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<document id="E38BB52F7EF1D34BF4D8078D30BF736C" ID-DOI="10.11646/zootaxa.4790.1.1" ID-ISSN="1175-5326" ID-Zenodo-Dep="3887775" ID-ZooBank="AA12710D-D4FB-4437-ABD5-D85373CEDF73" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1591774854251" checkinUser="plazi" docAuthor="Serri, Sayeh, Bologna, Marco A. &amp; Riccieri, Alessandra" docDate="2020" docId="039F878DFFC3FF83FF76FE90FAB814CC" docLanguage="en" docName="zootaxa.4790.1.1.pdf" docOrigin="Zootaxa 4790 (1)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Hycleus pirata Bologna &amp; Turco 2007" docType="treatment" docVersion="5" lastPageNumber="24" masterDocId="FFA6FFF5FFD5FF94FFE1FFB8FFDE1667" masterDocTitle="Revision of the Hycleus sexmaculatus species group (Coleoptera: Meloidae Mylabrini)" masterLastPageNumber="42" masterPageNumber="1" pageNumber="23" updateTime="1698773399993" updateUser="plazi" zenodo-license-document="CLOSED">
<mods:mods id="8D1F81E0ACA0D6F0C11DFB64A4ABBF04" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="C32F3193F45147811B548E0CE2DAB1A6">
<mods:title id="A19E364D828075727F75780F6FC8147A">Revision of the Hycleus sexmaculatus species group (Coleoptera: Meloidae Mylabrini)</mods:title>
</mods:titleInfo>
<mods:name id="E7B5517388F17165814209EE53A3774E" type="personal">
<mods:role id="D5C777D01AD4488BCAF38B4B0E4F70C3">
<mods:roleTerm id="039E5DE3595BE03002B9BF1FF3DB621E">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="14C222744FD170D628399A9E21F0DCB7">Serri, Sayeh</mods:namePart>
<mods:affiliation id="73C801FE3624D7AB1BA9E3EBA87604EF">Hayk Mirzayans Insect Museum (HMIM), Iranian Research Institute of Plant Protection, Agricultural Research, Education and Extension</mods:affiliation>
</mods:name>
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<mods:role id="C634F6E431CBA4B3D2E4DFBD04CA2823">
<mods:roleTerm id="762849DD1F314EE209715AE346D52F21">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DFB403FDB81764E4330E5ABF00567473">Bologna, Marco A.</mods:namePart>
<mods:affiliation id="5C462AB7F6A62EB08B41FAD40FFA7899">Dipartimento di Scienze, Università Roma Tre, Viale G. Marconi 446, 00146 Roma, Italy, alessandra. riccieri @ uniroma 3. it https: // orcid. org / 0000 - 0003 - 3719 - 9941. &amp; Corresponding Author, marcoalberto. bologna @ uniroma 3. it https: // orcid. org / 0000 - 0003 - 2498 - 8917</mods:affiliation>
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<mods:roleTerm id="E6CFF7BE8C7E5CBF7D457477E17D399B">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="DEC52B7C099282A36C3D8B22C886C353">Riccieri, Alessandra</mods:namePart>
<mods:affiliation id="A4A591DA30F510CC2A7FF2935EA78A47">Dipartimento di Scienze, Università Roma Tre, Viale G. Marconi 446, 00146 Roma, Italy, alessandra. riccieri @ uniroma 3. it https: // orcid. org / 0000 - 0003 - 3719 - 9941.</mods:affiliation>
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<mods:typeOfResource id="2BFF184446F2B2C213120087E6FBEB30">text</mods:typeOfResource>
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<mods:titleInfo id="FE5D667F17EBAC4764E9E52252959611">
<mods:title id="26D933C34227CEAD78A79DFB29C5D895">Zootaxa</mods:title>
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<mods:part id="8B860177CDD8F7D67731965F4F887659">
<mods:date id="656FE4C9880864C05C327B6082BCAB58">2020</mods:date>
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<mods:number id="2C20354C2F3D8A91F257BD91A3557854">2020-06-10</mods:number>
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<mods:number id="497FDD3B307E257BF7D90D3F27DDBF99">4790</mods:number>
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<mods:number id="6EF1C1801BF72AFD3D23A948FA23C86D">1</mods:number>
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<mods:identifier id="3FB438B7C34799CDFF4FC743743E72C2" type="ISSN">1175-5326</mods:identifier>
<mods:identifier id="BB5D0796D5005F47F063313EDB4BF7F4" type="Zenodo-Dep">3887775</mods:identifier>
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<treatment id="039F878DFFC3FF83FF76FE90FAB814CC" ID-DOI="http://doi.org/10.5281/zenodo.5103757" ID-Zenodo-Dep="5103757" LSID="urn:lsid:plazi:treatment:039F878DFFC3FF83FF76FE90FAB814CC" httpUri="http://treatment.plazi.org/id/039F878DFFC3FF83FF76FE90FAB814CC" lastPageId="23" lastPageNumber="24" pageId="22" pageNumber="23">
<subSubSection id="C32C6510FFC3FF82FF76FE90FE851700" pageId="22" pageNumber="23" type="nomenclature">
<paragraph id="8B89369BFFC3FF82FF76FE90FDAE1725" blockId="22.[151,624,296,359]" box="[151,624,296,323]" pageId="22" pageNumber="23">
<heading id="D0C181F7FFC3FF82FF76FE90FDAE1725" bold="true" box="[151,624,296,323]" fontSize="11" level="1" pageId="22" pageNumber="23" reason="1">
<taxonomicName id="4C364D18FFC3FF82FF76FE90FDAE1725" authority="Bologna &amp; Turco, 2007" authorityName="Bologna &amp; Turco" authorityYear="2007" box="[151,624,296,323]" class="Insecta" family="Meloidae" genus="Hycleus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="pirata">
<emphasis id="B942EA89FFC3FF82FF76FE90FDAE1725" bold="true" box="[151,624,296,323]" pageId="22" pageNumber="23">
<emphasis id="B942EA89FFC3FF82FF76FE90FE961724" bold="true" box="[151,328,296,323]" italics="true" pageId="22" pageNumber="23">Hycleus pirata</emphasis>
<bibRefCitation id="EFA74B6AFFC3FF82FEAEFE91FDAE1725" author="Bologna, M. A. &amp; Turco, F." box="[335,624,296,323]" pageId="22" pageNumber="23" pagination="1 - 33" refId="ref32438" refString="Bologna, M. A. &amp; Turco, F. (2007) The Meloidae (Coleoptera) of the United Arab Emirates with an updated Arabian checklist. Zootaxa, 1625 (1), 1 - 33. https: // doi. org / 10.11646 / zootaxa. 1625.1.1" type="journal article" year="2007">Bologna &amp; Turco, 2007</bibRefCitation>
</emphasis>
</taxonomicName>
</heading>
</paragraph>
<paragraph id="8B89369BFFC3FF82FF76FEF4FE851700" blockId="22.[151,624,296,359]" box="[151,347,332,359]" pageId="22" pageNumber="23">Figs 8, 20, 77–83</paragraph>
</subSubSection>
<subSubSection id="C32C6510FFC3FF82FF76FE29FD8F17CE" box="[151,593,400,425]" pageId="22" pageNumber="23" type="reference_group">
<paragraph id="8B89369BFFC3FF82FF76FE29FD8F17CE" blockId="22.[151,593,400,425]" box="[151,593,400,425]" pageId="22" pageNumber="23">
<treatmentCitationGroup id="AB2611B5FFC3FF82FF76FE29FD8F17CE" box="[151,593,400,425]" pageId="22" pageNumber="23">
<treatmentCitation id="0A97108AFFC3FF82FF76FE29FD9317CE" author="Bologna, M. A. &amp; Turco, F." box="[151,589,400,425]" httpUri="http://treatment.plazi.org/id/03FB4051D648DC1D229FFEC4B37365B0" page="16" pageId="22" pageNumber="23" year="2007">
<taxonomicName id="4C364D18FFC3FF82FF76FE29FD9317CE" authority="Bologna &amp; Turco, 2007: 16" authorityName="Bologna &amp; Turco" authorityPageNumber="16" authorityYear="2007" box="[151,589,400,425]" class="Insecta" family="Meloidae" genus="Hycleus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="pirata">
<emphasis id="B942EA89FFC3FF82FF76FE29FEF317CF" box="[151,301,401,425]" italics="true" pageId="22" pageNumber="23">Hycleus pirata</emphasis>
<bibRefCitation id="EFA74B6AFFC3FF82FED2FE28FD9317CE" author="Bologna, M. A. &amp; Turco, F." box="[307,589,400,425]" pageId="22" pageNumber="23" pagination="1 - 33" refId="ref32438" refString="Bologna, M. A. &amp; Turco, F. (2007) The Meloidae (Coleoptera) of the United Arab Emirates with an updated Arabian checklist. Zootaxa, 1625 (1), 1 - 33. https: // doi. org / 10.11646 / zootaxa. 1625.1.1" type="journal article" year="2007">Bologna &amp; Turco, 2007: 16</bibRefCitation>
</taxonomicName>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="C32C6510FFC3FF82FF76FE6FFC4111A5" pageId="22" pageNumber="23" type="materials_examined">
<paragraph id="8B89369BFFC3FF82FF76FE6FFD4614E6" blockId="22.[151,1436,470,642]" pageId="22" pageNumber="23">
<emphasis id="B942EA89FFC3FF82FF76FE6FFE9A1796" bold="true" box="[151,324,470,497]" pageId="22" pageNumber="23">Type material.</emphasis>
<materialsCitation id="3B5E3CC6FFC3FF82FEABFE6FFD4614E6" collectingDate="2005-03-12" collectingDateMax="2005-03-15" collectingDateMin="2005-03-12" collectionCode="MABC, HMIM" collectorName="M. Bologna &amp; F. Turco" country="United Arab Emirates" elevation="195" latitude="24.20491" location="Al Bateen" longLatPrecision="1" longitude="55.61263" pageId="22" pageNumber="23" specimenCount="12" specimenCount-female="6" specimenCount-male="6" stateProvince="Abu Zaby" typeStatus="holotype">
<typeStatus id="548D8839FFC3FF82FEABFE6FFE6C1796" box="[330,434,471,497]" pageId="22" pageNumber="23" type="holotype">Holotype</typeStatus>
<specimenCount id="9D30FD12FFC3FF82FE58FE60FE101795" box="[441,462,472,498]" pageId="22" pageNumber="23" type="male">♂</specimenCount>
and
<specimenCount id="9D30FD12FFC3FF82FDE2FE6FFDF21795" box="[515,556,471,498]" pageId="22" pageNumber="23" type="female">2 ♀</specimenCount>
<typeStatus id="548D8839FFC3FF82FDD3FE6FFD7A1796" box="[562,676,471,497]" pageId="22" pageNumber="23" type="paratype">paratypes</typeStatus>
:”
<collectingCountry id="F321760BFFC3FF82FD59FE6FFC711796" box="[696,943,471,497]" name="United Arab Emirates" pageId="22" pageNumber="23">United Arab Emirates</collectingCountry>
:
<collectingRegion id="49F2F879FFC3FF82FC56FE6FFB4B1796" box="[951,1173,471,497]" country="United Arab Emirates" name="Abu Zaby" pageId="22" pageNumber="23">Abu Dhabi Emirate</collectingRegion>
, EA4,
<location id="8EE96040FFC3FF82FB3FFE6FFA901795" LSID="urn:lsid:plazi:treatment:039F878DFFC3FF83FF76FE90FAB814CC:8EE96040FFC3FF82FB3FFE6FFA901795" box="[1246,1358,471,498]" country="United Arab Emirates" latitude="24.20491" longLatPrecision="1" longitude="55.61263" name="Al Bateen" pageId="22" pageNumber="23" stateProvince="Abu Zaby">Al Bateen</location>
dunes, 4 Km W Al Ain on the road to
<collectingRegion id="49F2F879FFC3FF82FE12FE43FDAC1472" box="[499,626,507,533]" country="United Arab Emirates" name="Abu Zaby" pageId="22" pageNumber="23">Abu Dhabi</collectingRegion>
,
<elevation id="001BD1A8FFC3FF82FD9FFE42FCDF1472" box="[638,769,506,534]" metricMagnitude="2" metricUnit="m" metricValue="1.95" pageId="22" pageNumber="23" unit="m" value="195.0">
<quantity id="4CCE9B7EFFC3FF82FD9FFE42FD1B1471" box="[638,709,506,534]" metricMagnitude="2" metricUnit="m" metricValue="1.95" pageId="22" pageNumber="23" unit="m" value="195.0">195 m</quantity>
a.s.l.
</elevation>
/
<geoCoordinate id="EE02505CFFC3FF82FCF6FE43FC701471" box="[791,942,507,534]" degrees="24, 20491" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="1" value="24.20491">24, 20491° N</geoCoordinate>
<geoCoordinate id="EE02505CFFC3FF82FC57FE42FB921471" box="[950,1100,506,534]" degrees="55, 61263" direction="east" orientation="longitude" pageId="22" pageNumber="23" precision="1" value="55.61263">55, 61263° E</geoCoordinate>
,
<date id="FF88105BFFC3FF82FBB8FE42FB071472" box="[1113,1241,506,533]" pageId="22" pageNumber="23" value="2005-03-15">
<collectingDate id="EFCCE9B3FFC3FF82FBB8FE42FB071472" box="[1113,1241,506,533]" pageId="22" pageNumber="23" value="2005-03-15">15.III.2005</collectingDate>
</date>
, leg.
<collectorName id="26C3534DFFC3FF82FAF7FE44FA421472" box="[1302,1436,507,534]" pageId="22" pageNumber="23">M. Bologna</collectorName>
&amp;
<collectorName id="26C3534DFFC3FF82FF56FD98FEC9145D" box="[183,279,544,570]" pageId="22" pageNumber="23">F. Turco</collectorName>
(
<collectionCode id="ED27AE5EFFC3FF82FEC8FDA7FE5A145E" box="[297,388,543,569]" pageId="22" pageNumber="23">MABC</collectionCode>
)”;
<specimenCount id="9D30FD12FFC3FF82FE45FDA7FE0F145D" box="[420,465,543,570]" pageId="22" pageNumber="23" type="male">4 ♂</specimenCount>
<specimenCount id="9D30FD12FFC3FF82FE3AFDA7FDD6145D" box="[475,520,543,570]" pageId="22" pageNumber="23" type="female">4 ♀</specimenCount>
<typeStatus id="548D8839FFC3FF82FDF3FD98FDA3145D" box="[530,637,544,570]" pageId="22" pageNumber="23" type="paratype">paratypes</typeStatus>
(
<collectionCode id="ED27AE5EFFC3FF82FD6EFDA7FD39145E" box="[655,743,543,569]" pageId="22" pageNumber="23">MABC</collectionCode>
),
<specimenCount id="9D30FD12FFC3FF82FD1CFDA7FCF4145D" box="[765,810,543,570]" pageId="22" pageNumber="23" type="male">1 ♂</specimenCount>
<typeStatus id="548D8839FFC3FF82FCD5FD98FC4A145D" box="[820,916,544,570]" pageId="22" pageNumber="23" type="paratype">paratype</typeStatus>
(
<collectionCode id="ED27AE5EFFC3FF82FC46FDA7FC23145E" box="[935,1021,543,569]" country="Spain" name="Jardí Botànic Marimurtra" pageId="22" pageNumber="23" type="Herbarium">HMIM</collectionCode>
): “United Arab Emirate,
<collectingRegion id="49F2F879FFC3FF82FAFFFDA7FF2D143A" country="United Arab Emirates" name="Abu Zaby" pageId="22" pageNumber="23">Abu Dhabi Emirate</collectingRegion>
, EA10, Al Aïn, 3–4 Km E/ Al Selimat on road Al Aïn-Abu Dhabi,
<geoCoordinate id="EE02505CFFC3FF82FC1EFDFBFB441439" box="[1023,1178,579,606]" degrees="24, 19263" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="1" value="24.19263">24, 19263° N</geoCoordinate>
<geoCoordinate id="EE02505CFFC3FF82FB42FDFAFAE31439" box="[1187,1341,578,606]" degrees="55, 58861" direction="east" orientation="longitude" pageId="22" pageNumber="23" precision="1" value="55.58861">55, 58861° E</geoCoordinate>
,
<quantity id="4CCE9B7EFFC3FF82FAADFDFBFA491439" box="[1356,1431,579,606]" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="22" pageNumber="23" unit="m" value="220.0">
<elevation id="001BD1A8FFC3FF82FAADFDFBFA491439" box="[1356,1431,579,606]" metricMagnitude="2" metricUnit="m" metricValue="2.2" pageId="22" pageNumber="23" unit="m" value="220.0">220 m</elevation>
</quantity>
,
<date id="FF88105BFFC3FF82FF76FDDEFE9F14E6" box="[151,321,614,641]" pageId="22" pageNumber="23" value="2005-03-12" valueMax="2005-03-14" valueMin="2005-03-12">
<collectingDate id="EFCCE9B3FFC3FF82FF76FDDEFE9F14E6" box="[151,321,614,641]" pageId="22" pageNumber="23" value="2005-03-12" valueMax="2005-03-14" valueMin="2005-03-12">12–14.III.2005</collectingDate>
</date>
, leg.
<collectorName id="26C3534DFFC3FF82FE9CFDD0FDDD14E6" box="[381,515,615,642]" pageId="22" pageNumber="23">M. Bologna</collectorName>
&amp;
<collectorName id="26C3534DFFC3FF82FDC6FDD0FD5414E6" box="[551,650,615,642]" pageId="22" pageNumber="23">F. Turco</collectorName>
”.
</materialsCitation>
</paragraph>
<caption id="DF496613FFC3FF82FF76F9A4FB70103F" ID-DOI="http://doi.org/10.5281/zenodo.3887797" ID-Zenodo-Dep="3887797" httpUri="https://zenodo.org/record/3887797/files/figure.png" pageId="22" pageNumber="23" startId="22.[151,264,1564,1589]" targetBox="[181,1404,667,1533]" targetPageId="22">
<paragraph id="8B89369BFFC3FF82FF76F9A4FB70103F" blockId="22.[151,1437,1564,1625]" pageId="22" pageNumber="23">
<emphasis id="B942EA89FFC3FF82FF76F9A4FE861052" bold="true" box="[151,344,1564,1589]" pageId="22" pageNumber="23">FIGURES 77–83.</emphasis>
<taxonomicName id="4C364D18FFC3FF82FE80F9A5FD1E1053" authority="Bologna &amp; Turco" authorityName="Bologna &amp; Turco" authorityYear="2007" box="[353,704,1564,1589]" class="Insecta" family="Meloidae" genus="Hycleus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="pirata">
<emphasis id="B942EA89FFC3FF82FE80F9A5FE241052" box="[353,506,1565,1589]" italics="true" pageId="22" pageNumber="23">Hycleus pirata</emphasis>
Bologna &amp; Turco
</taxonomicName>
: 77. male antenna; 78. female antenna; 79. tegmen, ventral view; 80. tegmen, lateral view; 81. aedeagus, lateral view; 82. mesosternum; 83. elytral pattern. Scale bar: 1 mm.
</paragraph>
</caption>
<paragraph id="8B89369BFFC3FF82FF26F930FD8110A2" blockId="22.[151,1437,1671,1986]" pageId="22" pageNumber="23">
<materialsCitation id="3B5E3CC6FFC3FF82FF26F930FD8510A2" country="United Arab Emirates" location="Abu Dhabi Emirate" pageId="22" pageNumber="23" specimenCount="1" stateProvince="Abu Zaby" typeStatus="holotype">
<typeStatus id="548D8839FFC3FF82FF26F930FF2110C5" box="[199,255,1672,1698]" pageId="22" pageNumber="23">Type</typeStatus>
locality: “
<collectingCountry id="F321760BFFC3FF82FE93F93FFDB010C6" box="[370,622,1671,1697]" name="United Arab Emirates" pageId="22" pageNumber="23">United Arab Emirates</collectingCountry>
:
<collectingRegion id="49F2F879FFC3FF82FD97F93FFC8B10C6" box="[630,853,1671,1697]" country="United Arab Emirates" name="Abu Zaby" pageId="22" pageNumber="23">Abu Dhabi Emirate</collectingRegion>
, EA4, Al Bateen dunes, 4 Km W Al Ain on the road to
<collectingRegion id="49F2F879FFC3FF82FF53F913FEEF10A2" box="[178,305,1707,1733]" country="United Arab Emirates" name="Abu Zaby" pageId="22" pageNumber="23">Abu Dhabi</collectingRegion>
” (
<bibRefCitation id="EFA74B6AFFC3FF82FEA8F913FD8D10A2" author="Bologna, M. A. &amp; Turco, F." box="[329,595,1707,1734]" pageId="22" pageNumber="23" pagination="1 - 33" refId="ref32438" refString="Bologna, M. A. &amp; Turco, F. (2007) The Meloidae (Coleoptera) of the United Arab Emirates with an updated Arabian checklist. Zootaxa, 1625 (1), 1 - 33. https: // doi. org / 10.11646 / zootaxa. 1625.1.1" type="journal article" year="2007">Bologna &amp; Turco 2007</bibRefCitation>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B89369BFFC3FF82FF26F976FB2D108E" blockId="22.[151,1437,1671,1986]" box="[199,1267,1742,1770]" pageId="22" pageNumber="23">
<emphasis id="B942EA89FFC3FF82FF26F976FDEA108E" bold="true" box="[199,564,1742,1769]" pageId="22" pageNumber="23">Additional material examined.</emphasis>
<materialsCitation id="3B5E3CC6FFC3FF82FDDAF977FC89108E" box="[571,855,1742,1770]" collectionCode="V" country="Oman" location="Wadi Air" pageId="22" pageNumber="23" specimenCount="1" specimenCount-female="1">
<specimenCount id="9D30FD12FFC3FF82FDDAF977FDBB108D" box="[571,613,1743,1770]" pageId="22" pageNumber="23" type="female">1 ♀</specimenCount>
<collectingCountry id="F321760BFFC3FF82FD8DF977FD6D108E" box="[620,691,1743,1769]" name="Oman" pageId="22" pageNumber="23">Oman</collectingCountry>
,
<location id="8EE96040FFC3FF82FD5FF977FCFB108E" LSID="urn:lsid:plazi:treatment:039F878DFFC3FF83FF76FE90FAB814CC:8EE96040FFC3FF82FD5FF977FCFB108E" box="[702,805,1743,1769]" country="Oman" name="Wadi Air" pageId="22" pageNumber="23">Wadi Air</location>
,
<date id="FF88105BFFC3FF82FCD1F976FC89108E" box="[816,855,1742,1769]" pageId="22" pageNumber="23" value="1957-05-07">
7.
<collectionCode id="ED27AE5EFFC3FF82FCA0F976FC89108E" box="[833,855,1742,1769]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="22" pageNumber="23" type="Museum">V</collectionCode>
</date>
</materialsCitation>
<materialsCitation id="3B5E3CC6FFC3FF82FCB9F976FB31108E" box="[856,1263,1742,1770]" collectionCode="MABC" collectorName="Popov &amp; Wood" country="Oman" pageId="22" pageNumber="23" specimenCount="1">
.1957, leg.
<collectorName id="26C3534DFFC3FF82FC32F968FBC5108D" box="[979,1051,1744,1770]" pageId="22" pageNumber="23">Popov</collectorName>
&amp;
<collectorName id="26C3534DFFC3FF82FBDFF977FB5E108E" box="[1086,1152,1743,1769]" pageId="22" pageNumber="23">Wood</collectorName>
(
<collectionCode id="ED27AE5EFFC3FF82FB6EF977FB36108E" box="[1167,1256,1743,1769]" pageId="22" pageNumber="23">MABC</collectionCode>
)
</materialsCitation>
.
</paragraph>
<paragraph id="8B89369BFFC3FF82FF26F94BFC4111A5" blockId="22.[151,1437,1671,1986]" pageId="22" pageNumber="23">
<emphasis id="B942EA89FFC3FF82FF26F94BFD9F116A" bold="true" box="[199,577,1778,1805]" pageId="22" pageNumber="23">
Records from literature.
<collectingCountry id="F321760BFFC3FF82FE0EF94AFDE2116A" box="[495,572,1778,1805]" name="Oman" pageId="22" pageNumber="23">Oman</collectingCountry>
.
</emphasis>
Oman (
<bibRefCitation id="EFA74B6AFFC3FF82FD7BF94BFC48116A" author="Kocak, A. O. &amp; Kemal, M." box="[666,918,1778,1805]" pageId="22" pageNumber="23" pagination="1 - 150" refId="ref34929" refString="Kocak, A. O. &amp; Kemal, M. (2015) Initial list of the Pterygota species of Malatya Prov., based upon the info-system of the Cesa (East Turkey). Cesa News, 115, 1 - 150." type="journal article" year="2015">Koçak &amp; Kemal 2015</bibRefCitation>
).
<emphasis id="B942EA89FFC3FF82FC49F94AFC34116A" bold="true" box="[936,1002,1778,1805]" pageId="22" pageNumber="23">
<collectingCountry id="F321760BFFC3FF82FC49F94AFC3D116A" box="[936,995,1778,1805]" name="United Arab Emirates" pageId="22" pageNumber="23">UAE</collectingCountry>
.
</emphasis>
UAE (
<bibRefCitation id="EFA74B6AFFC3FF82FBD9F94BFAFC116A" author="Gillet, M. P. T. &amp; Gillet, C." box="[1080,1314,1778,1805]" pageId="22" pageNumber="23" pagination="169 - 196" refId="ref33890" refString="Gillet, M. P. T. &amp; Gillet, C. (2005) Insects and other arthropods. In: Hellyer, P. &amp; Aspinall, S. (Eds.), The Emirates. A Natural History. Trident Press Limited, Abu Dhabi, pp. 169 - 196." type="book chapter" year="2005">Gillet &amp; Gillet, 2005</bibRefCitation>
as
<emphasis id="B942EA89FFC3FF82FAA6F94BFF121156" italics="true" pageId="22" pageNumber="23">
<taxonomicName id="4C364D18FFC3FF82FAA6F94BFF1A1156" authorityName="Olivier" authorityYear="1811" class="Insecta" family="Meloidae" genus="Mylabris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="bipunctata">bipunctata</taxonomicName>
;
</emphasis>
<bibRefCitation id="EFA74B6AFFC3FF82FF35F8AFFE111156" author="Kocak, A. O. &amp; Kemal, M." box="[212,463,1814,1841]" pageId="22" pageNumber="23" pagination="1 - 150" refId="ref34929" refString="Kocak, A. O. &amp; Kemal, M. (2015) Initial list of the Pterygota species of Malatya Prov., based upon the info-system of the Cesa (East Turkey). Cesa News, 115, 1 - 150." type="journal article" year="2015">Koçak &amp; Kemal 2015</bibRefCitation>
;
<bibRefCitation id="EFA74B6AFFC3FF82FE3DF8AFFC881156" author="Ghahari, H. &amp; Campos-Soldini, M. P." box="[476,854,1815,1841]" pageId="22" pageNumber="23" pagination="59 - 138" refId="ref33808" refString="Ghahari, H. &amp; Campos-Soldini, M. P. (2019) An annotated catalogue of blister-beetles (Coleoptera: Tenebrionoidea: Meloidae) of Iran. Entomofauna, 40, 59 - 138." type="journal article" year="2019">Ghahari &amp; Campos-Soldini 2019</bibRefCitation>
);
<collectingRegion id="49F2F879FFC3FF82FC89F8AFFB961156" box="[872,1096,1815,1841]" country="United Arab Emirates" name="Abu Zaby" pageId="22" pageNumber="23">Abu Dhabi Emirate</collectingRegion>
, EA4, Al Bateen dunes, 4 Km W Al Aïn on the road to
<collectingRegion id="49F2F879FFC3FF82FE46F883FDFB1132" box="[423,549,1851,1877]" country="United Arab Emirates" name="Abu Zaby" pageId="22" pageNumber="23">Abu Dhabi</collectingRegion>
,
<geoCoordinate id="EE02505CFFC3FF82FDD1F883FD181131" box="[560,710,1851,1878]" degrees="24, 20491" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="1" value="24.20491">24, 20491° N</geoCoordinate>
<geoCoordinate id="EE02505CFFC3FF82FD2CF882FCBE1131" box="[717,864,1850,1878]" degrees="55, 61263" direction="east" orientation="longitude" pageId="22" pageNumber="23" precision="1" value="55.61263">55, 61263° E</geoCoordinate>
(
<bibRefCitation id="EFA74B6AFFC3FF82FC8FF883FBAB1132" author="Bologna, M. A. &amp; Turco, F." box="[878,1141,1851,1878]" pageId="22" pageNumber="23" pagination="1 - 33" refId="ref32438" refString="Bologna, M. A. &amp; Turco, F. (2007) The Meloidae (Coleoptera) of the United Arab Emirates with an updated Arabian checklist. Zootaxa, 1625 (1), 1 - 33. https: // doi. org / 10.11646 / zootaxa. 1625.1.1" type="journal article" year="2007">Bologna &amp; Turco 2007</bibRefCitation>
,
<typeStatus id="548D8839FFC3FF82FB61F884FB6E1131" box="[1152,1200,1852,1878]" pageId="22" pageNumber="23">type</typeStatus>
locality);
<collectingRegion id="49F2F879FFC3FF82FAC0F883FF2D111E" country="United Arab Emirates" name="Abu Zaby" pageId="22" pageNumber="23">Abu Dhabi Emirate</collectingRegion>
, EA10, Al Aïn, 3–4 Km E Al Selimat on road
<collectingRegion id="49F2F879FFC3FF82FD1DF8E7FC0E111E" box="[764,976,1887,1913]" inferredBySuffix="true" pageId="22" pageNumber="23">Al Aïn-Abu Dhabi</collectingRegion>
,
<geoCoordinate id="EE02505CFFC3FF82FC3DF8E7FBAD111D" box="[988,1139,1887,1914]" degrees="24, 19263" direction="north" orientation="latitude" pageId="22" pageNumber="23" precision="1" value="24.19263">24, 19263° N</geoCoordinate>
<geoCoordinate id="EE02505CFFC3FF82FB9AF8E6FAD1111D" box="[1147,1295,1886,1914]" degrees="55, 58861" direction="east" orientation="longitude" pageId="22" pageNumber="23" precision="1" value="55.58861">55, 58861° E</geoCoordinate>
(
<bibRefCitation id="EFA74B6AFFC3FF82FAFFF8E7FEC311FA" author="Bologna, M. A. &amp; Turco, F." pageId="22" pageNumber="23" pagination="1 - 33" refId="ref32438" refString="Bologna, M. A. &amp; Turco, F. (2007) The Meloidae (Coleoptera) of the United Arab Emirates with an updated Arabian checklist. Zootaxa, 1625 (1), 1 - 33. https: // doi. org / 10.11646 / zootaxa. 1625.1.1" type="journal article" year="2007">Bologna &amp; Turco 2007</bibRefCitation>
); Jebel Hatif (
<bibRefCitation id="EFA74B6AFFC3FF82FE5AF83BFD1C11FA" author="Gillet, M. P. T. &amp; Howarth, B." box="[443,706,1923,1949]" pageId="22" pageNumber="23" pagination="94 - 143" refId="ref33947" refString="Gillet, M. P. T. &amp; Howarth, B. (2004) Insects. In: Aspinall, S. &amp; Hellyer, P. (Eds.), Jebel Hafit, a natural history. Emirates Natural History Group, Abu Dhabi, pp. 94 - 143." type="book chapter" year="2004">Gillet &amp; Howarth 2004</bibRefCitation>
as
<taxonomicName id="4C364D18FFC3FF82FD09F83BFCBE11FA" authorityName="Olivier" authorityYear="1811" box="[744,864,1923,1949]" class="Insecta" family="Meloidae" genus="Mylabris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="bipunctata">
<emphasis id="B942EA89FFC3FF82FD09F83BFCBE11FA" box="[744,864,1923,1949]" italics="true" pageId="22" pageNumber="23">bipunctata</emphasis>
</taxonomicName>
); Al Aïn area (
<bibRefCitation id="EFA74B6AFFC3FF82FBE9F83BFB3311FA" author="Gillet, M. P. T. &amp; Gillet, J." box="[1032,1261,1923,1949]" pageId="22" pageNumber="23" pagination="20 - 21" refId="ref33845" refString="Gillet, M. P. T. &amp; Gillet, J. (1996) A large blister beetle Mylabris maculiventris Klug, 1845 from Al Ain and neighbouring Oman (Coleoptera: Meloidae). Tribulus, 6, 20 - 21." type="journal article" year="1996">Gillet &amp; Gillet 1996</bibRefCitation>
as
<taxonomicName id="4C364D18FFC3FF82FAF2F83BFA5511FA" authorityName="Olivier" authorityYear="1811" box="[1299,1419,1923,1949]" class="Insecta" family="Meloidae" genus="Mylabris" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="bipunctata">
<emphasis id="B942EA89FFC3FF82FAF2F83BFA5511FA" box="[1299,1419,1923,1949]" italics="true" pageId="22" pageNumber="23">bipunctata</emphasis>
</taxonomicName>
); Wadi Bih; Wadi Shawkah (Batelka &amp; Geistardt 2009 as
<taxonomicName id="4C364D18FFC3FF82FCEDF81FFC5111A6" baseAuthorityName="Olivier" baseAuthorityYear="1811" box="[780,911,1959,1985]" class="Insecta" family="Meloidae" genus="Hycleus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="22" pageNumber="23" phylum="Arthropoda" rank="species" species="bipunctatus">
<emphasis id="B942EA89FFC3FF82FCEDF81FFC5111A6" box="[780,911,1959,1985]" italics="true" pageId="22" pageNumber="23">bipunctatus</emphasis>
</taxonomicName>
).
</paragraph>
</subSubSection>
<subSubSection id="C32C6510FFC2FF83FF26FF20FC1D1404" pageId="23" pageNumber="24" type="description">
<paragraph id="8B89369BFFC2FF83FF26FF20FD9917B3" blockId="23.[151,1437,152,683]" pageId="23" pageNumber="24">
<emphasis id="B942EA89FFC2FF83FF26FF20FE8716D4" bold="true" box="[199,345,152,179]" pageId="23" pageNumber="24">Description.</emphasis>
Body length 12–16.5 mm. Body black, elytra yellowish bright brown, sometimes red, with black pattern. Head subquadrate; male antennomere III (
<figureCitation id="130D2A1EFFC2FF83FD26FF05FCC616B0" box="[711,792,189,215]" captionStart="FIGURES 77–83" captionStartId="22.[151,264,1564,1589]" captionTargetBox="[181,1404,667,1533]" captionTargetId="figure@22.[177,1410,663,1539]" captionTargetPageId="22" captionText="FIGURES 77–83. Hycleus pirata Bologna &amp; Turco: 77. male antenna; 78. female antenna; 79. tegmen, ventral view; 80. tegmen, lateral view; 81. aedeagus, lateral view; 82. mesosternum; 83. elytral pattern. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3887797" httpUri="https://zenodo.org/record/3887797/files/figure.png" pageId="23" pageNumber="24">Fig. 77</figureCitation>
) less than half longer than IV, V–VIII transverse and widened, IX–X narrower than VIII, male XI narrower and longer than X; in female (
<figureCitation id="130D2A1EFFC2FF83FBFEFF59FBAE169C" box="[1055,1136,225,251]" captionStart="FIGURES 77–83" captionStartId="22.[151,264,1564,1589]" captionTargetBox="[181,1404,667,1533]" captionTargetId="figure@22.[177,1410,663,1539]" captionTargetPageId="22" captionText="FIGURES 77–83. Hycleus pirata Bologna &amp; Turco: 77. male antenna; 78. female antenna; 79. tegmen, ventral view; 80. tegmen, lateral view; 81. aedeagus, lateral view; 82. mesosternum; 83. elytral pattern. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3887797" httpUri="https://zenodo.org/record/3887797/files/figure.png" pageId="23" pageNumber="24">Fig. 78</figureCitation>
) shorter than X. Pronotum (
<figureCitation id="130D2A1EFFC2FF83FF41FEBDFF2F1778" box="[160,241,261,287]" captionStart="FIGURES 13–24" captionStartId="6.[151,262,1296,1321]" captionTargetBox="[223,1367,188,1264]" captionTargetId="figure@6.[220,1368,181,1271]" captionTargetPageId="6" captionText="FIGURES 13–24. Head and pronotum of: 13. Hycleus apicipennis; 14. H. balteatus; 15 H. bipunctatus; 16. H. cingulatus; 17. H. colligatus; 18. H. hayki; 19. H. lindbergi; 20. H. pirata; 21. H. schauffelei; 22. H. sexmaculatus; 23. H. tenuepictus; 24. H. trianguliferus." figureDoi="http://doi.org/10.5281/zenodo.3887777" httpUri="https://zenodo.org/record/3887777/files/figure.png" pageId="23" pageNumber="24">Fig. 20</figureCitation>
) longer than wide, with anterior depression; mesosternum as in
<figureCitation id="130D2A1EFFC2FF83FC20FEBDFBCA1778" box="[961,1044,261,287]" captionStart="FIGURES 77–83" captionStartId="22.[151,264,1564,1589]" captionTargetBox="[181,1404,667,1533]" captionTargetId="figure@22.[177,1410,663,1539]" captionTargetPageId="22" captionText="FIGURES 77–83. Hycleus pirata Bologna &amp; Turco: 77. male antenna; 78. female antenna; 79. tegmen, ventral view; 80. tegmen, lateral view; 81. aedeagus, lateral view; 82. mesosternum; 83. elytral pattern. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3887797" httpUri="https://zenodo.org/record/3887797/files/figure.png" pageId="23" pageNumber="24">Fig. 82</figureCitation>
. Each elytron with two black spots on anterior third, one sinuate middle fascia, and one black oblique fascia on apical part (
<figureCitation id="130D2A1EFFC2FF83FB90FE91FB1A1724" box="[1137,1220,297,323]" captionStart="FIGURES 77–83" captionStartId="22.[151,264,1564,1589]" captionTargetBox="[181,1404,667,1533]" captionTargetId="figure@22.[177,1410,663,1539]" captionTargetPageId="22" captionText="FIGURES 77–83. Hycleus pirata Bologna &amp; Turco: 77. male antenna; 78. female antenna; 79. tegmen, ventral view; 80. tegmen, lateral view; 81. aedeagus, lateral view; 82. mesosternum; 83. elytral pattern. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3887797" httpUri="https://zenodo.org/record/3887797/files/figure.png" pageId="23" pageNumber="24">Fig. 83</figureCitation>
). Male last ventrite regularly concave and depressed, posterior margin with a wide and deep middle emargination; aedeagus (
<figureCitation id="130D2A1EFFC2FF83FAA3FEF5FA4A1700" box="[1346,1428,333,359]" captionStart="FIGURES 77–83" captionStartId="22.[151,264,1564,1589]" captionTargetBox="[181,1404,667,1533]" captionTargetId="figure@22.[177,1410,663,1539]" captionTargetPageId="22" captionText="FIGURES 77–83. Hycleus pirata Bologna &amp; Turco: 77. male antenna; 78. female antenna; 79. tegmen, ventral view; 80. tegmen, lateral view; 81. aedeagus, lateral view; 82. mesosternum; 83. elytral pattern. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3887797" httpUri="https://zenodo.org/record/3887797/files/figure.png" pageId="23" pageNumber="24">Fig. 81</figureCitation>
) with endophallic hook narrow, distal aedeagal hook curved, far from proximal one and close to apex, proximal hook slightly curved; gonoforceps (
<figureCitation id="130D2A1EFFC2FF83FE0FFE2DFDAF17C8" box="[494,625,405,431]" captionStart="FIGURES 77–83" captionStartId="22.[151,264,1564,1589]" captionTargetBox="[181,1404,667,1533]" captionTargetId="figure@22.[177,1410,663,1539]" captionTargetPageId="22" captionText="FIGURES 77–83. Hycleus pirata Bologna &amp; Turco: 77. male antenna; 78. female antenna; 79. tegmen, ventral view; 80. tegmen, lateral view; 81. aedeagus, lateral view; 82. mesosternum; 83. elytral pattern. Scale bar: 1 mm." figureDoi="http://doi.org/10.5281/zenodo.3887797" httpUri="https://zenodo.org/record/3887797/files/figure.png" pageId="23" pageNumber="24">Figs 79–80</figureCitation>
) narrow and elongate both in ventral and lateral view (see
<bibRefCitation id="EFA74B6AFFC2FF83FAFFFE2DFEC617B4" author="Bologna, M. A. &amp; Turco, F." pageId="23" pageNumber="24" pagination="1 - 33" refId="ref32438" refString="Bologna, M. A. &amp; Turco, F. (2007) The Meloidae (Coleoptera) of the United Arab Emirates with an updated Arabian checklist. Zootaxa, 1625 (1), 1 - 33. https: // doi. org / 10.11646 / zootaxa. 1625.1.1" type="journal article" year="2007">Bologna &amp; Turco 2007</bibRefCitation>
for a detailed description).
</paragraph>
<paragraph id="8B89369BFFC2FF83FF26FE64FC1D1404" blockId="23.[151,1437,152,683]" pageId="23" pageNumber="24">
<emphasis id="B942EA89FFC2FF83FF26FE64FE601790" bold="true" box="[199,446,476,503]" pageId="23" pageNumber="24">Taxonomic remarks.</emphasis>
This species resembles
<taxonomicName id="4C364D18FFC2FF83FD2FFE66FCAC1790" baseAuthorityName="Reiche" baseAuthorityYear="1866" box="[718,882,478,503]" class="Insecta" family="Meloidae" genus="Hycleus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="apicipennis">
<emphasis id="B942EA89FFC2FF83FD2FFE66FCAC1790" box="[718,882,478,503]" italics="true" pageId="23" pageNumber="24">H. apicipennis</emphasis>
</taxonomicName>
but it can be distinguished by the elongate shape of antennomere IV, which is subequal in length to V, the shape of last male antennomere, which is strikingly curved, and by the male genitalia. The shape of last antennomere is also very similar to that of
<taxonomicName id="4C364D18FFC2FF83FBBFFD9EFB631458" authorityName="Serri &amp; Bologna &amp; Riccieri" authorityYear="2020" box="[1118,1213,549,575]" class="Insecta" family="Meloidae" genus="Hycleus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="hayki">
<emphasis id="B942EA89FFC2FF83FBBFFD9EFB631458" box="[1118,1213,549,575]" italics="true" pageId="23" pageNumber="24">H. hayki</emphasis>
</taxonomicName>
and of
<taxonomicName id="4C364D18FFC2FF83FAF0FD9EFF131404" baseAuthorityName="Olivier" baseAuthorityYear="1811" class="Insecta" family="Meloidae" genus="Hycleus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="sexmaculatus">
<emphasis id="B942EA89FFC2FF83FAF0FD9EFF131404" italics="true" pageId="23" pageNumber="24">H. sexmaculatus</emphasis>
</taxonomicName>
, but
<taxonomicName id="4C364D18FFC2FF83FEE7FDF2FEB31404" authorityName="Bologna &amp; Turco" authorityYear="2007" box="[262,365,586,611]" class="Insecta" family="Meloidae" genus="Hycleus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="pirata">
<emphasis id="B942EA89FFC2FF83FEE7FDF2FEB31404" box="[262,365,586,611]" italics="true" pageId="23" pageNumber="24">H. pirata</emphasis>
</taxonomicName>
differs because of the pronotum and aedeagus shape.
</paragraph>
</subSubSection>
<subSubSection id="C32C6510FFC2FF83FF26FDD4FAB814CC" pageId="23" pageNumber="24" type="distribution">
<paragraph id="8B89369BFFC2FF83FF26FDD4FAB814CC" blockId="23.[151,1437,152,683]" pageId="23" pageNumber="24">
<emphasis id="B942EA89FFC2FF83FF26FDD4FEBF14E0" bold="true" box="[199,353,620,647]" pageId="23" pageNumber="24">Distribution.</emphasis>
This species seems to have restricted distribution in the
<collectingCountry id="F321760BFFC2FF83FC3DFDD5FB0F14E0" box="[988,1233,621,647]" name="United Arab Emirates" pageId="23" pageNumber="24">United Arab Emirates</collectingCountry>
and
<collectingCountry id="F321760BFFC2FF83FAE6FDD5FA9514E0" box="[1287,1355,621,647]" name="Oman" pageId="23" pageNumber="24">Oman</collectingCountry>
(
<bibRefCitation id="EFA74B6AFFC2FF83FABAFDD5FEB714CC" author="Bologna, M. A. &amp; Turco, F." pageId="23" pageNumber="24" pagination="1 - 33" refId="ref32438" refString="Bologna, M. A. &amp; Turco, F. (2007) The Meloidae (Coleoptera) of the United Arab Emirates with an updated Arabian checklist. Zootaxa, 1625 (1), 1 - 33. https: // doi. org / 10.11646 / zootaxa. 1625.1.1" type="journal article" year="2007">Bologna &amp; Turco 2007</bibRefCitation>
). The record of this species from south
<collectingCountry id="F321760BFFC2FF83FCC3FD29FC9114CC" box="[802,847,657,683]" name="Iran" pageId="23" pageNumber="24">Iran</collectingCountry>
(
<bibRefCitation id="EFA74B6AFFC2FF83FCBFFD29FC0914CC" author="Serri, S." box="[862,983,656,683]" pageId="23" pageNumber="24" pagination="232 - 237" refId="ref37243" refString="Serri, S. (2015) Blister beetles (Coleoptera: Meloidae) of Southwest Iran. Proceedings of the 1 st Iranian International Congress of Entomology, 2015, 232 - 237." type="journal article" year="2015">Serri 2015</bibRefCitation>
) refers to the new species
<taxonomicName id="4C364D18FFC2FF83FAE1FD2AFA8114CC" authorityName="Serri &amp; Bologna &amp; Riccieri" authorityYear="2020" box="[1280,1375,657,683]" class="Insecta" family="Meloidae" genus="Hycleus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="23" pageNumber="24" phylum="Arthropoda" rank="species" species="hayki">
<emphasis id="B942EA89FFC2FF83FAE1FD2AFA8114CC" box="[1280,1375,657,683]" italics="true" pageId="23" pageNumber="24">H. hayki</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>
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