lastBlockId="12.[159,762,311,580]" lastPageId="12" lastPageNumber="304" pageId="11" pageNumber="303">
Protonymph — Dorsal shields weakly sclerotized. Podonotal shield with 11 pairs of setae (
<emphasis id="B90BCD5731725B56FA9784C4A232F8DB" italics="true" pageId="11" pageNumber="303">j1- j6</emphasis>
,
<emphasis id="B90BCD5731725B56FCBD84E9A20AF8DB" box="[861,886,1784,1808]" italics="true" pageId="11" pageNumber="303">z2</emphasis>
,
<emphasis id="B90BCD5731725B56FC6584E9A2E3F8DB" box="[901,927,1784,1808]" italics="true" pageId="11" pageNumber="303">z4</emphasis>
,
<emphasis id="B90BCD5731725B56FC4D84E8A2BBF8DB" box="[941,967,1785,1808]" italics="true" pageId="11" pageNumber="303">z5</emphasis>
,
<emphasis id="B90BCD5731725B56FC3584E9A291F8DB" box="[981,1005,1784,1808]" italics="true" pageId="11" pageNumber="303">s4</emphasis>
,
<emphasis id="B90BCD5731725B56FC1C84E8A568F8DB" box="[1020,1044,1785,1808]" italics="true" pageId="11" pageNumber="303">s5</emphasis>
) similar in moderate length and simple form, except
<emphasis id="B90BCD5731725B56FBDB850AA52CF8F8" box="[1083,1104,1819,1843]" italics="true" pageId="11" pageNumber="303">j1</emphasis>
shorter palmate-spiculate, flanked by four pairs of setae (
<emphasis id="B90BCD5731725B56FB47852FA5C3F89D" box="[1191,1215,1854,1878]" italics="true" pageId="11" pageNumber="303">r2</emphasis>
,
<emphasis id="B90BCD5731725B56FB2E852FA599F89D" box="[1230,1253,1854,1878]" italics="true" pageId="11" pageNumber="303">r3</emphasis>
,
<emphasis id="B90BCD5731725B56FB14852EA470F89D" box="[1268,1292,1855,1878]" italics="true" pageId="11" pageNumber="303">r5</emphasis>
,
<emphasis id="B90BCD5731725B56FAFB852FA44FF89D" box="[1307,1331,1854,1878]" italics="true" pageId="11" pageNumber="303">s6</emphasis>
) on soft lateral cuticle; pygidial shield with eight pairs of setae (
<emphasis id="B90BCD5731725B56FC8E8594A2DFF856" box="[878,931,1925,1949]" italics="true" pageId="11" pageNumber="303">J3-J5</emphasis>
,
<emphasis id="B90BCD5731725B56FC528594A285F856" box="[946,1017,1925,1949]" italics="true" pageId="11" pageNumber="303">Z3-Z5</emphasis>
,
<emphasis id="B90BCD5731725B56FBE98594A559F856" box="[1033,1061,1925,1949]" italics="true" pageId="11" pageNumber="303">S4</emphasis>
,
<emphasis id="B90BCD5731725B56FBD48594A52CF857" box="[1076,1104,1925,1948]" italics="true" pageId="11" pageNumber="303">S5</emphasis>
) of similar size and simple form except
<emphasis id="B90BCD5731755B51FED38326A035FE85" box="[307,329,311,334]" italics="true" pageId="12" pageNumber="304">J5</emphasis>
shorter, slightly spiculate, and three pairs of pore-like structures discernible (two pairs of gland pores, one pair of poroids); soft cuticle between shields with seven pairs of dorsal setae (
<emphasis id="B90BCD5731755B51FD5B83B1A3ACFE73" box="[699,720,416,440]" italics="true" pageId="12" pageNumber="304">J1</emphasis>
,
<emphasis id="B90BCD5731755B51FD3D83B1A38EFE73" box="[733,754,416,440]" italics="true" pageId="12" pageNumber="304">J2</emphasis>
,
<emphasis id="B90BCD5731755B51FF7F83D2A1C1FE10" box="[159,189,451,475]" italics="true" pageId="12" pageNumber="304">Z1</emphasis>
,
<emphasis id="B90BCD5731755B51FF2B83D2A196FE10" box="[203,234,451,475]" italics="true" pageId="12" pageNumber="304">Z2</emphasis>
,
<emphasis id="B90BCD5731755B51FF1783D2A06FFE10" box="[247,275,451,475]" italics="true" pageId="12" pageNumber="304">S2</emphasis>
,
<emphasis id="B90BCD5731755B51FEC083D2A040FE10" box="[288,316,451,475]" italics="true" pageId="12" pageNumber="304">S3</emphasis>
,
<emphasis id="B90BCD5731755B51FEAA83D2A015FE10" box="[330,361,451,475]" italics="true" pageId="12" pageNumber="304">R1</emphasis>
). Peritrematal shields fragmented and free, and peritremes short, alongside coxae III and IV; poroids
<emphasis id="B90BCD5731755B51FE828018A0F6FDEA" box="[354,394,521,545]" italics="true" pageId="12" pageNumber="304">gp2</emphasis>
on small peritrematal platelet,
<emphasis id="B90BCD5731755B51FF7F803DA1BBFD8F" box="[159,199,556,580]" italics="true" pageId="12" pageNumber="304">gp3</emphasis>
,
<emphasis id="B90BCD5731755B51FF34803DA18AFD8F" box="[212,246,556,580]" italics="true" pageId="12" pageNumber="304">ip3</emphasis>
on soft cuticle behind the stigma.
</paragraph>
<paragraph id="8BC0114531755B51FF258062A5DBFE73" blockId="12.[159,762,627,1949]" lastBlockId="12.[826,1429,311,1949]" pageId="12" pageNumber="304">
Idiosomatic venter — Adult female. Tritosternum with pilose laciniae free for most of length, their fused section and base without elaborations (
<figureCitation id="13440DC031755B51FF4780CDA076FD3F" box="[167,266,732,756]" captionStart="FIGURE" captionStartId="3.[159,171,1799,1818]" captionTargetBox="[268,1314,401,1625]" captionTargetId="figure-6@3.[265,1323,396,1767]" captionTargetPageId="3" captionText="FIGURE 1: Calyptoseius longinoi n. sp., adult female: A – Idiosoma, dorsal view; B – Detail of denoted dorsal setae; C – Gnathotectum; D – Subcapitulum; E – Tritosternum; F – Palp, dorsolateral view; dorsal setae in solid black color; G – Detail of palptarsus." figureDoi="http://doi.org/10.5281/zenodo.5394922" httpUri="https://zenodo.org/record/5394922/files/figure.png" pageId="12" pageNumber="304">Fig. 1E</figureCitation>
). Ventral shields unornamented, weakly sclerotized. Presternal region without platelets. Sternal shield entire, with well developed endopodal extensions between coxae I-II bearing gland pore
<emphasis id="B90BCD5731755B51FF3C8179A07FFC4B" box="[220,259,872,896]" italics="true" pageId="12" pageNumber="304">gvb</emphasis>
distally, and with those poorly developed between coxae II-III; shield with three pairs of setae,
<emphasis id="B90BCD5731755B51FF3481BEA189FC0C" box="[212,245,943,967]" italics="true" pageId="12" pageNumber="304">st3</emphasis>
the longest, and two pairs of lyrifissures; setae
<emphasis id="B90BCD5731755B51FF0681C3A07AFC21" box="[230,262,978,1002]" italics="true" pageId="12" pageNumber="304">st4</emphasis>
and lyrifissures
<emphasis id="B90BCD5731755B51FE3B81C3A081FC21" box="[475,509,978,1002]" italics="true" pageId="12" pageNumber="304">iv3</emphasis>
on soft cuticle (
<figureCitation id="13440DC031755B51FD2D81C3A1B8FBC6" captionStart="FIGURE" captionStartId="5.[159,171,1700,1719]" captionTargetBox="[265,1319,477,1635]" captionTargetId="figure-6@5.[265,1323,467,1668]" captionTargetPageId="5" captionText="FIGURE 2: Calyptoseius longinoi n. sp., adult female: A – Idiosoma, ventral view; B – Detail of proximal region of peritrematal shield and its fusion with exopodal strip; C – Chelicera, anterolateral view; D – Fixed cheliceral digit, lateral detail of distal region; E – Dorsum of fixed digit, with detail of dorsal cheliceral seta." figureDoi="http://doi.org/10.5281/zenodo.5394924" httpUri="https://zenodo.org/record/5394924/files/figure.png" pageId="12" pageNumber="304">Fig. 2A</figureCitation>
). Endopodal strips between coxae III and IV free, weakly defined, distant from sternal shield. Epigynal shield with its convex hyaline anterior margin between legs II-III abutting or barely overlapping posterior edge of sternal shield, its posterior margin slightly convex; setae
<emphasis id="B90BCD5731755B51FDF586B5A34AFB70" box="[533,566,1188,1211]" italics="true" pageId="12" pageNumber="304">st5</emphasis>
on shield’s lateral margins, paragenital poroids
<emphasis id="B90BCD5731755B51FDDB86D9A321FB14" box="[571,605,1224,1247]" italics="true" pageId="12" pageNumber="304">iv5</emphasis>
on soft cuticle; postgenital furrow present, its strip of transverse platelets hardly discernible. Soft opisthogastric integument with small, undivided metapodal platelets, five pairs of setae (
<emphasis id="B90BCD5731755B51FDE88745A34EFAA7" box="[520,562,1364,1388]" italics="true" pageId="12" pageNumber="304">JV1</emphasis>
,
<emphasis id="B90BCD5731755B51FDA68745A313FAA7" box="[582,623,1364,1388]" italics="true" pageId="12" pageNumber="304">JV2</emphasis>
,
<emphasis id="B90BCD5731755B51FD638745A3D0FAA0" box="[643,684,1364,1387]" italics="true" pageId="12" pageNumber="304">JV5</emphasis>
,
<emphasis id="B90BCD5731755B51FD218745A38FFAA7" box="[705,755,1364,1388]" italics="true" pageId="12" pageNumber="304">ZV1</emphasis>
,
<emphasis id="B90BCD5731755B51FF7F8766A1ADFA44" box="[159,209,1399,1423]" italics="true" pageId="12" pageNumber="304">ZV2</emphasis>
), and four pairs of poroids, flanked by posteriormost two pairs of
<emphasis id="B90BCD5731755B51FE48878BA0C6FA7A" box="[424,442,1434,1457]" italics="true" pageId="12" pageNumber="304">R</emphasis>
-setae. Opisthogaster with anal shield ovoid, with paranal setae inserted at midlevel of anus, and similar in size to postanal seta; anal valves with euanal poroids; shield with a well-developed cribrum behind level of postanal seta; gland pores
<emphasis id="B90BCD5731755B51FE948458A0E7F9AA" box="[372,411,1609,1633]" italics="true" pageId="12" pageNumber="304">gv3</emphasis>
on soft cuticle well removed from shield. Peritrematal shield connected with exopodal strips curving behind coxa IV, with two poroids (
<emphasis id="B90BCD5731755B51FEEA84A3A050F901" box="[266,300,1714,1738]" italics="true" pageId="12" pageNumber="304">ip3</emphasis>
,
<emphasis id="B90BCD5731755B51FEDB84A3A01BF901" box="[315,359,1714,1738]" italics="true" pageId="12" pageNumber="304">ips1</emphasis>
) and one gland pore (
<emphasis id="B90BCD5731755B51FD9084A3A3E4F901" box="[624,664,1714,1738]" italics="true" pageId="12" pageNumber="304">gp3</emphasis>
) in area behind stigma; shield with poroid
<emphasis id="B90BCD5731755B51FDB084C4A30DF926" box="[592,625,1749,1773]" italics="true" pageId="12" pageNumber="304">ip2</emphasis>
and gland pore
<emphasis id="B90BCD5731755B51FF3E84E9A07AF8DB" box="[222,262,1784,1808]" italics="true" pageId="12" pageNumber="304">gp2</emphasis>
along widened area below level of setae
<emphasis id="B90BCD5731755B51FF7F850AA1ABF8F8" box="[159,215,1819,1843]" italics="true" pageId="12" pageNumber="304">r3-r4</emphasis>
(
<figureCitation id="13440DC031755B51FF0A850AA036F8F8" box="[234,330,1819,1843]" captionStart="FIGURE" captionStartId="3.[159,171,1799,1818]" captionTargetBox="[268,1314,401,1625]" captionTargetId="figure-6@3.[265,1323,396,1767]" captionTargetPageId="3" captionText="FIGURE 1: Calyptoseius longinoi n. sp., adult female: A – Idiosoma, dorsal view; B – Detail of denoted dorsal setae; C – Gnathotectum; D – Subcapitulum; E – Tritosternum; F – Palp, dorsolateral view; dorsal setae in solid black color; G – Detail of palptarsus." figureDoi="http://doi.org/10.5281/zenodo.5394922" httpUri="https://zenodo.org/record/5394922/files/figure.png" pageId="12" pageNumber="304">Figs 1A</figureCitation>
,
<figureCitation id="13440DC031755B51FEBB850AA000F8F8" box="[347,380,1819,1843]" captionStart="FIGURE" captionStartId="5.[159,171,1700,1719]" captionTargetBox="[265,1319,477,1635]" captionTargetId="figure-6@5.[265,1323,467,1668]" captionTargetPageId="5" captionText="FIGURE 2: Calyptoseius longinoi n. sp., adult female: A – Idiosoma, ventral view; B – Detail of proximal region of peritrematal shield and its fusion with exopodal strip; C – Chelicera, anterolateral view; D – Fixed cheliceral digit, lateral detail of distal region; E – Dorsum of fixed digit, with detail of dorsal cheliceral seta." figureDoi="http://doi.org/10.5281/zenodo.5394924" httpUri="https://zenodo.org/record/5394924/files/figure.png" pageId="12" pageNumber="304">2B</figureCitation>
). Exopodal strip weakly developed freely alongside peritrematal shield by coxae II-IV, with small extensions between I-II, II-III, III-IV (not shown in figures). Spermathecal apparatus of phytoseioid-type, with minor duct emanating from embolus near base of sclerotized calyx, and with major duct leading from calyx to solenostome between coxae III-IV (
<figureCitation id="13440DC031755B51FBFF83B1A5E6FE73" box="[1055,1178,416,440]" captionStart="FIGURE" captionStartId="11.[159,171,1042,1061]" captionTargetBox="[265,1323,302,1010]" captionTargetId="figure-6@11.[265,1323,302,1010]" captionTargetPageId="11" captionText="FIGURE 8: Calyptoseius longinoi n. sp.: A, B – Female, spermathecal structures, different views, arrow pointing to diverticular lobe; C-E – Male, spermatodactyl, different views." figureDoi="http://doi.org/10.5281/zenodo.5394937" httpUri="https://zenodo.org/record/5394937/files/figure.png" pageId="12" pageNumber="304">Figs 8A, B</figureCitation>
).
</paragraph>
<paragraph id="8BC0114531755B51FC8083DEA2ECFB95" blockId="12.[826,1429,311,1949]" pageId="12" pageNumber="304">
Adult male. Aspects of tritosternum and presternal area as in female. Ventral shields unornamented, weakly sclerotized. Sternitigenital shield with five pairs of setae and three pairs of poroids, and fully contiguous with well developed endopodal extensions between coxae I-II, II-III, and III-IV (
<figureCitation id="13440DC031755B51FC6E80B0A285FD72" box="[910,1017,673,697]" captionStart="FIGURE" captionStartId="6.[159,171,1799,1818]" captionTargetBox="[265,1323,396,1751]" captionTargetId="figure-5@6.[265,1323,396,1767]" captionTargetPageId="6" captionText="FIGURE 3: Calyptoseius longinoi n. sp., adult male: A – Idiosoma, ventral view; B – Idiosoma, dorsal view; C – Subcapitulum; D – Chelicera and spermatodactyl, lateral view; E – Detail of cheliceral digits and spermatodactyl." figureDoi="http://doi.org/10.5281/zenodo.5394926" httpUri="https://zenodo.org/record/5394926/files/figure.png" pageId="12" pageNumber="304">Fig. 3A</figureCitation>
); relative lengths of sternal setae as in female. Opisthogaster with discrete ventral shield bearing one or two pairs of setae (
<emphasis id="B90BCD5731755B51FA8280F9A4E9FCCB" box="[1378,1429,744,768]" italics="true" pageId="12" pageNumber="304">ZV1</emphasis>
laterally and
<emphasis id="B90BCD5731755B51FC00811AA575FCE8" box="[992,1033,779,803]" italics="true" pageId="12" pageNumber="304">JV1</emphasis>
on or barely off posterior margin); soft opisthogastric integument with metapodal platelets as in female, four or five pairs of setae (
<emphasis id="B90BCD5731755B51FC8D8165A2EAFC47" box="[877,918,884,908]" italics="true" pageId="12" pageNumber="304">JV2</emphasis>
,
<emphasis id="B90BCD5731755B51FC458165A2B2FC40" box="[933,974,884,907]" italics="true" pageId="12" pageNumber="304">JV5</emphasis>
,
<emphasis id="B90BCD5731755B51FC3D8165A573FC47" box="[989,1039,884,908]" italics="true" pageId="12" pageNumber="304">ZV2</emphasis>
,
<emphasis id="B90BCD5731755B51FBFE8165A52CFC47" box="[1054,1104,884,908]" italics="true" pageId="12" pageNumber="304">ZV3</emphasis>
, sometimes
<emphasis id="B90BCD5731755B51FB048165A471FC47" box="[1252,1293,884,908]" italics="true" pageId="12" pageNumber="304">JV1</emphasis>
), and three pairs of poroids, flanked by posteriormost two pairs of
<emphasis id="B90BCD5731755B51FCB981AAA217FC19" box="[857,875,955,978]" italics="true" pageId="12" pageNumber="304">R</emphasis>
-setae. Opisthogaster with discrete anal shield, with aspects of its form, three circumanal setae, and cribrum as in female. Posteriorly and laterally, peritrematal shield, peritreme, and exopodal strip as in female.
</paragraph>
<paragraph id="8BC0114531755B51FC808664A40EF99E" blockId="12.[826,1429,311,1949]" pageId="12" pageNumber="304">
Deutonymph. Tritosternum and presternal area as in female. Ventral shields unornamented, weakly sclerotized. Sternal shield with four pairs of setae and three pairs of poroids (
<emphasis id="B90BCD5731755B51FB5A86CFA5A1FB3D" box="[1210,1245,1246,1270]" italics="true" pageId="12" pageNumber="304">iv3</emphasis>
present); setae
<emphasis id="B90BCD5731755B51FCDA8713A226FAD2" box="[826,858,1282,1305]" italics="true" pageId="12" pageNumber="304">st5</emphasis>
and poroids
<emphasis id="B90BCD5731755B51FBEA8713A550FAD2" box="[1034,1068,1282,1305]" italics="true" pageId="12" pageNumber="304">iv5</emphasis>
on soft cuticle; shield without endopodal extensions, but endopodal fragment well developed, apically with gland pore
<emphasis id="B90BCD5731755B51FAD88756A423FA94" box="[1336,1375,1351,1375]" italics="true" pageId="12" pageNumber="304">gvb</emphasis>
, between coxae I and II; endopodal fragments between coxae II-III indiscernible (
<figureCitation id="13440DC031755B51FB82879FA5CBFA6D" box="[1122,1207,1422,1446]" captionStart="FIGURE" captionStartId="9.[295,307,1617,1636]" captionTargetBox="[268,1317,623,1499]" captionTargetId="figure-6@9.[265,1323,605,1585]" captionTargetPageId="9" captionText="FIGURE 6: Calyptoseius longinoi n. sp. deutonymph: A – Idiosoma, dorsal view; B – Idiosoma, ventral view." figureDoi="http://doi.org/10.5281/zenodo.5394933" httpUri="https://zenodo.org/record/5394933/files/figure.png" pageId="12" pageNumber="304">Fig. 6B</figureCitation>
); narrow rim of exopodal plate behind coxa IV with gland pore
<emphasis id="B90BCD5731755B51FA8687A0A4F2FA02" box="[1382,1422,1457,1481]" italics="true" pageId="12" pageNumber="304">gv2</emphasis>
. Opisthogaster similar to female; soft integument with five pairs of setae, three pairs of poroids, and gland pore
<emphasis id="B90BCD5731755B51FC5E840BA29AF9F9" box="[958,998,1562,1586]" italics="true" pageId="12" pageNumber="304">gv3</emphasis>
. Anal shield with aspects of its form, three circumanal setae and cribrum as in female.
</paragraph>
<paragraph id="8BC0114531755B50FC80847DA365FE7C" blockId="12.[826,1429,311,1949]" lastBlockId="13.[159,762,311,440]" lastPageId="13" lastPageNumber="305" pageId="12" pageNumber="304">
Protonymph. Tritosternum and presternal area as in deutonymph. Sternal shield weakly delineated, with three pairs of setae and one pair of poroids (
<emphasis id="B90BCD5731755B51FC3384C4A289F926" box="[979,1013,1749,1773]" italics="true" pageId="12" pageNumber="304">iv1</emphasis>
indiscernible); endopodal extensions between coxae I-II hardly discernible but
<emphasis id="B90BCD5731755B51FA8E84E9A4E9F8DB" box="[1390,1429,1784,1808]" italics="true" pageId="12" pageNumber="304">gvb</emphasis>
present (
<figureCitation id="13440DC031755B51FC7F850AA288F8F8" box="[927,1012,1819,1843]" captionStart="FIGURE" captionStartId="10.[159,171,1799,1818]" captionTargetBox="[268,1314,399,1661]" captionTargetId="figure-5@10.[265,1323,396,1767]" captionTargetPageId="10" captionText="FIGURE 7: Calyptoseius longinoi n. sp. protonymph: A – Idiosoma, dorsal view; B – Idiosoma, ventral view; C – Detail of corniculi and salivary styli; D – Detail of apex of subcapitulum." figureDoi="http://doi.org/10.5281/zenodo.5394935" httpUri="https://zenodo.org/record/5394935/files/figure.png" pageId="12" pageNumber="304">Fig. 7B</figureCitation>
); intercoxal soft cuticle with tiny setae
<emphasis id="B90BCD5731755B51FC83852EA2F8F89D" box="[867,900,1855,1878]" italics="true" pageId="12" pageNumber="304">st5</emphasis>
between bases of legs IV. Opisthogaster with well delineated anal shield surrounded by soft integument with four pairs of setae (
<emphasis id="B90BCD5731755B51FB068594A473F856" box="[1254,1295,1925,1949]" italics="true" pageId="12" pageNumber="304">JV1</emphasis>
,
<emphasis id="B90BCD5731755B51FAC18594A436F856" box="[1313,1354,1925,1949]" italics="true" pageId="12" pageNumber="304">JV2</emphasis>
,
<emphasis id="B90BCD5731755B51FABC8594A4F2F856" box="[1372,1422,1925,1949]" italics="true" pageId="12" pageNumber="304">ZV2</emphasis>
,
<emphasis id="B90BCD5731745B50FF7F8326A1B4FE85" box="[159,200,311,334]" italics="true" pageId="13" pageNumber="305">JV5</emphasis>
) and two pairs of discernible pore-like structures including
<emphasis id="B90BCD5731745B50FE83834BA0F6FEB9" box="[355,394,346,370]" italics="true" pageId="13" pageNumber="305">gv3</emphasis>
. Anal shield with structures much as in deutonymph, except postanal seta somewhat shorter than paranal setae.
</paragraph>
<paragraph id="8BC0114531745B50FF2583C4A1BFF973" blockId="13.[159,762,469,1720]" pageId="13" pageNumber="305">
Gnathosoma — Gnathotectum with three denticulate branches (
<figureCitation id="13440DC031745B50FE9283E9A0BBFDDB" box="[370,455,504,528]" captionStart="FIGURE" captionStartId="3.[159,171,1799,1818]" captionTargetBox="[268,1314,401,1625]" captionTargetId="figure-6@3.[265,1323,396,1767]" captionTargetPageId="3" captionText="FIGURE 1: Calyptoseius longinoi n. sp., adult female: A – Idiosoma, dorsal view; B – Detail of denoted dorsal setae; C – Gnathotectum; D – Subcapitulum; E – Tritosternum; F – Palp, dorsolateral view; dorsal setae in solid black color; G – Detail of palptarsus." figureDoi="http://doi.org/10.5281/zenodo.5394922" httpUri="https://zenodo.org/record/5394922/files/figure.png" pageId="13" pageNumber="305">Fig. 1C</figureCitation>
), median branch not elongated. Chelicera slender, its shaft moderately elongated (but dimorphically shorter on male), without any conspicuous process along antiaxial or paraxial lateral surfaces near bases of the digits; fixed digit with setiform pilus dentilis and row of few small teeth along apical one-third of masticatory surface, and an offset tooth (gabelzahn) subapically (
<figureCitation id="13440DC031745B50FF478101A058FCE2" box="[167,292,784,809]" captionStart="FIGURE" captionStartId="5.[159,171,1700,1719]" captionTargetBox="[265,1319,477,1635]" captionTargetId="figure-6@5.[265,1323,467,1668]" captionTargetPageId="5" captionText="FIGURE 2: Calyptoseius longinoi n. sp., adult female: A – Idiosoma, ventral view; B – Detail of proximal region of peritrematal shield and its fusion with exopodal strip; C – Chelicera, anterolateral view; D – Fixed cheliceral digit, lateral detail of distal region; E – Dorsum of fixed digit, with detail of dorsal cheliceral seta." figureDoi="http://doi.org/10.5281/zenodo.5394924" httpUri="https://zenodo.org/record/5394924/files/figure.png" pageId="13" pageNumber="305">Figs 2C, D</figureCitation>
); movable cheliceral digit of female and nymphs bidentate, with margin of arthrodial envelope smooth; movable digit of male unidentate with spermatodactyl digitiform, free to its connection at base, directed anteriorly and somewhat ventrally. Deutosternum with usually seven (variably eight) transverse rows of denticles, anterior five rows similarly moderately wide, sixth and seventh rows somewhat wider; all rows similarly spaced from one another, multidenticulate and indistinctly connected by lateral edges. Corniculi normal in form, entire, well separated from base to apex, with blunt-tipped salivary styli appressed to dorsal cornicular faces; male with blunt lobe medial to insertion of corniculus (
<figureCitation id="13440DC031745B50FE6086EAA0A0FADF" box="[384,476,1275,1300]" captionStart="FIGURE" captionStartId="6.[159,171,1799,1818]" captionTargetBox="[265,1323,396,1751]" captionTargetId="figure-5@6.[265,1323,396,1767]" captionTargetPageId="6" captionText="FIGURE 3: Calyptoseius longinoi n. sp., adult male: A – Idiosoma, ventral view; B – Idiosoma, dorsal view; C – Subcapitulum; D – Chelicera and spermatodactyl, lateral view; E – Detail of cheliceral digits and spermatodactyl." figureDoi="http://doi.org/10.5281/zenodo.5394926" httpUri="https://zenodo.org/record/5394926/files/figure.png" pageId="13" pageNumber="305">Fig. 3C</figureCitation>
); internal malae normal in form, somewhat longer than corniculi, fimbriated laterally, not subdivided. Subcapitular setae similarly smooth, attenuate,
<emphasis id="B90BCD5731745B50FE2E8774A08AFAB6" box="[462,502,1381,1405]" italics="true" pageId="13" pageNumber="305">hp1</emphasis>
slightly longer, more whip-like than
<emphasis id="B90BCD5731745B50FEAF8799A00BFA6B" box="[335,375,1416,1440]" italics="true" pageId="13" pageNumber="305">hp3</emphasis>
and
<emphasis id="B90BCD5731745B50FE4E8799A0BAFA54" box="[430,454,1416,1439]" italics="true" pageId="13" pageNumber="305">pc</emphasis>
, which clearly longer than lateral
<emphasis id="B90BCD5731745B50FF1487BAA061FA08" box="[244,285,1451,1475]" italics="true" pageId="13" pageNumber="305">hp2</emphasis>
(
<figureCitation id="13440DC031745B50FED087BAA0E4FA08" box="[304,408,1451,1475]" captionStart="FIGURE" captionStartId="3.[159,171,1799,1818]" captionTargetBox="[268,1314,401,1625]" captionTargetId="figure-6@3.[265,1323,396,1767]" captionTargetPageId="3" captionText="FIGURE 1: Calyptoseius longinoi n. sp., adult female: A – Idiosoma, dorsal view; B – Detail of denoted dorsal setae; C – Gnathotectum; D – Subcapitulum; E – Tritosternum; F – Palp, dorsolateral view; dorsal setae in solid black color; G – Detail of palptarsus." figureDoi="http://doi.org/10.5281/zenodo.5394922" httpUri="https://zenodo.org/record/5394922/files/figure.png" pageId="13" pageNumber="305">Fig. 1D</figureCitation>
). Palpi with normal setation as described for Gamasina by
<bibRefCitation id="EFEE6CB431745B50FDF187DFA3D0FA2D" author="Evans G. O." box="[529,684,1486,1510]" pageId="13" pageNumber="305" pagination="513 - 527" refId="ref16939" refString="Evans G. O. 1964 - Some observations on the chaetotaxy of the pedipalps in the Mesostigmata (Acari) - Ann. Mag. Nat. Hist. Series 13, 6: 513 - 527." type="journal article" year="1964">Evans (1964)</bibRefCitation>
; palptrochanter with inner seta longer, more whip-like, than outer seta; palpfemur with seta
<emphasis id="B90BCD5731745B50FD8E8405A3F5F9E7" box="[622,649,1556,1580]" italics="true" pageId="13" pageNumber="305">pd</emphasis>
palmate, spiculate, like most dorsal setae on legs, and with
<emphasis id="B90BCD5731745B50FF7F844BA1CEF9B9" box="[159,178,1626,1650]" italics="true" pageId="13" pageNumber="305">al</emphasis>
spatulate; palpgenual setae
<emphasis id="B90BCD5731745B50FDE4844BA352F9B9" box="[516,558,1626,1650]" italics="true" pageId="13" pageNumber="305">al-1</emphasis>
and
<emphasis id="B90BCD5731745B50FD8B844BA3EFF9B9" box="[619,659,1626,1650]" italics="true" pageId="13" pageNumber="305">al-2</emphasis>
more or less spatulate; palptarsal apotele two-tined (
<figureCitation id="13440DC031745B50FD41846CA1CAF973" captionStart="FIGURE" captionStartId="3.[159,171,1799,1818]" captionTargetBox="[268,1314,401,1625]" captionTargetId="figure-6@3.[265,1323,396,1767]" captionTargetPageId="3" captionText="FIGURE 1: Calyptoseius longinoi n. sp., adult female: A – Idiosoma, dorsal view; B – Detail of denoted dorsal setae; C – Gnathotectum; D – Subcapitulum; E – Tritosternum; F – Palp, dorsolateral view; dorsal setae in solid black color; G – Detail of palptarsus." figureDoi="http://doi.org/10.5281/zenodo.5394922" httpUri="https://zenodo.org/record/5394922/files/figure.png" pageId="13" pageNumber="305">Figs 1F, G</figureCitation>
).
</paragraph>
<paragraph id="8BC0114531745B50FF2584C7A538FB70" blockId="13.[159,762,1749,1948]" lastBlockId="13.[826,1429,311,1574]" pageId="13" pageNumber="305">
Legs — Legs I to IV with pretarsi bearing paired claws, those of II-IV with blunt paradactyli, pulvillae with three blunt lobes, and one slender ventral acuminate lobe (
<figureCitation id="13440DC031745B50FEBE852FA0A4F89D" box="[350,472,1854,1878]" captionStart="FIGURE" captionStartId="8.[159,171,1587,1606]" captionTargetBox="[265,1316,580,1499]" captionTargetId="figure-5@8.[265,1323,580,1556]" captionTargetPageId="8" captionText="FIGURE 5: Calyptoseius longinoi n. sp., adult female: A – Tarsus I, detail of distal seta &quot;s&quot;; B – Tarsus I, dorsal view; C – Tarsus II, dorsal view; D – detail of pretarsus, ventral view; E – Detail of basitarsus III; F – Tarsus IV, anterolateral view and detail of dorsal strap-like seta. Fig. B, solid black circles refer to ventral setae; Figs. C-F, dorsal setae in solid black color." figureDoi="http://doi.org/10.5281/zenodo.5394930" httpUri="https://zenodo.org/record/5394930/files/figure.png" pageId="13" pageNumber="305">Figs 5C, D</figureCitation>
). Legs I the thinnest, legs IV especially elongate and stronger; legs II to IV with tarsus two to three times as long as tibia. Distal margins of coxae I-IV lacking serrations or spurlike processes. Tarsus I without markedly elongated setae apically, and with sensilla
<emphasis id="B90BCD5731745B50FAFD836FA45BFE5E" box="[1309,1319,382,405]" italics="true" pageId="13" pageNumber="305">s</emphasis>
conspicuous, its apex slightly lanceolate (
<figureCitation id="13440DC031745B50FB3D83B1A438FE73" box="[1245,1348,416,440]" captionStart="FIGURE" captionStartId="8.[159,171,1587,1606]" captionTargetBox="[265,1316,580,1499]" captionTargetId="figure-5@8.[265,1323,580,1556]" captionTargetPageId="8" captionText="FIGURE 5: Calyptoseius longinoi n. sp., adult female: A – Tarsus I, detail of distal seta &quot;s&quot;; B – Tarsus I, dorsal view; C – Tarsus II, dorsal view; D – detail of pretarsus, ventral view; E – Detail of basitarsus III; F – Tarsus IV, anterolateral view and detail of dorsal strap-like seta. Fig. B, solid black circles refer to ventral setae; Figs. C-F, dorsal setae in solid black color." figureDoi="http://doi.org/10.5281/zenodo.5394930" httpUri="https://zenodo.org/record/5394930/files/figure.png" pageId="13" pageNumber="305">Fig. 5A</figureCitation>
). Tarsi II-IV each with apical setal processes
<emphasis id="B90BCD5731745B50FAE183D2A44DFE10" box="[1281,1329,451,475]" italics="true" pageId="13" pageNumber="305">ad-1</emphasis>
,
<emphasis id="B90BCD5731745B50FAA283D2A40EFE10" box="[1346,1394,451,475]" italics="true" pageId="13" pageNumber="305">pd-1</emphasis>
as long as pretarsi, and with four elongated, strap- or whip-like setae,
<emphasis id="B90BCD5731745B50FBE08018A555FDEA" box="[1024,1065,521,545]" italics="true" pageId="13" pageNumber="305">al-2</emphasis>
and
<emphasis id="B90BCD5731745B50FB898018A5EFFDEA" box="[1129,1171,521,545]" italics="true" pageId="13" pageNumber="305">pl-2</emphasis>
nearly as long as
<emphasis id="B90BCD5731745B50FA938018A23BFD8F" italics="true" pageId="13" pageNumber="305">ad- 2</emphasis>
and
<emphasis id="B90BCD5731745B50FC67803DA2CBFD8F" box="[903,951,556,580]" italics="true" pageId="13" pageNumber="305">pd-2</emphasis>
. Complement of setae on segments of legs I-II-III-IV differs in several respects (indicated by boldface) from that of other
<taxonomicName id="4C7F6AC631745B50FB4F8063A440FD41" box="[1199,1340,626,650]" class="Arachnida" family="Ascidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Mesostigmata" pageId="13" pageNumber="305" phylum="Arthropoda" rank="subFamily" subFamily="Platyseiinae">Platyseiinae</taxonomicName>
as presented by
<bibRefCitation id="EFEE6CB431745B50FC538084A59DFD66" author="Lindquist E. E. &amp; Evans G. O." box="[947,1249,661,686]" pageId="13" pageNumber="305" pagination="1 - 64" refId="ref17619" refString="Lindquist E. E., Evans G. O. 1965 - Taxonomic concepts in the Ascidae, with a modified setal nomenclature for the idiosoma of the Gamasina (Acarina: Mesostigmata) - Mem. Entomol. Soc. Canada, 47: 1 - 64." type="journal article" year="1965">Lindquist &amp; Evans (1965)</bibRefCitation>
: coxae, 2-2-2-1; trochanters, 6-5-5-5; femora, 12 (2 3/1 2/
<emphasis id="B90BCD5731745B50FAF880A9A459FD1B" bold="true" box="[1304,1317,696,720]" pageId="13" pageNumber="305">2</emphasis>
2) – 11 (2 3/1 2/
<emphasis id="B90BCD5731745B50FC6880CAA2E9FD38" bold="true" box="[904,917,731,755]" pageId="13" pageNumber="305">2</emphasis>
1) – 6 (1 2/0 1/1 1) – 6 (1 2/0 1/1 1); genua, 13 (2 3/1 3/2 2) – 11 (2 3/1 2/
<emphasis id="B90BCD5731745B50FB4080EFA5D1FCDD" bold="true" box="[1184,1197,766,790]" pageId="13" pageNumber="305">1</emphasis>
2) – 9 (2 2/1 2/
<emphasis id="B90BCD5731745B50FA8A80EFA40BFCDD" bold="true" box="[1386,1399,766,790]" pageId="13" pageNumber="305">1</emphasis>
1) – 9 (2 2/1 3/0 1); tibiae, 13 (2 3/1 3/2 2) – 10 (2 2/1 2/1 2) – 9 (2 1/1 2/1 2) – 10 (2 1/1 3/1 2); femora I and II each with a third ventral seta (
<emphasis id="B90BCD5731745B50FAED8179A442FC4B" box="[1293,1342,872,896]" italics="true" pageId="13" pageNumber="305">pv-2</emphasis>
, sometimes denoted as
<emphasis id="B90BCD5731745B50FBE5819AA555FC68" box="[1029,1065,907,931]" italics="true" pageId="13" pageNumber="305">v-3</emphasis>
) added to deutonymph; genua II and III each with deutonymphal seta
<emphasis id="B90BCD5731745B50FAF981BFA436FC0D" box="[1305,1354,942,966]" italics="true" pageId="13" pageNumber="305">pv-1</emphasis>
, genu IV lacking
<emphasis id="B90BCD5731745B50FC5D81C0A292FC22" box="[957,1006,977,1001]" italics="true" pageId="13" pageNumber="305">pv-1</emphasis>
; tibia II with 10 setae,
<emphasis id="B90BCD5731745B50FAE481C0A449FC22" box="[1284,1333,977,1001]" italics="true" pageId="13" pageNumber="305">pd-2</emphasis>
present (
<figureCitation id="13440DC031745B50FCA281E5A2F5FBC7" box="[834,905,1012,1036]" captionStart="FIGURE" captionStartId="7.[159,171,1615,1634]" captionTargetBox="[270,1319,589,1549]" captionTargetId="figure-6@7.[265,1323,579,1583]" captionTargetPageId="7" captionText="FIGURE 4: Calyptoseius longinoi n. sp., legs I-IV excluding tarsus, adult female: A – Leg I, dorsal view; B – Leg II, dorsal view; C – Leg III, anterolateral view; D – Leg IV, anterolateral view. Dorsal setae in solid black color." figureDoi="http://doi.org/10.5281/zenodo.5394928" httpUri="https://zenodo.org/record/5394928/files/figure.png" pageId="13" pageNumber="305">Figs 4</figureCitation>
A-D). Other than on tarsi, dorsal and lateral leg setae collectively palmate-spiculate, and ventral setae mostly smooth. Legs of male without conspicuous dimorphically modified setae; dorsal setae of tibiae I and II smooth; dorsal setae of genua I and II less palmate-spiculate.
</paragraph>
<paragraph id="8BC0114531745B50FC8086C2A43DF9ED" blockId="13.[826,1429,311,1574]" pageId="13" pageNumber="305">
Immature instars. Structures of legs and shapes of setae in nymphal instars, and leg chaetotaxy of deutonymph as in adults. Leg chaetotaxy of protonymph typical of general pattern presented for free-living Gamasina by
<bibRefCitation id="EFEE6CB431745B50FB67874EA45CFABD" author="Evans G. O." box="[1159,1312,1374,1398]" pageId="13" pageNumber="305" pagination="275 - 303" refId="ref16887" refString="Evans G. O. 1963 - Observations on the chaetotaxy of the legs in the free-living Gamasina (Acari: Mesostigmata) - Bull. Br. Mus. Nat. Hist. Zool., 10: 275 - 303." type="journal article" year="1963">Evans (1963)</bibRefCitation>
: coxae, 2- 2-2-1; trochanters, 4-4-4-4; femora, 10 (2 3/1 1/1 2) – 8 (1 2/1 2/1 1) – 5 (1 2/1 1/0 0) – 4 (1 2/0 1/0 0); genua, 8 (1 2/1 2/1 1) – 6 (1 2/0 2/0 1) – 6 (1 2/0 2/0 1) – 5 (1 2/0 2/0 0); tibiae, 8 (1 2/1 2/1 1) – 7 (1 1/1 2/1 1) – 7 (1 1/1 2/1 1) – 7 (1 1/1 2/1 1).
</paragraph>
</subSubSection>
<subSubSection id="C36542CE31745B50FBC68442A5FFF857" pageId="13" pageNumber="305" type="etymology">
<paragraph id="8BC0114531745B50FBC68442A5D5F9A0" blockId="13.[1062,1193,1619,1643]" box="[1062,1193,1619,1643]" pageId="13" pageNumber="305">
<emphasis id="B90BCD5731745B50FBC68442A5D5F9A0" bold="true" box="[1062,1193,1619,1643]" pageId="13" pageNumber="305">Etymology</emphasis>
</paragraph>
<paragraph id="8BC0114531745B50FCDA849EA5FFF857" blockId="13.[826,1429,1679,1948]" pageId="13" pageNumber="305">
The genus name is a Latinized combination of the term
<emphasis id="B90BCD5731745B50FC9A84A3A2BBF901" box="[890,967,1714,1738]" italics="true" pageId="13" pageNumber="305">calypto</emphasis>
, based on the Greek
<emphasis id="B90BCD5731745B50FB2684A3A463F901" box="[1222,1311,1714,1738]" italics="true" pageId="13" pageNumber="305">kalyptos</emphasis>
, meaning ’concealed’ or ’covered’, and
<emphasis id="B90BCD5731745B50FB6F84C7A5B8F926" box="[1167,1220,1750,1773]" italics="true" pageId="13" pageNumber="305">seius</emphasis>
or
<emphasis id="B90BCD5731745B50FB0A84C7A463F926" box="[1258,1311,1750,1773]" italics="true" pageId="13" pageNumber="305">sejus</emphasis>
, a Roman surname used by many authors to form names for genera of mesostigmatic mites. The name is masculine in gender, and is intended to refer to the hidden niche in which these mites are found associated with ants in
<taxonomicName id="4C7F6AC631745B50FC2C8594A555F857" box="[972,1065,1925,1948]" class="Magnoliopsida" family="Urticaceae" genus="Cecropia" kingdom="Plantae" order="Rosales" pageId="13" pageNumber="305" phylum="Tracheophyta" rank="genus">
<emphasis id="B90BCD5731745B50FC2C8594A555F857" box="[972,1065,1925,1948]" italics="true" pageId="13" pageNumber="305">Cecropia</emphasis>
</taxonomicName>
trunks.
</paragraph>
</subSubSection>
</treatment>
</document>
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