267 lines
33 KiB
XML
267 lines
33 KiB
XML
<document ID-DOI="http://doi.org/10.5281/zenodo.3664826" ID-GBIF-Dataset="dd032848-7430-45d0-85d9-e528f25c83bb" ID-GBIF-Taxon="162054122" ID-Zenodo-Dep="3664826" checkinTime="1581022300054" checkinUser="plazi" docAuthor="Perrichot, Vincent, Wang, Bo & Barden, Phillip" docDate="2020" docId="03A3850A9F53FFAEFF87FF2E0D467EC7" docLanguage="en" docName="createceousResearch.109.104.381.pdf" docOrigin="Cretaceous Research 109" docStyle="DocumentStyle{}" docTitle="Dhagnathos autokrator Perrichot & Wang & Barden 2020, sp. nov." docType="treatment" docVersion="5" lastPageId="6" lastPageNumber="7" masterDocId="FF9AFD729F57FFA8FFF6FFD90E4F796E" masterDocTitle="New remarkable hell ants (Formicidae: Haidomyrmecinae stat. nov.) from mid-Cretaceous amber of northern Myanmar" masterLastPageNumber="18" masterPageNumber="1" pageId="4" pageNumber="5" updateTime="1643457689448" updateUser="ExternalLinkService">
|
||
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
|
||
<mods:titleInfo>
|
||
<mods:title>New remarkable hell ants (Formicidae: Haidomyrmecinae stat. nov.) from mid-Cretaceous amber of northern Myanmar</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Perrichot, Vincent</mods:namePart>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Wang, Bo</mods:namePart>
|
||
</mods:name>
|
||
<mods:name type="personal">
|
||
<mods:role>
|
||
<mods:roleTerm>Author</mods:roleTerm>
|
||
</mods:role>
|
||
<mods:namePart>Barden, Phillip</mods:namePart>
|
||
</mods:name>
|
||
<mods:typeOfResource>text</mods:typeOfResource>
|
||
<mods:relatedItem type="host">
|
||
<mods:titleInfo>
|
||
<mods:title>Cretaceous Research</mods:title>
|
||
</mods:titleInfo>
|
||
<mods:part>
|
||
<mods:date>2020</mods:date>
|
||
<mods:detail type="pubDate">
|
||
<mods:number>2020-01-10</mods:number>
|
||
</mods:detail>
|
||
<mods:detail type="volume">
|
||
<mods:number>109</mods:number>
|
||
</mods:detail>
|
||
<mods:extent unit="page">
|
||
<mods:start>1</mods:start>
|
||
<mods:end>18</mods:end>
|
||
</mods:extent>
|
||
</mods:part>
|
||
</mods:relatedItem>
|
||
<mods:classification>journal article</mods:classification>
|
||
<mods:identifier type="DOI">10.1016/j.cretres.2020.104381</mods:identifier>
|
||
<mods:identifier type="GBIF-Dataset">dd032848-7430-45d0-85d9-e528f25c83bb</mods:identifier>
|
||
<mods:identifier type="Zenodo-Dep">3653072</mods:identifier>
|
||
</mods:mods>
|
||
<treatment ID-DOI="http://doi.org/10.5281/zenodo.3664826" ID-GBIF-Taxon="162054122" ID-Zenodo-Dep="3664826" LSID="urn:lsid:plazi:treatment:03A3850A9F53FFAEFF87FF2E0D467EC7" httpUri="http://treatment.plazi.org/id/03A3850A9F53FFAEFF87FF2E0D467EC7" lastPageId="6" lastPageNumber="7" pageId="4" pageNumber="5">
|
||
<subSubSection box="[113,443,247,268]" pageId="4" pageNumber="5" type="nomenclature">
|
||
<paragraph blockId="4.[113,658,247,351]" box="[113,443,247,268]" pageId="4" pageNumber="5">
|
||
<taxonomicName authority="Perrichot & Wang & Barden, 2020" authorityName="Perrichot & Wang & Barden" authorityYear="2020" box="[113,359,247,267]" class="Insecta" family="Formicidae" genus="Dhagnathos" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="autokrator" status="sp. nov.">
|
||
<emphasis bold="true" box="[113,359,247,267]" italics="true" pageId="4" pageNumber="5">
|
||
Dhagnathos
|
||
<taxonomicName box="[244,359,247,267]" pageId="4" pageNumber="5" rank="species" species="autokrator" status="sp. nov.">autokrator</taxonomicName>
|
||
</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[366,443,249,268]" pageId="4" pageNumber="5" rank="species">sp. nov.</taxonomicNameLabel>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="4" pageNumber="5" type="multiple">
|
||
<paragraph blockId="4.[113,658,247,351]" pageId="4" pageNumber="5">
|
||
<uri pageId="4" pageNumber="5">
|
||
urn:lsid:zoobank.org:act:
|
||
<uuid pageId="4" pageNumber="5">
|
||
EC8760
|
||
<collectionCode box="[441,460,276,295]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="4" pageNumber="5" type="Herbarium">A</collectionCode>
|
||
9-9
|
||
<collectionCode box="[489,505,276,295]" country="Denmark" httpUri="http://grbio.org/cool/na0h-6vfh" name="University of Copenhagen" pageId="4" pageNumber="5" type="Herbarium">C</collectionCode>
|
||
00-44
|
||
<collectionCode box="[564,583,276,295]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="4" pageNumber="5" type="Herbarium">A</collectionCode>
|
||
9-9311- 2EC5
|
||
<collectionCode box="[164,196,304,323]" country="Hungary" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15158" name="Debrecen University" pageId="4" pageNumber="5" type="Herbarium">DE</collectionCode>
|
||
24
|
||
<collectionCode box="[219,236,304,323]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="4" pageNumber="5" type="Herbarium">A</collectionCode>
|
||
4
|
||
<collectionCode box="[247,261,304,323]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15534" name="Botanischer Garten und Botanisches Museum Berlin-Dahlem, Zentraleinrichtung der Freien Universitaet" pageId="4" pageNumber="5" type="Herbarium">B</collectionCode>
|
||
1
|
||
</uuid>
|
||
</uri>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[113,291,332,351]" pageId="4" pageNumber="5" type="description">
|
||
<paragraph blockId="4.[113,658,247,351]" box="[113,291,332,351]" pageId="4" pageNumber="5">
|
||
<figureCitation box="[113,222,332,351]" captionStart="Fig" captionStartId="5.[88,113,1850,1864]" captionTargetBox="[176,1386,178,1821]" captionTargetId="figure@5.[176,1386,178,1821]" captionTargetPageId="5" captionText="Fig. 2. Representative alate females of Dhagnathos autokrator gen. et sp. nov. (AeC, holotype IGR.BU-003), Chonidris insolita gen. et sp. nov. (DeF), and Aquilomyrmex huangi gen. et sp. nov. (GeH). Habitus (A), head in lateral (B) and frontal (C) views. Habitus of holotype NIGP171998 (D), head of paratype NIGP172003 (E) in anterolateral view, head of specimen HA04 (F) in frontal view. Habitus of holotype NIGP172000 (G), head of specimen TJ41-020 (H) in lateral view. Scale bars: 1 mm (A, D, G), 0.5 mm (B, C, H), 0.2 mm (E, F)." figureDoi="http://doi.org/10.5281/zenodo.3653076" httpUri="https://zenodo.org/record/3653076/files/figure.png" pageId="4" pageNumber="5">Figs. 2AeC</figureCitation>
|
||
,
|
||
<figureCitation box="[232,245,332,351]" captionStart="Fig" captionStartId="6.[113,138,1494,1508]" captionTargetBox="[202,1412,178,1465]" captionTargetId="figure@6.[202,1412,178,1465]" captionTargetPageId="6" captionText="Fig. 3. Overview and details of alate females (gynes) of Dhagnathos autokrator gen. et sp. nov. Specimen HA03 (A), specimen XA01 (B), specimen RM1 (CeD), and holotype IGR. BU- 003 (EeG). Habitus (A, B, C), fore wing (D), mandibles, clypeus and labrum in lateral views (E, F), wings, propodeum and petiole (G). Abbreviations: cc, lateral clypeal carina; cd, clypeal denticles; cp, clypeal process (horn); lss, labral spine-like setae; mib, mandibular medioventral blade; mp, mesal process of 1st gastral sternite; pdr, propodeal ridge; spp, sub-petiolar process. Scale bars: 2 mm (AeC), 1 mm (D, G), 0.5 mm (E, F)." figureDoi="http://doi.org/10.5281/zenodo.3653078" httpUri="https://zenodo.org/record/3653078/files/figure.png" pageId="4" pageNumber="5">3</figureCitation>
|
||
,
|
||
<figureCitation box="[257,287,332,351]" captionStart="Fig" captionStartId="14.[113,138,1215,1229]" captionTargetBox="[282,1334,179,1185]" captionTargetId="figure@14.[278,1337,178,1185]" captionTargetPageId="14" captionText="Fig. 8. Artistic reconstructions of the heads of hell ants in lateral view, with mandibles in closed position.A, Haidomyrmex. B, Protoceratomyrmex gen.nov.C,Aquilomyrmex gen.nov. D, Linguamyrmex. E, Ceratomyrmex. F, Chonidris gen. nov. G, Dhagnathos gen. nov. Scale bars: 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.3653088" httpUri="https://zenodo.org/record/3653088/files/figure.png" pageId="4" pageNumber="5">
|
||
8
|
||
<collectionCode box="[269,287,332,351]" country="Switzerland" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15706" name="Conservatoire et Jardin botaniques de la Ville de Gen<65>ve" pageId="4" pageNumber="5" type="Herbarium">G</collectionCode>
|
||
</figureCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="4" pageNumber="5" type="etymology">
|
||
<paragraph blockId="4.[113,783,373,784]" pageId="4" pageNumber="5">
|
||
<emphasis box="[113,214,373,393]" italics="true" pageId="4" pageNumber="5">Etymology</emphasis>
|
||
. The specific epithet refers to
|
||
<taxonomicName authorityName="Perrichot & Wang & Barden" authorityYear="2020" box="[510,613,373,393]" class="Insecta" family="Formicidae" genus="Dhagnathos" kingdom="Animalia" order="Hymenoptera" pageId="4" pageNumber="5" phylum="Arthropoda" rank="species" species="autokrator">
|
||
<emphasis box="[510,613,373,393]" italics="true" pageId="4" pageNumber="5">autokrator</emphasis>
|
||
</taxonomicName>
|
||
(Greek, meaning ‘self-ruler’), an individual who exercises absolute power, unrestrained by superiors; in reference to the highly powerful aspect of this ant.
|
||
</paragraph>
|
||
<paragraph blockId="4.[113,783,373,784]" box="[113,668,484,505]" pageId="4" pageNumber="5">
|
||
<materialsCitation ID-GBIF-Occurrence="2564234954" box="[113,664,484,505]" collectionCode="IGR" pageId="4" pageNumber="5" specimenCode="BU-003" specimenCount="1" specimenCount-female="1" typeStatus="holotype">
|
||
<emphasis box="[113,200,484,504]" italics="true" pageId="4" pageNumber="5">
|
||
<typeStatus box="[113,200,484,504]" pageId="4" pageNumber="5">Holotype</typeStatus>
|
||
</emphasis>
|
||
.
|
||
<collectionCode box="[212,254,486,505]" collectionName="IGR" pageId="4" pageNumber="5">IGR</collectionCode>
|
||
.
|
||
<specimenCode box="[256,331,486,505]" pageId="4" pageNumber="5">BU-003</specimenCode>
|
||
, alate
|
||
<specimenCount box="[396,464,485,504]" pageId="4" pageNumber="5" type="female">female</specimenCount>
|
||
(
|
||
<figureCitation box="[478,587,485,505]" captionStart="Fig" captionStartId="5.[88,113,1850,1864]" captionTargetBox="[176,1386,178,1821]" captionTargetId="figure@5.[176,1386,178,1821]" captionTargetPageId="5" captionText="Fig. 2. Representative alate females of Dhagnathos autokrator gen. et sp. nov. (AeC, holotype IGR.BU-003), Chonidris insolita gen. et sp. nov. (DeF), and Aquilomyrmex huangi gen. et sp. nov. (GeH). Habitus (A), head in lateral (B) and frontal (C) views. Habitus of holotype NIGP171998 (D), head of paratype NIGP172003 (E) in anterolateral view, head of specimen HA04 (F) in frontal view. Habitus of holotype NIGP172000 (G), head of specimen TJ41-020 (H) in lateral view. Scale bars: 1 mm (A, D, G), 0.5 mm (B, C, H), 0.2 mm (E, F)." figureDoi="http://doi.org/10.5281/zenodo.3653076" httpUri="https://zenodo.org/record/3653076/files/figure.png" pageId="4" pageNumber="5">Figs. 2AeC</figureCitation>
|
||
,
|
||
<figureCitation box="[597,657,485,505]" captionStart="Fig" captionStartId="6.[113,138,1494,1508]" captionTargetBox="[202,1412,178,1465]" captionTargetId="figure@6.[202,1412,178,1465]" captionTargetPageId="6" captionText="Fig. 3. Overview and details of alate females (gynes) of Dhagnathos autokrator gen. et sp. nov. Specimen HA03 (A), specimen XA01 (B), specimen RM1 (CeD), and holotype IGR. BU- 003 (EeG). Habitus (A, B, C), fore wing (D), mandibles, clypeus and labrum in lateral views (E, F), wings, propodeum and petiole (G). Abbreviations: cc, lateral clypeal carina; cd, clypeal denticles; cp, clypeal process (horn); lss, labral spine-like setae; mib, mandibular medioventral blade; mp, mesal process of 1st gastral sternite; pdr, propodeal ridge; spp, sub-petiolar process. Scale bars: 2 mm (AeC), 1 mm (D, G), 0.5 mm (E, F)." figureDoi="http://doi.org/10.5281/zenodo.3653078" httpUri="https://zenodo.org/record/3653078/files/figure.png" pageId="4" pageNumber="5">3EeG</figureCitation>
|
||
)
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="4" pageNumber="5" type="materials_examined">
|
||
<paragraph blockId="4.[113,783,373,784]" pageId="4" pageNumber="5">
|
||
<emphasis box="[113,323,512,532]" italics="true" pageId="4" pageNumber="5">Additional specimens</emphasis>
|
||
.
|
||
<collectionCode box="[338,373,513,532]" country="Ecuador" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14290" name="Universidad del Azuay" pageId="4" pageNumber="5" type="Herbarium">HA</collectionCode>
|
||
03, XA01 and
|
||
<collectionCode box="[522,561,513,532]" country="Canada" httpUri="http://grbio.org/cool/0a5t-r74w" name="McGill University, Redpath Museum" pageId="4" pageNumber="5" type="Museum">RM</collectionCode>
|
||
1, three alate females (
|
||
<figureCitation box="[121,246,541,560]" captionStart="Fig" captionStartId="6.[113,138,1494,1508]" captionTargetBox="[202,1412,178,1465]" captionTargetId="figure@6.[202,1412,178,1465]" captionTargetPageId="6" captionText="Fig. 3. Overview and details of alate females (gynes) of Dhagnathos autokrator gen. et sp. nov. Specimen HA03 (A), specimen XA01 (B), specimen RM1 (CeD), and holotype IGR. BU- 003 (EeG). Habitus (A, B, C), fore wing (D), mandibles, clypeus and labrum in lateral views (E, F), wings, propodeum and petiole (G). Abbreviations: cc, lateral clypeal carina; cd, clypeal denticles; cp, clypeal process (horn); lss, labral spine-like setae; mib, mandibular medioventral blade; mp, mesal process of 1st gastral sternite; pdr, propodeal ridge; spp, sub-petiolar process. Scale bars: 2 mm (AeC), 1 mm (D, G), 0.5 mm (E, F)." figureDoi="http://doi.org/10.5281/zenodo.3653078" httpUri="https://zenodo.org/record/3653078/files/figure.png" pageId="4" pageNumber="5">Figs. 3Ae3D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph blockId="4.[113,783,373,784]" pageId="4" pageNumber="5">
|
||
<emphasis box="[113,319,568,588]" italics="true" pageId="4" pageNumber="5">Horizon and locality.</emphasis>
|
||
Upper Cretaceous, upper Albianelower Cenomanian (ca. 99 Ma); in amber from the Hukawng Valley,
|
||
<collectingRegion country="Myanmar" name="Kachin" pageId="4" pageNumber="5">Kachin State</collectingRegion>
|
||
,
|
||
<collectingCountry box="[175,273,625,644]" name="Myanmar" pageId="4" pageNumber="5">Myanmar</collectingCountry>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection box="[113,533,652,672]" pageId="4" pageNumber="5" type="diagnosis">
|
||
<paragraph blockId="4.[113,783,373,784]" box="[113,533,652,672]" pageId="4" pageNumber="5">
|
||
<emphasis box="[113,206,652,672]" italics="true" pageId="4" pageNumber="5">Diagnosis</emphasis>
|
||
. As for the genus, by monotypy.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection lastPageId="6" lastPageNumber="7" pageId="4" pageNumber="5" type="description">
|
||
<paragraph blockId="4.[113,783,373,784]" pageId="4" pageNumber="5">
|
||
<emphasis box="[113,223,679,699]" italics="true" pageId="4" pageNumber="5">Description</emphasis>
|
||
(gyne). Body length ca.
|
||
<quantity box="[507,588,681,700]" metricMagnitude="-2" metricUnit="m" metricValue="1.4" pageId="4" pageNumber="5" unit="mm" value="14.0">14 mm</quantity>
|
||
. Cuticle generally smooth, without distinct sculpturing, sparsely covered by thin, long, erect setae, the head additionally densely covered by short, adpressed setae on vertex and genae.
|
||
</paragraph>
|
||
<paragraph blockId="4.[113,784,820,1983]" lastBlockId="4.[831,1501,179,1984]" pageId="4" pageNumber="5">
|
||
Head only slightly longer than high and wide. Vertex and posteroventral surface rounded, anterior surface relatively flat, and genae shorter than eyes and projecting anteroventrally above mandible insertion into a cheek-like lobe. Ocelli present near top of vertex, conspicuous, ocellar diameter slightly larger than width of first antennomere; interocellar distance about half of ocellar diameter. Compound eyes bulging, reniform, 2.4 as long as wide, situated posteriorly on head (
|
||
<collectionCode box="[408,442,1016,1035]" pageId="4" pageNumber="5">EPI</collectionCode>
|
||
440). Antennae inserted between compound eyes around their midlength, closely flanking lateral edges of clypeus; base of antenna with basal bulb exposed, inserted within thick annular torulus. Antenna geniculate, filiform; scape short, 0.5 head length, weakly arched and broadened apically; first funicular article (pedicel) very short, 0.22 scape length, less than twice as long as wide, broadened apically; flagellomeres unusually slender, funicular article II (antennomere
|
||
<collectionCode box="[610,632,1211,1230]" pageId="4" pageNumber="5">III</collectionCode>
|
||
) about 22 as long as wide; following antennomeres gradually decreasing in length and width. Posterior clypeal margin apparently fused, while horn is the result of
|
||
<emphasis box="[321,397,1293,1313]" italics="true" pageId="4" pageNumber="5">de novo</emphasis>
|
||
medial margin/ridge; anterior clypeal margin broadly rounded. Clypeal horn directed upward for its basal quarter, then bent at a right angle and directed forward for remaining length; horn gently rounded apically, without expanded lobe; dorsal surface of horn convex; ventral surface emarginate, its lateral margins prominent and prolonged basally into raised frontal carinae diverging anteriorly to reach the anterior margin of head, just above insertion of mandibles. Setation of horn consisting, on ventral surface, of a dense brush of short, peg-like denticles at apex; similar peg-like denticles widely spaced and arranged in a single row on each lateral margin, and becoming progressively denser and arranged in 2e3 longitudinal rows along lateral clypeal carinae; dorsal and ventral surfaces of horn sparsely covered by thin, long, erect setae. Labrum well exposed, large, nearly trapezoid, with anterior margin convex, posterior margin slightly emarginate medially, sides unsutured to clypeus so that anterior part of labrum is apparently movable; dorsal surface of labrum rimmed laterally by a longitudinal brush of stiff, spine-like setae, also densely coated by thin, erect setae becoming progressively longer and stiffer along lateral and posterior margins. Mandibles long (
|
||
<collectionCode box="[607,652,1824,1843]" country="Malaysia" httpUri="http://grbio.org/cool/nf70-xac8" name="Malaysian Agricultural Research and Development Institute (MARDI)" pageId="4" pageNumber="5" type="Herbarium">MDI</collectionCode>
|
||
97), scytheshaped, widely spaced basally and converging apically, with tips curved and acute, nearly reaching the rounded portion of the horn as preserved; basal portion linear, short; apical portion 5 as long as basal portion, curved dorsally and posteriorly, with dorsal surface concave and rimmed on each margin by row of acute teeth and thin, erect setae directed backwards; medioventral blade between basal and apical portions forming a large, isosceles, blunt tooth perpendicular to apical portion. Palps long (visible on specimen
|
||
<collectionCode box="[831,867,263,282]" country="Ecuador" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14290" name="Universidad del Azuay" pageId="4" pageNumber="5" type="Herbarium">HA</collectionCode>
|
||
03), coated dorsally in fine, tapered setae, maxillary palp with 6 segments, as long as head capsule when combined; labial palp with 5 segments.
|
||
</paragraph>
|
||
<paragraph blockId="4.[831,1501,179,1984]" pageId="4" pageNumber="5">
|
||
Mesosoma. Pronotal colar pronounced, concealing propleuron in dorsal view, separated from remaining pronotal dorsum by a distinct transverse ridge; pronotal dorsum strongly concave immediately anterior to ridge, nearly flat posterior to ridge; promesonotal suture deeply impressed. Mesoscutum as long as pronotum (excluding neck) in dorsal view, about as broad as long; mesoscutal dorsal outline feebly convex, with long parapsidal furrows almost reaching anterior mesonotal margin, converging posteriorly but not touching. Mesoscutellum posteriorly expanded, in dorsal view concealing median portion of metanotum; dorsal and posterior mesoscutellar surfaces concave, their junction forming a sharp angle; dorsal mesoscutellar surface with a deep, broad, transverse groove immediately posterior to scuto-scutellar suture. Metanotum medially as high as long, with posterior surface forming distinct angle with pronotal dorsum. Propodeum 1.25 as high as long, dorsal and declivitous surfaces meet at pronounced right angle, forming conspicuous ridge; dorsal surface nearly flat, declivitous surface faintly concave; propodeal spiracle slit-like, opening posteriad, at junction of propodeal dorsum and sides; metapleural gland orifice opening laterally, protected by guard setae. Legs long and robust (mostly visible on specimen
|
||
<collectionCode box="[831,868,933,952]" country="Ecuador" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14290" name="Universidad del Azuay" pageId="4" pageNumber="5" type="Herbarium">HA</collectionCode>
|
||
03); mesocoxa distinctly shorter than pro-and metacoxae; small trochantellus present on mid- and hind legs; all femora distinctly swollen in their basal half, tibiae swollen in their apical half; ventral margin of protibia apically with large calcar gently curved, protibia possessing small subapical point, and two straight, stout setae less than half as long as calcar; mesotibia apically with two long, straight, pectinate spurs, and two short, stout setae; metatibia apically with one long, pectinate spur and one long, simple spur; tarsomeres IeIV of all legs with pairs of stout setae along entire ventral surface (8e10 pairs on tI, 4e5 pairs on tII, 3 pairs on tIII, 2 pairs on tIV), and apically with 2 pairs of stout setae each flanking a spatulate spine; additionally the ventral surface of tarsomeres IeIV covered by dense brush of thin, erect setae; pretarsal claws strong, with a distinct subapical tooth.
|
||
</paragraph>
|
||
<paragraph blockId="4.[831,1501,179,1984]" pageId="4" pageNumber="5">
|
||
Fore wing with Rs∙f2, basal portion of Rs∙f3,
|
||
<collectionCode box="[1347,1367,1323,1342]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung M<>nchen" pageId="4" pageNumber="5" type="Herbarium">M</collectionCode>
|
||
∙f4, and Cu1 nebulous, all other veins tubular; pterostigma elongate, ca. 6 as long as broad; a short stub of cross-vein 1r-rs present, nebulous; Rs∙f1 half as long as
|
||
<collectionCode box="[1043,1063,1407,1426]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung M<>nchen" pageId="4" pageNumber="5" type="Herbarium">M</collectionCode>
|
||
∙f1, both distinctly arched; Rs∙f2 and Rs∙f3 nearly at right angle, Rs∙f2 half as long as
|
||
<collectionCode box="[1281,1301,1435,1454]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung M<>nchen" pageId="4" pageNumber="5" type="Herbarium">M</collectionCode>
|
||
∙f2; 2rs-m present, situated beyond apex of pterostigma; discal and subdiscal cells pentagonal; cu-a arising from
|
||
<collectionCode box="[1137,1157,1491,1510]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung M<>nchen" pageId="4" pageNumber="5" type="Herbarium">M</collectionCode>
|
||
<emphasis bold="true" box="[1158,1175,1490,1509]" pageId="4" pageNumber="5">þ</emphasis>
|
||
Cu and proximal to
|
||
<collectionCode box="[1375,1395,1491,1510]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung M<>nchen" pageId="4" pageNumber="5" type="Herbarium">M</collectionCode>
|
||
∙f1 (Cu∙f1 short); vein Cu with both Cu1 and Cu2 present. Hind wing with jugal lobe present; anterior margin with 5 median and 22 distal hamuli; vein
|
||
<collectionCode box="[979,992,1574,1593]" country="Denmark" httpUri="http://grbio.org/cool/na0h-6vfh" name="University of Copenhagen" pageId="4" pageNumber="5" type="Herbarium">C</collectionCode>
|
||
present; vein
|
||
<collectionCode box="[1159,1173,1574,1593]" country="Chile" httpUri="http://grbio.org/cool/az7k-p4g9" name="Departamento de Geologia, Universidad de Chile" pageId="4" pageNumber="5">R</collectionCode>
|
||
present, reaching distal wing margin; Rs∙f1 more than twice as long as 1rs-m; cu-a arising from
|
||
<collectionCode box="[831,851,1630,1649]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15637" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="4" pageNumber="5" type="Herbarium">M</collectionCode>
|
||
<emphasis bold="true" box="[851,868,1630,1649]" pageId="4" pageNumber="5">þ</emphasis>
|
||
Cu, proximal to fork of
|
||
<collectionCode box="[1098,1118,1630,1649]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15637" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="4" pageNumber="5" type="Herbarium">M</collectionCode>
|
||
∙f1 and Cu (Cu∙f1 short); Rs∙f2,
|
||
<collectionCode box="[1443,1463,1630,1649]" country="Germany" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15637" lsid="urn:lsid:biocol.org:col:15637" name="Botanische Staatssammlung München" pageId="4" pageNumber="5" type="Herbarium">M</collectionCode>
|
||
∙f2, Cu, and
|
||
<collectionCode box="[913,927,1658,1677]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15406" lsid="urn:lsid:biocol.org:col:15406" name="Harvard University - Arnold Arboretum" pageId="4" pageNumber="5" type="Herbarium">A</collectionCode>
|
||
∙f2 present, not reaching wing margin.
|
||
</paragraph>
|
||
<paragraph blockId="4.[831,1501,179,1984]" lastBlockId="6.[113,784,1635,1794]" lastPageId="6" lastPageNumber="7" pageId="4" pageNumber="5">
|
||
Metasoma with petiole short-pedunculate, almost 0.6 as high as long; petiolar tergite a broadly convex node, with anterior surface approximately twice as long as posterior face; subpetiolar process present, in profile forming a high, transverse, lamella pointing ventrally, with anterior face concave, posterior face vertical; not fused tergosternally, suture visible; attaching broadly to gaster. Gaster elongate. First gastral tergite with helcium pronounced, forming a post-petiolar peduncle, with anterior surface behind helcium high, oblique, and dorsal surface strongly convex, short; anteriormost part of first gastral sternite with a distinct mesal process (keel) pointing anteroventrally below helcium. Second gastral segment distinctly longer than first, with presclerite largely exposed to form a deep, broad constriction between first and second gastral segments (abdominal segments
|
||
<collectionCode box="[667,687,1692,1711]" pageId="6" pageNumber="7">III</collectionCode>
|
||
and IV). Gastral segments unfused with deep lateral suture. Following segments poorly preserved, pygidium apparently broadly acute towards sting shaft.
|
||
</paragraph>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.3653076" ID-Zenodo-Dep="3653076" httpUri="https://zenodo.org/record/3653076/files/figure.png" pageId="5" pageNumber="6" startId="5.[88,113,1850,1864]" targetBox="[176,1386,178,1821]" targetPageId="5">
|
||
<paragraph blockId="5.[88,1474,1850,1910]" pageId="5" pageNumber="6">
|
||
<emphasis bold="true" box="[88,135,1850,1864]" pageId="5" pageNumber="6">Fig. 2.</emphasis>
|
||
Representative alate females of
|
||
<taxonomicName authority="Perrichot & Wang & Barden, 2020" authorityName="Perrichot & Wang & Barden" authorityYear="2020" box="[389,562,1850,1865]" class="Insecta" family="Formicidae" genus="Dhagnathos" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="autokrator" status="gen. et sp. nov.">
|
||
<emphasis box="[389,562,1850,1865]" italics="true" pageId="5" pageNumber="6">Dhagnathos autokrator</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[566,682,1851,1865]" pageId="5" pageNumber="6" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
(AeC, holotype IGR.BU-003),
|
||
<taxonomicName authority="Perrichot & Wang & Barden, 2020" authorityName="Perrichot & Wang & Barden" authorityYear="2020" box="[912,1042,1850,1865]" class="Insecta" family="Formicidae" genus="Chonidris" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="insolita" status="gen. et sp. nov.">
|
||
<emphasis box="[912,1042,1850,1865]" italics="true" pageId="5" pageNumber="6">Chonidris insolita</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[1046,1162,1851,1865]" pageId="5" pageNumber="6" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
(DeF), and
|
||
<taxonomicName authority="Perrichot & Wang & Barden, 2020" authorityName="Perrichot & Wang & Barden" authorityYear="2020" box="[1253,1417,1850,1865]" class="Insecta" family="Formicidae" genus="Aquilomyrmex" kingdom="Animalia" order="Hymenoptera" pageId="5" pageNumber="6" phylum="Arthropoda" rank="species" species="huangi" status="gen. et sp. nov.">
|
||
<emphasis box="[1253,1417,1850,1865]" italics="true" pageId="5" pageNumber="6">Aquilomyrmex huangi</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel pageId="5" pageNumber="6" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
(GeH). Habitus (A), head in lateral (B) and frontal (C) views. Habitus of holotype NIGP171998 (D), head of paratype NIGP172003 (E) in anterolateral view, head of specimen HA04 (F) in frontal view. Habitus of holotype NIGP172000 (G), head of specimen TJ41-020 (H) in lateral view. Scale bars: 1 mm (A, D, G), 0.5 mm (B, C, H), 0.2 mm (E, F).
|
||
</paragraph>
|
||
</caption>
|
||
<caption ID-DOI="http://doi.org/10.5281/zenodo.3653078" ID-Zenodo-Dep="3653078" httpUri="https://zenodo.org/record/3653078/files/figure.png" pageId="6" pageNumber="7" startId="6.[113,138,1494,1508]" targetBox="[202,1412,178,1465]" targetPageId="6">
|
||
<paragraph blockId="6.[113,1501,1494,1577]" pageId="6" pageNumber="7">
|
||
<emphasis bold="true" box="[113,161,1494,1508]" pageId="6" pageNumber="7">Fig. 3.</emphasis>
|
||
Overview and details of alate females (gynes) of
|
||
<taxonomicName authority="Perrichot & Wang & Barden, 2020" authorityName="Perrichot & Wang & Barden" authorityYear="2020" box="[545,718,1494,1509]" class="Insecta" family="Formicidae" genus="Dhagnathos" kingdom="Animalia" order="Hymenoptera" pageId="6" pageNumber="7" phylum="Arthropoda" rank="species" species="autokrator" status="gen. et sp. nov.">
|
||
<emphasis box="[545,718,1494,1509]" italics="true" pageId="6" pageNumber="7">Dhagnathos autokrator</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel box="[722,839,1495,1509]" pageId="6" pageNumber="7" rank="species">gen. et sp. nov.</taxonomicNameLabel>
|
||
Specimen HA03 (A), specimen XA01 (B), specimen RM1 (CeD), and holotype IGR.BU- 003 (EeG). Habitus (A, B, C), fore wing (D), mandibles, clypeus and labrum in lateral views (E, F), wings, propodeum and petiole (G). Abbreviations: cc, lateral clypeal carina; cd, clypeal denticles; cp, clypeal process (horn); lss, labral spine-like setae; mib, mandibular medioventral blade; mp, mesal process of 1st gastral sternite; pdr, propodeal ridge; spp, sub-petiolar process. Scale bars: 2 mm (AeC), 1 mm (D, G), 0.5 mm (E, F).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph blockId="6.[113,783,1830,1962]" pageId="6" pageNumber="7">
|
||
<emphasis box="[113,254,1830,1850]" italics="true" pageId="6" pageNumber="7">Measurements</emphasis>
|
||
(
|
||
<typeStatus box="[266,356,1831,1850]" pageId="6" pageNumber="7">holotype</typeStatus>
|
||
<collectionCode box="[361,403,1831,1850]" pageId="6" pageNumber="7">IGR</collectionCode>
|
||
.BU-003; in mm).
|
||
<collectionCode box="[573,600,1831,1850]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13245" lsid="urn:lsid:biocol.org:col:13245" name="Houghton Lake Wildlife Research Station" pageId="6" pageNumber="7" type="Herbarium">HL</collectionCode>
|
||
2.50; HoL ca.1.70; EL 1.20; ocelli diameter 0.20; MDbL 0.40, MDtL 0.55, MDaL 2.00; length/width of antennomeres:
|
||
<collectionCode box="[434,441,1887,1906]" country="Romania" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14415" lsid="urn:lsid:biocol.org:col:14415" name="&quot;Alexandru Ioan Cuza&quot; University" pageId="6" pageNumber="7" type="Herbarium">I</collectionCode>
|
||
(scape) 1.15/0.16, II (pedicel) 0.26/ 0.14, III 2.16/0.10, IV 1.50/0.07,
|
||
<collectionCode box="[424,438,1915,1934]" country="Canada" httpUri="http://biocol.org/urn:lsid:biocol.org:col:13946" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="6" pageNumber="7" type="Museum">V</collectionCode>
|
||
1.34/0.07;
|
||
<collectionCode box="[553,587,1915,1934]" country="China" httpUri="http://biocol.org/urn:lsid:biocol.org:col:14766" lsid="urn:lsid:biocol.org:col:14766" name="Wolong Nature Reserve" pageId="6" pageNumber="7" type="Herbarium">WL</collectionCode>
|
||
3.85; FWL (as preserved) 6.35 (7.90 on specimen
|
||
<collectionCode box="[435,488,1943,1962]" pageId="6" pageNumber="7">DHA</collectionCode>
|
||
4);
|
||
<collectionCode box="[516,539,1943,1962]" country="Czech Republic" httpUri="http://grbio.org/cool/7jyr-w2tp" name="Západoceské muzeum v Plzni" pageId="6" pageNumber="7" type="Museum">PL</collectionCode>
|
||
1.84,
|
||
<collectingCountry box="[599,628,1943,1962]" name="Philippines" pageId="6" pageNumber="7">
|
||
<collectionCode box="[599,628,1943,1962]" country="USA" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15723" lsid="urn:lsid:biocol.org:col:15723" name="Academy of Natural Sciences, Botany Department" pageId="6" pageNumber="7" type="Herbarium">PH</collectionCode>
|
||
</collectingCountry>
|
||
1.00,
|
||
<collectionCode box="[688,723,1943,1962]" country="Thailand" httpUri="http://grbio.org/cool/vkzx-f5mv" name="Paleontological Collections" pageId="6" pageNumber="7">PW</collectionCode>
|
||
0.67.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |