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<document id="3CBF94CFFE9CBA431A9B08AB8D3F8BA2" ID-CLB-Dataset="198863" ID-DOI="10.11646/zootaxa.5214.2.1" ID-GBIF-Dataset="14abb8a4-9e30-4d40-83b4-c725db688c72" ID-ISSN="1175-5326" ID-Zenodo-Dep="7384985" ID-ZooBank="8E7D1FC9-FD58-4DB3-ACA6-5AE38BDB6B0A" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1669872199474" checkinUser="plazi" docAuthor="Giłka, Wojciech &amp; Gadawski, Piotr" docDate="2022" docId="03C087D47F3DFFBBFF7EE52516407A2A" docLanguage="en" docName="zootaxa.5214.2.1.pdf" docOrigin="Zootaxa 5214 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Tanytarsus occultus Brundin 1949" docType="treatment" docVersion="7" lastPageNumber="161" masterDocId="FFF9FFAC7F37FFB1FFE9E46514417E1E" masterDocTitle="Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea" masterLastPageNumber="175" masterPageNumber="151" pageNumber="161" updateTime="1699412675151" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
<mods:mods id="F5D2DA72F5EB8925FFB17F9FECE002FE" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="25C0A45E16124FA59766E057AD96CCF8">
<mods:title id="0E62BDC47C6087073D1FCAA1B807A20C">Tanytarsini (Diptera: Chironomidae) collected during Polish expeditions to North Korea</mods:title>
</mods:titleInfo>
<mods:name id="D78F4CEE27BBE8A5891C57A0C73A084C" type="personal">
<mods:role id="F241C71E09F577D83D654F1D014C1562">
<mods:roleTerm id="A21289BFB123C5D612AD7E6641F4C7B4">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="80E48E89F79FAFA5731B25CD9C8ECF60">Giłka, Wojciech</mods:namePart>
<mods:nameIdentifier id="C6AF0971A971F1954B0F1F6FDF932F3B" type="ORCID">0000-0002-8403-5432</mods:nameIdentifier>
<mods:affiliation id="9DB6007F56432EE53A4876CD6FB55501">University of Gdańsk, Faculty of Biology, Department of Invertebrate Zoology and Parasitology, Laboratory of Systematic Zoology; Wita Stwosza 59, 80 - 308 Gdańsk, Poland. &amp; wojciech. gilka @ ug. edu. pl; https: // orcid. org / 0000 - 0002 - 8403 - 5432</mods:affiliation>
<mods:nameIdentifier id="D1A816E2972477869A0FA662A5102748" type="email">wojciech.gilka@ug.edu.pl</mods:nameIdentifier>
</mods:name>
<mods:name id="B668D5DBE7B6E915FF449541F12A0F08" type="personal">
<mods:role id="3992B762D294C38F1979BC5978A131B7">
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<mods:namePart id="15FCFB39AE3BED44DBAB6802BDDAFEAC">Gadawski, Piotr</mods:namePart>
<mods:nameIdentifier id="2AB74B3E56EE95BC29E1A49FC5996E4B" type="ORCID">0000-0001-9334-1960</mods:nameIdentifier>
<mods:affiliation id="FE2DBD827DBEF4C2388BBE5878D31615">University of Lódź, Faculty of Biology and Environmental Protection, Department of Invertebrate Zoology and Hydrobiology; Banacha 12 / 16, 90 - 237 Łódź, Poland. &amp; piotr. gadawski @ biol. uni. lodz. pl; https: // orcid. org / 0000 - 0001 - 9334 - 1960</mods:affiliation>
<mods:nameIdentifier id="5099259355CE232FD1DF40DF756B68E0" type="email">piotr.gadawski@biol.uni.lodz.pl</mods:nameIdentifier>
</mods:name>
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<mods:title id="448C8B6E2B09F1BEA72456F9F8C1B80C">Zootaxa</mods:title>
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<mods:date id="569C06C557A637FDF547C6D382AB7ECD">2022</mods:date>
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<mods:number id="82946600382BE8220EFCE7AA000EE062">2022-12-01</mods:number>
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<mods:detail id="0F777EC984A8EC6E88EC4B71DFCF3C3C" type="volume">
<mods:number id="398DF701B3E8406236B593898C5876D0">5214</mods:number>
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<mods:identifier id="50C17DC4E9D97FA3A1539FCF6D741242" type="CLB-Dataset">198863</mods:identifier>
<mods:identifier id="1DA600FDC2B542EB28C511E8F41A9629" type="DOI">10.11646/zootaxa.5214.2.1</mods:identifier>
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<treatment id="03C087D47F3DFFBBFF7EE52516407A2A" ID-DOI="http://doi.org/10.5281/zenodo.7386143" ID-GBIF-Taxon="205341219" ID-Zenodo-Dep="7386143" LSID="urn:lsid:plazi:treatment:03C087D47F3DFFBBFF7EE52516407A2A" httpUri="http://treatment.plazi.org/id/03C087D47F3DFFBBFF7EE52516407A2A" lastPageNumber="161" pageId="10" pageNumber="161">
<subSubSection id="C37365497F3DFFBBFF7EE525166E7F44" box="[151,559,319,346]" pageId="10" pageNumber="161" type="nomenclature">
<paragraph id="8BD636C27F3DFFBBFF7EE525166E7F44" blockId="10.[151,1437,319,469]" box="[151,559,319,346]" pageId="10" pageNumber="161">
<heading id="D09E81AE7F3DFFBBFF7EE525166E7F44" bold="true" box="[151,559,319,346]" fontSize="11" level="1" pageId="10" pageNumber="161" reason="1">
<taxonomicName id="4C694D417F3DFFBBFF7EE525166E7F44" ID-CoL="54QTV" authority="Brundin, 1949" authorityName="Brundin" authorityYear="1949" box="[151,559,319,346]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="occultus">
<emphasis id="B91DEAD07F3DFFBBFF7EE525166E7F44" bold="true" box="[151,559,319,346]" pageId="10" pageNumber="161">
<emphasis id="B91DEAD07F3DFFBBFF7EE525153C7F44" bold="true" box="[151,381,319,346]" italics="true" pageId="10" pageNumber="161">Tanytarsus occultus</emphasis>
<bibRefCitation id="EFF84B337F3DFFBBFE6DE525166E7F44" author="Brundin, L." box="[388,559,320,346]" pageId="10" pageNumber="161" pagination="1 - 915" refId="ref10404" refString="Brundin, L. (1949) Chironomiden und andere Bodentiere der suedschwedischen Urgebirgsseen. Ein Beitrag zur Kenntnis der bodenfaunistischen Charakterzuege schwedischer oligotropher Seen. Institute of Freshwater Research, Drottningholm, Lund, 30, 1 - 915." type="journal article" year="1949">Brundin, 1949</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="C37365497F3DFFBBFF7EE50411467FCA" pageId="10" pageNumber="161" type="reference_group">
<paragraph id="8BD636C27F3DFFBBFF7EE50417C67F66" blockId="10.[151,1437,319,469]" box="[151,903,352,377]" pageId="10" pageNumber="161">
<treatmentCitationGroup id="AB7911EC7F3DFFBBFF7EE50417C67F66" box="[151,903,352,377]" pageId="10" pageNumber="161">
<taxonomicName id="4C694D417F3DFFBBFF7EE50415827F66" ID-CoL="898FW" authority="Edw." authorityName="Edw." box="[151,451,352,376]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="holochlorus">
<emphasis id="B91DEAD07F3DFFBBFF7EE50415C47F66" box="[151,389,352,376]" italics="true" pageId="10" pageNumber="161">Tanytarsus holochlorus</emphasis>
Edw.
</taxonomicName>
:
<treatmentCitation id="0AC810D37F3DFFBBFE25E505163F7F66" author="Brundin, L." box="[460,638,352,377]" page="67" pageId="10" pageNumber="161" year="1947">
<bibRefCitation id="EFF84B337F3DFFBBFE25E505163F7F66" author="Brundin, L." box="[460,638,352,377]" pageId="10" pageNumber="161" pagination="1 - 95" refId="ref10381" refString="Brundin, L. (1947) Zur Kenntnis der Swedischen Chironomiden. Arkiv for Zoologi, 39, 1 - 95." type="journal article" year="1947">Brundin 1947: 67</bibRefCitation>
</treatmentCitation>
(male, fig. 100;
<collectingCountry id="F37E76527F3DFFBBFCCEE505173D7F66" box="[807,892,352,376]" name="Sweden" pageId="10" pageNumber="161">Sweden</collectingCountry>
).
</treatmentCitationGroup>
</paragraph>
<paragraph id="8BD636C27F3DFFBBFF7EE5E511467FCA" blockId="10.[151,1437,319,469]" pageId="10" pageNumber="161">
<treatmentCitationGroup id="AB7911EC7F3DFFBBFF7EE5E511467FCA" pageId="10" pageNumber="161">
<treatmentCitation id="0AC810D37F3DFFBBFF7EE5E516567F89" author="Brundin, L." box="[151,535,383,407]" page="847" pageId="10" pageNumber="161" year="1949">
<taxonomicName id="4C694D417F3DFFBBFF7EE5E516567F89" ID-CoL="54QTV" authority="Brundin, 1949: 847" authorityName="Brundin" authorityPageNumber="847" authorityYear="1949" box="[151,535,383,407]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="occultus">
<emphasis id="B91DEAD07F3DFFBBFF7EE5E515177F89" box="[151,342,383,407]" italics="true" pageId="10" pageNumber="161">Tanytarsus occultus</emphasis>
<bibRefCitation id="EFF84B337F3DFFBBFEB2E51A16567F89" author="Brundin, L." box="[347,535,383,407]" pageId="10" pageNumber="161" pagination="1 - 915" refId="ref10404" refString="Brundin, L. (1949) Chironomiden und andere Bodentiere der suedschwedischen Urgebirgsseen. Ein Beitrag zur Kenntnis der bodenfaunistischen Charakterzuege schwedischer oligotropher Seen. Institute of Freshwater Research, Drottningholm, Lund, 30, 1 - 915." type="journal article" year="1949">Brundin, 1949: 847</bibRefCitation>
</taxonomicName>
</treatmentCitation>
(male,
<collectingCountry id="F37E76527F3DFFBBFD89E51A16F17F89" box="[608,688,383,407]" name="Sweden" pageId="10" pageNumber="161">Sweden</collectingCountry>
);
<treatmentCitation id="0AC810D37F3DFFBBFD28E51A17F47F89" author="Reiss, F. &amp; Fittkau, E. J." box="[705,949,383,407]" page="89" pageId="10" pageNumber="161" year="1971">
<bibRefCitation id="EFF84B337F3DFFBBFD28E51A17F47F89" author="Reiss, F. &amp; Fittkau, E. J." box="[705,949,383,407]" pageId="10" pageNumber="161" pagination="75 - 200" refId="ref13257" refString="Reiss, F. &amp; Fittkau, E. J. (1971) Taxonomie und Okologie europaisch verbreiteter Tanytarsus-Arten (Chironomidae, Diptera). Archiv fur Hydrobiologie, supplement 40, 75 - 200." type="journal article" year="1971">Reiss &amp; Fittkau 1971: 89</bibRefCitation>
</treatmentCitation>
, 123 (male, fig. 35a–c, in key);
<bibRefCitation id="EFF84B337F3DFFBBFB01E51A11DC7F89" author="Ekrem, T. &amp; Sublette, M. F. &amp; Sublette, J. E." box="[1256,1437,383,407]" pageId="10" pageNumber="161" pagination="265 - 328" refId="ref10794" refString="Ekrem, T., Sublette, M. F. &amp; Sublette, J. E. (2003) North American Tanytarsus I. Descriptions and keys to species in the eminulus, gregarius, lugens and mendax species groups (Diptera: Chironomidae). Annals of the Entomological Society of America, 96, 265 - 328. https: // doi. org / 10.1603 / 0013 - 8746 (2003) 096 % 5 b 0265: NATIDA % 5 d 2.0. CO; 2" type="journal article" year="2003">
Ekrem
<emphasis id="B91DEAD07F3DFFBBFAC6E5E511217F89" box="[1327,1376,383,407]" italics="true" pageId="10" pageNumber="161">et al.</emphasis>
2003:
</bibRefCitation>
315 (male, pupa, figs 248–250; Europe, North America);
<bibRefCitation id="EFF84B337F3DFFBBFD18E5FB17DB7FA8" author="Ekrem, T." box="[753,922,414,438]" pageId="10" pageNumber="161" pagination="97 - 146" refId="ref10729" refString="Ekrem, T. (2004) Immature stages of European Tanytarsus species I. The eminulus -, gregarius -, lugens - and mendax species groups (Diptera, Chironomidae). Mitteilungen aus dem Museum fur Naturkunde in Berlin, Deutsche Entomologische Zeitschrift, 51, 97 - 146. https: // doi. org / 10.1002 / mmnd. 20040510110" type="journal article" year="2004">Ekrem 2004: 137</bibRefCitation>
(pupa, figs 183–187);
<bibRefCitation id="EFF84B337F3DFFBBFB9CE5FB11DD7FA8" author="Gilka, W. &amp; Paasivirta, L." box="[1141,1436,414,438]" pageId="10" pageNumber="161" pagination="107 - 113" refId="ref11665" refString="Gilka, W. &amp; Paasivirta, L. (2007) Two new species of the genus Tanytarsus van der Wulp (Diptera: Chironomidae) from Fennoscandia. In: Andersen, T. (Ed.), Contributions to the Systematics and Ecology of Aquatic Diptera - A Tribute to Ole A. Saether. The Caddis Press, Columbus, Ohio, pp. 107 - 113." type="book chapter" year="2007">Giłka &amp; Paasivirta 2007: 107–</bibRefCitation>
110 (male, figs 2–4, 13, 16; Fennoscandia);
<bibRefCitation id="EFF84B337F3DFFBBFD83E5D917287FCA" author="Gilka, W. &amp; Paasivirta, L. &amp; Gadawski, P. &amp; Grabowski, M." box="[618,873,444,469]" pageId="10" pageNumber="161" pagination="569 - 579" refId="ref11849" refString="Gilka, W., Paasivirta, L., Gadawski, P. &amp; Grabowski, M. (2018) Morphology and molecules say: Tanytarsus latens, sp. nov. from Finland (Diptera: Chironomidae). Zootaxa, 4471 (3), 569 - 579. https: // doi. org / 10.11646 / zootaxa. 4471.3.8" type="journal article" year="2018">
Giłka
<emphasis id="B91DEAD07F3DFFBBFD4CE5D816947FCA" box="[677,725,444,468]" italics="true" pageId="10" pageNumber="161">et al.</emphasis>
2018: 573–576
</bibRefCitation>
(male, diagnostics, barcoding, phylogeny).
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C37365497F3DFFBBFF7EE598110E7C09" box="[151,1359,509,536]" pageId="10" pageNumber="161" type="materials_examined">
<paragraph id="8BD636C27F3DFFBBFF7EE598110E7C09" blockId="10.[151,1437,509,1076]" box="[151,1359,509,536]" pageId="10" pageNumber="161">
<emphasis id="B91DEAD07F3DFFBBFF7EE59815C07C09" bold="true" box="[151,385,509,535]" pageId="10" pageNumber="161">Material examined.</emphasis>
Phjŏngjang,
<date id="FFD710027F3DFFBBFDFCE59816ED7C09" box="[533,684,509,536]" pageId="10" pageNumber="161" value="1981-06-13">13 June 1981</date>
,
<specimenCount id="9D6FFD4B7F3DFFBBFD51E59817537C09" box="[696,786,509,535]" pageId="10" pageNumber="161" type="male">2 males</specimenCount>
; Sokam,
<date id="FFD710027F3DFFBBFC90E59817BF7C09" box="[889,1022,509,535]" pageId="10" pageNumber="161" value="1981-07-08">8 July 1981</date>
,
<specimenCount id="9D6FFD4B7F3DFFBBFBE3E59810257C09" box="[1034,1124,509,535]" pageId="10" pageNumber="161" type="male">3 males</specimenCount>
; leg. W. Krzemiński.
</paragraph>
</subSubSection>
<subSubSection id="C37365497F3DFFBBFF2EE64416407A2A" pageId="10" pageNumber="161" type="discussion">
<paragraph id="8BD636C27F3DFFBBFF2EE64416407A2A" blockId="10.[151,1437,509,1076]" pageId="10" pageNumber="161">
<emphasis id="B91DEAD07F3DFFBBFF2EE644157A7C25" bold="true" box="[199,315,545,571]" pageId="10" pageNumber="161">Remarks.</emphasis>
A diagnostic character of the
<taxonomicName id="4C694D417F3DFFBBFD79E647172C7C25" authorityName="Brundin" authorityYear="1949" box="[656,877,545,571]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="occultus">
<emphasis id="B91DEAD07F3DFFBBFD79E647172C7C25" box="[656,877,545,571]" italics="true" pageId="10" pageNumber="161">Tanytarsus occultus</emphasis>
</taxonomicName>
male is the hypopygial anal point, usually with a square or concave apex, which the shape, however, may be affected by strong intraspecific variations.
<taxonomicName id="4C694D417F3DFFBBFACAE62311DD7C41" authorityName="Brundin" authorityYear="1949" box="[1315,1436,581,607]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="occultus">
<emphasis id="B91DEAD07F3DFFBBFACAE62311DD7C41" box="[1315,1436,581,607]" italics="true" pageId="10" pageNumber="161">T. occultus</emphasis>
</taxonomicName>
was described on the basis of a males’ series by
<bibRefCitation id="EFF84B337F3DFFBBFD58E60C17217C9D" author="Brundin, L." box="[689,864,617,643]" pageId="10" pageNumber="161" pagination="1 - 915" refId="ref10404" refString="Brundin, L. (1949) Chironomiden und andere Bodentiere der suedschwedischen Urgebirgsseen. Ein Beitrag zur Kenntnis der bodenfaunistischen Charakterzuege schwedischer oligotropher Seen. Institute of Freshwater Research, Drottningholm, Lund, 30, 1 - 915." type="journal article" year="1949">Brundin (1949)</bibRefCitation>
, who referred the description to his illustration of a specimen’s hypopygium presented two years earlier [
<bibRefCitation id="EFF84B337F3DFFBBFD35E6E817387CB9" author="Brundin, L." box="[732,889,653,679]" pageId="10" pageNumber="161" pagination="1 - 95" refId="ref10381" refString="Brundin, L. (1947) Zur Kenntnis der Swedischen Chironomiden. Arkiv for Zoologi, 39, 1 - 95." type="journal article" year="1947">Brundin 1947</bibRefCitation>
, originally erroneously as
<taxonomicName id="4C694D417F3DFFBBFB73E6EB11DD7CB9" box="[1178,1436,653,679]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="holochlorus">
<emphasis id="B91DEAD07F3DFFBBFB73E6EB11DD7CB9" box="[1178,1436,653,679]" italics="true" pageId="10" pageNumber="161">Tanytarsus holochlorus</emphasis>
</taxonomicName>
(=
<taxonomicName id="4C694D417F3DFFBBFF59E6D715827CD2" authority="Kieffer, 1925" authorityName="Kieffer" authorityYear="1925" box="[176,451,689,716]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="mendax">
<emphasis id="B91DEAD07F3DFFBBFF59E6D715637CD5" box="[176,290,689,715]" italics="true" pageId="10" pageNumber="161">T. mendax</emphasis>
Kieffer, 1925
</taxonomicName>
)]. The illustration was apparently based on a specimen with a hypopygial anal point of a structure rarely observed in
<taxonomicName id="4C694D417F3DFFBBFE25E6B316037CF1" authorityName="Brundin" authorityYear="1949" box="[460,578,725,751]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="occultus">
<emphasis id="B91DEAD07F3DFFBBFE25E6B316037CF1" box="[460,578,725,751]" italics="true" pageId="10" pageNumber="161">T. occultus</emphasis>
</taxonomicName>
. That choice could lead to confusion until
<bibRefCitation id="EFF84B337F3DFFBBFBF0E6B011577CF1" author="Reiss, F. &amp; Fittkau, E. J." box="[1049,1302,725,751]" pageId="10" pageNumber="161" pagination="75 - 200" refId="ref13257" refString="Reiss, F. &amp; Fittkau, E. J. (1971) Taxonomie und Okologie europaisch verbreiteter Tanytarsus-Arten (Chironomidae, Diptera). Archiv fur Hydrobiologie, supplement 40, 75 - 200." type="journal article" year="1971">Reiss &amp; Fittkau (1971)</bibRefCitation>
redescribed the male on the basis of Brundin’s series, indicating variations in the shape of the anal point, thus stabilizing the status of the species. Recently, one more close species was described,
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<emphasis id="B91DEAD07F3DFFBBFC48E77B10207D29" box="[929,1121,797,823]" italics="true" pageId="10" pageNumber="161">Tanytarsus latens</emphasis>
<bibRefCitation id="EFF84B337F3DFFBBFB80E77815317D45" author="Gilka, W. &amp; Paasivirta, L. &amp; Gadawski, P. &amp; Grabowski, M." pageId="10" pageNumber="161" pagination="569 - 579" refId="ref11849" refString="Gilka, W., Paasivirta, L., Gadawski, P. &amp; Grabowski, M. (2018) Morphology and molecules say: Tanytarsus latens, sp. nov. from Finland (Diptera: Chironomidae). Zootaxa, 4471 (3), 569 - 579. https: // doi. org / 10.11646 / zootaxa. 4471.3.8" type="journal article" year="2018">
Giłka, Paasivirta, Gadawski
<emphasis id="B91DEAD07F3DFFBBFF7EE72714EA7D45" box="[151,171,834,859]" italics="true" pageId="10" pageNumber="161">et</emphasis>
Grabowski, 2018
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, in which the anal point is similar or nearly identical (variable) to that illustrated from the atypical specimen of
<taxonomicName id="4C694D417F3DFFBBFECAE70315D87D61" authorityName="Brundin" authorityYear="1949" box="[291,409,869,895]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="occultus">
<emphasis id="B91DEAD07F3DFFBBFECAE70315D87D61" box="[291,409,869,895]" italics="true" pageId="10" pageNumber="161">T. occultus</emphasis>
</taxonomicName>
by
<bibRefCitation id="EFF84B337F3DFFBBFE2BE700162E7D61" author="Brundin, L." box="[450,623,869,895]" pageId="10" pageNumber="161" pagination="1 - 95" refId="ref10381" refString="Brundin, L. (1947) Zur Kenntnis der Swedischen Chironomiden. Arkiv for Zoologi, 39, 1 - 95." type="journal article" year="1947">Brundin (1947)</bibRefCitation>
. In case of these two species, the anal point structure can be thus critically misleading in their delimitation. The characters best separating males of the two species is the shape/length of the superior volsella and the digitus, and the arrangement of anal tergite bands (cf.
<bibRefCitation id="EFF84B337F3DFFBBFBE1E7C810867DD9" author="Gilka, W. &amp; Paasivirta, L. &amp; Gadawski, P. &amp; Grabowski, M." box="[1032,1223,941,967]" pageId="10" pageNumber="161" pagination="569 - 579" refId="ref11849" refString="Gilka, W., Paasivirta, L., Gadawski, P. &amp; Grabowski, M. (2018) Morphology and molecules say: Tanytarsus latens, sp. nov. from Finland (Diptera: Chironomidae). Zootaxa, 4471 (3), 569 - 579. https: // doi. org / 10.11646 / zootaxa. 4471.3.8" type="journal article" year="2018">
Giłka
<emphasis id="B91DEAD07F3DFFBBFBA4E7CB10C47DD9" box="[1101,1157,941,967]" italics="true" pageId="10" pageNumber="161">et al.</emphasis>
2018
</bibRefCitation>
).
<taxonomicName id="4C694D417F3DFFBBFB32E7CB11137DD9" authorityName="Brundin" authorityYear="1949" box="[1243,1362,941,967]" class="Insecta" family="Chironomidae" genus="Tanytarsus" kingdom="Animalia" order="Diptera" pageId="10" pageNumber="161" phylum="Arthropoda" rank="species" species="occultus">
<emphasis id="B91DEAD07F3DFFBBFB32E7CB11137DD9" box="[1243,1362,941,967]" italics="true" pageId="10" pageNumber="161">T. occultus</emphasis>
</taxonomicName>
is one of the best-defined species in the genus in term of morphological variability and diagnostics supported by molecular analyses (
<bibRefCitation id="EFF84B337F3DFFBBFEEDE79015B67A11" author="Reiss, F. &amp; Fittkau, E. J." box="[260,503,1013,1039]" pageId="10" pageNumber="161" pagination="75 - 200" refId="ref13257" refString="Reiss, F. &amp; Fittkau, E. J. (1971) Taxonomie und Okologie europaisch verbreiteter Tanytarsus-Arten (Chironomidae, Diptera). Archiv fur Hydrobiologie, supplement 40, 75 - 200." type="journal article" year="1971">Reiss &amp; Fittkau 1971</bibRefCitation>
;
<bibRefCitation id="EFF84B337F3DFFBBFDEAE790168F7A11" author="Ekrem, T. &amp; Sublette, M. F. &amp; Sublette, J. E." box="[515,718,1013,1039]" pageId="10" pageNumber="161" pagination="265 - 328" refId="ref10794" refString="Ekrem, T., Sublette, M. F. &amp; Sublette, J. E. (2003) North American Tanytarsus I. Descriptions and keys to species in the eminulus, gregarius, lugens and mendax species groups (Diptera: Chironomidae). Annals of the Entomological Society of America, 96, 265 - 328. https: // doi. org / 10.1603 / 0013 - 8746 (2003) 096 % 5 b 0265: NATIDA % 5 d 2.0. CO; 2" type="journal article" year="2003">
Ekrem
<emphasis id="B91DEAD07F3DFFBBFDBCE79316CC7A11" box="[597,653,1013,1039]" italics="true" pageId="10" pageNumber="161">et al.</emphasis>
2003
</bibRefCitation>
,
<bibRefCitation id="EFF84B337F3DFFBBFD32E79017267A11" author="Ekrem, T." box="[731,871,1013,1039]" pageId="10" pageNumber="161" pagination="97 - 146" refId="ref10729" refString="Ekrem, T. (2004) Immature stages of European Tanytarsus species I. The eminulus -, gregarius -, lugens - and mendax species groups (Diptera, Chironomidae). Mitteilungen aus dem Museum fur Naturkunde in Berlin, Deutsche Entomologische Zeitschrift, 51, 97 - 146. https: // doi. org / 10.1002 / mmnd. 20040510110" type="journal article" year="2004">Ekrem 2004</bibRefCitation>
;
<bibRefCitation id="EFF84B337F3DFFBBFC9DE79010C57A11" author="Gilka, W. &amp; Paasivirta, L." box="[884,1156,1013,1039]" pageId="10" pageNumber="161" pagination="107 - 113" refId="ref11665" refString="Gilka, W. &amp; Paasivirta, L. (2007) Two new species of the genus Tanytarsus van der Wulp (Diptera: Chironomidae) from Fennoscandia. In: Andersen, T. (Ed.), Contributions to the Systematics and Ecology of Aquatic Diptera - A Tribute to Ole A. Saether. The Caddis Press, Columbus, Ohio, pp. 107 - 113." type="book chapter" year="2007">Giłka &amp; Paasivirta 2007</bibRefCitation>
,
<bibRefCitation id="EFF84B337F3DFFBBFB78E79011107A11" author="Gilka, W. &amp; Paasivirta, L. &amp; Gadawski, P. &amp; Grabowski, M." box="[1169,1361,1013,1040]" pageId="10" pageNumber="161" pagination="569 - 579" refId="ref11849" refString="Gilka, W., Paasivirta, L., Gadawski, P. &amp; Grabowski, M. (2018) Morphology and molecules say: Tanytarsus latens, sp. nov. from Finland (Diptera: Chironomidae). Zootaxa, 4471 (3), 569 - 579. https: // doi. org / 10.11646 / zootaxa. 4471.3.8" type="journal article" year="2018">
Giłka
<emphasis id="B91DEAD07F3DFFBBFB3FE79311467A11" box="[1238,1287,1013,1039]" italics="true" pageId="10" pageNumber="161">et al</emphasis>
. 2018
</bibRefCitation>
). Only now recorded from
<collectingCountry id="F37E76527F3DFFBBFE99E07C15BC7A2A" box="[368,509,1049,1076]" name="North Korea" pageId="10" pageNumber="161">North Korea</collectingCountry>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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