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<document id="A8E7B6867C11C8EE17773276504C09CA" ID="10.11646/zootaxa.4162.2.5" ID-DOI="10.11646/zootaxa.4162.2.5" ID-GBIF-Dataset="b2f6eebe-ccaf-4b7b-914f-eca28acf2c28" ID-ISSN="1175-5326" ID-Zenodo-Dep="164959" ID-ZooBank="1F76AD06-AA68-42C7-BBAF-DA6CF276E332" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1473446029981" checkinUser="plazi" docAuthor="Chatzimanolis, Stylianos" docDate="2016" docId="03C78788A9062E71FF460B9EDBF2A59A" docLanguage="en" docName="zootaxa.4162.2.5.pdf" docOrigin="Zootaxa 4162 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Smilax" docType="key" docVersion="11" lastPageNumber="300" masterDocId="FFFEFFF0A9172E60FFD10D35DE69A26B" masterDocTitle="A revision of the myrmecophilous genus Smilax Laporte (Coleoptera: Staphylinidae: Staphylininae)" masterLastPageNumber="303" masterPageNumber="283" pageNumber="300" updateTime="1698384781401" updateUser="plazi">
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<mods:title id="04E806F323DE8905DA23D95B88028151">A revision of the myrmecophilous genus Smilax Laporte (Coleoptera: Staphylinidae: Staphylininae)</mods:title>
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<mods:namePart id="257A846CDBF174005254CC6BA1FF1776">Chatzimanolis, Stylianos</mods:namePart>
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<mods:date id="0AF248C514B0CD48ED667EE85468BBE1">2016</mods:date>
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<treatment id="03C78788A9062E71FF460B9EDBF2A59A" ID-DOI="http://doi.org/10.5281/zenodo.6089857" ID-GBIF-Taxon="124483640" ID-Zenodo-Dep="6089857" LSID="urn:lsid:plazi:treatment:03C78788A9062E71FF460B9EDBF2A59A" httpUri="http://treatment.plazi.org/id/03C78788A9062E71FF460B9EDBF2A59A" lastPageNumber="300" pageId="17" pageNumber="300">
<subSubSection id="C3746515A9062E71FF460B9EDF98A4AF" box="[151,497,1706,1733]" pageId="17" pageNumber="300" type="nomenclature">
<paragraph id="8BD1369EA9062E71FF460B9EDF98A4AF" blockId="17.[151,497,1706,1733]" box="[151,497,1706,1733]" pageId="17" pageNumber="300">
<heading id="D09981F2A9062E71FF460B9EDF98A4AF" bold="true" box="[151,497,1706,1733]" fontSize="11" level="1" pageId="17" pageNumber="300" reason="1">
<emphasis id="B91AEA8CA9062E71FF460B9EDF98A4AF" bold="true" box="[151,497,1706,1733]" pageId="17" pageNumber="300">
Key to the species of
<taxonomicName id="4C6E4D1DA9062E71FE4D0B9FDF98A4AF" ID-CoL="8W3MK" box="[412,497,1706,1732]" class="Insecta" family="Staphylinidae" genus="Smilax" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="300" phylum="Arthropoda" rank="genus">
<emphasis id="B91AEA8CA9062E71FE4D0B9FDF98A4AF" bold="true" box="[412,497,1706,1732]" italics="true" pageId="17" pageNumber="300">Smilax</emphasis>
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C3746515A9062E71FF460BC5DBF2A59A" pageId="17" pageNumber="300" type="key">
<keyStep id="309A2E3BA9062E71FF460BC5DBF5A52A" pageId="17" pageNumber="300">
<paragraph id="8BD1369EA9062E71FF460BC5DBF5A548" blockId="17.[151,1436,1776,2033]" pageId="17" pageNumber="300">
<keyLead id="309F95ABA9062E71FF460BC5DBF5A548" pageId="17" pageNumber="300">
1. Pronotum with metallic green-golden coloration (
<figureCitation id="13552A1BA9062E71FD6A0BC5DC97A56D" box="[699,766,1776,1798]" captionStart="FIGURES 3 4" captionStartId="4.[151,264,1738,1761]" captionTargetBox="[328,1248,185,1711]" captionTargetId="figure@4.[326,1261,181,1717]" captionTargetPageId="4" captionText="FIGURES 3 4. Habitus photographs of Smilax Laporte. 3. Smilax kuntzeni (Scheerpeltz), total length = 19.1 mm. 4. Smilax lynchi (Bruch), total length = 18.5 mm." httpUri="https://zenodo.org/record/268255/files/figure.png" pageId="17" pageNumber="300">Figs. 3</figureCitation>
,
<figureCitation id="13552A1BA9062E71FCD80BC5DD4DA56D" box="[777,804,1776,1798]" captionStart="FIGURES 9 12" captionStartId="6.[151,264,1220,1243]" captionTargetBox="[219,1369,447,1179]" captionTargetId="figure@6.[217,1369,447,1180]" captionTargetPageId="6" captionText="FIGURES 9 12. Pronotum of Smilax Laporte. 9. Smilax deneinephyto Chatzimanolis. 10. Smilax kuntzeni (Scheerpeltz). 11. Smilax lynchi (Bruch). 12. Smilax pilosa (F.). Not to scale." httpUri="https://zenodo.org/record/268257/files/figure.png" pageId="17" pageNumber="300">10</figureCitation>
); aedeagus with paramere having sensory spinules (
<figureCitation id="13552A1BA9062E71FAC70BC5DBE5A56D" box="[1302,1420,1776,1798]" captionStart="FIGURES 26 28" captionStartId="10.[151,264,1770,1793]" captionTargetBox="[196,1383,517,1726]" captionTargetId="figure@10.[195,1383,517,1739]" captionTargetPageId="10" captionText="FIGURES 26 28. Aedeagus of S. kuntzeni (Scheerpeltz). 26. Lateral view. 27. Dorsal view. 28. Detail of paramere, ventral view." httpUri="https://zenodo.org/record/164967/files/figure.png" pageId="17" pageNumber="300">Figs. 26, 28</figureCitation>
)............................................................................................ ..
<taxonomicName id="4C6E4D1DA9062E71FAE50A3BDBF5A548" ID-CoL="4XWKJ" box="[1332,1436,1806,1827]" class="Insecta" family="Staphylinidae" genus="Smilax" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="300" phylum="Arthropoda" rank="species" species="kuntzeni">
<emphasis id="B91AEA8CA9062E71FAE50A3BDBF5A548" box="[1332,1436,1806,1827]" italics="true" pageId="17" pageNumber="300">S. kuntzeni</emphasis>
</taxonomicName>
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</paragraph>
<paragraph id="8BD1369EA9062E71FF460A19DBF5A52A" blockId="17.[151,1436,1776,2033]" box="[151,1436,1835,1857]" pageId="17" pageNumber="300">
<keyLead id="309F95ABA9062E71FF460A19DBF5A52A" box="[151,1436,1835,1857]" pageId="17" pageNumber="300">
<emphasis id="B91AEA8CA9062E71FF460A19DEF6A52A" box="[151,159,1836,1857]" italics="true" pageId="17" pageNumber="300">-</emphasis>
Pronotum with metallic dark blue-brown coloration (
<figureCitation id="13552A1BA9062E71FD0D0A1EDD00A52A" box="[732,873,1835,1857]" captionStart="FIGURES 9 12" captionStartId="6.[151,264,1220,1243]" captionTargetBox="[219,1369,447,1179]" captionTargetId="figure@6.[217,1369,447,1180]" captionTargetPageId="6" captionText="FIGURES 9 12. Pronotum of Smilax Laporte. 9. Smilax deneinephyto Chatzimanolis. 10. Smilax kuntzeni (Scheerpeltz). 11. Smilax lynchi (Bruch). 12. Smilax pilosa (F.). Not to scale." httpUri="https://zenodo.org/record/268257/files/figure.png" pageId="17" pageNumber="300">Figs. 9, 1011</figureCitation>
); aedeagus without sensory spinules................. 2
</keyLead>
</paragraph>
</keyStep>
<keyStep id="309A2E3BA9062E71FF460A7DDBF5A5DD" pageId="17" pageNumber="300">
<paragraph id="8BD1369EA9062E71FF460A7DDBF2A510" blockId="17.[151,1436,1776,2033]" pageId="17" pageNumber="300">
<keyLead id="309F95ABA9062E71FF460A7DDBF2A510" pageId="17" pageNumber="300">
2. Posterior-lateral margins of head straight (
<figureCitation id="13552A1BA9062E71FDAE0A7DDCAAA535" box="[639,707,1864,1886]" captionStart="FIGURES 1 2" captionStartId="3.[151,264,1978,2001]" captionTargetBox="[281,1333,523,1944]" captionTargetId="figure@3.[253,1333,517,1961]" captionTargetPageId="3" captionText="FIGURES 1 2. Habitus photographs of Smilax Laporte. 1. Smilax deneinephyto Chatzimanolis, total length = 18.5 mm. 2. Smilax pilosa (F.), total length = 16.6 mm." httpUri="https://zenodo.org/record/164960/files/figure.png" pageId="17" pageNumber="300">Figs. 1</figureCitation>
,
<figureCitation id="13552A1BA9062E71FD000A7DDCB7A535" box="[721,734,1864,1886]" captionStart="FIGURES 5 8" captionStartId="5.[151,264,1518,1541]" captionTargetBox="[217,1369,452,1497]" captionTargetId="figure@5.[217,1369,445,1498]" captionTargetPageId="5" captionText="FIGURES 5 8. Head of Smilax Laporte. 5. Smilax deneinephyto Chatzimanolis. 6. Smilax kuntzeni (Scheerpeltz). 7. Smilax lynchi (Bruch). 8. Smilax pilosa (F.). Not to scale." httpUri="https://zenodo.org/record/268256/files/figure.png" pageId="17" pageNumber="300">5</figureCitation>
), not rounded; pronotum explanate (
<figureCitation id="13552A1BA9062E71FB960A7DDAE2A535" box="[1095,1163,1864,1886]" captionStart="FIGURES 1 2" captionStartId="3.[151,264,1978,2001]" captionTargetBox="[281,1333,523,1944]" captionTargetId="figure@3.[253,1333,517,1961]" captionTargetPageId="3" captionText="FIGURES 1 2. Habitus photographs of Smilax Laporte. 1. Smilax deneinephyto Chatzimanolis, total length = 18.5 mm. 2. Smilax pilosa (F.), total length = 16.6 mm." httpUri="https://zenodo.org/record/164960/files/figure.png" pageId="17" pageNumber="300">Figs. 1</figureCitation>
,
<figureCitation id="13552A1BA9062E71FB480A7DDACEA535" box="[1177,1191,1864,1886]" captionStart="FIGURES 9 12" captionStartId="6.[151,264,1220,1243]" captionTargetBox="[219,1369,447,1179]" captionTargetId="figure@6.[217,1369,447,1180]" captionTargetPageId="6" captionText="FIGURES 9 12. Pronotum of Smilax Laporte. 9. Smilax deneinephyto Chatzimanolis. 10. Smilax kuntzeni (Scheerpeltz). 11. Smilax lynchi (Bruch). 12. Smilax pilosa (F.). Not to scale." httpUri="https://zenodo.org/record/268257/files/figure.png" pageId="17" pageNumber="300">9</figureCitation>
); overall body shape not robust; aedeagus with paramere straight (
<figureCitation id="13552A1BA9062E71FDBD0A50DCD8A510" box="[620,689,1893,1915]" captionStart="FIGURES 24 25" captionStartId="9.[151,264,1619,1642]" captionTargetBox="[329,1249,195,1573]" captionTargetId="figure@9.[329,1249,195,1588]" captionTargetPageId="9" captionText="FIGURES 24 25. Aedeagus of S. deneinephyto Chatzimanolis. 24. Lateral view. 25. Dorsal view." httpUri="https://zenodo.org/record/164966/files/figure.png" pageId="17" pageNumber="300">Fig. 24</figureCitation>
)................................................
<taxonomicName id="4C6E4D1DA9062E71FAD20A53DBF2A510" ID-CoL="6YRVH" box="[1283,1435,1894,1915]" class="Insecta" family="Staphylinidae" genus="Smilax" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="300" phylum="Arthropoda" rank="species" species="deneinephyto">
<emphasis id="B91AEA8CA9062E71FAD20A53DBF2A510" box="[1283,1435,1894,1915]" italics="true" pageId="17" pageNumber="300">S. deneinephyto</emphasis>
</taxonomicName>
</keyLead>
</paragraph>
<paragraph id="8BD1369EA9062E71FF460AB1DBF5A5DD" blockId="17.[151,1436,1776,2033]" pageId="17" pageNumber="300">
<keyLead id="309F95ABA9062E71FF460AB1DBF5A5DD" pageId="17" pageNumber="300">
- Posterior-lateral margins of head rounded (
<figureCitation id="13552A1BA9062E71FD520AB6DCA1A5F2" box="[643,712,1923,1945]" captionStart="FIGURES 1 2" captionStartId="3.[151,264,1978,2001]" captionTargetBox="[281,1333,523,1944]" captionTargetId="figure@3.[253,1333,517,1961]" captionTargetPageId="3" captionText="FIGURES 1 2. Habitus photographs of Smilax Laporte. 1. Smilax deneinephyto Chatzimanolis, total length = 18.5 mm. 2. Smilax pilosa (F.), total length = 16.6 mm." httpUri="https://zenodo.org/record/164960/files/figure.png" pageId="17" pageNumber="300">Figs. 2</figureCitation>
,
<figureCitation id="13552A1BA9062E71FD050AB6DC8BA5F2" box="[724,738,1923,1945]" captionStart="FIGURES 3 4" captionStartId="4.[151,264,1738,1761]" captionTargetBox="[328,1248,185,1711]" captionTargetId="figure@4.[326,1261,181,1717]" captionTargetPageId="4" captionText="FIGURES 3 4. Habitus photographs of Smilax Laporte. 3. Smilax kuntzeni (Scheerpeltz), total length = 19.1 mm. 4. Smilax lynchi (Bruch), total length = 18.5 mm." httpUri="https://zenodo.org/record/268255/files/figure.png" pageId="17" pageNumber="300">4</figureCitation>
,
<figureCitation id="13552A1BA9062E71FD3C0AB6DD7DA5F2" box="[749,788,1923,1945]" captionStart="FIGURES 5 8" captionStartId="5.[151,264,1518,1541]" captionTargetBox="[217,1369,452,1497]" captionTargetId="figure@5.[217,1369,445,1498]" captionTargetPageId="5" captionText="FIGURES 5 8. Head of Smilax Laporte. 5. Smilax deneinephyto Chatzimanolis. 6. Smilax kuntzeni (Scheerpeltz). 7. Smilax lynchi (Bruch). 8. Smilax pilosa (F.). Not to scale." httpUri="https://zenodo.org/record/268256/files/figure.png" pageId="17" pageNumber="300">78</figureCitation>
); pronotum disc broadly and strongly concave (
<figureCitation id="13552A1BA9062E71FB360AB6DB44A5F2" box="[1255,1325,1923,1945]" captionStart="FIGURES 1 2" captionStartId="3.[151,264,1978,2001]" captionTargetBox="[281,1333,523,1944]" captionTargetId="figure@3.[253,1333,517,1961]" captionTargetPageId="3" captionText="FIGURES 1 2. Habitus photographs of Smilax Laporte. 1. Smilax deneinephyto Chatzimanolis, total length = 18.5 mm. 2. Smilax pilosa (F.), total length = 16.6 mm." httpUri="https://zenodo.org/record/164960/files/figure.png" pageId="17" pageNumber="300">Figs. 2</figureCitation>
,
<figureCitation id="13552A1BA9062E71FAE90AB6DB2FA5F2" box="[1336,1350,1923,1945]" captionStart="FIGURES 3 4" captionStartId="4.[151,264,1738,1761]" captionTargetBox="[328,1248,185,1711]" captionTargetId="figure@4.[326,1261,181,1717]" captionTargetPageId="4" captionText="FIGURES 3 4. Habitus photographs of Smilax Laporte. 3. Smilax kuntzeni (Scheerpeltz), total length = 19.1 mm. 4. Smilax lynchi (Bruch), total length = 18.5 mm." httpUri="https://zenodo.org/record/268255/files/figure.png" pageId="17" pageNumber="300">4</figureCitation>
,
<figureCitation id="13552A1BA9062E71FA800AB6DBF9A5F2" box="[1361,1424,1923,1945]" captionStart="FIGURES 9 12" captionStartId="6.[151,264,1220,1243]" captionTargetBox="[219,1369,447,1179]" captionTargetId="figure@6.[217,1369,447,1180]" captionTargetPageId="6" captionText="FIGURES 9 12. Pronotum of Smilax Laporte. 9. Smilax deneinephyto Chatzimanolis. 10. Smilax kuntzeni (Scheerpeltz). 11. Smilax lynchi (Bruch). 12. Smilax pilosa (F.). Not to scale." httpUri="https://zenodo.org/record/268257/files/figure.png" pageId="17" pageNumber="300">1112</figureCitation>
); overall body shape robust; aedeagus with paramere strongly to slightly concave (
<figureCitation id="13552A1BA9062E71FC390A95DA51A5DD" box="[1000,1080,1952,1974]" captionStart="FIGURES 29 30" captionStartId="12.[151,264,1997,2020]" captionTargetBox="[299,1285,409,1949]" captionTargetId="figure@12.[299,1285,409,1966]" captionTargetPageId="12" captionText="FIGURES 29 30. Aedeagus of S. lynchi (Bruch). 29. Lateral view. 30. Dorsal view." httpUri="https://zenodo.org/record/164968/files/figure.png" pageId="17" pageNumber="300">Figs. 29</figureCitation>
,
<figureCitation id="13552A1BA9062E71FB920A95DA35A5DD" box="[1091,1116,1952,1974]" captionStart="FIGURES 31 32" captionStartId="16.[151,264,1879,1902]" captionText="FIGURES 31 32. Aedeagus of S. pilosa (F.). 31. Lateral view. 32. Dorsal view." pageId="17" pageNumber="300">31</figureCitation>
)......................... 3
</keyLead>
</paragraph>
</keyStep>
<paragraph id="8BD1369EA9062E71FF460A88DBF2A59A" blockId="17.[151,1436,1776,2033]" pageId="17" pageNumber="300">
<keyLead id="309F95ABA9062E71FF460A88DBF2A59A" pageId="17" pageNumber="300">
<keyStep id="309A2E3BA9062E71FF460A88DBF2A59A" pageId="17" pageNumber="300">
3. Head with reddish-brown coloration (
<figureCitation id="13552A1BA9062E71FD9B0A88DCE7A5B8" box="[586,654,1981,2003]" captionStart="FIGURES 3 4" captionStartId="4.[151,264,1738,1761]" captionTargetBox="[328,1248,185,1711]" captionTargetId="figure@4.[326,1261,181,1717]" captionTargetPageId="4" captionText="FIGURES 3 4. Habitus photographs of Smilax Laporte. 3. Smilax kuntzeni (Scheerpeltz), total length = 19.1 mm. 4. Smilax lynchi (Bruch), total length = 18.5 mm." httpUri="https://zenodo.org/record/268255/files/figure.png" pageId="17" pageNumber="300">Figs. 4</figureCitation>
,
<figureCitation id="13552A1BA9062E71FD490A8BDCCFA5B8" box="[664,678,1982,2003]" captionStart="FIGURES 5 8" captionStartId="5.[151,264,1518,1541]" captionTargetBox="[217,1369,452,1497]" captionTargetId="figure@5.[217,1369,445,1498]" captionTargetPageId="5" captionText="FIGURES 5 8. Head of Smilax Laporte. 5. Smilax deneinephyto Chatzimanolis. 6. Smilax kuntzeni (Scheerpeltz). 7. Smilax lynchi (Bruch). 8. Smilax pilosa (F.). Not to scale." httpUri="https://zenodo.org/record/268256/files/figure.png" pageId="17" pageNumber="300">7</figureCitation>
); anterolateral and lateral sides of pronotum reddish-brown (
<figureCitation id="13552A1BA9062E71FB3C0A88DB5BA5B8" box="[1261,1330,1981,2003]" captionStart="FIGURES 3 4" captionStartId="4.[151,264,1738,1761]" captionTargetBox="[328,1248,185,1711]" captionTargetId="figure@4.[326,1261,181,1717]" captionTargetPageId="4" captionText="FIGURES 3 4. Habitus photographs of Smilax Laporte. 3. Smilax kuntzeni (Scheerpeltz), total length = 19.1 mm. 4. Smilax lynchi (Bruch), total length = 18.5 mm." httpUri="https://zenodo.org/record/268255/files/figure.png" pageId="17" pageNumber="300">Figs. 4</figureCitation>
,
<figureCitation id="13552A1BA9062E71FAED0A88DB3CA5B8" box="[1340,1365,1981,2003]" captionStart="FIGURES 9 12" captionStartId="6.[151,264,1220,1243]" captionTargetBox="[219,1369,447,1179]" captionTargetId="figure@6.[217,1369,447,1180]" captionTargetPageId="6" captionText="FIGURES 9 12. Pronotum of Smilax Laporte. 9. Smilax deneinephyto Chatzimanolis. 10. Smilax kuntzeni (Scheerpeltz). 11. Smilax lynchi (Bruch). 12. Smilax pilosa (F.). Not to scale." httpUri="https://zenodo.org/record/268257/files/figure.png" pageId="17" pageNumber="300">11</figureCitation>
); aedeagus with paramere strongly concave (
<figureCitation id="13552A1BA9062E71FD820AEEDCF1A59A" box="[595,664,2011,2033]" captionStart="FIGURES 29 30" captionStartId="12.[151,264,1997,2020]" captionTargetBox="[299,1285,409,1949]" captionTargetId="figure@12.[299,1285,409,1966]" captionTargetPageId="12" captionText="FIGURES 29 30. Aedeagus of S. lynchi (Bruch). 29. Lateral view. 30. Dorsal view." httpUri="https://zenodo.org/record/164968/files/figure.png" pageId="17" pageNumber="300">Fig. 29</figureCitation>
)........................................................
<taxonomicName id="4C6E4D1DA9062E71FA990AE9DBF2A59A" ID-CoL="4XWM3" box="[1352,1435,2012,2033]" class="Insecta" family="Staphylinidae" genus="Smilax" kingdom="Animalia" order="Coleoptera" pageId="17" pageNumber="300" phylum="Arthropoda" rank="species" species="lynchi">
<emphasis id="B91AEA8CA9062E71FA990AE9DBF2A59A" box="[1352,1435,2012,2033]" italics="true" pageId="17" pageNumber="300">S. lynchi</emphasis>
</taxonomicName>
</keyStep>
</keyLead>
</paragraph>
</subSubSection>
</treatment>
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