230 lines
34 KiB
XML
230 lines
34 KiB
XML
<document id="26740585A450051EE41E30D2B981715D" ID-DOI="10.5281/zenodo.190516" ID-GBIF-Dataset="83063e17-1519-4a48-a51f-effa8b3c2f48" ID-ISSN="1175-5326" ID-Zenodo-Dep="190516" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461114628907" checkinUser="plazi" docAuthor="Ortuño, Vicente M. & Arillo, Antonio" docDate="2009" docId="03D6700AFF89E65445D9FD49FCFCFE9D" docLanguage="en" docName="zt02239p061.pdf" docOrigin="Zootaxa 2239" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Calathus elpis Ortuño & Arillo, 2009, n. sp." docType="treatment" docVersion="7" lastPageNumber="59" masterDocId="FFEF0872FF88E651454EFF8FFFD2FFEB" masterDocTitle="Fossil carabids from Baltic amber – I – A new species of the genus Calathus Bonelli, 1810 (Coleoptera: Carabidae: Pterostichinae)" masterLastPageNumber="61" masterPageNumber="55" pageNumber="56" updateTime="1698594811055" updateUser="plazi">
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<mods:titleInfo id="1DAEF35972B196768912513376BFFC12">
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<mods:title id="ECF27D85C0B7EDCA2CB32A0D7CB104C5">Fossil carabids from Baltic amber – I – A new species of the genus Calathus Bonelli, 1810 (Coleoptera: Carabidae: Pterostichinae)</mods:title>
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</mods:titleInfo>
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<mods:name id="A62EC5EF47A0BF746CB5B234177256C4" type="personal">
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<mods:roleTerm id="3095D6B078EBE18FA22DBDFCFE2474B5">Author</mods:roleTerm>
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<mods:namePart id="90F7B914C400692A2A9C970B4C61E057">Ortuño, Vicente M.</mods:namePart>
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<mods:name id="723F0E82FC70F725724A5667F43203F1" type="personal">
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<mods:roleTerm id="BE1B850A6B2F6FCE64C84D51784E550A">Author</mods:roleTerm>
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</mods:role>
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<mods:namePart id="6101E4226A52299E70EA135CCF61471E">Arillo, Antonio</mods:namePart>
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</mods:name>
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<mods:typeOfResource id="8AF9B25130F3FFB15A41B176EED4E213">text</mods:typeOfResource>
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<mods:titleInfo id="6C5493CB358D2ADB4BD86101D0F3CDE0">
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<mods:title id="A6DDBC40B829A40E8B5C0DB9977D2420">Zootaxa</mods:title>
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</mods:titleInfo>
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<mods:part id="028265C88CA69595859492B637A8F48C">
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<mods:date id="E81877922B3B5C471D70AC670A0CACBA">2009</mods:date>
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<mods:detail id="A763D240F91E62CE706F46C65F94663F" type="volume">
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<mods:number id="4C3216867760DFAA384927E951897D3A">2239</mods:number>
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<mods:start id="8068D31EB240C1959575B9295C1A6DD2">55</mods:start>
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<mods:end id="BFA2DD830F3F49E32085E841A4574038">61</mods:end>
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<mods:classification id="D3F64DB9A7ABF5D74EA62509837DC030">journal article</mods:classification>
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<mods:identifier id="0ACD59689F69AC4DA2E6FB892E7DA1DE" type="DOI">10.5281/zenodo.190516</mods:identifier>
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<mods:identifier id="53BAFE9D7BEAFAFD50121DD2781F7719" type="GBIF-Dataset">83063e17-1519-4a48-a51f-effa8b3c2f48</mods:identifier>
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<mods:identifier id="64C94D5C31CB312AA2C0A40379E12EBA" type="ISSN">1175-5326</mods:identifier>
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<mods:identifier id="50CC696FB4147325711320C95F8B00BF" type="Zenodo-Dep">190516</mods:identifier>
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<treatment id="03D6700AFF89E65445D9FD49FCFCFE9D" ID-DOI="http://doi.org/10.5281/zenodo.6217017" ID-GBIF-Taxon="119624709" ID-Zenodo-Dep="6217017" LSID="urn:lsid:plazi:treatment:03D6700AFF89E65445D9FD49FCFCFE9D" httpUri="http://treatment.plazi.org/id/03D6700AFF89E65445D9FD49FCFCFE9D" lastPageId="5" lastPageNumber="59" pageId="1" pageNumber="56">
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<subSubSection id="C3659297FF89E65045D9FD49FECAFCE8" pageId="1" pageNumber="56" type="nomenclature">
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<paragraph id="8BC0C11CFF89E65045D9FD49FE5EFD0B" blockId="1.[151,396,710,771]" box="[151,396,710,736]" pageId="1" pageNumber="56">
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<heading id="D0887670FF89E65045D9FD49FE5EFD0B" bold="true" box="[151,396,710,736]" fontSize="11" level="1" pageId="1" pageNumber="56" reason="1">
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<emphasis id="B90B1D0EFF89E65045D9FD49FE5EFD0B" bold="true" box="[151,396,710,736]" pageId="1" pageNumber="56">
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<taxonomicName id="4C7FBA9FFF89E65045D9FD49FE90FD0B" ID-CoL="PGDW" box="[151,322,710,736]" class="Insecta" family="Carabidae" genus="Calathus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="56" phylum="Arthropoda" rank="species" species="elpis" status="sp. nov.">
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<emphasis id="B90B1D0EFF89E65045D9FD49FE90FD0B" bold="true" box="[151,322,710,736]" italics="true" pageId="1" pageNumber="56">Calathus elpis</emphasis>
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</taxonomicName>
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<taxonomicNameLabel id="A238A075FF89E6504404FD49FE5EFD0B" box="[330,396,710,736]" pageId="1" pageNumber="56" rank="species">n. sp.</taxonomicNameLabel>
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</emphasis>
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</heading>
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</paragraph>
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<paragraph id="8BC0C11CFF89E65045D9FD66FECAFCE8" blockId="1.[151,396,710,771]" box="[151,280,745,771]" pageId="1" pageNumber="56">
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(
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<figureCitation id="1344DD99FF89E65045D1FD66FEDCFCE8" box="[159,270,745,771]" captionStart-0="FIGURE 1" captionStart-1="FIGURE 2" captionStart-2="FIGURE 3" captionStartId-0="2.[151,255,1974,1998]" captionStartId-1="3.[151,255,1952,1976]" captionStartId-2="4.[151,255,1538,1562]" captionTargetBox-0="[192,1396,183,1954]" captionTargetBox-1="[185,1393,190,1922]" captionTargetBox-2="[353,1223,191,1498]" captionTargetId-0="figure@2.[192,1398,183,1955]" captionTargetId-1="figure@3.[177,1411,156,1938]" captionTargetId-2="figure@4.[330,1257,176,1514]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionTargetPageId-2="4" captionText-0="FIGURE 1. Calathus elpis n. sp. Photographs: a. piece of amber; b. dorsal view; c. ventral view. Drawing: d. specimen in dorsal view (scale: 2 mm)." captionText-1="FIGURE 2. Calathus elpis n. sp. camera lucida drawings. a. prosternal coxae; b. metaepisternum; c. mesosternal coxae; d. metasternal coxae. (scale: 0.5 mm)." captionText-2="FIGURE 3. Calathus elpis n. sp. camera lucida drawings. a. right antennomeres; b. right prothoracic tibia and tarsus; c. right mesothoracic tibia and tarsus; d. right metathoracic tarsus; e. onychium in ventral view. (scale: 0.5 mm)." httpUri-0="https://zenodo.org/record/190517/files/figure.png" httpUri-1="https://zenodo.org/record/190519/files/figure.png" httpUri-2="https://zenodo.org/record/190520/files/figure.png" pageId="1" pageNumber="56">Figs. 1–3</figureCitation>
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)
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</paragraph>
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</subSubSection>
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<subSubSection id="C3659297FF89E65545D9FCB9FCC2F8C8" lastPageId="4" lastPageNumber="59" pageId="1" pageNumber="56" type="description">
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<paragraph id="8BC0C11CFF89E65045D9FCB9FB2CFC75" blockId="1.[151,1437,822,1984]" pageId="1" pageNumber="56">
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<emphasis id="B90B1D0EFF89E65045D9FCB9FEE0FCBB" bold="true" box="[151,306,822,848]" pageId="1" pageNumber="56">Description.</emphasis>
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<typeStatus id="54C47FBEFF89E6504475FCB9FE7DFCBB" box="[315,431,822,848]" pageId="1" pageNumber="56" type="holotype">Holotype</typeStatus>
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:
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<specimenCount id="9D790A95FF89E65044F5FCB9FDF1FCBB" box="[443,547,822,848]" pageId="1" pageNumber="56" type="female">1 female</specimenCount>
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preserved in a piece of Baltic amber (
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<quantity id="4C876CF9FF89E65046BCFCB9FB95FCBB" box="[1010,1095,822,848]" metricMagnitude="-2" metricUnit="m" metricValue="1.6" pageId="1" pageNumber="56" unit="mm" value="16.0">16 mm</quantity>
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<date id="FFC1E7DCFF89E650411EFCB9FBA4FCBB" box="[1104,1142,822,848]" pageId="1" pageNumber="56">
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x
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<quantity id="4C876CF9FF89E6504129FCB9FBA4FCBB" box="[1127,1142,822,848]" metricMagnitude="-3" metricUnit="m" metricValue="5.0" pageId="1" pageNumber="56" unit="mm" value="5.0">5</quantity>
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</date>
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mm / diameter x width) (Eocene) deposited in the Álava Museum of Natural Sciences (Vitoria,
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<collectingCountry id="F368818CFF89E65046BCFCD2FBEAFC9C" box="[1010,1080,861,887]" name="Spain" pageId="1" pageNumber="56">Spain</collectingCountry>
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) MCNA-13638 (the genitalia and other soft tissues are not preserved but the specimen is a female as protarsi are not swollen).
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</paragraph>
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<paragraph id="8BC0C11CFF89E650458BFC25FB68FC00" blockId="1.[151,1437,822,1984]" pageId="1" pageNumber="56">
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Length (from head to elytra apex): 7.5 mm. Color brown-reddish (
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<figureCitation id="1344DD99FF89E6504144FC25FB81FC2F" box="[1034,1107,938,964]" captionStart="FIGURE 1" captionStartId="2.[151,255,1974,1998]" captionTargetBox="[192,1396,183,1954]" captionTargetId="figure@2.[192,1398,183,1955]" captionTargetPageId="2" captionText="FIGURE 1. Calathus elpis n. sp. Photographs: a. piece of amber; b. dorsal view; c. ventral view. Drawing: d. specimen in dorsal view (scale: 2 mm)." httpUri="https://zenodo.org/record/190517/files/figure.png" pageId="1" pageNumber="56">Fig. 1</figureCitation>
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b and 1c). Disk of elytron depigmented at its posterior part (probably due to taphonomic processes) (
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<figureCitation id="1344DD99FF89E65046B0FC5EFB92FC00" box="[1022,1088,977,1003]" captionStart="FIGURE 1" captionStartId="2.[151,255,1974,1998]" captionTargetBox="[192,1396,183,1954]" captionTargetId="figure@2.[192,1398,183,1955]" captionTargetPageId="2" captionText="FIGURE 1. Calathus elpis n. sp. Photographs: a. piece of amber; b. dorsal view; c. ventral view. Drawing: d. specimen in dorsal view (scale: 2 mm)." httpUri="https://zenodo.org/record/190517/files/figure.png" pageId="1" pageNumber="56">Fig. 1</figureCitation>
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b and 1d).
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</paragraph>
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<paragraph id="8BC0C11CFF89E650458BFC77FEACFB17" blockId="1.[151,1437,822,1984]" pageId="1" pageNumber="56">
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Head (
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<figureCitation id="1344DD99FF89E6504456FC77FEB2FBF9" box="[280,352,1016,1042]" captionStart="FIGURE 1" captionStartId="2.[151,255,1974,1998]" captionTargetBox="[192,1396,183,1954]" captionTargetId="figure@2.[192,1398,183,1955]" captionTargetPageId="2" captionText="FIGURE 1. Calathus elpis n. sp. Photographs: a. piece of amber; b. dorsal view; c. ventral view. Drawing: d. specimen in dorsal view (scale: 2 mm)." httpUri="https://zenodo.org/record/190517/files/figure.png" pageId="1" pageNumber="56">Fig. 1</figureCitation>
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b, 1c, 1d, and 3a): Large and convex compound eyes. Long and oblique tempora. Two supraocular setae, being the posterior one at the level of posterior edge of eyes. Two setae on the clypeus. Rectangular labrum with six antero-marginal setae. Mandibles without setae on the lateral sulcus. Slender antennae with eleven antennomeres, pubescent from 4th to 11th (the 4th one lacking pubescence in its basal third part). Antennomeres bearing a crown of distal setae except the first one (with only one long seta) and the second one (shorter than the rest and without setae). Mentum tooth not visible. Maxillary and labial palps typical of
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<taxonomicName id="4C7FBA9FFF89E6504443FB6CFEA5FB17" box="[269,375,1251,1276]" class="Insecta" family="Carabidae" genus="Calathus" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="56" phylum="Arthropoda" rank="genus">
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<emphasis id="B90B1D0EFF89E6504443FB6CFEA5FB17" box="[269,375,1251,1276]" italics="true" pageId="1" pageNumber="56">Calathus</emphasis>
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</taxonomicName>
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.
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</paragraph>
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<paragraph id="8BC0C11CFF89E650458BFA86FEB8F9E0" blockId="1.[151,1437,822,1984]" pageId="1" pageNumber="56">
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Pronotum (
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<figureCitation id="1344DD99FF89E6504401FA86FE44FAC8" box="[335,406,1289,1315]" captionStart="FIGURE 1" captionStartId="2.[151,255,1974,1998]" captionTargetBox="[192,1396,183,1954]" captionTargetId="figure@2.[192,1398,183,1955]" captionTargetPageId="2" captionText="FIGURE 1. Calathus elpis n. sp. Photographs: a. piece of amber; b. dorsal view; c. ventral view. Drawing: d. specimen in dorsal view (scale: 2 mm)." httpUri="https://zenodo.org/record/190517/files/figure.png" pageId="1" pageNumber="56">Fig. 1</figureCitation>
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b and 1d) quadrangular, not transverse. Anterior margin slightly concave, being its anterior angles protuding but round shaped. Basal margin straight. Lateral margins curved, with its maximum width at the level of the anterior marginal setae. Posterior angles obtuse and blunt. Lateral channel barely evident being wider at its base. Basal fossae not very deep. Prosternum (
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<figureCitation id="1344DD99FF89E6504155FAF2FBB0FA7C" box="[1051,1122,1405,1431]" captionStart="FIGURE 1" captionStartId="2.[151,255,1974,1998]" captionTargetBox="[192,1396,183,1954]" captionTargetId="figure@2.[192,1398,183,1955]" captionTargetPageId="2" captionText="FIGURE 1. Calathus elpis n. sp. Photographs: a. piece of amber; b. dorsal view; c. ventral view. Drawing: d. specimen in dorsal view (scale: 2 mm)." httpUri="https://zenodo.org/record/190517/files/figure.png" pageId="1" pageNumber="56">Fig. 1</figureCitation>
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c and 2a) with acuminate linguiform intercoxal apophysis. Intercoxal apophysis of mesosternum (
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<figureCitation id="1344DD99FF89E650414DFA2BFB9BFA55" box="[1027,1097,1444,1470]" captionStart="FIGURE 1" captionStartId="2.[151,255,1974,1998]" captionTargetBox="[192,1396,183,1954]" captionTargetId="figure@2.[192,1398,183,1955]" captionTargetPageId="2" captionText="FIGURE 1. Calathus elpis n. sp. Photographs: a. piece of amber; b. dorsal view; c. ventral view. Drawing: d. specimen in dorsal view (scale: 2 mm)." httpUri="https://zenodo.org/record/190517/files/figure.png" pageId="1" pageNumber="56">Fig. 1</figureCitation>
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c and 2c) W-shaped, that of metasternum (
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<figureCitation id="1344DD99FF89E6504471FA45FE50FA0F" box="[319,386,1482,1508]" captionStart="FIGURE 1" captionStartId="2.[151,255,1974,1998]" captionTargetBox="[192,1396,183,1954]" captionTargetId="figure@2.[192,1398,183,1955]" captionTargetPageId="2" captionText="FIGURE 1. Calathus elpis n. sp. Photographs: a. piece of amber; b. dorsal view; c. ventral view. Drawing: d. specimen in dorsal view (scale: 2 mm)." httpUri="https://zenodo.org/record/190517/files/figure.png" pageId="1" pageNumber="56">Fig. 1</figureCitation>
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c and 2d) subtriangular. Long metaepisternum (
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<figureCitation id="1344DD99FF89E65046FAFA45FC29FA0F" box="[948,1019,1482,1508]" captionStart="FIGURE 2" captionStartId="3.[151,255,1952,1976]" captionTargetBox="[185,1393,190,1922]" captionTargetId="figure@3.[177,1411,156,1938]" captionTargetPageId="3" captionText="FIGURE 2. Calathus elpis n. sp. camera lucida drawings. a. prosternal coxae; b. metaepisternum; c. mesosternal coxae; d. metasternal coxae. (scale: 0.5 mm)." httpUri="https://zenodo.org/record/190519/files/figure.png" pageId="1" pageNumber="56">Fig. 2</figureCitation>
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b). All thoracic esternal pieces with smooth tegument.
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</paragraph>
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<paragraph id="8BC0C11CFF89E650458BF997FD3FF8C8" blockId="1.[151,1437,822,1984]" pageId="1" pageNumber="56">
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Elytra (
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<figureCitation id="1344DD99FF89E6504450F997FEB3F9D9" box="[286,353,1560,1586]" captionStart="FIGURE 1" captionStartId="2.[151,255,1974,1998]" captionTargetBox="[192,1396,183,1954]" captionTargetId="figure@2.[192,1398,183,1955]" captionTargetPageId="2" captionText="FIGURE 1. Calathus elpis n. sp. Photographs: a. piece of amber; b. dorsal view; c. ventral view. Drawing: d. specimen in dorsal view (scale: 2 mm)." httpUri="https://zenodo.org/record/190517/files/figure.png" pageId="1" pageNumber="56">Fig. 1</figureCitation>
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b and 1d) lateral margins are subparallel, sharped towards the apex. Epipleura not crossed. Rounded shoulders, without humeral teeth. All the striae are visible, from the base to the apex. Striae from 1st to 7th clearly stippled, 8th one sulciform. Lateral umbilicate serie (
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<figureCitation id="1344DD99FF89E65046E6F9E4FC3EF96E" box="[936,1004,1643,1669]" captionStart="FIGURE 1" captionStartId="2.[151,255,1974,1998]" captionTargetBox="[192,1396,183,1954]" captionTargetId="figure@2.[192,1398,183,1955]" captionTargetPageId="2" captionText="FIGURE 1. Calathus elpis n. sp. Photographs: a. piece of amber; b. dorsal view; c. ventral view. Drawing: d. specimen in dorsal view (scale: 2 mm)." httpUri="https://zenodo.org/record/190517/files/figure.png" pageId="1" pageNumber="56">Fig. 1</figureCitation>
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d) is continuous, with 15 setigerous pores over the 8th striae. Interstriae flat and with micropunctures. Discal setae (
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<figureCitation id="1344DD99FF89E650410AF91DFB55F947" box="[1092,1159,1682,1708]" captionStart="FIGURE 1" captionStartId="2.[151,255,1974,1998]" captionTargetBox="[192,1396,183,1954]" captionTargetId="figure@2.[192,1398,183,1955]" captionTargetPageId="2" captionText="FIGURE 1. Calathus elpis n. sp. Photographs: a. piece of amber; b. dorsal view; c. ventral view. Drawing: d. specimen in dorsal view (scale: 2 mm)." httpUri="https://zenodo.org/record/190517/files/figure.png" pageId="1" pageNumber="56">Fig. 1</figureCitation>
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d) barely visible due to the presence of a whitish veil partially covering the dorsum of the specimen; one seta visible in the third interval, adjacent to striae 2, two setae adjacent to striae 5 and one seta close to the apex, over the 7th striae. One apical seta is preserved over the right elytrum.
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</paragraph>
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<paragraph id="8BC0C11CFF89E650458BF8BDFC92F871" blockId="1.[151,1437,822,1984]" pageId="1" pageNumber="56">
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Both elytra are not equal in length (
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<figureCitation id="1344DD99FF89E6504732F8BDFD13F8A7" box="[636,705,1842,1868]" captionStart="FIGURE 1" captionStartId="2.[151,255,1974,1998]" captionTargetBox="[192,1396,183,1954]" captionTargetId="figure@2.[192,1398,183,1955]" captionTargetPageId="2" captionText="FIGURE 1. Calathus elpis n. sp. Photographs: a. piece of amber; b. dorsal view; c. ventral view. Drawing: d. specimen in dorsal view (scale: 2 mm)." httpUri="https://zenodo.org/record/190517/files/figure.png" pageId="1" pageNumber="56">Fig. 1</figureCitation>
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d) so striae 1st to 4th make up different anastomosis on each elytrum: setae 1–4, 2–3 on the left elytrum and setae 1–2, 3–4 on the right elytrum. This kind of light malformations are quite frequent on recent carabidofauna.
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</paragraph>
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<paragraph id="8BC0C11CFF89E650458BF829FE03F82B" blockId="1.[151,1437,822,1984]" box="[197,465,1958,1984]" pageId="1" pageNumber="56">Wings well developed.</paragraph>
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<caption id="DF009194FF8AE65345D9F839FE1DF81B" httpUri="https://zenodo.org/record/190517/files/figure.png" pageId="2" pageNumber="57" targetBox="[192,1396,183,1954]" targetPageId="2">
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<paragraph id="8BC0C11CFF8AE65345D9F839FE1DF81B" blockId="2.[151,1436,1974,2032]" pageId="2" pageNumber="57">
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<emphasis id="B90B1D0EFF8AE65345D9F839FECBF825" bold="true" box="[151,281,1974,1998]" pageId="2" pageNumber="57">FIGURE 1.</emphasis>
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<taxonomicName id="4C7FBA9FFF8AE653446EF839FE6BF826" box="[288,441,1974,1997]" class="Insecta" family="Carabidae" genus="Calathus" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="57" phylum="Arthropoda" rank="species" species="elpis" status="sp. nov.">
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<emphasis id="B90B1D0EFF8AE653446EF839FE6BF826" box="[288,441,1974,1997]" italics="true" pageId="2" pageNumber="57">Calathus elpis</emphasis>
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</taxonomicName>
|
||
<taxonomicNameLabel id="A238A075FF8AE653448EF839FE2EF825" box="[448,508,1974,1998]" pageId="2" pageNumber="57" rank="species">
|
||
<emphasis id="B90B1D0EFF8AE653448EF839FE2EF825" bold="true" box="[448,508,1974,1998]" pageId="2" pageNumber="57">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
Photographs:
|
||
<emphasis id="B90B1D0EFF8AE65347D9F839FD79F825" bold="true" box="[663,683,1974,1998]" pageId="2" pageNumber="57">a.</emphasis>
|
||
piece of amber;
|
||
<emphasis id="B90B1D0EFF8AE6534611F839FCA6F825" bold="true" box="[863,884,1974,1998]" pageId="2" pageNumber="57">b.</emphasis>
|
||
dorsal view;
|
||
<emphasis id="B90B1D0EFF8AE653414BF839FBCAF825" bold="true" box="[1029,1048,1974,1998]" pageId="2" pageNumber="57">c.</emphasis>
|
||
ventral view. Drawing:
|
||
<emphasis id="B90B1D0EFF8AE6534055F839FAE2F825" bold="true" box="[1307,1328,1974,1998]" pageId="2" pageNumber="57">d.</emphasis>
|
||
specimen in dorsal view (scale: 2 mm).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF009194FF8BE65245D9F82FFDFCF831" httpUri="https://zenodo.org/record/190519/files/figure.png" pageId="3" pageNumber="58" targetBox="[185,1393,190,1922]" targetPageId="3">
|
||
<paragraph id="8BC0C11CFF8BE65245D9F82FFDFCF831" blockId="3.[151,1436,1952,2010]" pageId="3" pageNumber="58">
|
||
<emphasis id="B90B1D0EFF8BE65245D9F82FFECBF853" bold="true" box="[151,281,1952,1976]" pageId="3" pageNumber="58">FIGURE 2.</emphasis>
|
||
<taxonomicName id="4C7FBA9FFF8BE6524451F82FFE6BF85C" box="[287,441,1952,1975]" class="Insecta" family="Carabidae" genus="Calathus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="58" phylum="Arthropoda" rank="species" species="elpis" status="sp. nov.">
|
||
<emphasis id="B90B1D0EFF8BE6524451F82FFE6BF85C" box="[287,441,1952,1975]" italics="true" pageId="3" pageNumber="58">Calathus elpis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A238A075FF8BE65244F1F82FFE29F853" box="[447,507,1952,1976]" pageId="3" pageNumber="58" rank="species">
|
||
<emphasis id="B90B1D0EFF8BE65244F1F82FFE29F853" bold="true" box="[447,507,1952,1976]" pageId="3" pageNumber="58">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
camera lucida drawings.
|
||
<emphasis id="B90B1D0EFF8BE6524642F82FFCCDF853" bold="true" box="[780,799,1952,1976]" pageId="3" pageNumber="58">a.</emphasis>
|
||
prosternal coxae;
|
||
<emphasis id="B90B1D0EFF8BE65246ADF82FFC2AF853" bold="true" box="[995,1016,1952,1976]" pageId="3" pageNumber="58">b.</emphasis>
|
||
metaepisternum;
|
||
<emphasis id="B90B1D0EFF8BE65241F9F82FFB1BF853" bold="true" box="[1207,1225,1952,1976]" pageId="3" pageNumber="58">c.</emphasis>
|
||
mesosternal coxae;
|
||
<emphasis id="B90B1D0EFF8BE65245D9F84DFF7EF831" bold="true" box="[151,172,1986,2010]" pageId="3" pageNumber="58">d.</emphasis>
|
||
metasternal coxae. (scale: 0.5 mm).
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF009194FF8CE65545D9F98DFAF8F9D7" httpUri="https://zenodo.org/record/190520/files/figure.png" pageId="4" pageNumber="59" targetBox="[353,1223,191,1498]" targetPageId="4">
|
||
<paragraph id="8BC0C11CFF8CE65545D9F98DFAF8F9D7" blockId="4.[151,1435,1538,1597]" pageId="4" pageNumber="59">
|
||
<emphasis id="B90B1D0EFF8CE65545D9F98DFECBF9F1" bold="true" box="[151,281,1538,1562]" pageId="4" pageNumber="59">FIGURE 3.</emphasis>
|
||
<taxonomicName id="4C7FBA9FFF8CE655446EF98CFE6BF9F1" box="[288,441,1539,1562]" class="Insecta" family="Carabidae" genus="Calathus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="59" phylum="Arthropoda" rank="species" species="elpis" status="sp. nov.">
|
||
<emphasis id="B90B1D0EFF8CE655446EF98CFE6BF9F1" box="[288,441,1539,1562]" italics="true" pageId="4" pageNumber="59">Calathus elpis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A238A075FF8CE655448EF98DFE2EF9F1" box="[448,508,1538,1562]" pageId="4" pageNumber="59" rank="species">
|
||
<emphasis id="B90B1D0EFF8CE655448EF98DFE2EF9F1" bold="true" box="[448,508,1538,1562]" pageId="4" pageNumber="59">n. sp.</emphasis>
|
||
</taxonomicNameLabel>
|
||
camera lucida drawings.
|
||
<emphasis id="B90B1D0EFF8CE6554640F98DFCF0F9F1" bold="true" box="[782,802,1538,1562]" pageId="4" pageNumber="59">a.</emphasis>
|
||
right antennomeres;
|
||
<emphasis id="B90B1D0EFF8CE655414AF98DFBCBF9F1" bold="true" box="[1028,1049,1538,1562]" pageId="4" pageNumber="59">b.</emphasis>
|
||
right prothoracic tibia and tarsus;
|
||
<emphasis id="B90B1D0EFF8CE65540C7F98DFA49F9F1" bold="true" box="[1417,1435,1538,1562]" pageId="4" pageNumber="59">c.</emphasis>
|
||
right mesothoracic tibia and tarsus;
|
||
<emphasis id="B90B1D0EFF8CE655475AF9AAFDFBF9D6" bold="true" box="[532,553,1573,1597]" pageId="4" pageNumber="59">d.</emphasis>
|
||
right metathoracic tarsus;
|
||
<emphasis id="B90B1D0EFF8CE6554608F9AAFC8AF9D6" bold="true" box="[838,856,1573,1597]" pageId="4" pageNumber="59">e.</emphasis>
|
||
onychium in ventral view. (scale: 0.5 mm).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8BC0C11CFF8CE655458BF9E0FCC2F8C8" blockId="4.[151,1436,1647,2020]" pageId="4" pageNumber="59">
|
||
Legs moderately long and gracile (
|
||
<figureCitation id="1344DD99FF8CE6554723F9E0FD60F962" box="[621,690,1647,1673]" captionStart="FIGURE 1" captionStartId="2.[151,255,1974,1998]" captionTargetBox="[192,1396,183,1954]" captionTargetId="figure@2.[192,1398,183,1955]" captionTargetPageId="2" captionText="FIGURE 1. Calathus elpis n. sp. Photographs: a. piece of amber; b. dorsal view; c. ventral view. Drawing: d. specimen in dorsal view (scale: 2 mm)." httpUri="https://zenodo.org/record/190517/files/figure.png" pageId="4" pageNumber="59">Fig. 1</figureCitation>
|
||
b, 1c and 1d). Spherical prothoracic (
|
||
<figureCitation id="1344DD99FF8CE6554135F9E0FB6DF962" box="[1147,1215,1647,1673]" captionStart="FIGURE 1" captionStartId="2.[151,255,1974,1998]" captionTargetBox="[192,1396,183,1954]" captionTargetId="figure@2.[192,1398,183,1955]" captionTargetPageId="2" captionText="FIGURE 1. Calathus elpis n. sp. Photographs: a. piece of amber; b. dorsal view; c. ventral view. Drawing: d. specimen in dorsal view (scale: 2 mm)." httpUri="https://zenodo.org/record/190517/files/figure.png" pageId="4" pageNumber="59">Fig. 1</figureCitation>
|
||
d and 2a), conical mesothoracic (
|
||
<figureCitation id="1344DD99FF8CE655440AF91AFE54F944" box="[324,390,1685,1711]" captionStart="FIGURE 1" captionStartId="2.[151,255,1974,1998]" captionTargetBox="[192,1396,183,1954]" captionTargetId="figure@2.[192,1398,183,1955]" captionTargetPageId="2" captionText="FIGURE 1. Calathus elpis n. sp. Photographs: a. piece of amber; b. dorsal view; c. ventral view. Drawing: d. specimen in dorsal view (scale: 2 mm)." httpUri="https://zenodo.org/record/190517/files/figure.png" pageId="4" pageNumber="59">Fig. 1</figureCitation>
|
||
d and 2c) and transverse metathoracic (aliform) coxae (
|
||
<figureCitation id="1344DD99FF8CE6554158F91AFB8AF944" box="[1046,1112,1685,1711]" captionStart="FIGURE 1" captionStartId="2.[151,255,1974,1998]" captionTargetBox="[192,1396,183,1954]" captionTargetId="figure@2.[192,1398,183,1955]" captionTargetPageId="2" captionText="FIGURE 1. Calathus elpis n. sp. Photographs: a. piece of amber; b. dorsal view; c. ventral view. Drawing: d. specimen in dorsal view (scale: 2 mm)." httpUri="https://zenodo.org/record/190517/files/figure.png" pageId="4" pageNumber="59">Fig. 1</figureCitation>
|
||
d and 2d). Protibial cleaner organ with internal notch and two guide setae (
|
||
<figureCitation id="1344DD99FF8CE6554783F933FCC7F93D" box="[717,789,1724,1750]" captionStart="FIGURE 3" captionStartId="4.[151,255,1538,1562]" captionTargetBox="[353,1223,191,1498]" captionTargetId="figure@4.[330,1257,176,1514]" captionTargetPageId="4" captionText="FIGURE 3. Calathus elpis n. sp. camera lucida drawings. a. right antennomeres; b. right prothoracic tibia and tarsus; c. right mesothoracic tibia and tarsus; d. right metathoracic tarsus; e. onychium in ventral view. (scale: 0.5 mm)." httpUri="https://zenodo.org/record/190520/files/figure.png" pageId="4" pageNumber="59">Fig. 3</figureCitation>
|
||
b). Mesotibial cleaner organ (
|
||
<figureCitation id="1344DD99FF8CE6554136F933FB6DF93D" box="[1144,1215,1724,1750]" captionStart="FIGURE 3" captionStartId="4.[151,255,1538,1562]" captionTargetBox="[353,1223,191,1498]" captionTargetId="figure@4.[330,1257,176,1514]" captionTargetPageId="4" captionText="FIGURE 3. Calathus elpis n. sp. camera lucida drawings. a. right antennomeres; b. right prothoracic tibia and tarsus; c. right mesothoracic tibia and tarsus; d. right metathoracic tarsus; e. onychium in ventral view. (scale: 0.5 mm)." httpUri="https://zenodo.org/record/190520/files/figure.png" pageId="4" pageNumber="59">Fig. 3</figureCitation>
|
||
c) similar to comb model (
|
||
<emphasis id="B90B1D0EFF8CE65545BEF96CFEE3F917" box="[240,305,1763,1788]" italics="true" pageId="4" pageNumber="59">sensu</emphasis>
|
||
<bibRefCitation id="EFEEBCEDFF8CE6554476F96DFE02F917" author="Ortuno" box="[312,464,1762,1788]" pageId="4" pageNumber="61" refString="Ortuno, V. M. (1988) Importancia de los caracteres tibiales en la sistematica de Harpalidae (Col. Caraboidea). Actas del III Congreso Iberico de Entomologia, Granada, pp. 263 - 270." type="book chapter" year="1988">Ortuño 1988</bibRefCitation>
|
||
). Tarsi smooth, long and dorsally glabrous (
|
||
<figureCitation id="1344DD99FF8CE6554683F96DFBC0F917" box="[973,1042,1762,1788]" captionStart="FIGURE 3" captionStartId="4.[151,255,1538,1562]" captionTargetBox="[353,1223,191,1498]" captionTargetId="figure@4.[330,1257,176,1514]" captionTargetPageId="4" captionText="FIGURE 3. Calathus elpis n. sp. camera lucida drawings. a. right antennomeres; b. right prothoracic tibia and tarsus; c. right mesothoracic tibia and tarsus; d. right metathoracic tarsus; e. onychium in ventral view. (scale: 0.5 mm)." httpUri="https://zenodo.org/record/190520/files/figure.png" pageId="4" pageNumber="59">Fig. 3</figureCitation>
|
||
b, 3c and 3d); onychium ventrally setose with tarsal claws pectinate (
|
||
<figureCitation id="1344DD99FF8CE6554762F886FDA3F8C8" box="[556,625,1801,1827]" captionStart="FIGURE 3" captionStartId="4.[151,255,1538,1562]" captionTargetBox="[353,1223,191,1498]" captionTargetId="figure@4.[330,1257,176,1514]" captionTargetPageId="4" captionText="FIGURE 3. Calathus elpis n. sp. camera lucida drawings. a. right antennomeres; b. right prothoracic tibia and tarsus; c. right mesothoracic tibia and tarsus; d. right metathoracic tarsus; e. onychium in ventral view. (scale: 0.5 mm)." httpUri="https://zenodo.org/record/190520/files/figure.png" pageId="4" pageNumber="59">Fig. 3</figureCitation>
|
||
b, 3c and 3e).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C3659297FF8CE6544588F8BFFCFCFE9D" lastPageId="5" lastPageNumber="60" pageId="4" pageNumber="59" type="etymology">
|
||
<paragraph id="8BC0C11CFF8CE6554588F8BFFBF8F87C" blockId="4.[151,1436,1647,2020]" pageId="4" pageNumber="59">
|
||
<emphasis id="B90B1D0EFF8CE6554588F8BFFE85F8A1" bold="true" box="[198,343,1840,1866]" pageId="4" pageNumber="59">Etymology:</emphasis>
|
||
Specific epithet is after
|
||
<taxonomicName id="4C7FBA9FFF8CE655473AF8BFFD62F8A1" box="[628,688,1840,1866]" class="Insecta" family="Carabidae" genus="Calathus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="60" phylum="Arthropoda" rank="species" species="elpis" status="sp. nov.">Elpis</taxonomicName>
|
||
(λπίς), who was in the Greek mythology the personification of hope that remained in the Pandora’s box to help people to bear the misfortunes.
|
||
<taxonomicName id="4C7FBA9FFF8CE6554106F8D8FB56F89A" box="[1096,1156,1879,1905]" class="Insecta" family="Carabidae" genus="Calathus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="60" phylum="Arthropoda" rank="species" species="elpis" status="sp. nov.">Elpis</taxonomicName>
|
||
(Esperanza in Spanish) is after our colleague, the anthropologist Dra. Esperanza Gutiérrez Redomero.
|
||
</paragraph>
|
||
<paragraph id="8BC0C11CFF8CE6544588F82BFCFCFE9D" blockId="4.[151,1436,1647,2020]" lastBlockId="5.[151,1437,152,374]" lastPageId="5" lastPageNumber="60" pageId="4" pageNumber="59">
|
||
<emphasis id="B90B1D0EFF8CE6554588F82BFE1DF855" bold="true" box="[198,463,1956,1982]" pageId="4" pageNumber="59">State of preservation.</emphasis>
|
||
Although the specimen is slightly damaged, the state of preservation is quite good, so morphological and taxonomic studies are possible. Most of soft tissues are digested, being the tegument quite transparent. Dorsal side of the fossil is partially covered by a withish coat (
|
||
<figureCitation id="1344DD99FF8DE654411BFF17FB4AFF59" box="[1109,1176,152,178]" captionStart="FIGURE 1" captionStartId="2.[151,255,1974,1998]" captionTargetBox="[192,1396,183,1954]" captionTargetId="figure@2.[192,1398,183,1955]" captionTargetPageId="2" captionText="FIGURE 1. Calathus elpis n. sp. Photographs: a. piece of amber; b. dorsal view; c. ventral view. Drawing: d. specimen in dorsal view (scale: 2 mm)." httpUri="https://zenodo.org/record/190517/files/figure.png" pageId="5" pageNumber="60">Fig. 1</figureCitation>
|
||
b and 1d) but most of diagnostic characters of the genus are visible. Abdominal sternites are partially destroyed (
|
||
<figureCitation id="1344DD99FF8DE65441F8FF31FB25FF33" box="[1206,1271,190,216]" captionStart="FIGURE 1" captionStartId="2.[151,255,1974,1998]" captionTargetBox="[192,1396,183,1954]" captionTargetId="figure@2.[192,1398,183,1955]" captionTargetPageId="2" captionText="FIGURE 1. Calathus elpis n. sp. Photographs: a. piece of amber; b. dorsal view; c. ventral view. Drawing: d. specimen in dorsal view (scale: 2 mm)." httpUri="https://zenodo.org/record/190517/files/figure.png" pageId="5" pageNumber="60">Fig. 1</figureCitation>
|
||
c) allowing to observe the folded wings under the elytra. Left prothoracic leg is not complete; only the coxa, the trochanter and the base of the femur are preserved (
|
||
<figureCitation id="1344DD99FF8DE6544739FE83FD68FECD" box="[631,698,268,294]" captionStart="FIGURE 1" captionStartId="2.[151,255,1974,1998]" captionTargetBox="[192,1396,183,1954]" captionTargetId="figure@2.[192,1398,183,1955]" captionTargetPageId="2" captionText="FIGURE 1. Calathus elpis n. sp. Photographs: a. piece of amber; b. dorsal view; c. ventral view. Drawing: d. specimen in dorsal view (scale: 2 mm)." httpUri="https://zenodo.org/record/190517/files/figure.png" pageId="5" pageNumber="60">Fig. 1</figureCitation>
|
||
c). Left mesothoracic leg has lost the tarsi and most part of the tibia (
|
||
<figureCitation id="1344DD99FF8DE6544592FEBDFECDFEA7" box="[220,287,306,332]" captionStart="FIGURE 1" captionStartId="2.[151,255,1974,1998]" captionTargetBox="[192,1396,183,1954]" captionTargetId="figure@2.[192,1398,183,1955]" captionTargetPageId="2" captionText="FIGURE 1. Calathus elpis n. sp. Photographs: a. piece of amber; b. dorsal view; c. ventral view. Drawing: d. specimen in dorsal view (scale: 2 mm)." httpUri="https://zenodo.org/record/190517/files/figure.png" pageId="5" pageNumber="60">Fig. 1</figureCitation>
|
||
b, 1c and 1d). Right metathoracic leg has lost the tarsi (
|
||
<figureCitation id="1344DD99FF8DE65446FCFEBDFC24FEA7" box="[946,1014,306,332]" captionStart="FIGURE 1" captionStartId="2.[151,255,1974,1998]" captionTargetBox="[192,1396,183,1954]" captionTargetId="figure@2.[192,1398,183,1955]" captionTargetPageId="2" captionText="FIGURE 1. Calathus elpis n. sp. Photographs: a. piece of amber; b. dorsal view; c. ventral view. Drawing: d. specimen in dorsal view (scale: 2 mm)." httpUri="https://zenodo.org/record/190517/files/figure.png" pageId="5" pageNumber="60">Fig. 1</figureCitation>
|
||
c). Left antenna is incomplete, only preserved up to the 4th antennomere (
|
||
<figureCitation id="1344DD99FF8DE6544706FED3FD58FE9D" box="[584,650,348,374]" captionStart="FIGURE 1" captionStartId="2.[151,255,1974,1998]" captionTargetBox="[192,1396,183,1954]" captionTargetId="figure@2.[192,1398,183,1955]" captionTargetPageId="2" captionText="FIGURE 1. Calathus elpis n. sp. Photographs: a. piece of amber; b. dorsal view; c. ventral view. Drawing: d. specimen in dorsal view (scale: 2 mm)." httpUri="https://zenodo.org/record/190517/files/figure.png" pageId="5" pageNumber="60">Fig. 1</figureCitation>
|
||
b, 1c and 1d).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |