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<document id="10426751616ACD6161CCC5D9C7116F43" ID-DOI="10.5281/zenodo.191317" ID-GBIF-Dataset="94e38b1e-c437-4315-8538-daaf32535555" ID-ISSN="1175-5326" ID-Zenodo-Dep="191317" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1461126606189" checkinUser="plazi" docAuthor="Elberg, Kaupo, Rozkošný, Rudolf &amp; Knutson, Lloyd" docDate="2009" docId="039F8602D56DD00F5F930C689619FC40" docLanguage="en" docName="zt02288p060.pdf" docOrigin="Zootaxa 2288" docStyle="DocumentStyle:890A69B780ED73D6DB8551B71C8AC79E.4:Zootaxa.2009-2012.journal_article" docStyleId="890A69B780ED73D6DB8551B71C8AC79E" docStyleName="Zootaxa.2009-2012.journal_article" docStyleVersion="4" docTitle="Sepedon hecate Elberg, Rozkošný &amp; Knutson, 2009, sp.nov." docType="treatment" docVersion="9" lastPageNumber="55" masterDocId="FFA6FE7AD56CD00B5F040B349741FF9E" masterDocTitle="A review of of the Holarctic Sepedon fuscipennis and S. spinipes groups with description of a new species (Diptera: Sciomyzidae)" masterLastPageNumber="60" masterPageNumber="51" pageNumber="52" updateTime="1698596924154" updateUser="plazi">
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<mods:title id="F496F2DCC9DB02A2C18521B582FA76E4">A review of of the Holarctic Sepedon fuscipennis and S. spinipes groups with description of a new species (Diptera: Sciomyzidae)</mods:title>
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<mods:namePart id="2F67C1E8252BB0D35A42983379E25564">Elberg, Kaupo</mods:namePart>
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<mods:namePart id="2E0AFB8D5610DC856BD37E5A3A139131">Rozkošný, Rudolf</mods:namePart>
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<mods:namePart id="A8CB88A735A14B145BC7526F45F771CD">Knutson, Lloyd</mods:namePart>
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<mods:date id="0253469EB63CEF8442B2F6877D9D8D46">2009</mods:date>
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<treatment id="039F8602D56DD00F5F930C689619FC40" ID-DOI="http://doi.org/10.5281/zenodo.6216436" ID-GBIF-Taxon="119626346" ID-Zenodo-Dep="6216436" LSID="urn:lsid:plazi:treatment:039F8602D56DD00F5F930C689619FC40" httpUri="http://treatment.plazi.org/id/039F8602D56DD00F5F930C689619FC40" lastPageId="4" lastPageNumber="55" pageId="1" pageNumber="52">
<subSubSection id="C32C649FD56DD00A5F930C68967CF806" pageId="1" pageNumber="52" type="nomenclature">
<paragraph id="8B893714D56DD00A5F930C6896F1F8EB" blockId="1.[151,432,1883,1944]" box="[151,432,1883,1910]" pageId="1" pageNumber="52">
<heading id="D0C18078D56DD00A5F930C6896F1F8EB" bold="true" box="[151,432,1883,1910]" fontSize="11" level="1" pageId="1" pageNumber="52" reason="1">
<emphasis id="B942EB06D56DD00A5F930C6896F1F8EB" bold="true" box="[151,432,1883,1910]" pageId="1" pageNumber="52">
<taxonomicName id="4C364C97D56DD00A5F930C689612F8E8" ID-CoL="9HMWJ" box="[151,339,1884,1910]" class="Insecta" family="Sciomyzidae" genus="Sepedon" kingdom="Animalia" order="Diptera" pageId="1" pageNumber="52" phylum="Arthropoda" rank="species" species="hecate" status="sp. nov.">
<emphasis id="B942EB06D56DD00A5F930C689612F8E8" bold="true" box="[151,339,1884,1910]" italics="true" pageId="1" pageNumber="52">Sepedon hecate</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A271567DD56DD00A5E5E0C6F96F1F8EB" box="[346,432,1883,1909]" pageId="1" pageNumber="52" rank="species">sp.nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B893714D56DD00A5F930C4A967CF806" blockId="1.[151,432,1883,1944]" box="[151,317,1918,1944]" pageId="1" pageNumber="52">
(
<figureCitation id="130D2B91D56DD00A5F9B0C4A9649F806" box="[159,264,1918,1944]" captionStart-0="FIGURES 1 3" captionStart-1="FIGURES 4 7" captionStartId-0="2.[151,269,1183,1207]" captionStartId-1="3.[151,270,1886,1910]" captionTargetBox-0="[190,1402,308,1131]" captionTargetBox-1="[205,1398,1066,1837]" captionTargetId-0="figure@2.[151,1436,280,1159]" captionTargetId-1="figure@3.[151,1436,1037,1862]" captionTargetPageId-0="2" captionTargetPageId-1="3" captionText-0="FIGURES 1 3. Sepedon hecate sp. nov., male. 1. Head in dorsal view. 2. Head in lateral view. 3. Left hind leg in lateral view. Scales: 1.0 mm." captionText-1="FIGURES 4 7. Sepedon hecate sp. nov., male terminalia. 4. Lateral view. 5. Hypandrium in lateral view. 6. Aedeagal complex in lateral view. 7. Epiphallus in caudal view. Scales: 0.3 mm. aa aedeagal apodeme, ae aedeagus, ce cercus, dph distiphallus, ea ejaculatory apodeme, ep epandrium, eph epiphallus, hy hypandrium, po postgonite, pr pregonite, ss surstylus." httpUri-0="https://zenodo.org/record/191318/files/figure.png" httpUri-1="https://zenodo.org/record/191319/files/figure.png" pageId="1" pageNumber="52">Figs 17</figureCitation>
,
<figureCitation id="130D2B91D56DD00A5E110C4A9675F806" box="[277,308,1918,1944]" captionStart="FIGURES 28 29" captionStartId="8.[151,269,686,710]" captionTargetBox="[432,1143,514,655]" captionTargetId="figure@8.[430,1146,514,658]" captionTargetPageId="8" captionText="FIGURES 28 29. Wings of Sepedon species. 28. S. hecate sp. nov. 29. S. spinipes." httpUri="https://zenodo.org/record/191322/files/figure.png" pageId="1" pageNumber="52">28</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="C32C649FD56DD0095F930CFF97AEFF61" lastPageId="2" lastPageNumber="53" pageId="1" pageNumber="52" type="diagnosis">
<paragraph id="8B893714D56DD0095F930CFF97AEFF61" blockId="1.[151,1436,1995,2022]" lastBlockId="2.[151,1436,152,255]" lastPageId="2" lastPageNumber="53" pageId="1" pageNumber="52">
<emphasis id="B942EB06D56DD00A5F930CFF9659F87B" bold="true" box="[151,280,1995,2021]" pageId="1" pageNumber="52">Diagnosis.</emphasis>
A brown species with an elongate, subtriangular, black orbital spot (
<figureCitation id="130D2B91D56DD00A5B490CF893D2F878" box="[1101,1171,1996,2022]" captionStart="FIGURES 1 3" captionStartId="2.[151,269,1183,1207]" captionTargetBox="[190,1402,308,1131]" captionTargetId="figure@2.[151,1436,280,1159]" captionTargetPageId="2" captionText="FIGURES 1 3. Sepedon hecate sp. nov., male. 1. Head in dorsal view. 2. Head in lateral view. 3. Left hind leg in lateral view. Scales: 1.0 mm." httpUri="https://zenodo.org/record/191318/files/figure.png" pageId="1" pageNumber="52">Fig. 1</figureCitation>
); moderately concave face (
<figureCitation id="130D2B91D56ED0095FD90BAC966AFF2C" box="[221,299,152,178]" captionStart="FIGURES 1 3" captionStartId="2.[151,269,1183,1207]" captionTargetBox="[190,1402,308,1131]" captionTargetId="figure@2.[151,1436,280,1159]" captionTargetPageId="2" captionText="FIGURES 1 3. Sepedon hecate sp. nov., male. 1. Head in dorsal view. 2. Head in lateral view. 3. Left hind leg in lateral view. Scales: 1.0 mm." httpUri="https://zenodo.org/record/191318/files/figure.png" pageId="2" pageNumber="53">Fig. 2</figureCitation>
); pedicel index (see p. 2) about 5.0; hypandrium produced basally in lateral view (
<figureCitation id="130D2B91D56ED0095A3A0BAC92CDFF2C" box="[1342,1420,152,178]" captionStart="FIGURES 4 7" captionStartId="3.[151,270,1886,1910]" captionTargetBox="[205,1398,1066,1837]" captionTargetId="figure@3.[151,1436,1037,1862]" captionTargetPageId="3" captionText="FIGURES 4 7. Sepedon hecate sp. nov., male terminalia. 4. Lateral view. 5. Hypandrium in lateral view. 6. Aedeagal complex in lateral view. 7. Epiphallus in caudal view. Scales: 0.3 mm. aa aedeagal apodeme, ae aedeagus, ce cercus, dph distiphallus, ea ejaculatory apodeme, ep epandrium, eph epiphallus, hy hypandrium, po postgonite, pr pregonite, ss surstylus." httpUri="https://zenodo.org/record/191319/files/figure.png" pageId="2" pageNumber="53">Fig. 5</figureCitation>
); aedeagus: 12 distinct midprojections; distiphallus shorter than epiphallus (
<figureCitation id="130D2B91D56ED0095B400B8A93D0FF46" box="[1092,1169,190,216]" captionStart="FIGURES 4 7" captionStartId="3.[151,270,1886,1910]" captionTargetBox="[205,1398,1066,1837]" captionTargetId="figure@3.[151,1436,1037,1862]" captionTargetPageId="3" captionText="FIGURES 4 7. Sepedon hecate sp. nov., male terminalia. 4. Lateral view. 5. Hypandrium in lateral view. 6. Aedeagal complex in lateral view. 7. Epiphallus in caudal view. Scales: 0.3 mm. aa aedeagal apodeme, ae aedeagus, ce cercus, dph distiphallus, ea ejaculatory apodeme, ep epandrium, eph epiphallus, hy hypandrium, po postgonite, pr pregonite, ss surstylus." httpUri="https://zenodo.org/record/191319/files/figure.png" pageId="2" pageNumber="53">Fig. 6</figureCitation>
); and epiphallus oval (
<figureCitation id="130D2B91D56ED0095FA40BD197A0FF61" box="[160,225,229,255]" captionStart="FIGURES 4 7" captionStartId="3.[151,270,1886,1910]" captionTargetBox="[205,1398,1066,1837]" captionTargetId="figure@3.[151,1436,1037,1862]" captionTargetPageId="3" captionText="FIGURES 4 7. Sepedon hecate sp. nov., male terminalia. 4. Lateral view. 5. Hypandrium in lateral view. 6. Aedeagal complex in lateral view. 7. Epiphallus in caudal view. Scales: 0.3 mm. aa aedeagal apodeme, ae aedeagus, ce cercus, dph distiphallus, ea ejaculatory apodeme, ep epandrium, eph epiphallus, hy hypandrium, po postgonite, pr pregonite, ss surstylus." httpUri="https://zenodo.org/record/191319/files/figure.png" pageId="2" pageNumber="53">Fig.7</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="DF49679CD56ED0095F930FAB96C0FB47" httpUri="https://zenodo.org/record/191318/files/figure.png" pageId="2" pageNumber="53" targetBox="[190,1402,308,1131]" targetPageId="2">
<paragraph id="8B893714D56ED0095F930FAB96C0FB47" blockId="2.[151,1436,1183,1241]" pageId="2" pageNumber="53">
<emphasis id="B942EB06D56ED0095F930FAB9603FB29" bold="true" box="[151,322,1183,1207]" pageId="2" pageNumber="53">FIGURES 13.</emphasis>
<taxonomicName id="4C364C97D56ED0095E4D0FAB96B1FB28" box="[329,496,1183,1206]" class="Insecta" family="Sciomyzidae" genus="Sepedon" kingdom="Animalia" order="Diptera" pageId="2" pageNumber="53" phylum="Arthropoda" rank="species" species="hecate" status="sp. nov.">
<emphasis id="B942EB06D56ED0095E4D0FAB96B1FB28" box="[329,496,1183,1206]" italics="true" pageId="2" pageNumber="53">Sepedon hecate</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A271567DD56ED0095EF20FAB950DFB29" box="[502,588,1183,1207]" pageId="2" pageNumber="53" rank="species">
<emphasis id="B942EB06D56ED0095EF20FAB950DFB29" bold="true" box="[502,588,1183,1207]" pageId="2" pageNumber="53">sp. nov.</emphasis>
</taxonomicNameLabel>
, male. 1. Head in dorsal view. 2. Head in lateral view. 3. Left hind leg in lateral view. Scales: 1.0 mm.
</paragraph>
</caption>
<subSubSection id="C32C649FD56ED0085FC20E33947DFC40" lastPageId="3" lastPageNumber="54" pageId="2" pageNumber="53" type="description">
<paragraph id="8B893714D56ED0095FC20E339385F954" blockId="2.[151,1437,1287,2009]" pageId="2" pageNumber="53">
<emphasis id="B942EB06D56ED0095FC20E33961EFABF" bold="true" box="[198,351,1287,1313]" pageId="2" pageNumber="53">Description.</emphasis>
Male: Body length (including porrect antenna): 7.48.1 mm; wing length 5.86.7 mm. Head (
<figureCitation id="130D2B91D56ED0095F9B0E1A9649FAD6" box="[159,264,1326,1352]" captionStart="FIGURES 1 3" captionStartId="2.[151,269,1183,1207]" captionTargetBox="[190,1402,308,1131]" captionTargetId="figure@2.[151,1436,280,1159]" captionTargetPageId="2" captionText="FIGURES 1 3. Sepedon hecate sp. nov., male. 1. Head in dorsal view. 2. Head in lateral view. 3. Left hind leg in lateral view. Scales: 1.0 mm." httpUri="https://zenodo.org/record/191318/files/figure.png" pageId="2" pageNumber="53">Figs 12</figureCitation>
) conspicuously produced in front of and below eye in lateral view. Eye obliquely oval, somewhat higher than gena. Frons broader than long, broadly excavated in front of ocellar tubercle. Velvety black orbital spot along eye margin elongate subtriangular, gradually tapering toward level of orbital seta. Velvety black parafacial spot at eye margin short oval. Antenna long and slender. Pedicel rod-like, about 5.0 times as long as broad in middle and slightly dilated apically. Postpedicel subtriangular, tapered in distal half but with rounded apex. Pedicel and postpedicel darkened, with greyish pollination, postpedicel almost black along lower margin and in distal half. Arista inserted in middle of postpedicel, with yellowish basal segment, more brown in thickened basal third and contrastingly white and finely pubescent in rest of its length. Face with sharp mid keel in upper half, distinctly concave in lateral view. Some parts of head surface (especially occiput, space around parafacial spot, upper mid face and a genal stripe below lower eye margin) with diffuse whitish pollination. Proboscis and palpus brown, palpus lanceolate, dilated in more than distal half.
</paragraph>
<paragraph id="8B893714D56ED0095FC10DE39497F82D" blockId="2.[151,1437,1287,2009]" pageId="2" pageNumber="53">Thorax brown, densely grey pollinated on scutum, scutellum and greater part of pleura. Scutum with two pairs of brown longitudinal stripes: each submedian stripe broader than pollinated space between them and more lateral stripes very narrow, almost linear. Shining postpronotal callus and broad notopleuron separated from rest of scutum by more or less visible brown stripe continuing to wing base. Pleura densely pale grey pollinated, only anterior part of katepisternum and katatergite (in front of and above posterior spiracle) more shining. Prosternum with more or less distinct setulae, katatergite bare.</paragraph>
<paragraph id="8B893714D56ED0085FC10C8B95C8FF2C" blockId="2.[151,1437,1287,2009]" lastBlockId="3.[151,1437,152,990]" lastPageId="3" lastPageNumber="54" pageId="2" pageNumber="53">
Wing (
<figureCitation id="130D2B91D56ED0095E1E0C8B9636F847" box="[282,375,1983,2009]" captionStart="FIGURES 28 29" captionStartId="8.[151,269,686,710]" captionTargetBox="[432,1143,514,655]" captionTargetId="figure@8.[430,1146,514,658]" captionTargetPageId="8" captionText="FIGURES 28 29. Wings of Sepedon species. 28. S. hecate sp. nov. 29. S. spinipes." httpUri="https://zenodo.org/record/191322/files/figure.png" pageId="2" pageNumber="53">Fig. 28</figureCitation>
) pale, brownish infumated with tendency to darkening of both crossveins. Posterior crossvein distinctly sinuate. Halter yellow.
</paragraph>
<paragraph id="8B893714D56FD0085FC10B8A93F9FEB8" blockId="3.[151,1437,152,990]" pageId="3" pageNumber="54">
Legs almost uniformly yellowish to reddish brown. Mid femur with short but distinct seta in middle of anterior surface.
<collectingCountry id="F3217784D56FD0085E5F0BD196D7FF61" box="[347,406,229,255]" name="India" pageId="3" pageNumber="54">Hind</collectingCountry>
femur (
<figureCitation id="130D2B91D56FD0085EF60BD19579FF61" box="[498,568,229,255]" captionStart="FIGURES 1 3" captionStartId="2.[151,269,1183,1207]" captionTargetBox="[190,1402,308,1131]" captionTargetId="figure@2.[151,1436,280,1159]" captionTargetPageId="2" captionText="FIGURES 1 3. Sepedon hecate sp. nov., male. 1. Head in dorsal view. 2. Head in lateral view. 3. Left hind leg in lateral view. Scales: 1.0 mm." httpUri="https://zenodo.org/record/191318/files/figure.png" pageId="3" pageNumber="54">Fig. 3</figureCitation>
) conspicuously swollen, with a double row of irregular short black spines.
<collectingCountry id="F3217784D56FD0085F930A389793FEB8" box="[151,210,268,294]" name="India" pageId="3" pageNumber="54">Hind</collectingCountry>
tibia distinctly curved following ventral outline of hind femur in its distal two thirds.
</paragraph>
<paragraph id="8B893714D56FD0085FC10A06956DFDC5" blockId="3.[151,1437,152,990]" pageId="3" pageNumber="54">
Abdomen shining yellowish brown, though sternites in basal half usually matt, greyish pollinated. Male terminalia (
<figureCitation id="130D2B91D56FD0085E270A6D96D5FEED" box="[291,404,345,371]" captionStart="FIGURES 4 7" captionStartId="3.[151,270,1886,1910]" captionTargetBox="[205,1398,1066,1837]" captionTargetId="figure@3.[151,1436,1037,1862]" captionTargetPageId="3" captionText="FIGURES 4 7. Sepedon hecate sp. nov., male terminalia. 4. Lateral view. 5. Hypandrium in lateral view. 6. Aedeagal complex in lateral view. 7. Epiphallus in caudal view. Scales: 0.3 mm. aa aedeagal apodeme, ae aedeagus, ce cercus, dph distiphallus, ea ejaculatory apodeme, ep epandrium, eph epiphallus, hy hypandrium, po postgonite, pr pregonite, ss surstylus." httpUri="https://zenodo.org/record/191319/files/figure.png" pageId="3" pageNumber="54">Figs 47</figureCitation>
): Cerci long but not reaching ventral margin of epandrium, surstylus relatively long, slightly pointed in lateral view. Hypandrium with markedly prominent basal part (
<figureCitation id="130D2B91D56FD0085B650AB493E8FE04" box="[1121,1193,384,410]" captionStart="FIGURES 4 7" captionStartId="3.[151,270,1886,1910]" captionTargetBox="[205,1398,1066,1837]" captionTargetId="figure@3.[151,1436,1037,1862]" captionTargetPageId="3" captionText="FIGURES 4 7. Sepedon hecate sp. nov., male terminalia. 4. Lateral view. 5. Hypandrium in lateral view. 6. Aedeagal complex in lateral view. 7. Epiphallus in caudal view. Scales: 0.3 mm. aa aedeagal apodeme, ae aedeagus, ce cercus, dph distiphallus, ea ejaculatory apodeme, ep epandrium, eph epiphallus, hy hypandrium, po postgonite, pr pregonite, ss surstylus." httpUri="https://zenodo.org/record/191319/files/figure.png" pageId="3" pageNumber="54">Fig. 5</figureCitation>
). Aedeagal complex (
<figureCitation id="130D2B91D56FD0085F9B0A9297A4FE5E" box="[159,229,422,448]" captionStart="FIGURES 4 7" captionStartId="3.[151,270,1886,1910]" captionTargetBox="[205,1398,1066,1837]" captionTargetId="figure@3.[151,1436,1037,1862]" captionTargetPageId="3" captionText="FIGURES 4 7. Sepedon hecate sp. nov., male terminalia. 4. Lateral view. 5. Hypandrium in lateral view. 6. Aedeagal complex in lateral view. 7. Epiphallus in caudal view. Scales: 0.3 mm. aa aedeagal apodeme, ae aedeagus, ce cercus, dph distiphallus, ea ejaculatory apodeme, ep epandrium, eph epiphallus, hy hypandrium, po postgonite, pr pregonite, ss surstylus." httpUri="https://zenodo.org/record/191319/files/figure.png" pageId="3" pageNumber="54">Fig. 6</figureCitation>
) consisting of aedeagus, pregonite, postgonite, ejaculatory apodeme and aedeagal apodeme as in other species of this family but the shape of aedeagus especially species-specific. Epiphallus (
<figureCitation id="130D2B91D56FD0085BAA0AF993B4FE79" box="[1198,1269,461,487]" captionStart="FIGURES 4 7" captionStartId="3.[151,270,1886,1910]" captionTargetBox="[205,1398,1066,1837]" captionTargetId="figure@3.[151,1436,1037,1862]" captionTargetPageId="3" captionText="FIGURES 4 7. Sepedon hecate sp. nov., male terminalia. 4. Lateral view. 5. Hypandrium in lateral view. 6. Aedeagal complex in lateral view. 7. Epiphallus in caudal view. Scales: 0.3 mm. aa aedeagal apodeme, ae aedeagus, ce cercus, dph distiphallus, ea ejaculatory apodeme, ep epandrium, eph epiphallus, hy hypandrium, po postgonite, pr pregonite, ss surstylus." httpUri="https://zenodo.org/record/191319/files/figure.png" pageId="3" pageNumber="54">Fig. 7</figureCitation>
) much higher than distiphallus in lateral view (cf.
<figureCitation id="130D2B91D56FD0085D3A0AC095C2FD90" box="[574,643,500,526]" captionStart="FIGURES 4 7" captionStartId="3.[151,270,1886,1910]" captionTargetBox="[205,1398,1066,1837]" captionTargetId="figure@3.[151,1436,1037,1862]" captionTargetPageId="3" captionText="FIGURES 4 7. Sepedon hecate sp. nov., male terminalia. 4. Lateral view. 5. Hypandrium in lateral view. 6. Aedeagal complex in lateral view. 7. Epiphallus in caudal view. Scales: 0.3 mm. aa aedeagal apodeme, ae aedeagus, ce cercus, dph distiphallus, ea ejaculatory apodeme, ep epandrium, eph epiphallus, hy hypandrium, po postgonite, pr pregonite, ss surstylus." httpUri="https://zenodo.org/record/191319/files/figure.png" pageId="3" pageNumber="54">Fig. 6</figureCitation>
) and at least one spine-like process distinct between them (actually this process is flat, leaf-shaped in caudal view). Epiphallus (
<figureCitation id="130D2B91D56FD0085C66092E94E8FDAA" box="[866,937,538,564]" captionStart="FIGURES 4 7" captionStartId="3.[151,270,1886,1910]" captionTargetBox="[205,1398,1066,1837]" captionTargetId="figure@3.[151,1436,1037,1862]" captionTargetPageId="3" captionText="FIGURES 4 7. Sepedon hecate sp. nov., male terminalia. 4. Lateral view. 5. Hypandrium in lateral view. 6. Aedeagal complex in lateral view. 7. Epiphallus in caudal view. Scales: 0.3 mm. aa aedeagal apodeme, ae aedeagus, ce cercus, dph distiphallus, ea ejaculatory apodeme, ep epandrium, eph epiphallus, hy hypandrium, po postgonite, pr pregonite, ss surstylus." httpUri="https://zenodo.org/record/191319/files/figure.png" pageId="3" pageNumber="54">Fig. 7</figureCitation>
) with produced oval apical part, in lateral view rectangularly curved distally.
</paragraph>
<paragraph id="8B893714D56FD0085FC1095C9315FC0E" blockId="3.[151,1437,152,990]" pageId="3" pageNumber="54">Female. Body length 8.39.3 mm; wing length 6.37.4 mm. No significant differences in external characters (except for abdominal tip) were found between the male and the female. In general (as far as it may be estimated from the limited specimens available) the female seems to be larger and slightly darker and the wing is somewhat broader than in the male. Tergites and sternites 6 and 7 are much broader than long, sternite 6 almost twice as broad as long and sternite 7 more than 2.5 times as long. Hypoproct transverse, somewhat produced posteriorly in middle third, distinctly narrower than sternite 7. Cercus short and oval, rounded posteriorly and more pointed proximally. Both spermathecae spherical, their diameter not more than the length of hypoproct in lateral part. One spermatheca indistinctly larger than other.</paragraph>
<paragraph id="8B893714D56FD0085FC108A9947DFC40" blockId="3.[151,1437,152,990]" pageId="3" pageNumber="54">
Etymology. We use here Elberg´s original collection name “
<taxonomicName id="4C364C97D56FD0085C8B08A9949DFC28" box="[911,988,925,950]" class="Insecta" family="Sciomyzidae" genus="Sepedon" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="hecate">
<emphasis id="B942EB06D56FD0085C8B08A9949DFC28" box="[911,988,925,950]" italics="true" pageId="3" pageNumber="54">hecate</emphasis>
</taxonomicName>
“ which refers to “
<taxonomicName id="4C364C97D56FD0085BB308A99250FC29" box="[1207,1297,925,951]" class="Insecta" family="Sciomyzidae" genus="Sepedon" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="59" phylum="Arthropoda" rank="species" species="hecate" status="sp. nov.">Hecate</taxonomicName>
“, the Greek goddess of magic, witchcraft, the night, moon and ghosts.
</paragraph>
</subSubSection>
<caption id="DF49679CD56FD0085F930C6A97BCF843" httpUri="https://zenodo.org/record/191319/files/figure.png" pageId="3" pageNumber="54" targetBox="[205,1398,1066,1837]" targetPageId="3">
<paragraph id="8B893714D56FD0085F930C6A97BCF843" blockId="3.[151,1436,1886,2013]" pageId="3" pageNumber="54">
<emphasis id="B942EB06D56FD0085F930C6A9605F8E8" bold="true" box="[151,324,1886,1910]" pageId="3" pageNumber="54">FIGURES 47.</emphasis>
<taxonomicName id="4C364C97D56FD0085E480C6A96B5F8EB" box="[332,500,1886,1909]" class="Insecta" family="Sciomyzidae" genus="Sepedon" kingdom="Animalia" order="Diptera" pageId="3" pageNumber="54" phylum="Arthropoda" rank="species" species="hecate" status="sp. nov.">
<emphasis id="B942EB06D56FD0085E480C6A96B5F8EB" box="[332,500,1886,1909]" italics="true" pageId="3" pageNumber="54">Sepedon hecate</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A271567DD56FD0085EF80C6A9512F8E8" box="[508,595,1886,1910]" pageId="3" pageNumber="54" rank="species">
<emphasis id="B942EB06D56FD0085EF80C6A9512F8E8" bold="true" box="[508,595,1886,1910]" pageId="3" pageNumber="54">sp. nov.</emphasis>
</taxonomicNameLabel>
, male terminalia. 4. Lateral view. 5. Hypandrium in lateral view. 6. Aedeagal complex in lateral view. 7. Epiphallus in caudal view. Scales: 0.3 mm. aa aedeagal apodeme, ae aedeagus, ce cercus, dph distiphallus, ea ejaculatory apodeme, ep epandrium, eph epiphallus, hy hypandrium, po postgonite, pr pregonite, ss surstylus.
</paragraph>
</caption>
<subSubSection id="C32C649FD568D00F5FC20BA39573FE5E" pageId="4" pageNumber="55" type="discussion">
<paragraph id="8B893714D568D00F5FC20BA39573FE5E" blockId="4.[151,1437,151,990]" pageId="4" pageNumber="55">
<emphasis id="B942EB06D568D00F5FC20BA39601FF2F" bold="true" box="[198,320,151,177]" pageId="4" pageNumber="55">Remarks.</emphasis>
The bare katatergite of
<taxonomicName id="4C364C97D568D00F5D5D0BAC9582FF2F" box="[601,707,152,178]" class="Insecta" family="Sciomyzidae" genus="Sepedon" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="hecate">
<emphasis id="B942EB06D568D00F5D5D0BAC9529FF2F" box="[601,616,152,177]" italics="true" pageId="4" pageNumber="55">S</emphasis>
.
<emphasis id="B942EB06D568D00F5D730BAC9582FF2F" box="[631,707,152,177]" italics="true" pageId="4" pageNumber="55">hecate</emphasis>
</taxonomicName>
shows that it belongs undoubtedly to the
<taxonomicName id="4C364C97D568D00F5BAA0BAC9211FF2F" box="[1198,1360,152,178]" class="Insecta" family="Sciomyzidae" genus="Sepedon" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="fuscipennis">
<emphasis id="B942EB06D568D00F5BAA0BAC93FCFF2F" box="[1198,1213,152,177]" italics="true" pageId="4" pageNumber="55">S</emphasis>
.
<emphasis id="B942EB06D568D00F5BC80BAC9211FF2F" box="[1228,1360,152,177]" italics="true" pageId="4" pageNumber="55">fuscipennis</emphasis>
</taxonomicName>
group although its face is bare, whereas so-called medifacial setulae may occur in some Nearctic members of this group. On the other hand
<taxonomicName id="4C364C97D568D00F5ECB0BD1957DFF60" box="[463,572,229,255]" class="Insecta" family="Sciomyzidae" genus="Sepedon" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="hecate">
<emphasis id="B942EB06D568D00F5ECB0BD1969FFF60" box="[463,478,229,254]" italics="true" pageId="4" pageNumber="55">S</emphasis>
.
<emphasis id="B942EB06D568D00F5EEA0BD1957DFF60" box="[494,572,229,254]" italics="true" pageId="4" pageNumber="55">hecate</emphasis>
</taxonomicName>
conspicuously differs from the Nearctic species by the species-specific shape of the hypandrium with the markedly developed basal frame appearing as a prominent basal lobe in lateral view (
<figureCitation id="130D2B91D568D00F5E360A06963AFED2" box="[306,379,306,332]" captionStart="FIGURES 4 7" captionStartId="3.[151,270,1886,1910]" captionTargetBox="[205,1398,1066,1837]" captionTargetId="figure@3.[151,1436,1037,1862]" captionTargetPageId="3" captionText="FIGURES 4 7. Sepedon hecate sp. nov., male terminalia. 4. Lateral view. 5. Hypandrium in lateral view. 6. Aedeagal complex in lateral view. 7. Epiphallus in caudal view. Scales: 0.3 mm. aa aedeagal apodeme, ae aedeagus, ce cercus, dph distiphallus, ea ejaculatory apodeme, ep epandrium, eph epiphallus, hy hypandrium, po postgonite, pr pregonite, ss surstylus." httpUri="https://zenodo.org/record/191319/files/figure.png" pageId="4" pageNumber="55">Fig. 5</figureCitation>
). According to the slender pedicel
<taxonomicName id="4C364C97D568D00F5C1E0A0694C4FED5" box="[794,901,306,332]" class="Insecta" family="Sciomyzidae" genus="Sepedon" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="hecate">
<emphasis id="B942EB06D568D00F5C1E0A069468FED5" box="[794,809,306,331]" italics="true" pageId="4" pageNumber="55">S</emphasis>
.
<emphasis id="B942EB06D568D00F5C3C0A0694C4FED5" box="[824,901,306,331]" italics="true" pageId="4" pageNumber="55">hecate</emphasis>
</taxonomicName>
resembles
<taxonomicName id="4C364C97D568D00F5B0E0A069381FED5" box="[1034,1216,306,332]" class="Insecta" family="Sciomyzidae" genus="Sepedon" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="gracilicornis">
<emphasis id="B942EB06D568D00F5B0E0A069358FED5" box="[1034,1049,306,331]" italics="true" pageId="4" pageNumber="55">S</emphasis>
.
<emphasis id="B942EB06D568D00F5B2C0A069381FED5" box="[1064,1216,306,331]" italics="true" pageId="4" pageNumber="55">gracilicornis</emphasis>
</taxonomicName>
and
<taxonomicName id="4C364C97D568D00F5BFE0A0692DDFED5" box="[1274,1436,306,332]" class="Insecta" family="Sciomyzidae" genus="Sepedon" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="tenuicornis">
<emphasis id="B942EB06D568D00F5BFE0A069249FED5" box="[1274,1288,306,331]" italics="true" pageId="4" pageNumber="55">S</emphasis>
.
<emphasis id="B942EB06D568D00F5A1C0A0692DDFED5" box="[1304,1436,306,331]" italics="true" pageId="4" pageNumber="55">tenuicornis</emphasis>
</taxonomicName>
but its specific structures of the male terminalia are distinctly different from all Nearctic species of the group (see
<figureCitation id="130D2B91D568D00F5FD40AB4960DFE04" box="[208,332,384,410]" captionStart="FIGURES 8 19" captionStartId="5.[151,270,1420,1444]" captionTargetBox="[185,1388,691,1404]" captionTargetId="figure@5.[151,1436,682,1414]" captionTargetPageId="5" captionText="FIGURES 8 19. Nearctic species of Sepedon fuscipennis group, details of male terminalia. 8, 12, 16. S. floridensis. 9, 13, 17. S. fuscipennis. 10, 14, 18. S. gracilicornis. 11, 15, 19. S. tenuicornis. 8 11. Hypandrium in lateral view. 12 15. Aedeagus in lateral view. 16 19. Epiphallus in caudal view. Scale: 0.3 mm." httpUri="https://zenodo.org/record/191320/files/figure.png" pageId="4" pageNumber="55">Figs 819</figureCitation>
). We did not succeed in finding reliable differences in the female terminalia for separating species of the
<taxonomicName id="4C364C97D568D00F5E390A93969EFE5E" box="[317,479,422,448]" class="Insecta" family="Sciomyzidae" genus="Sepedon" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="fuscipennis">
<emphasis id="B942EB06D568D00F5E390A93960DFE5E" box="[317,332,423,448]" italics="true" pageId="4" pageNumber="55">S</emphasis>
.
<emphasis id="B942EB06D568D00F5E5F0A93969EFE5E" box="[347,479,423,448]" italics="true" pageId="4" pageNumber="55">fuscipennis</emphasis>
</taxonomicName>
group.
</paragraph>
</subSubSection>
<subSubSection id="C32C649FD568D00F5FC20AF9967AFD1C" pageId="4" pageNumber="55" type="distribution">
<paragraph id="8B893714D568D00F5FC20AF9967AFD1C" blockId="4.[151,1437,151,990]" pageId="4" pageNumber="55">
<emphasis id="B942EB06D568D00F5FC20AF99626FE79" bold="true" box="[198,359,461,487]" pageId="4" pageNumber="55">Distribution.</emphasis>
<taxonomicName id="4C364C97D568D00F5E6B0AF9969AFE78" box="[367,475,461,487]" class="Insecta" family="Sciomyzidae" genus="Sepedon" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="hecate">
<emphasis id="B942EB06D568D00F5E6B0AF9963FFE78" box="[367,382,461,486]" italics="true" pageId="4" pageNumber="55">S</emphasis>
.
<emphasis id="B942EB06D568D00F5E8A0AF9969AFE78" box="[398,475,461,486]" italics="true" pageId="4" pageNumber="55">hecate</emphasis>
</taxonomicName>
is presumably of Nearctic origin although became established exclusively in the Palaearctic Region. The present knowledge of distribution of the new species indicates its penetration from eastern Siberia to Central Europe. Possibly a careful review of
<taxonomicName id="4C364C97D568D00F5C8B092E94B5FDAD" box="[911,1012,538,563]" class="Insecta" family="Sciomyzidae" genus="Sepedon" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="genus">
<emphasis id="B942EB06D568D00F5C8B092E94B5FDAD" box="[911,1012,538,563]" italics="true" pageId="4" pageNumber="55">Sepedon</emphasis>
</taxonomicName>
specimens in European collections will provide further evidence of this species in Europe in addition to the unique known record from the
<collectingCountry id="F3217784D568D00F5F93095C97B4FD1C" box="[151,245,616,642]" name="Germany" pageId="4" pageNumber="55">German</collectingCountry>
Alps.
</paragraph>
</subSubSection>
<subSubSection id="C32C649FD568D00F5FC209BA9619FC40" pageId="4" pageNumber="55" type="materials_examined">
<paragraph id="8B893714D568D00F5FC209BA9617FD68" blockId="4.[151,1437,151,990]" pageId="4" pageNumber="55">
<emphasis id="B942EB06D568D00F5FC209BA96C0FD36" bold="true" box="[198,385,654,680]" pageId="4" pageNumber="55">
<typeStatus id="548D89B6D568D00F5FC209BA9642FD36" box="[198,259,654,680]" pageId="4" pageNumber="55">Type</typeStatus>
material:
</emphasis>
<typeStatus id="548D89B6D568D00F5E8809BA96BFFD36" box="[396,510,654,680]" pageId="4" pageNumber="55" type="holotype">Holotype</typeStatus>
: 3,
<collectingCountry id="F3217784D568D00F5D3709BA95E1FD36" box="[563,672,654,680]" name="Russia" pageId="4" pageNumber="55">RUSSIA</collectingCountry>
: Amur region, Klimoucy,
<quantity id="4CCE9AF1D568D00F5CDB09BA936DFD36" box="[991,1068,654,680]" metricMagnitude="4" metricUnit="m" metricValue="4.0" pageId="4" pageNumber="55" unit="km" value="40.0">40 km</quantity>
W of Svobodniy,
<date id="FF8811D4D568D00F5A0B09BA92D7FD36" box="[1295,1430,654,680]" pageId="4" pageNumber="55" value="1958-07-15">15.vii.1958</date>
, Borisova leg., provided with a red label „
<typeStatus id="548D89B6D568D00F5D8A09819440FD51" box="[654,769,693,719]" pageId="4" pageNumber="55" type="holotype">Holotype</typeStatus>
<taxonomicName id="4C364C97D568D00F5C0E09819489FD50" box="[778,968,693,718]" class="Insecta" family="Sciomyzidae" genus="Sepedon" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="hecate" status="sp. nov.">
<emphasis id="B942EB06D568D00F5C0E09819489FD50" box="[778,968,693,718]" italics="true" pageId="4" pageNumber="55">Sepedon hecate</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A271567DD568D00F5CD409819367FD51" box="[976,1062,693,719]" pageId="4" pageNumber="55" rank="species">sp.nov.</taxonomicNameLabel>
Elberg, Rozkošný et Knutson, 2009“ (
<collectionCode id="ED27AFD1D568D00F5FEA09E89608FD68" LSID="urn:lsid:biocol.org:col:34871" box="[238,329,732,758]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="55">USNM</collectionCode>
).
</paragraph>
<paragraph id="8B893714D568D00F5FC208369619FC40" blockId="4.[151,1437,151,990]" pageId="4" pageNumber="55">
<typeStatus id="548D89B6D568D00F5FC20836967CFC82" box="[198,317,770,796]" pageId="4" pageNumber="55" type="paratype">Paratypes</typeStatus>
:
<date id="FF8811D4D568D00F5E4F083696E8FC82" box="[331,425,770,796]" pageId="4" pageNumber="55" value="1911-07-03">
7 3,
<specimenCount id="9D30FC9DD568D00F5E89083696E8FC82" box="[397,425,770,796]" pageId="4" pageNumber="55" type="generic">11</specimenCount>
</date>
Ƥ. 1 Ƥ, as
<typeStatus id="548D89B6D568D00F5D45083695E6FC82" box="[577,679,770,796]" pageId="4" pageNumber="55" type="holotype">holotype</typeStatus>
but
<date id="FF8811D4D568D00F5DE608369433FC82" box="[738,882,770,796]" pageId="4" pageNumber="55" value="1958-08-31">31.viii.1958</date>
, Zinoviev (
<collectionCode id="ED27AFD1D568D00F5CFB0836931AFC82" LSID="urn:lsid:biocol.org:col:34871" box="[1023,1115,770,796]" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34871" name="Smithsonian Institution, National Museum of Natural History" pageId="4" pageNumber="55">USNM</collectionCode>
).
<collectingCountry id="F3217784D568D00F5B750836939FFC82" box="[1137,1246,770,796]" name="Russia" pageId="4" pageNumber="55">RUSSIA</collectingCountry>
: Magadan area, Seimchan,
<date id="FF8811D4D568D00F5E1A081D96F2FCDD" box="[286,435,809,835]" pageId="4" pageNumber="55" value="1972-08-16">16.viii.1972</date>
, 5 Ƥ; Khabarovsk area, P. Osipenko,
<date id="FF8811D4D568D00F5C7A081D934DFCDD" box="[894,1036,809,835]" pageId="4" pageNumber="55" value="1973-07-29">29.vii.1973</date>
, 1 3,
<date id="FF8811D4D568D00F5B53081D9323FCDD" box="[1111,1122,809,835]" pageId="4" pageNumber="55" value="1973-08-01">1</date>
.viii.1973, 1 Ƥ; Manoma,
<date id="FF8811D4D568D00F5F9308649674FCF4" box="[151,309,848,874]" pageId="4" pageNumber="55" value="1973-08-29">29.viii.1973</date>
, 1 3, all K. Elberg (
<collectionCode id="ED27AFD1D568D00F5D38086495CAFCF4" box="[572,651,848,874]" pageId="4" pageNumber="55">IZHT</collectionCode>
). Amur region, Zeya,
<date id="FF8811D4D568D00F5CA20864937BFCF4" box="[934,1082,848,874]" pageId="4" pageNumber="55" value="1981-07-16">16.vii.1981</date>
, 1 3,
<date id="FF8811D4D568D00F5B8E086493EAFCF4" box="[1162,1195,848,874]" pageId="4" pageNumber="55" value="1981-09-12">12</date>
.ix.1981, 1 3, 1 Ƥ,
<date id="FF8811D4D568D00F5F9308429656FC0E" box="[151,279,886,912]" pageId="4" pageNumber="55" value="1981-09-13">13.ix.1981</date>
, 2 3, 3 Ƥ, all A. Shatalkin (
<collectionCode id="ED27AFD1D568D00F5D6008429587FC0E" box="[612,710,886,912]" httpUri="http://grbio.org/cool/k3qp-615s" name="Zoological Museum, Moscow Lomonosov State University" pageId="4" pageNumber="55">ZMMU</collectionCode>
).
<collectingCountry id="F3217784D568D00F5DDE08429431FC0E" box="[730,880,886,912]" name="Germany" pageId="4" pageNumber="55">GERMANY</collectingCountry>
: Oberbayern, Schluifeldt Moos,
<date id="FF8811D4D568D00F5BF4084292C1FC0E" box="[1264,1408,886,912]" pageId="4" pageNumber="55" value="1935-08-27">27.viii.1935</date>
, 1 3 (
<collectionCode id="ED27AFD1D568D00F5FB908A99640FC29" box="[189,257,925,951]" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="4" pageNumber="55">CNC</collectionCode>
). Each
<typeStatus id="548D89B6D568D00F5E5E08A99680FC29" box="[346,449,925,951]" pageId="4" pageNumber="55" type="paratype">paratype</typeStatus>
provided with a red label „
<typeStatus id="548D89B6D568D00F5C0C08A99432FC29" box="[776,883,925,951]" pageId="4" pageNumber="55" type="paratype">Paratype</typeStatus>
<taxonomicName id="4C364C97D568D00F5C7808A99379FC28" box="[892,1080,925,950]" class="Insecta" family="Sciomyzidae" genus="Sepedon" kingdom="Animalia" order="Diptera" pageId="4" pageNumber="55" phylum="Arthropoda" rank="species" species="hecate" status="sp. nov.">
<emphasis id="B942EB06D568D00F5C7808A99379FC28" box="[892,1080,925,950]" italics="true" pageId="4" pageNumber="55">Sepedon hecate</emphasis>
</taxonomicName>
<taxonomicNameLabel id="A271567DD568D00F5B4408A993DEFC29" box="[1088,1183,925,951]" pageId="4" pageNumber="55" rank="species">
<emphasis id="B942EB06D568D00F5B4408A993DEFC29" bold="true" box="[1088,1183,925,951]" pageId="4" pageNumber="55">sp. nov.</emphasis>
</taxonomicNameLabel>
Elberg, Rozkošný et Knutson, 2009“.
</paragraph>
</subSubSection>
</treatment>
</document>