573 lines
97 KiB
XML
573 lines
97 KiB
XML
<document id="5B9DC1423D2EB3C80CDA3E7F547B4502" ID-ISSN="1536-9307" IM.materialsCitations_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1638258019449" checkinUser="felipe" docAuthor="Kovařík, František, Lowe, Graeme, Just, Pavel, Awale, Ahmed Ibrahim & Sh, Hassan" docDate="2018" docId="03842B1A795DCB06FEE6FB67EC1DFC30" docLanguage="en" docName="Euscorpius.259.1-41.pdf" docOrigin="Euscorpius 259" docStyle="DocumentStyle:05AD4CDCECA6C90C60787CEF3EC4672A.3:Euscorpius.2002-2019.journal_article" docStyleId="05AD4CDCECA6C90C60787CEF3EC4672A" docStyleName="Euscorpius.2002-2019.journal_article" docStyleVersion="3" docTitle="Gint amoudensis Kovarik 2018, sp. n." docType="treatment" docVersion="5" lastPageNumber="18" masterDocId="FFBD53627950CB15FFE8FFBDE816FFCB" masterDocTitle="Scorpions of the Horn of Africa (Arachnida: Scorpiones) Part XV Review of the genus Gint Kova ík et al 2013, with description of three new species from Somaliland (Scorpiones Buthidae)" masterLastPageNumber="41" masterPageNumber="1" pageNumber="12" updateTime="1699436189298" updateUser="plazi" zenodo-license-document="CC-BY-4.0" zenodo-license-figures="CC-BY-4.0">
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<mods:title id="ACD59C2042C4AEC1342F4118BA5DBD49">Scorpions of the Horn of Africa (Arachnida: Scorpiones) Part XV Review of the genus Gint Kova ík et al 2013, with description of three new species from Somaliland (Scorpiones Buthidae)</mods:title>
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<mods:namePart id="55F1FB3E8E2BAB93AD43D58111FA37C0">Kovařík, František</mods:namePart>
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<mods:namePart id="AC7FFDDF4B5708B2BF694225880C6ABF">Lowe, Graeme</mods:namePart>
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<mods:namePart id="5C182D1EC9D3779D316A186BA9FCED54">Just, Pavel</mods:namePart>
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<mods:namePart id="374C03AE88F64C02BDACC1EB7A096072">Awale, Ahmed Ibrahim</mods:namePart>
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<mods:namePart id="AFA8D2B884AFDD26C85098E5773A3CFD">Sh, Hassan</mods:namePart>
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<mods:title id="76B51BB3535CE454CE54C5C0864E6ED2">Euscorpius</mods:title>
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<mods:date id="BE66C8E89251C941061D46A1B6BF3095">2018</mods:date>
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<taxonomicName id="4C2DE18F795DCB18FEE6FB67EA92FB3F" ID-CoL="3G3HX" authority="Kovarik et al." authorityName="Kovarik" authorityYear="2018" box="[270,644,1241,1268]" class="Arachnida" family="Buthidae" genus="Gint" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="species" species="amoudensis" status="sp. nov.">
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<emphasis id="B959461E795DCB18FEE6FB67E9CCFB3F" bold="true" box="[270,474,1241,1268]" italics="true" pageId="13" pageNumber="12">Gint amoudensis</emphasis>
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Kovařík et al.
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,
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.
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<emphasis id="B959461E795DCB18FD54FB67EADAFB3F" bold="true" box="[700,716,1242,1268]" pageId="13" pageNumber="12">n</emphasis>
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.
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<paragraph id="8B929A0C795DCB18FF1EFB46EB0AFA9B" blockId="13.[197,796,1241,1391]" pageId="13" pageNumber="12">
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(
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<figureCitation id="13168689795DCB18FF17FB46E979FAD9" box="[255,367,1275,1298]" captionStart-0="Figures 1–2" captionStart-1="Figures 3–6" captionStart-2="Figures 7–10" captionStart-3="Figures 11–20" captionStart-4="Figures 21–28" captionStart-5="Figures 29–45" captionStart-6="Figures 46–47" captionStart-7="Figure 48" captionStartId-0="4.[192,278,1854,1877]" captionStartId-1="5.[192,278,1766,1789]" captionStartId-2="6.[192,278,1561,1584]" captionStartId-3="7.[192,278,1254,1277]" captionStartId-4="8.[192,278,1653,1676]" captionStartId-5="9.[192,278,1694,1717]" captionStartId-6="10.[192,278,1843,1866]" captionStartId-7="11.[192,268,1329,1352]" captionTargetBox-0="[209,1417,191,1825]" captionTargetBox-1="[212,1410,199,1716]" captionTargetBox-2="[207,1427,230,1503]" captionTargetBox-3="[191,1437,217,1225]" captionTargetBox-4="[207,1436,194,1589]" captionTargetBox-5="[201,1435,218,1644]" captionTargetBox-6="[191,1440,191,1813]" captionTargetBox-7="[191,1917,227,1297]" captionTargetId-0="figure-12@4.[210,1416,192,1824]" captionTargetId-1="figure-6@5.[192,1439,182,1728]" captionTargetId-2="figure-12@6.[192,1439,192,1531]" captionTargetId-3="figure-6@7.[192,1437,218,1224]" captionTargetId-4="figure-12@8.[192,1439,192,1623]" captionTargetId-5="figure-6@9.[192,1439,212,1656]" captionTargetId-6="figure-12@10.[192,1439,192,1812]" captionTargetId-7="figure-6@11.[192,1916,228,1296]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionTargetPageId-4="8" captionTargetPageId-5="9" captionTargetPageId-6="10" captionTargetPageId-7="11" captionText-0="Figures 1–2: Gint amoudensis sp. n., paratypes from locality No. 17SF (Laas Gel), in vivo habitus. Figure 1. Female and two males. Figure 2. Male." captionText-1="Figures 3–6: Gint amoudensis sp. n. from type locality. Figures 3–4. Holotype male, dorsal (3) and ventral (4) views. Figures 5–6. Paratype female, dorsal (5) and ventral (6) views. Scale bar: 10 mm." captionText-2="Figures 7–10: Gint amoudensis sp. n., paratypes from Laas Gel. Figures 7–8. Male, dorsal (7) and ventral (8) views. Figures 9–10. Female, dorsal (9) and ventral (10) views. Scale bar: 10 mm." captionText-3="Figures 11–20: Gint amoudensis sp. n. Figures 11, 13, 17–20. Holotype male, chelicerae, carapace and tergites I–III (11), sternopectinal region and sternites III–V (13), distal segments of right legs I–IV, retrolateral views (17–20 respectively). Figures 12, 14. Paratype female from type locality, chelicerae, carapace and tergites I–III (12), sternopectinal region and sternites III–VI (14). Figure 15. Paratype male from Laas Gel, chelicerae, carapace and tergites I–III. Figure 16. Paratype female from Laas Gel, chelicerae, carapace and tergites I–III." captionText-4="Figures 21–28: Gint amoudensis sp. n. Figures 21, 23–25. Holotype male, metasoma V and telson lateral view (21), metasoma and telson, lateral (23), dorsal (24), and ventral (25) views. Figures 22, 26–28. Paratype female from type locality, metasoma V and telson lateral view (22), metasoma and telson, lateral (26), dorsal (27), and ventral (28) views. Scale bar: 10 mm (23–28)." captionText-5="Figures 29–45: Gint amoudensis sp. n. from type locality. Figures 29–38. Holotype male. Pedipalp chela, dorsoexternal (29), external (30), and ventrointernal (31) views. Pedipalp patella, dorsal (32), external (33), and ventral (34) views. Pedipalp femur and trochanter, internal (35) and dorsal (36) views. Pedipalp chela, movable (37) and fixed (38) finger dentate margins. Figures 39–45. Paratype female. Pedipalp chela, dorsoexternal (39), external (40), and ventrointernal (41) views. Pedipalp patella, dorsal (42), external (43), and ventral (44) views. Pedipalp femur and trochanter dorsal (45) view. The trichobothrial pattern is indicated in Figures 29–36 (white circles)." captionText-6="Figures 46–47: Gint amoudensis sp. n., paratype male from type locality, in vivo habitus (46) and the type locality (47)." captionText-7="Figure 48: Gint amoudensis sp. n., locality No. 17SF, Laas Gel, 50 km NE Hargeisa." pageId="13" pageNumber="12">Figs. 1–48</figureCitation>
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,
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<figureCitation id="13168689795DCB18FE95FB46E9CBFAD9" box="[381,477,1275,1298]" captionStart="Figures 121–128" captionStartId="30.[192,278,1662,1685]" captionTargetBox="[213,1423,188,1621]" captionTargetId="figure-12@30.[192,1439,173,1632]" captionTargetPageId="30" captionText="Figures 121–128: Metasoma and telson, ventral views, comparison of males of Gint species. Figure 121. Gint puntlandus, holotype. Figure 122. Gint dabakalo, holotype. Figure 123. Gint gubanensis sp. n., holotype. Figure 124. Gint maidensis sp. n., holotype. Figure 125. Gint calviceps, Somaliland, south of Qool-Cadday, between Hargeisa and Salahle, Woqooyi Galbeed, 09°11'56"N 44°09'50"E. Figure 126. Gint gaitako, Ethiopia, Oromia State, Sidamo Province, Wachile env., 115 km N Moyale, 04°32'34"N 39° 03'08"E, 1073 m a. s. l. Figures 127–128. Gint amoudensis sp. n., from Laas Gel (127) and holotype from type locality (128)." pageId="13" pageNumber="12">127–128</figureCitation>
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,
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<figureCitation id="13168689795DCB18FE0FFB46EA53FAD9" box="[487,581,1275,1298]" captionStart="Figures 129–147" captionStartId="31.[192,278,1694,1717]" captionTargetBox="[199,1437,219,1638]" captionTargetId="figure-6@31.[192,1440,202,1656]" captionTargetPageId="31" captionText="Figures 129–147: Telson, lateral views of both sexes and metasoma IV ventral views of males, comparison of Gint species. 129–131. Gint amoudensis sp. n., holotype (129, 131) and female from type locality (130). 132–134. Gint calviceps, male from Somaliland, south of Qool-Cadday, between Hargeisa and Salahle, Woqooyi Galbeed, 09°11'56"N 44°09'50"E (132, 134) and female from locality 17SE (133). 135–137. Gint dabakalo, holotype (135, 137) and paratype (136). 138–140. Gint gaitako, male (138, 140) and female (139) from Ethiopia, Oromia State, Sidamo Province, Wachile env., 115 km N Moyale, 04°32'34"N 39° 03'08"E, 1073 m a. s. l. 141–142. Gint gubanensis sp. n., holotype. 143–145. Gint maidensis sp. n., holotype (143, 145) and paratype (144). 146–147. Gint puntlandus, holotype." pageId="13" pageNumber="12">129–131</figureCitation>
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,
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<figureCitation id="13168689795DCB18FDBAFB46EAB9FAD9" box="[594,687,1275,1298]" captionStart="Figures 148–163" captionStartId="33.[192,278,1689,1712]" captionTargetBox="[191,1440,191,1660]" captionTargetId="figure-6@33.[192,1439,192,1659]" captionTargetPageId="33" captionText="Figures 148–163: Figures 148–151: Gint amoudensis sp. n., paratype male, No. 1327 from Laas Gel (Locality No. 17SF), right hemispermatophore. 148. Whole hemispermatophore. 149–151. Capsule region. Views of posterior (flagellar) margin (149), convex side (150), and anterior (sperm duct) margin (151). Scale bars: 1 mm (148), 200 µm (149–151). Figures 152–163: Gint amoudensis sp. n., capsule region of right hemispermatophores from four additional paratype males, showing morphological variation. Nos. 1291 (158–160), 1292 (152–154) and 1293 (161–163) from Laas Gel (Locality No. 17SF). No. 1325 (155–157) from Borama, Armoud University (Locality No. 17SR). Views of posterior (flagellar) margin (152, 155, 158, 161), convex side (153, 156, 159, 162), and anterior (sperm duct) margin (154, 157, 160, 163). Scale bars: 200 µm (152–154; 155–157; 158–160; 161–163)." pageId="13" pageNumber="12">148–163</figureCitation>
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,
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<figureCitation id="13168689795DCB18FD55FB46EAF3FAD9" box="[701,741,1275,1298]" captionStart="Figures 195–201" captionStartId="37.[192,278,1197,1220]" captionTargetBox="[191,1440,191,1168]" captionTargetId="figure-6@37.[192,1440,192,1167]" captionTargetPageId="37" captionText="Figures 195–201: Figures 195–197: Profiles in distal view, along the longitudinal axis, of the convex surfaces at the level of the basal lobe of the right hemispermatophores of three species of Gint, showing different development of the basal lobe. 195. Gint gaitako Kovařík et al., 2013, No. 958 from Wachile (Locality No. 16EH). 196. Gint amoudensis sp. n., paratype male, No. 1292 from Laas Gel (Locality No. 17SF). 197. Gint maidensis sp. n., No. 1321 paratype male from Maid (Type locality No. 17SN). Abbreviations: b, basal lobe, f, flagellum. Figures 198–201: Mitotic metaphases of Gint spp. males. 198. G. dabakalo from locality 17SH (2n=23). 199. G. amoudensis sp. n., paratype from Laas Gel (2n=35). 200. G. amoudensis sp. n., paratype from type locality (2n=36). 201. G. maidensis sp. n., paratype (2n=34). Scale bar: 5 µm." pageId="13" pageNumber="12">196</figureCitation>
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,
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<figureCitation id="13168689795DCB18FE72FAA7E9D3FAFA" box="[410,453,1306,1329]" captionStart="Figure 202" captionStartId="38.[192,268,1190,1213]" captionTargetBox="[196,1396,196,1156]" captionTargetId="figure-12@38.[192,1399,192,1160]" captionTargetPageId="38" captionText="Figure 202: Map showing confirmed distribution of Gint spp. Points indicate sites sampled during 2011–2017 expeditions and type locality of G. puntlandus. Scorpion shown in the photo inset is a female paratype of G. maidensis sp. n." pageId="13" pageNumber="12">202</figureCitation>
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,
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<tableCitation id="C6AFAFB7795DCB18FE21FAA7EA32FAFA" box="[457,548,1306,1329]" captionStart="Table 1" captionStartId="12.[192,257,1044,1067]" captionText="Table 1: Comparative measurements of adults of Gint amoundensis sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to" pageId="13" pageNumber="12">Tables 1</tableCitation>
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,
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<tableCitation id="C6AFAFB7795DCB18FDD9FAA7EA28FAFA" box="[561,574,1306,1329]" captionStart="Table 3" captionStartId="29.[192,257,1072,1095]" captionText="Table 3: Comparison among Gint species (specimens), based upon selected morphometric ratios of adult males. Abbreviations: length (L), width (W), depth (D)." pageId="13" pageNumber="12">3</tableCitation>
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)
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<emphasis id="B959461E795DCB18FF2DFA84EB0AFA9B" bold="true" box="[197,796,1337,1360]" pageId="13" pageNumber="12">http://zoobank.org/urn:lsid:zoobank.org:act:51A060</emphasis>
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<emphasis id="B959461E795DCB18FEFBFAE5EAD9FAA4" bold="true" box="[275,719,1368,1391]" pageId="13" pageNumber="12">CE-A0CA-4B30-94D7-B6861DC5D364</emphasis>
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<paragraph id="8B929A0C795DCB18FF28FA28EA59FA67" blockId="13.[192,591,1429,1452]" box="[192,591,1429,1452]" pageId="13" pageNumber="12">
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<taxonomicName id="4C2DE18F795DCB18FF28FA28E8E6FA67" authority="Kovařík & Lowe & Just & Awale & Sh, 2018" authorityName="Kovařík & Lowe & Just & Awale & Sh" authorityYear="2018" box="[192,240,1429,1452]" class="Arachnida" family="Buthidae" genus="Gint" kingdom="Animalia" order="Scorpiones" pageId="13" pageNumber="12" phylum="Arthropoda" rank="genus" status="sp. nov.">
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<emphasis id="B959461E795DCB18FF28FA28E8E6FA67" box="[192,240,1429,1452]" italics="true" pageId="13" pageNumber="12">Gint</emphasis>
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<taxonomicNameLabel id="A26AFB65795DCB18FF1EFA28E939FA67" box="[246,303,1429,1452]" pageId="13" pageNumber="12" rank="species">sp. n.</taxonomicNameLabel>
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: Kovařík et al., 2017b: 18.
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<emphasis id="B959461E795DCB18FD72FA6FEB0DFA22" bold="true" box="[666,795,1490,1513]" pageId="13" pageNumber="12">Somaliland</emphasis>
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, Borama, Amoud University campus,
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;
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.
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<collectionCode id="ED3C02C9795DCB18FF28F9F0EA1DF9A8" box="[192,523,1613,1636]" pageId="13" pageNumber="12">TYPE MATERIAL EXAMINED</collectionCode>
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.
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<collectingCounty id="62F3E280795DCB18FDC4F9F0EAB0F9AF" box="[556,678,1613,1636]" pageId="13" pageNumber="12">Somaliland</collectingCounty>
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campus,
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<quantity id="4CD537E9795DCB18FF28F937E900F96A" box="[192,278,1674,1697]" metricMagnitude="3" metricUnit="m" metricValue="1.394" pageId="13" pageNumber="12" unit="m" value="1394.0">1394 m</quantity>
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a.s.l.
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(
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<figureCitation id="13168689795DCB18FE8CF937E9A0F96A" box="[356,438,1674,1697]" captionStart="Figure 48" captionStartId="11.[192,268,1329,1352]" captionTargetBox="[191,1917,227,1297]" captionTargetId="figure-6@11.[192,1916,228,1296]" captionTargetPageId="11" captionText="Figure 48: Gint amoudensis sp. n., locality No. 17SF, Laas Gel, 50 km NE Hargeisa." pageId="13" pageNumber="12">Fig. 48</figureCitation>
|
||
,
|
||
<collectorName id="26D8FFDA795DCB18FE2FF937EA45F96A" box="[455,595,1674,1697]" pageId="13" pageNumber="12">Locality No.</collectorName>
|
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<emphasis id="B959461E795DCB18FDB5F937EA8CF96A" bold="true" box="[605,666,1674,1697]" pageId="13" pageNumber="12">17SR</emphasis>
|
||
=17
|
||
<collectionCode id="ED3C02C9795DCB18FD27F937EAE2F96A" box="[719,756,1674,1697]" country="France" name="Museum national d'Histoire Naturelle, Laboratiore de Paleontologie" pageId="13" pageNumber="12">SA</collectionCode>
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)
|
||
</materialsCitation>
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,
|
||
<date id="FF93BCCC795DCB18FCE3F937E92AF90B" pageId="13" pageNumber="12" value="2017-09-09" valueMax="2017-09-13" valueMin="2017-09-09">9- 13.IX.2017</date>
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,
|
||
<specimenCount id="9D2B5185795DCB18FEA2F914E97DF90B" box="[330,363,1705,1728]" pageId="13" pageNumber="12" type="male">7♂</specimenCount>
|
||
<specimenCount id="9D2B5185795DCB18FE83F914E9B8F90B" box="[363,430,1705,1728]" pageId="13" pageNumber="12" type="female">3♀3♀</specimenCount>
|
||
juvs.
|
||
<specimenCount id="9D2B5185795DCB18FE09F914EA14F90B" box="[481,514,1705,1728]" pageId="13" pageNumber="12" type="male">2♂</specimenCount>
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juvs. (
|
||
<typeStatus id="549624AE795DCB18FDAFF914EAB1F90B" box="[583,679,1705,1728]" pageId="13" pageNumber="12" type="holotype">holotype</typeStatus>
|
||
,
|
||
<figureCitation id="13168689795DCB18FD5DF914EB0DF90B" box="[693,795,1705,1728]" captionStart="Figures 3–6" captionStartId="5.[192,278,1766,1789]" captionTargetBox="[212,1410,199,1716]" captionTargetId="figure-6@5.[192,1439,182,1728]" captionTargetPageId="5" captionText="Figures 3–6: Gint amoudensis sp. n. from type locality. Figures 3–4. Holotype male, dorsal (3) and ventral (4) views. Figures 5–6. Paratype female, dorsal (5) and ventral (6) views. Scale bar: 10 mm." pageId="13" pageNumber="12">Figs. 3–4</figureCitation>
|
||
,
|
||
<figureCitation id="13168689795DCB18FF28F97AE9BAF915" box="[192,428,1735,1758]" captionStart-0="Figures 11–20" captionStart-1="Figures 21–28" captionStartId-0="7.[192,278,1254,1277]" captionStartId-1="8.[192,278,1653,1676]" captionTargetBox-0="[191,1437,217,1225]" captionTargetBox-1="[207,1436,194,1589]" captionTargetId-0="figure-6@7.[192,1437,218,1224]" captionTargetId-1="figure-12@8.[192,1439,192,1623]" captionTargetPageId-0="7" captionTargetPageId-1="8" captionText-0="Figures 11–20: Gint amoudensis sp. n. Figures 11, 13, 17–20. Holotype male, chelicerae, carapace and tergites I–III (11), sternopectinal region and sternites III–V (13), distal segments of right legs I–IV, retrolateral views (17–20 respectively). Figures 12, 14. Paratype female from type locality, chelicerae, carapace and tergites I–III (12), sternopectinal region and sternites III–VI (14). Figure 15. Paratype male from Laas Gel, chelicerae, carapace and tergites I–III. Figure 16. Paratype female from Laas Gel, chelicerae, carapace and tergites I–III." captionText-1="Figures 21–28: Gint amoudensis sp. n. Figures 21, 23–25. Holotype male, metasoma V and telson lateral view (21), metasoma and telson, lateral (23), dorsal (24), and ventral (25) views. Figures 22, 26–28. Paratype female from type locality, metasoma V and telson lateral view (22), metasoma and telson, lateral (26), dorsal (27), and ventral (28) views. Scale bar: 10 mm (23–28)." pageId="13" pageNumber="12">11, 13, 17–21, 23–25</figureCitation>
|
||
,
|
||
<figureCitation id="13168689795DCB18FE54F97AEA17F915" box="[444,513,1735,1758]" captionStart="Figures 29–45" captionStartId="9.[192,278,1694,1717]" captionTargetBox="[201,1435,218,1644]" captionTargetId="figure-6@9.[192,1439,212,1656]" captionTargetPageId="9" captionText="Figures 29–45: Gint amoudensis sp. n. from type locality. Figures 29–38. Holotype male. Pedipalp chela, dorsoexternal (29), external (30), and ventrointernal (31) views. Pedipalp patella, dorsal (32), external (33), and ventral (34) views. Pedipalp femur and trochanter, internal (35) and dorsal (36) views. Pedipalp chela, movable (37) and fixed (38) finger dentate margins. Figures 39–45. Paratype female. Pedipalp chela, dorsoexternal (39), external (40), and ventrointernal (41) views. Pedipalp patella, dorsal (42), external (43), and ventral (44) views. Pedipalp femur and trochanter dorsal (45) view. The trichobothrial pattern is indicated in Figures 29–36 (white circles)." pageId="13" pageNumber="12">29–38</figureCitation>
|
||
,
|
||
<figureCitation id="13168689795DCB18FDE7F97AEAB2F915" box="[527,676,1735,1758]" captionStart-0="Figures 121–128" captionStart-1="Figures 129–147" captionStartId-0="30.[192,278,1662,1685]" captionStartId-1="31.[192,278,1694,1717]" captionTargetBox-0="[213,1423,188,1621]" captionTargetBox-1="[199,1437,219,1638]" captionTargetId-0="figure-12@30.[192,1439,173,1632]" captionTargetId-1="figure-6@31.[192,1440,202,1656]" captionTargetPageId-0="30" captionTargetPageId-1="31" captionText-0="Figures 121–128: Metasoma and telson, ventral views, comparison of males of Gint species. Figure 121. Gint puntlandus, holotype. Figure 122. Gint dabakalo, holotype. Figure 123. Gint gubanensis sp. n., holotype. Figure 124. Gint maidensis sp. n., holotype. Figure 125. Gint calviceps, Somaliland, south of Qool-Cadday, between Hargeisa and Salahle, Woqooyi Galbeed, 09°11'56"N 44°09'50"E. Figure 126. Gint gaitako, Ethiopia, Oromia State, Sidamo Province, Wachile env., 115 km N Moyale, 04°32'34"N 39° 03'08"E, 1073 m a. s. l. Figures 127–128. Gint amoudensis sp. n., from Laas Gel (127) and holotype from type locality (128)." captionText-1="Figures 129–147: Telson, lateral views of both sexes and metasoma IV ventral views of males, comparison of Gint species. 129–131. Gint amoudensis sp. n., holotype (129, 131) and female from type locality (130). 132–134. Gint calviceps, male from Somaliland, south of Qool-Cadday, between Hargeisa and Salahle, Woqooyi Galbeed, 09°11'56"N 44°09'50"E (132, 134) and female from locality 17SE (133). 135–137. Gint dabakalo, holotype (135, 137) and paratype (136). 138–140. Gint gaitako, male (138, 140) and female (139) from Ethiopia, Oromia State, Sidamo Province, Wachile env., 115 km N Moyale, 04°32'34"N 39° 03'08"E, 1073 m a. s. l. 141–142. Gint gubanensis sp. n., holotype. 143–145. Gint maidensis sp. n., holotype (143, 145) and paratype (144). 146–147. Gint puntlandus, holotype." pageId="13" pageNumber="12">128–129, 131</figureCitation>
|
||
,
|
||
<typeStatus id="549624AE795DCB18FD5DF97AEB08F915" box="[693,798,1735,1758]" pageId="13" pageNumber="12" type="paratype">paratypes</typeStatus>
|
||
,
|
||
<figureCitation id="13168689795DCB18FF28F95BE931F936" box="[192,295,1766,1789]" captionStart="Figures 3–6" captionStartId="5.[192,278,1766,1789]" captionTargetBox="[212,1410,199,1716]" captionTargetId="figure-6@5.[192,1439,182,1728]" captionTargetPageId="5" captionText="Figures 3–6: Gint amoudensis sp. n. from type locality. Figures 3–4. Holotype male, dorsal (3) and ventral (4) views. Figures 5–6. Paratype female, dorsal (5) and ventral (6) views. Scale bar: 10 mm." pageId="13" pageNumber="12">Figs. 5–6</figureCitation>
|
||
,
|
||
<figureCitation id="13168689795DCB18FEDEF95BE96FF936" box="[310,377,1766,1789]" captionStart="Figures 11–20" captionStartId="7.[192,278,1254,1277]" captionTargetBox="[191,1437,217,1225]" captionTargetId="figure-6@7.[192,1437,218,1224]" captionTargetPageId="7" captionText="Figures 11–20: Gint amoudensis sp. n. Figures 11, 13, 17–20. Holotype male, chelicerae, carapace and tergites I–III (11), sternopectinal region and sternites III–V (13), distal segments of right legs I–IV, retrolateral views (17–20 respectively). Figures 12, 14. Paratype female from type locality, chelicerae, carapace and tergites I–III (12), sternopectinal region and sternites III–VI (14). Figure 15. Paratype male from Laas Gel, chelicerae, carapace and tergites I–III. Figure 16. Paratype female from Laas Gel, chelicerae, carapace and tergites I–III." pageId="13" pageNumber="12">12, 14</figureCitation>
|
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,
|
||
<figureCitation id="13168689795DCB18FE62F95BE9E3F936" box="[394,501,1766,1789]" captionStart="Figures 21–28" captionStartId="8.[192,278,1653,1676]" captionTargetBox="[207,1436,194,1589]" captionTargetId="figure-12@8.[192,1439,192,1623]" captionTargetPageId="8" captionText="Figures 21–28: Gint amoudensis sp. n. Figures 21, 23–25. Holotype male, metasoma V and telson lateral view (21), metasoma and telson, lateral (23), dorsal (24), and ventral (25) views. Figures 22, 26–28. Paratype female from type locality, metasoma V and telson lateral view (22), metasoma and telson, lateral (26), dorsal (27), and ventral (28) views. Scale bar: 10 mm (23–28)." pageId="13" pageNumber="12">22, 26–28</figureCitation>
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,
|
||
<figureCitation id="13168689795DCB18FDEEF95BEA5CF936" box="[518,586,1766,1789]" captionStart-0="Figures 29–45" captionStart-1="Figures 46–47" captionStartId-0="9.[192,278,1694,1717]" captionStartId-1="10.[192,278,1843,1866]" captionTargetBox-0="[201,1435,218,1644]" captionTargetBox-1="[191,1440,191,1813]" captionTargetId-0="figure-6@9.[192,1439,212,1656]" captionTargetId-1="figure-12@10.[192,1439,192,1812]" captionTargetPageId-0="9" captionTargetPageId-1="10" captionText-0="Figures 29–45: Gint amoudensis sp. n. from type locality. Figures 29–38. Holotype male. Pedipalp chela, dorsoexternal (29), external (30), and ventrointernal (31) views. Pedipalp patella, dorsal (32), external (33), and ventral (34) views. Pedipalp femur and trochanter, internal (35) and dorsal (36) views. Pedipalp chela, movable (37) and fixed (38) finger dentate margins. Figures 39–45. Paratype female. Pedipalp chela, dorsoexternal (39), external (40), and ventrointernal (41) views. Pedipalp patella, dorsal (42), external (43), and ventral (44) views. Pedipalp femur and trochanter dorsal (45) view. The trichobothrial pattern is indicated in Figures 29–36 (white circles)." captionText-1="Figures 46–47: Gint amoudensis sp. n., paratype male from type locality, in vivo habitus (46) and the type locality (47)." pageId="13" pageNumber="12">39–47</figureCitation>
|
||
,
|
||
<figureCitation id="13168689795DCB18FDB0F95BEA68F936" box="[600,638,1766,1789]" captionStart="Figures 129–147" captionStartId="31.[192,278,1694,1717]" captionTargetBox="[199,1437,219,1638]" captionTargetId="figure-6@31.[192,1440,202,1656]" captionTargetPageId="31" captionText="Figures 129–147: Telson, lateral views of both sexes and metasoma IV ventral views of males, comparison of Gint species. 129–131. Gint amoudensis sp. n., holotype (129, 131) and female from type locality (130). 132–134. Gint calviceps, male from Somaliland, south of Qool-Cadday, between Hargeisa and Salahle, Woqooyi Galbeed, 09°11'56"N 44°09'50"E (132, 134) and female from locality 17SE (133). 135–137. Gint dabakalo, holotype (135, 137) and paratype (136). 138–140. Gint gaitako, male (138, 140) and female (139) from Ethiopia, Oromia State, Sidamo Province, Wachile env., 115 km N Moyale, 04°32'34"N 39° 03'08"E, 1073 m a. s. l. 141–142. Gint gubanensis sp. n., holotype. 143–145. Gint maidensis sp. n., holotype (143, 145) and paratype (144). 146–147. Gint puntlandus, holotype." pageId="13" pageNumber="12">130</figureCitation>
|
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,
|
||
<figureCitation id="13168689795DCB18FD66F95BEAFAF936" box="[654,748,1766,1789]" captionStart="Figures 148–163" captionStartId="33.[192,278,1689,1712]" captionTargetBox="[191,1440,191,1660]" captionTargetId="figure-6@33.[192,1439,192,1659]" captionTargetPageId="33" captionText="Figures 148–163: Figures 148–151: Gint amoudensis sp. n., paratype male, No. 1327 from Laas Gel (Locality No. 17SF), right hemispermatophore. 148. Whole hemispermatophore. 149–151. Capsule region. Views of posterior (flagellar) margin (149), convex side (150), and anterior (sperm duct) margin (151). Scale bars: 1 mm (148), 200 µm (149–151). Figures 152–163: Gint amoudensis sp. n., capsule region of right hemispermatophores from four additional paratype males, showing morphological variation. Nos. 1291 (158–160), 1292 (152–154) and 1293 (161–163) from Laas Gel (Locality No. 17SF). No. 1325 (155–157) from Borama, Armoud University (Locality No. 17SR). Views of posterior (flagellar) margin (152, 155, 158, 161), convex side (153, 156, 159, 162), and anterior (sperm duct) margin (154, 157, 160, 163). Scale bars: 200 µm (152–154; 155–157; 158–160; 161–163)." pageId="13" pageNumber="12">155–157</figureCitation>
|
||
, No. 1325), leg. F. Kovařík;
|
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<materialsCitation id="3B459051795DCB18FE22F8B8ED48FF15" collectingDate="2017-08-28" collectingDateMax="2017-08-30" collectingDateMin="2017-08-28" collectionCode="SA" collectorName="Locality No. & Nos. & F. Kovarik & P. Just. All" elevation="1043" latitude="9.779722" location="Gel" longLatPrecision="21" longitude="44.44528" pageId="13" pageNumber="12" specimenCount="23" specimenCount-female="1" specimenCount-male="22" stateProvince="Mjini Magharibi" typeStatus="paratype">
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,
|
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NE Hargeisa
|
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|
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a.s.l.
|
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</elevation>
|
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(
|
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<collectorName id="26D8FFDA795DCB18FD78F89EEB34F8F1" box="[656,802,1827,1850]" pageId="13" pageNumber="12">Locality No.</collectorName>
|
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<emphasis id="B959461E795DCB18FF28F8FFE8ECF892" bold="true" box="[192,250,1858,1881]" pageId="13" pageNumber="12">17SF</emphasis>
|
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),
|
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<date id="FF93BCCC795DCB18FEFAF8FFE9D1F892" box="[274,455,1858,1881]" pageId="13" pageNumber="12" value="2017-08-28" valueMax="2017-08-30" valueMin="2017-08-28">
|
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<collectingDate id="EFD74524795DCB18FEFAF8FFE9D1F892" box="[274,455,1858,1881]" pageId="13" pageNumber="12" value="2017-08-28" valueMax="2017-08-30" valueMin="2017-08-28">28.-30.VIII.2017</collectingDate>
|
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</date>
|
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,
|
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<specimenCount id="9D2B5185795DCB18FE30F8FFEA13F892" box="[472,517,1858,1881]" pageId="13" pageNumber="12" type="male">22♂</specimenCount>
|
||
<specimenCount id="9D2B5185795DCB18FDEDF8FFEA30F892" box="[517,550,1858,1881]" pageId="13" pageNumber="12" type="female">1♀</specimenCount>
|
||
2juvs. (
|
||
<typeStatus id="549624AE795DCB18FD9EF8FFEAF7F892" box="[630,737,1858,1881]" pageId="13" pageNumber="12" type="paratype">paratypes</typeStatus>
|
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,
|
||
<figureCitation id="13168689795DCB18FD05F8FFE8FEF8B3" captionStart="Figures 1–2" captionStartId="4.[192,278,1854,1877]" captionTargetBox="[209,1417,191,1825]" captionTargetId="figure-12@4.[210,1416,192,1824]" captionTargetPageId="4" captionText="Figures 1–2: Gint amoudensis sp. n., paratypes from locality No. 17SF (Laas Gel), in vivo habitus. Figure 1. Female and two males. Figure 2. Male." pageId="13" pageNumber="12">Figs. 1–2</figureCitation>
|
||
,
|
||
<figureCitation id="13168689795DCB18FF15F8DCE925F8B3" box="[253,307,1889,1912]" captionStart="Figures 7–10" captionStartId="6.[192,278,1561,1584]" captionTargetBox="[207,1427,230,1503]" captionTargetId="figure-12@6.[192,1439,192,1531]" captionTargetPageId="6" captionText="Figures 7–10: Gint amoudensis sp. n., paratypes from Laas Gel. Figures 7–8. Male, dorsal (7) and ventral (8) views. Figures 9–10. Female, dorsal (9) and ventral (10) views. Scale bar: 10 mm." pageId="13" pageNumber="12">7–10</figureCitation>
|
||
,
|
||
<figureCitation id="13168689795DCB18FEAFF8DCE979F8B3" box="[327,367,1889,1912]" captionStart="Figures 121–128" captionStartId="30.[192,278,1662,1685]" captionTargetBox="[213,1423,188,1621]" captionTargetId="figure-12@30.[192,1439,173,1632]" captionTargetPageId="30" captionText="Figures 121–128: Metasoma and telson, ventral views, comparison of males of Gint species. Figure 121. Gint puntlandus, holotype. Figure 122. Gint dabakalo, holotype. Figure 123. Gint gubanensis sp. n., holotype. Figure 124. Gint maidensis sp. n., holotype. Figure 125. Gint calviceps, Somaliland, south of Qool-Cadday, between Hargeisa and Salahle, Woqooyi Galbeed, 09°11'56"N 44°09'50"E. Figure 126. Gint gaitako, Ethiopia, Oromia State, Sidamo Province, Wachile env., 115 km N Moyale, 04°32'34"N 39° 03'08"E, 1073 m a. s. l. Figures 127–128. Gint amoudensis sp. n., from Laas Gel (127) and holotype from type locality (128)." pageId="13" pageNumber="12">127</figureCitation>
|
||
,
|
||
<figureCitation id="13168689795DCB18FE6BF8DCEA45F8B3" box="[387,595,1889,1912]" captionStart="Figures 148–163" captionStartId="33.[192,278,1689,1712]" captionTargetBox="[191,1440,191,1660]" captionTargetId="figure-6@33.[192,1439,192,1659]" captionTargetPageId="33" captionText="Figures 148–163: Figures 148–151: Gint amoudensis sp. n., paratype male, No. 1327 from Laas Gel (Locality No. 17SF), right hemispermatophore. 148. Whole hemispermatophore. 149–151. Capsule region. Views of posterior (flagellar) margin (149), convex side (150), and anterior (sperm duct) margin (151). Scale bars: 1 mm (148), 200 µm (149–151). Figures 152–163: Gint amoudensis sp. n., capsule region of right hemispermatophores from four additional paratype males, showing morphological variation. Nos. 1291 (158–160), 1292 (152–154) and 1293 (161–163) from Laas Gel (Locality No. 17SF). No. 1325 (155–157) from Borama, Armoud University (Locality No. 17SR). Views of posterior (flagellar) margin (152, 155, 158, 161), convex side (153, 156, 159, 162), and anterior (sperm duct) margin (154, 157, 160, 163). Scale bars: 200 µm (152–154; 155–157; 158–160; 161–163)." pageId="13" pageNumber="12">148–154, 158–163</figureCitation>
|
||
,
|
||
<figureCitation id="13168689795DCB18FD80F8DCEA84F8B3" box="[616,658,1889,1912]" captionStart="Figures 195–201" captionStartId="37.[192,278,1197,1220]" captionTargetBox="[191,1440,191,1168]" captionTargetId="figure-6@37.[192,1440,192,1167]" captionTargetPageId="37" captionText="Figures 195–201: Figures 195–197: Profiles in distal view, along the longitudinal axis, of the convex surfaces at the level of the basal lobe of the right hemispermatophores of three species of Gint, showing different development of the basal lobe. 195. Gint gaitako Kovařík et al., 2013, No. 958 from Wachile (Locality No. 16EH). 196. Gint amoudensis sp. n., paratype male, No. 1292 from Laas Gel (Locality No. 17SF). 197. Gint maidensis sp. n., No. 1321 paratype male from Maid (Type locality No. 17SN). Abbreviations: b, basal lobe, f, flagellum. Figures 198–201: Mitotic metaphases of Gint spp. males. 198. G. dabakalo from locality 17SH (2n=23). 199. G. amoudensis sp. n., paratype from Laas Gel (2n=35). 200. G. amoudensis sp. n., paratype from type locality (2n=36). 201. G. maidensis sp. n., paratype (2n=34). Scale bar: 5 µm." pageId="13" pageNumber="12">196</figureCitation>
|
||
,
|
||
<collectorName id="26D8FFDA795DCB18FD4EF8DCEACEF8B3" box="[678,728,1889,1912]" pageId="13" pageNumber="12">Nos.</collectorName>
|
||
1291, 1292, 1293, 1327), leg.
|
||
<collectorName id="26D8FFDA795DCB18FBA0FF7AECA8FF15" box="[1096,1214,199,222]" pageId="13" pageNumber="12">F. Kovařík</collectorName>
|
||
et
|
||
<collectorName id="26D8FFDA795DCB18FB0CFF7AED48FF15" box="[1252,1374,199,222]" pageId="13" pageNumber="12">P. Just. All</collectorName>
|
||
</materialsCitation>
|
||
types are in
|
||
<collectionCode id="ED3C02C9795DCB18FC69FF5BEBD2FF36" box="[897,964,230,253]" pageId="13" pageNumber="12">FKCP</collectionCode>
|
||
collections.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C337C987795DCB18FCD6FE9EECB9FE92" pageId="13" pageNumber="12" type="etymology">
|
||
<paragraph id="8B929A0C795DCB18FCD6FE9EECB9FE92" blockId="13.[830,1440,291,345]" pageId="13" pageNumber="12">ETYMOLOGY. A patronym in honor of Amoud University of Republic of Somaliland.</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C337C987795DCB18FCD6FEC2EC71FB9C" pageId="13" pageNumber="12" type="diagnosis">
|
||
<paragraph id="8B929A0C795DCB18FCD6FEC2EC71FB9C" blockId="13.[830,1441,383,1111]" pageId="13" pageNumber="12">
|
||
DIAGNOSIS. Total length
|
||
<quantity id="4CD537E9795DCB18FB98FEC2ECA3FE5D" box="[1136,1205,383,406]" metricMagnitude="-2" metricUnit="m" metricValue="2.1" pageId="13" pageNumber="12" unit="mm" value="21.0">21mm</quantity>
|
||
(male) to
|
||
<quantity id="4CD537E9795DCB18FAA1FEC2EDB7FE5D" box="[1353,1441,383,406]" metricMagnitude="-2" metricUnit="m" metricValue="3.8" pageId="13" pageNumber="12" unit="mm" value="38.0">38 mm</quantity>
|
||
(female); chelicerae yellow with reticulation in anterior part or uniformly yellow; carapace densely granulated with only anterior median carinae developed; anterior margin of carapace straight; pectine teeth
|
||
<quantity id="4CD537E9795DCB18FAD0FE47EDB6FDDA" box="[1336,1440,506,529]" metricMagnitude="-1" metricUnit="m" metricValue="5.334" metricValueMax="5.588" metricValueMin="5.08" pageId="13" pageNumber="12" unit="in" value="21.0" valueMax="22.0" valueMin="20.0">20–22 in</quantity>
|
||
females and
|
||
<quantity id="4CD537E9795DCB18FC26FDA5EC27FDE4" box="[974,1073,536,559]" metricMagnitude="-1" metricUnit="m" metricValue="5.842" metricValueMax="6.35" metricValueMin="5.334" pageId="13" pageNumber="12" unit="in" value="23.0" valueMax="25.0" valueMin="21.0">21–25 in</quantity>
|
||
males; all sternites lacking carinae; sternite VII with four weakly indicated carinae, intercarinal surface weakly granulated (mainly in males); metasomal segment V length/width ratio
|
||
<quantity id="4CD537E9795DCB18FA8CFDC9EB85FD61" metricMagnitude="-2" metricUnit="m" metricValue="5.5753" metricValueMax="5.9182" metricValueMin="5.2324" pageId="13" pageNumber="12" unit="in" value="2.195" valueMax="2.33" valueMin="2.06">2.06– 2.33 in</quantity>
|
||
male; metasomal segment II–IV intercarinal surfaces granulated in both sexes; metasomal segment IV bears 8 carinae that are complete and granulate in both sexes; metasomal segment V of both sexes has only ventromedial and ventrolateral carinae that in posterior halves bear several lobate granules; dorsal surface of segment V smooth and lateral surface may be weakly granulated (more so in males); all metasomal segments sparsely setose; metasomal segment V bearing ca. 48 long setae in both sexes; telson rather elongate without sexual dimorphism, aculeus slightly shorter than vesicle in both sexes; legs I–III with tarsal bristle combs composed of 5 to 9 long, thin setae; movable finger of pedipalp with 8 rows of granules, with external and internal accessory granules.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C337C987795DCB06FC86FB3CE903F8D2" lastPageId="19" lastPageNumber="18" pageId="13" pageNumber="12" type="description">
|
||
<paragraph id="8B929A0C795DCB18FC86FB3CEB8FFAA5" blockId="13.[830,1447,1153,1912]" pageId="13" pageNumber="12">
|
||
DESCRIPTION. Adult males are
|
||
<quantity id="4CD537E9795DCB18FB2BFB3CED2FFB53" box="[1219,1337,1153,1176]" metricMagnitude="-2" metricUnit="m" metricValue="2.5" metricValueMax="2.9" metricValueMin="2.1" pageId="13" pageNumber="12" unit="mm" value="25.0" valueMax="29.0" valueMin="21.0">21–29 mm</quantity>
|
||
long and the adult females are
|
||
<quantity id="4CD537E9795DCB18FBC6FB22ECB5FB7D" box="[1070,1187,1183,1206]" metricMagnitude="-2" metricUnit="m" metricValue="3.25" metricValueMax="3.8" metricValueMin="2.7" pageId="13" pageNumber="12" unit="mm" value="32.5" valueMax="38.0" valueMin="27.0">27–38 mm</quantity>
|
||
long. For position and distribution of trichobothria of pedipalps see
|
||
<figureCitation id="13168689795DCB18FAD1FB03EB4FFB3F" captionStart="Figures 29–45" captionStartId="9.[192,278,1694,1717]" captionTargetBox="[201,1435,218,1644]" captionTargetId="figure-6@9.[192,1439,212,1656]" captionTargetPageId="9" captionText="Figures 29–45: Gint amoudensis sp. n. from type locality. Figures 29–38. Holotype male. Pedipalp chela, dorsoexternal (29), external (30), and ventrointernal (31) views. Pedipalp patella, dorsal (32), external (33), and ventral (34) views. Pedipalp femur and trochanter, internal (35) and dorsal (36) views. Pedipalp chela, movable (37) and fixed (38) finger dentate margins. Figures 39–45. Paratype female. Pedipalp chela, dorsoexternal (39), external (40), and ventrointernal (41) views. Pedipalp patella, dorsal (42), external (43), and ventral (44) views. Pedipalp femur and trochanter dorsal (45) view. The trichobothrial pattern is indicated in Figures 29–36 (white circles)." pageId="13" pageNumber="12">Figs. 29– 36</figureCitation>
|
||
. Sexual dimorphism is noticeable. Males are substantially smaller, but there are no differences in the shape of the telson. Pedipalp patella and femur are granulate and matte in males, smooth and glossy in females.
|
||
</paragraph>
|
||
<paragraph id="8B929A0C795DCB18FC86FACBEC59F96A" blockId="13.[830,1447,1153,1912]" pageId="13" pageNumber="12">
|
||
<materialsCitation id="3B459051795DCB18FC86FACBEC5FF96A" collectionCode="V" location="Gel" municipality="The" pageId="13" pageNumber="12" specimenCount="2" stateProvince="Mjini Magharibi">
|
||
COLORATION (
|
||
<figureCitation id="13168689795DCB18FBE4FACBEC69FA46" box="[1036,1151,1398,1421]" captionStart-0="Figures 1–2" captionStart-1="Figures 3–6" captionStart-2="Figures 7–10" captionStart-3="Figures 11–20" captionStartId-0="4.[192,278,1854,1877]" captionStartId-1="5.[192,278,1766,1789]" captionStartId-2="6.[192,278,1561,1584]" captionStartId-3="7.[192,278,1254,1277]" captionTargetBox-0="[209,1417,191,1825]" captionTargetBox-1="[212,1410,199,1716]" captionTargetBox-2="[207,1427,230,1503]" captionTargetBox-3="[191,1437,217,1225]" captionTargetId-0="figure-12@4.[210,1416,192,1824]" captionTargetId-1="figure-6@5.[192,1439,182,1728]" captionTargetId-2="figure-12@6.[192,1439,192,1531]" captionTargetId-3="figure-6@7.[192,1437,218,1224]" captionTargetPageId-0="4" captionTargetPageId-1="5" captionTargetPageId-2="6" captionTargetPageId-3="7" captionText-0="Figures 1–2: Gint amoudensis sp. n., paratypes from locality No. 17SF (Laas Gel), in vivo habitus. Figure 1. Female and two males. Figure 2. Male." captionText-1="Figures 3–6: Gint amoudensis sp. n. from type locality. Figures 3–4. Holotype male, dorsal (3) and ventral (4) views. Figures 5–6. Paratype female, dorsal (5) and ventral (6) views. Scale bar: 10 mm." captionText-2="Figures 7–10: Gint amoudensis sp. n., paratypes from Laas Gel. Figures 7–8. Male, dorsal (7) and ventral (8) views. Figures 9–10. Female, dorsal (9) and ventral (10) views. Scale bar: 10 mm." captionText-3="Figures 11–20: Gint amoudensis sp. n. Figures 11, 13, 17–20. Holotype male, chelicerae, carapace and tergites I–III (11), sternopectinal region and sternites III–V (13), distal segments of right legs I–IV, retrolateral views (17–20 respectively). Figures 12, 14. Paratype female from type locality, chelicerae, carapace and tergites I–III (12), sternopectinal region and sternites III–VI (14). Figure 15. Paratype male from Laas Gel, chelicerae, carapace and tergites I–III. Figure 16. Paratype female from Laas Gel, chelicerae, carapace and tergites I–III." pageId="13" pageNumber="12">Figs. 1–16</figureCitation>
|
||
,
|
||
<figureCitation id="13168689795DCB18FB78FACBECBFFA46" box="[1168,1193,1398,1421]" captionStart="Figures 46–47" captionStartId="10.[192,278,1843,1866]" captionTargetBox="[191,1440,191,1813]" captionTargetId="figure-12@10.[192,1439,192,1812]" captionTargetPageId="10" captionText="Figures 46–47: Gint amoudensis sp. n., paratype male from type locality, in vivo habitus (46) and the type locality (47)." pageId="13" pageNumber="12">46</figureCitation>
|
||
). Basic color is yellow to orange with strong dark patterning and spots (population from
|
||
<typeStatus id="549624AE795DCB18FC3CFA0EEC14FA01" box="[980,1026,1459,1482]" pageId="13" pageNumber="12">type</typeStatus>
|
||
locality), but expression of colors is quite variable. The dark spots can also occur on the legs, mainly on the femur. The carinae on the metasoma can be dark. Metasomal segment
|
||
<collectionCode id="ED3C02C9795DCB18FB6EF9B2EC8FF9ED" box="[1158,1177,1551,1574]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="13" pageNumber="12" type="Museum">V</collectionCode>
|
||
is darker than the other metasomal segments.
|
||
<collectingMunicipality id="6BF60076795DCB18FBD3F993EC73F98E" box="[1083,1125,1582,1605]" pageId="13" pageNumber="12">The</collectingMunicipality>
|
||
chelicerae are yellow with reticulation in the anterior part (population from
|
||
<typeStatus id="549624AE795DCB18FA91F9F0EDB1F9AF" box="[1401,1447,1613,1636]" pageId="13" pageNumber="12">type</typeStatus>
|
||
locality) or uniformly yellow (population from
|
||
<collectingRegion id="49E954EE795DCB18FA9DF9D6EDB1F949" box="[1397,1447,1643,1666]" country="Tanzania" name="Mjini Magharibi" pageId="13" pageNumber="12">Laas</collectingRegion>
|
||
<location id="8EF2CCD7795DCB18FCD6F937EB7FF96A" LSID="urn:lsid:plazi:treatment:03842B1A795DCB06FEE6FB67EC1DFC30:8EF2CCD7795DCB18FCD6F937EB7FF96A" box="[830,873,1674,1697]" municipality="The" name="Gel" pageId="13" pageNumber="12" stateProvince="Mjini Magharibi">Gel</location>
|
||
); dentition is reddish
|
||
</materialsCitation>
|
||
.
|
||
</paragraph>
|
||
<paragraph id="8B929A0C795DCB18FC86F914EC5AF8F1" blockId="13.[830,1447,1153,1912]" pageId="13" pageNumber="12">
|
||
CARAPACE (
|
||
<figureCitation id="13168689795DCB18FC13F914ECCDF90B" box="[1019,1243,1705,1728]" captionStart="Figures 11–20" captionStartId="7.[192,278,1254,1277]" captionTargetBox="[191,1437,217,1225]" captionTargetId="figure-6@7.[192,1437,218,1224]" captionTargetPageId="7" captionText="Figures 11–20: Gint amoudensis sp. n. Figures 11, 13, 17–20. Holotype male, chelicerae, carapace and tergites I–III (11), sternopectinal region and sternites III–V (13), distal segments of right legs I–IV, retrolateral views (17–20 respectively). Figures 12, 14. Paratype female from type locality, chelicerae, carapace and tergites I–III (12), sternopectinal region and sternites III–VI (14). Figure 15. Paratype male from Laas Gel, chelicerae, carapace and tergites I–III. Figure 16. Paratype female from Laas Gel, chelicerae, carapace and tergites I–III." pageId="13" pageNumber="12">Figs. 11–12, 15–16</figureCitation>
|
||
). The surface is densely granulated. The anterior margin is straight and bears six to eight macrosetae. Anterior median carinae are coarsely granular. There are 5 lateral eyes on each side (3 larger, 2 smaller).
|
||
</paragraph>
|
||
<paragraph id="8B929A0C795DCB05FC86F8FFE9F6F8A5" blockId="13.[830,1447,1153,1912]" lastBlockId="16.[192,802,1419,1902]" lastPageId="16" lastPageNumber="15" pageId="13" pageNumber="12">
|
||
MESOSOMA (
|
||
<figureCitation id="13168689795DCB18FC17F8FFEC95F892" box="[1023,1155,1858,1881]" captionStart="Figures 11–20" captionStartId="7.[192,278,1254,1277]" captionTargetBox="[191,1437,217,1225]" captionTargetId="figure-6@7.[192,1437,218,1224]" captionTargetPageId="7" captionText="Figures 11–20: Gint amoudensis sp. n. Figures 11, 13, 17–20. Holotype male, chelicerae, carapace and tergites I–III (11), sternopectinal region and sternites III–V (13), distal segments of right legs I–IV, retrolateral views (17–20 respectively). Figures 12, 14. Paratype female from type locality, chelicerae, carapace and tergites I–III (12), sternopectinal region and sternites III–VI (14). Figure 15. Paratype male from Laas Gel, chelicerae, carapace and tergites I–III. Figure 16. Paratype female from Laas Gel, chelicerae, carapace and tergites I–III." pageId="13" pageNumber="12">Figs. 11–16</figureCitation>
|
||
). The tergites bear three coarsely granular carinae, of which the lateral pair on tergites I–II are inconspicuous. All tergites with dense coarse and fine granulation. The pectinal tooth count is 21–25 (22.879) (±0.860) [58] in males and 20–22 (21.000) (±0.633) [16] in females. The marginal tips of the pectines extend from the third quarter to the end of sternite IV in females, and from the end of sternite IV to the first quarter of sternite V in males. The pectines have 3 marginal lamellae and 8–10 middle lamellae. The lamellae bear numerous dark setae, three to six on each fulcrum. Sternites III–VI lack carinae, their surfaces are smooth except for finely shagreened lateral areas on sternite III covered by the pectines. Sternite VII has two pairs of poorly indicated carinae and is weakly granulated in the area outside the lateral carinae, more so in males. All sternites bear many long macrosetae on their surfaces and margins.
|
||
</paragraph>
|
||
<caption id="DF52CA84795ECB1BFF28F8BFECF5F8F9" pageId="14" pageNumber="13" startId="14.[192,278,1794,1817]" targetBox="[191,1440,191,1765]" targetPageId="14">
|
||
<paragraph id="8B929A0C795ECB1BFF28F8BFECF5F8F9" blockId="14.[192,1440,1794,1842]" pageId="14" pageNumber="13">
|
||
<emphasis id="B959461E795ECB1BFF28F8BFE9C0F8D2" bold="true" box="[192,470,1794,1817]" pageId="14" pageNumber="13">Figures 49–50: Figure 49</emphasis>
|
||
.
|
||
<taxonomicName id="4C2DE18F795ECB1BFE0CF8B9EA66F8D2" baseAuthorityName="Pocock" baseAuthorityYear="1900" box="[484,624,1796,1817]" class="Arachnida" family="Buthidae" genus="Gint" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="calviceps">
|
||
<emphasis id="B959461E795ECB1BFE0CF8B9EA66F8D2" box="[484,624,1796,1817]" italics="true" pageId="14" pageNumber="13">Gint calviceps</emphasis>
|
||
</taxonomicName>
|
||
, female from locality No. 17SE (Somaliland, Toon village near Hargeisa), in vivo habitus.
|
||
<emphasis id="B959461E795ECB1BFEF9F8A0E965F8F9" bold="true" box="[273,371,1821,1842]" pageId="14" pageNumber="13">Figure 50</emphasis>
|
||
.
|
||
<taxonomicName id="4C2DE18F795ECB1BFE68F8A0EA1FF8F9" authorityName="Kovarik et Mazuch" authorityYear="2015" box="[384,521,1821,1842]" class="Arachnida" family="Buthidae" genus="Gint" kingdom="Animalia" order="Scorpiones" pageId="14" pageNumber="13" phylum="Arthropoda" rank="species" species="dabakalo">
|
||
<emphasis id="B959461E795ECB1BFE68F8A0EA1FF8F9" box="[384,521,1821,1842]" italics="true" pageId="14" pageNumber="13">Gint dabakalo</emphasis>
|
||
</taxonomicName>
|
||
, male from locality No. 17SC (Somaliland, Burao, airport), in vivo habitus.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF52CA84795FCB1AFF28F8ACE9ACF88A" pageId="15" pageNumber="14" startId="15.[192,278,1809,1832]" targetBox="[191,1440,219,1777]" targetPageId="15">
|
||
<paragraph id="8B929A0C795FCB1AFF28F8ACE9ACF88A" blockId="15.[192,1441,1809,1857]" pageId="15" pageNumber="14">
|
||
<emphasis id="B959461E795FCB1AFF28F8ACE97CF8E3" bold="true" box="[192,362,1809,1832]" pageId="15" pageNumber="14">Figures 51–52:</emphasis>
|
||
<taxonomicName id="4C2DE18F795FCB1AFE9DF8AEE9F8F8E3" box="[373,494,1811,1832]" class="Arachnida" family="Buthidae" genus="Gint" kingdom="Animalia" order="Scorpiones" pageId="15" pageNumber="14" phylum="Arthropoda" rank="species" species="gaitako">
|
||
<emphasis id="B959461E795FCB1AFE9DF8AEE9F8F8E3" box="[373,494,1811,1832]" italics="true" pageId="15" pageNumber="14">Gint gaitako</emphasis>
|
||
</taxonomicName>
|
||
, male (51) and female (52) from locality No. 16EH (Ethiopia, Oromia State, Sidamo Province, Wachile), in vivo habitus.
|
||
</paragraph>
|
||
</caption>
|
||
<caption id="DF52CA847940CB05FF28FA82ECFEFA9D" box="[192,1256,1343,1366]" pageId="16" pageNumber="15" startId="16.[192,278,1343,1366]" targetBox="[208,1419,208,1295]" targetPageId="16">
|
||
<paragraph id="8B929A0C7940CB05FF28FA82ECFEFA9D" blockId="16.[192,1256,1343,1366]" box="[192,1256,1343,1366]" pageId="16" pageNumber="15">
|
||
<emphasis id="B959461E7940CB05FF28FA82E97EFA9D" bold="true" box="[192,360,1343,1366]" pageId="16" pageNumber="15">Figures 53–54:</emphasis>
|
||
<taxonomicName id="4C2DE18F7940CB05FE86FAFCEA1DFA9D" authority="Kovařík & Lowe & Just & Awale & Sh, 2018" authorityName="Kovařík & Lowe & Just & Awale & Sh" authorityYear="2018" box="[366,523,1345,1366]" class="Arachnida" family="Buthidae" genus="Gint" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="species" species="gubanensis" status="sp. nov.">
|
||
<emphasis id="B959461E7940CB05FE86FAFCEA1DFA9D" box="[366,523,1345,1366]" italics="true" pageId="16" pageNumber="15">Gint gubanensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26AFB657940CB05FDF9FAFCEA50FA9D" box="[529,582,1345,1366]" pageId="16" pageNumber="15" rank="species">
|
||
<emphasis id="B959461E7940CB05FDF9FAFCEA31FA9D" bold="true" box="[529,551,1345,1366]" pageId="16" pageNumber="15">sp</emphasis>
|
||
.
|
||
<emphasis id="B959461E7940CB05FDDCFAFCEA57FA9D" bold="true" box="[564,577,1345,1366]" pageId="16" pageNumber="15">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, holotype male, dorsal (53) and ventral (54) views. Scale bar: 10 mm.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B929A0C7940CB05FC86FA33ECA8F8A5" blockId="16.[830,1441,1419,1902]" pageId="16" pageNumber="15">
|
||
HEMISPERMATOPHORE (
|
||
<figureCitation id="131686897940CB05FB9CFA36ED01FA69" box="[1140,1303,1419,1442]" captionStart="Figures 148–163" captionStartId="33.[192,278,1689,1712]" captionTargetBox="[191,1440,191,1660]" captionTargetId="figure-6@33.[192,1439,192,1659]" captionTargetPageId="33" captionText="Figures 148–163: Figures 148–151: Gint amoudensis sp. n., paratype male, No. 1327 from Laas Gel (Locality No. 17SF), right hemispermatophore. 148. Whole hemispermatophore. 149–151. Capsule region. Views of posterior (flagellar) margin (149), convex side (150), and anterior (sperm duct) margin (151). Scale bars: 1 mm (148), 200 µm (149–151). Figures 152–163: Gint amoudensis sp. n., capsule region of right hemispermatophores from four additional paratype males, showing morphological variation. Nos. 1291 (158–160), 1292 (152–154) and 1293 (161–163) from Laas Gel (Locality No. 17SF). No. 1325 (155–157) from Borama, Armoud University (Locality No. 17SR). Views of posterior (flagellar) margin (152, 155, 158, 161), convex side (153, 156, 159, 162), and anterior (sperm duct) margin (154, 157, 160, 163). Scale bars: 200 µm (152–154; 155–157; 158–160; 161–163)." pageId="16" pageNumber="15">Figs. 148–163</figureCitation>
|
||
,
|
||
<figureCitation id="131686897940CB05FAC5FA36ED43FA69" box="[1325,1365,1419,1442]" captionStart="Figures 195–201" captionStartId="37.[192,278,1197,1220]" captionTargetBox="[191,1440,191,1168]" captionTargetId="figure-6@37.[192,1440,192,1167]" captionTargetPageId="37" captionText="Figures 195–201: Figures 195–197: Profiles in distal view, along the longitudinal axis, of the convex surfaces at the level of the basal lobe of the right hemispermatophores of three species of Gint, showing different development of the basal lobe. 195. Gint gaitako Kovařík et al., 2013, No. 958 from Wachile (Locality No. 16EH). 196. Gint amoudensis sp. n., paratype male, No. 1292 from Laas Gel (Locality No. 17SF). 197. Gint maidensis sp. n., No. 1321 paratype male from Maid (Type locality No. 17SN). Abbreviations: b, basal lobe, f, flagellum. Figures 198–201: Mitotic metaphases of Gint spp. males. 198. G. dabakalo from locality 17SH (2n=23). 199. G. amoudensis sp. n., paratype from Laas Gel (2n=35). 200. G. amoudensis sp. n., paratype from type locality (2n=36). 201. G. maidensis sp. n., paratype (2n=34). Scale bar: 5 µm." pageId="16" pageNumber="15">196</figureCitation>
|
||
). Flagelliform, trunk long and slender, capsule relatively short. Flagellum approximately as long as trunk, with shorter pars recta bearing a fin-like expansion along proximal anterior margin, and longer, hyaline pars reflecta. Capsule with 3+1 lobe structure typical of ‘
|
||
<taxonomicName id="4C2DE18F7940CB05FCAFF9FEEB87F991" authorityName="Leach" authorityYear="1815" box="[839,913,1603,1626]" class="Arachnida" family="Buthidae" genus="Buthus" kingdom="Animalia" order="Scorpiones" pageId="16" pageNumber="15" phylum="Arthropoda" rank="genus">
|
||
<emphasis id="B959461E7940CB05FCAFF9FEEB87F991" box="[839,913,1603,1626]" italics="true" pageId="16" pageNumber="15">Buthus</emphasis>
|
||
</taxonomicName>
|
||
’ group (Fet et al., 2005;
|
||
<bibRefCitation id="EFBCE7FD7940CB05FB41F9FEED83F991" author="KOVARIK" box="[1193,1429,1603,1626]" pageId="16" pageNumber="15" pagination="1 - 133" refId="ref16815" refString="KOVARIK, F., G. LOWE, K. B. RANAWANA, D. HOFEREK, V. A. SANJEEWA JAYARATHNE, J. PLISKOVA & F. STAHLAVSKY. 2016 c. Scorpions of Sri Lanka (Arachnida, Scorpiones: Buthidae, Chaerilidae, Scorpionidae) with description of four new species of the genera Charmus Karsch, 1879 and Reddyanus Vachon, 1972 stat. n .. Euscorpius, 220: 1 - 133." type="journal article" year="2016">Kovařík et al., 2016c</bibRefCitation>
|
||
). Sperm hemiduct separated from flagellum, tripartite with posterior lobe largest, median lobe shortest and apically acuminate, anterior lobe of intermediate size and apically tapered. Posterior margin of median lobe overhanging the posterior lobe, the two lobes fused along a robust carina. Basal lobe well developed as a low, rounded scoop of variable size (
|
||
<figureCitation id="131686897940CB05FB10F8A4ED8FF8FB" box="[1272,1433,1817,1840]" captionStart="Figures 148–163" captionStartId="33.[192,278,1689,1712]" captionTargetBox="[191,1440,191,1660]" captionTargetId="figure-6@33.[192,1439,192,1659]" captionTargetPageId="33" captionText="Figures 148–163: Figures 148–151: Gint amoudensis sp. n., paratype male, No. 1327 from Laas Gel (Locality No. 17SF), right hemispermatophore. 148. Whole hemispermatophore. 149–151. Capsule region. Views of posterior (flagellar) margin (149), convex side (150), and anterior (sperm duct) margin (151). Scale bars: 1 mm (148), 200 µm (149–151). Figures 152–163: Gint amoudensis sp. n., capsule region of right hemispermatophores from four additional paratype males, showing morphological variation. Nos. 1291 (158–160), 1292 (152–154) and 1293 (161–163) from Laas Gel (Locality No. 17SF). No. 1325 (155–157) from Borama, Armoud University (Locality No. 17SR). Views of posterior (flagellar) margin (152, 155, 158, 161), convex side (153, 156, 159, 162), and anterior (sperm duct) margin (154, 157, 160, 163). Scale bars: 200 µm (152–154; 155–157; 158–160; 161–163)." pageId="16" pageNumber="15">Figs. 149–163</figureCitation>
|
||
,
|
||
<figureCitation id="131686897940CB05FCD6F885EB70F884" box="[830,870,1848,1871]" captionStart="Figures 195–201" captionStartId="37.[192,278,1197,1220]" captionTargetBox="[191,1440,191,1168]" captionTargetId="figure-6@37.[192,1440,192,1167]" captionTargetPageId="37" captionText="Figures 195–201: Figures 195–197: Profiles in distal view, along the longitudinal axis, of the convex surfaces at the level of the basal lobe of the right hemispermatophores of three species of Gint, showing different development of the basal lobe. 195. Gint gaitako Kovařík et al., 2013, No. 958 from Wachile (Locality No. 16EH). 196. Gint amoudensis sp. n., paratype male, No. 1292 from Laas Gel (Locality No. 17SF). 197. Gint maidensis sp. n., No. 1321 paratype male from Maid (Type locality No. 17SN). Abbreviations: b, basal lobe, f, flagellum. Figures 198–201: Mitotic metaphases of Gint spp. males. 198. G. dabakalo from locality 17SH (2n=23). 199. G. amoudensis sp. n., paratype from Laas Gel (2n=35). 200. G. amoudensis sp. n., paratype from type locality (2n=36). 201. G. maidensis sp. n., paratype (2n=34). Scale bar: 5 µm." pageId="16" pageNumber="15">196</figureCitation>
|
||
). Morphology was consistent across 10 hemispermatophores extracted from
|
||
<specimenCount id="9D2B51857940CB05FB8CF8EAECACF8A5" box="[1124,1210,1879,1902]" pageId="16" pageNumber="15" type="male">5 males</specimenCount>
|
||
.
|
||
</paragraph>
|
||
<caption id="DF52CA847941CB04FF28F99CEBCCF9A1" pageId="17" pageNumber="16" startId="17.[192,278,1569,1592]" targetBox="[191,1437,191,1539]" targetPageId="17">
|
||
<paragraph id="8B929A0C7941CB04FF28F99CEBCCF9A1" blockId="17.[192,1441,1569,1642]" pageId="17" pageNumber="16">
|
||
<emphasis id="B959461E7941CB04FF28F99CE97CF9F3" bold="true" box="[192,362,1569,1592]" pageId="17" pageNumber="16">Figures 55–62:</emphasis>
|
||
<taxonomicName id="4C2DE18F7941CB04FE9BF99EEA07F9F3" authority="Kovařík & Lowe & Just & Awale & Sh, 2018" authorityName="Kovařík & Lowe & Just & Awale & Sh" authorityYear="2018" box="[371,529,1571,1592]" class="Arachnida" family="Buthidae" genus="Gint" kingdom="Animalia" order="Scorpiones" pageId="17" pageNumber="16" phylum="Arthropoda" rank="species" species="gubanensis" status="sp. nov.">
|
||
<emphasis id="B959461E7941CB04FE9BF99EEA07F9F3" box="[371,529,1571,1592]" italics="true" pageId="17" pageNumber="16">Gint gubanensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26AFB657941CB04FDF1F99EEA47F9F3" box="[537,593,1571,1592]" pageId="17" pageNumber="16" rank="species">
|
||
<emphasis id="B959461E7941CB04FDF1F99EEA26F9F3" bold="true" box="[537,560,1571,1592]" pageId="17" pageNumber="16">sp</emphasis>
|
||
.
|
||
<emphasis id="B959461E7941CB04FDD6F99EEA5DF9F3" bold="true" box="[574,587,1571,1592]" pageId="17" pageNumber="16">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, holotype male, chelicerae, carapace and tergites I–IV (55), sternopectinal region and sternites III–VII (56), metasoma V and telson lateral view (57), metasoma and telson, lateral (58), dorsal (59), and ventral (60) views, left chelicera dorsal (61) and ventral (62) views. Scale bar: 10 mm (58–60).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B929A0C7941CB06FF18F923E90BFE7E" blockId="17.[192,804,1694,1901]" lastBlockId="19.[192,809,199,1050]" lastPageId="19" lastPageNumber="18" pageId="17" pageNumber="16">
|
||
METASOMA AND TELSON (
|
||
<figureCitation id="131686897941CB04FDF4F923EAB5F97E" box="[540,675,1694,1717]" captionStart="Figures 21–28" captionStartId="8.[192,278,1653,1676]" captionTargetBox="[207,1436,194,1589]" captionTargetId="figure-12@8.[192,1439,192,1623]" captionTargetPageId="8" captionText="Figures 21–28: Gint amoudensis sp. n. Figures 21, 23–25. Holotype male, metasoma V and telson lateral view (21), metasoma and telson, lateral (23), dorsal (24), and ventral (25) views. Figures 22, 26–28. Paratype female from type locality, metasoma V and telson lateral view (22), metasoma and telson, lateral (26), dorsal (27), and ventral (28) views. Scale bar: 10 mm (23–28)." pageId="17" pageNumber="16">Figs. 21–28</figureCitation>
|
||
,
|
||
<figureCitation id="131686897941CB04FD5DF923EB05F97E" box="[693,787,1694,1717]" captionStart="Figures 121–128" captionStartId="30.[192,278,1662,1685]" captionTargetBox="[213,1423,188,1621]" captionTargetId="figure-12@30.[192,1439,173,1632]" captionTargetPageId="30" captionText="Figures 121–128: Metasoma and telson, ventral views, comparison of males of Gint species. Figure 121. Gint puntlandus, holotype. Figure 122. Gint dabakalo, holotype. Figure 123. Gint gubanensis sp. n., holotype. Figure 124. Gint maidensis sp. n., holotype. Figure 125. Gint calviceps, Somaliland, south of Qool-Cadday, between Hargeisa and Salahle, Woqooyi Galbeed, 09°11'56"N 44°09'50"E. Figure 126. Gint gaitako, Ethiopia, Oromia State, Sidamo Province, Wachile env., 115 km N Moyale, 04°32'34"N 39° 03'08"E, 1073 m a. s. l. Figures 127–128. Gint amoudensis sp. n., from Laas Gel (127) and holotype from type locality (128)." pageId="17" pageNumber="16">127–128</figureCitation>
|
||
). Metasoma I–III bear 10 carinae, the ventromedial carinae on metasoma I are present but smooth. Median lateral carinae are complete or almost complete on I–III. Ventromedial and ventrolateral carinae on metasoma II– III are granulated, with larger granules posteriorly, and strong granulation in females. Metasoma IV bears 8 carinae that are complete and granulate in both sexes. Metasoma V of both sexes has only ventromedial and ventrolateral carinae, which in posterior halves bear several lobate granules. Intercarinal surfaces of segments II–IV are partly granulated in both sexes, with granules of approximately equal size. The ventral aspect of metasoma I is smooth in both sexes. Dorsal and lateral surfaces of this segment are granulated in both sexes. The anal arch consists of three lobes in both sexes. All segments are sparsely setose; the fifth segment has ca. 48 long setae in both sexes. The telson is rather elongate without sexual dimorphism. The aculeus is slightly shorter than the vesicle in both sexes. The surface of the telson is smooth, sparsely hirsute, without a subaculear tubercle.
|
||
</paragraph>
|
||
<caption id="DF52CA847942CB07FF28F8B6EC28F8A7" pageId="18" pageNumber="17" startId="18.[192,278,1803,1826]" targetBox="[230,1385,201,1763]" targetPageId="18">
|
||
<paragraph id="8B929A0C7942CB07FF28F8B6EC28F8A7" blockId="18.[192,1441,1803,1900]" pageId="18" pageNumber="17">
|
||
<emphasis id="B959461E7942CB07FF28F8B6E97EF8E9" bold="true" box="[192,360,1803,1826]" pageId="18" pageNumber="17">Figures 63–75:</emphasis>
|
||
<taxonomicName id="4C2DE18F7942CB07FE87F8B0EA1DF8E9" authority="Kovařík & Lowe & Just & Awale & Sh, 2018" authorityName="Kovařík & Lowe & Just & Awale & Sh" authorityYear="2018" box="[367,523,1805,1826]" class="Arachnida" family="Buthidae" genus="Gint" kingdom="Animalia" order="Scorpiones" pageId="18" pageNumber="17" phylum="Arthropoda" rank="species" species="gubanensis" status="sp. nov.">
|
||
<emphasis id="B959461E7942CB07FE87F8B0EA1DF8E9" box="[367,523,1805,1826]" italics="true" pageId="18" pageNumber="17">Gint gubanensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26AFB657942CB07FDFAF8B0EA51F8E9" box="[530,583,1805,1826]" pageId="18" pageNumber="17" rank="species">
|
||
<emphasis id="B959461E7942CB07FDFAF8B0EA3EF8E9" bold="true" box="[530,552,1805,1826]" pageId="18" pageNumber="17">sp</emphasis>
|
||
.
|
||
<emphasis id="B959461E7942CB07FDDCF8B0EA57F8E9" bold="true" box="[564,577,1805,1826]" pageId="18" pageNumber="17">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
, holotype male. Pedipalp chela, dorsal (63), external (64), and ventrointernal (65) views. Pedipalp patella, dorsal (66), external (67), and ventral (68) views. Pedipalp femur and trochanter, internal (69) and dorsal (70) views. Pedipalp chela, movable finger dentate margin (71). Distal segments of right legs I–IV, retrolateral views (72–75, respectively). The trichobothrial pattern is indicated in Figures 63–70 (labeled white circles).
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="8B929A0C7943CB06FF18FE01EACAFDA6" blockId="19.[192,809,199,1050]" pageId="19" pageNumber="18">
|
||
LEGS (
|
||
<figureCitation id="131686897943CB06FEDEFE01E9A3FE18" box="[310,437,444,467]" captionStart="Figures 11–20" captionStartId="7.[192,278,1254,1277]" captionTargetBox="[191,1437,217,1225]" captionTargetId="figure-6@7.[192,1437,218,1224]" captionTargetPageId="7" captionText="Figures 11–20: Gint amoudensis sp. n. Figures 11, 13, 17–20. Holotype male, chelicerae, carapace and tergites I–III (11), sternopectinal region and sternites III–V (13), distal segments of right legs I–IV, retrolateral views (17–20 respectively). Figures 12, 14. Paratype female from type locality, chelicerae, carapace and tergites I–III (12), sternopectinal region and sternites III–VI (14). Figure 15. Paratype male from Laas Gel, chelicerae, carapace and tergites I–III. Figure 16. Paratype female from Laas Gel, chelicerae, carapace and tergites I–III." pageId="19" pageNumber="18">Figs. 17–20</figureCitation>
|
||
). The tarsomeres bear two rows of macrosetae on the ventral surface and numerous macrosetae on the other surfaces, which on legs I–III form bristle combs with 5–9 bristles. The macrosetae are thin in both sexes. The femur and patella may bear four to six carinae. The femur bears only solitary macrosetae.
|
||
</paragraph>
|
||
<paragraph id="8B929A0C7943CB06FF18FDC9EA42FBD1" blockId="19.[192,809,199,1050]" pageId="19" pageNumber="18">
|
||
PEDIPALPS (
|
||
<figureCitation id="131686897943CB06FE9AFDC9E9E4FD40" box="[370,498,628,651]" captionStart="Figures 29–45" captionStartId="9.[192,278,1694,1717]" captionTargetBox="[201,1435,218,1644]" captionTargetId="figure-6@9.[192,1439,212,1656]" captionTargetPageId="9" captionText="Figures 29–45: Gint amoudensis sp. n. from type locality. Figures 29–38. Holotype male. Pedipalp chela, dorsoexternal (29), external (30), and ventrointernal (31) views. Pedipalp patella, dorsal (32), external (33), and ventral (34) views. Pedipalp femur and trochanter, internal (35) and dorsal (36) views. Pedipalp chela, movable (37) and fixed (38) finger dentate margins. Figures 39–45. Paratype female. Pedipalp chela, dorsoexternal (39), external (40), and ventrointernal (41) views. Pedipalp patella, dorsal (42), external (43), and ventral (44) views. Pedipalp femur and trochanter dorsal (45) view. The trichobothrial pattern is indicated in Figures 29–36 (white circles)." pageId="19" pageNumber="18">Figs. 29–45</figureCitation>
|
||
). The femur and patella are matte and granulated in males, and smooth and glossy in females. The femur bears three to four carinae; the ventroexternal carina is absent, the other carinae are granular. The patella bears seven coarsely granular carinae. The chela is smooth in both sexes, with only traces of incomplete obsolete carinae. All pedipalp segments including the trochanter are sparsely hirsute, with long, dark macrosetae in both sexes. The dentate margin of the movable finger has eight rows of granules, each with one external and one internal granule, and 5 terminal granules (4 terminal and one proximal terminal). The fixed finger has eight or nine rows of granules, each with one external and one internal granule.
|
||
</paragraph>
|
||
<paragraph id="8B929A0C7943CB06FF28FBFDE903F8D2" blockId="19.[192,802,1088,1817]" pageId="19" pageNumber="18">
|
||
AFFINITIES.
|
||
<taxonomicName id="4C2DE18F7943CB06FEA3FBFDE9E7FB9C" authority="Kovařík & Lowe & Just & Awale & Sh, 2018" authorityName="Kovařík & Lowe & Just & Awale & Sh" authorityYear="2018" box="[331,497,1088,1111]" class="Arachnida" family="Buthidae" genus="Gint" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="amoudensis" status="sp. nov.">
|
||
<emphasis id="B959461E7943CB06FEA3FBFDE9E7FB9C" box="[331,497,1088,1111]" italics="true" pageId="19" pageNumber="18">G. amoudensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26AFB657943CB06FE17FBFDEA55FB9C" box="[511,579,1088,1111]" pageId="19" pageNumber="18" rank="species">
|
||
<emphasis id="B959461E7943CB06FE17FBFDEA0EFB9C" bold="true" box="[511,536,1088,1111]" pageId="19" pageNumber="18">sp</emphasis>
|
||
.
|
||
<emphasis id="B959461E7943CB06FDC6FBFDEA2BFB9C" bold="true" box="[558,573,1088,1111]" pageId="19" pageNumber="18">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
is morphologically similar to
|
||
<taxonomicName id="4C2DE18F7943CB06FEDEFBE2E9BAFBBD" box="[310,428,1119,1142]" class="Arachnida" family="Buthidae" genus="Gint" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="gaitako">
|
||
<emphasis id="B959461E7943CB06FEDEFBE2E9BAFBBD" box="[310,428,1119,1142]" italics="true" pageId="19" pageNumber="18">G. gaitako</emphasis>
|
||
</taxonomicName>
|
||
. The DNA analysis of
|
||
<emphasis id="B959461E7943CB06FD55FBE2EAF9FBBD" box="[701,751,1119,1142]" italics="true" pageId="19" pageNumber="18">cox1</emphasis>
|
||
and
|
||
<emphasis id="B959461E7943CB06FF28FBC3E8FEFB5E" box="[192,232,1150,1173]" italics="true" pageId="19" pageNumber="18">16S</emphasis>
|
||
gene fragments (Just et al., in preparation) indicates that these populations represent separate lineages with separate areas of distribution (
|
||
<figureCitation id="131686897943CB06FDF0FB06EA6EFB19" box="[536,632,1211,1234]" captionStart="Figure 202" captionStartId="38.[192,268,1190,1213]" captionTargetBox="[196,1396,196,1156]" captionTargetId="figure-12@38.[192,1399,192,1160]" captionTargetPageId="38" captionText="Figure 202: Map showing confirmed distribution of Gint spp. Points indicate sites sampled during 2011–2017 expeditions and type locality of G. puntlandus. Scorpion shown in the photo inset is a female paratype of G. maidensis sp. n." pageId="19" pageNumber="18">Fig. 202</figureCitation>
|
||
). They can be also differentiated by morphology according to different metasomal granulation (see morphological key below) and according to the different chromosome numbers, which are 2n=35, 36 in
|
||
<taxonomicName id="4C2DE18F7943CB06FE2CFA8BEA74FA86" authority="Kovařík & Lowe & Just & Awale & Sh, 2018" authorityName="Kovařík & Lowe & Just & Awale & Sh" authorityYear="2018" box="[452,610,1334,1357]" class="Arachnida" family="Buthidae" genus="Gint" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="amoudensis" status="sp. nov.">
|
||
<emphasis id="B959461E7943CB06FE2CFA8BEA74FA86" box="[452,610,1334,1357]" italics="true" pageId="19" pageNumber="18">G. amoudensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26AFB657943CB06FD82FA8BEABEFA86" box="[618,680,1334,1357]" pageId="19" pageNumber="18" rank="species">
|
||
<emphasis id="B959461E7943CB06FD82FA8BEA95FA86" bold="true" box="[618,643,1334,1357]" pageId="19" pageNumber="18">sp</emphasis>
|
||
.
|
||
<emphasis id="B959461E7943CB06FD7AFA8BEAB7FA86" bold="true" box="[658,673,1334,1357]" pageId="19" pageNumber="18">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
and 2n=
|
||
<quantity id="4CD537E97943CB06FCEFFA8BE8C0FAA0" metricMagnitude="-1" metricUnit="m" metricValue="7.62" pageId="19" pageNumber="18" unit="in" value="30.0">30 in</quantity>
|
||
<taxonomicName id="4C2DE18F7943CB06FF09FAE9E941FAA0" box="[225,343,1364,1387]" class="Arachnida" family="Buthidae" genus="Gint" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="gaitako">
|
||
<emphasis id="B959461E7943CB06FF09FAE9E941FAA0" box="[225,343,1364,1387]" italics="true" pageId="19" pageNumber="18">G. gaitako</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<figureCitation id="131686897943CB06FE84FAE9EA1DFAA0" box="[364,523,1364,1387]" captionStart="Figures 148–163" captionStartId="33.[192,278,1689,1712]" captionTargetBox="[191,1440,191,1660]" captionTargetId="figure-6@33.[192,1439,192,1659]" captionTargetPageId="33" captionText="Figures 148–163: Figures 148–151: Gint amoudensis sp. n., paratype male, No. 1327 from Laas Gel (Locality No. 17SF), right hemispermatophore. 148. Whole hemispermatophore. 149–151. Capsule region. Views of posterior (flagellar) margin (149), convex side (150), and anterior (sperm duct) margin (151). Scale bars: 1 mm (148), 200 µm (149–151). Figures 152–163: Gint amoudensis sp. n., capsule region of right hemispermatophores from four additional paratype males, showing morphological variation. Nos. 1291 (158–160), 1292 (152–154) and 1293 (161–163) from Laas Gel (Locality No. 17SF). No. 1325 (155–157) from Borama, Armoud University (Locality No. 17SR). Views of posterior (flagellar) margin (152, 155, 158, 161), convex side (153, 156, 159, 162), and anterior (sperm duct) margin (154, 157, 160, 163). Scale bars: 200 µm (152–154; 155–157; 158–160; 161–163)." pageId="19" pageNumber="18">Figs. 150–151</figureCitation>
|
||
, see also figs.
|
||
<quantity id="4CD537E97943CB06FD56FAE9EB34FAA0" box="[702,802,1364,1387]" metricMagnitude="0" metricUnit="m" metricValue="1.5875" metricValueMax="1.6002" metricValueMin="1.5748" pageId="19" pageNumber="18" unit="in" value="62.5" valueMax="63.0" valueMin="62.0">62–63 in</quantity>
|
||
<bibRefCitation id="EFBCE7FD7943CB06FF28FACEE9DFFA41" author="KOVARIK" box="[192,457,1395,1418]" pageId="19" pageNumber="18" refId="ref17062" refString="KOVARIK, F. & A. A. OJANGUREN AFFILASTRO. 2013. Illustrated catalog of scorpions. Part II. Bothriuridae; Chaerilidae; Buthidae I. Genera Compsobuthus, Hottentotta, Isometrus, Lychas, and Sassanidotus. Prague: Clairon Production, 400 pp." type="book" year="2013">Kovařík et al., 2013: 16</bibRefCitation>
|
||
). We described
|
||
<taxonomicName id="4C2DE18F7943CB06FD68FACEEB34FA41" authority="Kovařík & Lowe & Just & Awale & Sh, 2018" authorityName="Kovařík & Lowe & Just & Awale & Sh" authorityYear="2018" box="[640,802,1395,1418]" class="Arachnida" family="Buthidae" genus="Gint" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="amoudensis" status="sp. nov.">
|
||
<emphasis id="B959461E7943CB06FD68FACEEB34FA41" box="[640,802,1395,1418]" italics="true" pageId="19" pageNumber="18">G. amoudensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26AFB657943CB06FF28FA2FE915FA62" box="[192,259,1426,1449]" pageId="19" pageNumber="18" rank="species">
|
||
<emphasis id="B959461E7943CB06FF28FA2FE8CFFA62" bold="true" box="[192,217,1426,1449]" pageId="19" pageNumber="18">sp</emphasis>
|
||
.
|
||
<emphasis id="B959461E7943CB06FF06FA2FE8EBFA62" bold="true" box="[238,253,1426,1449]" pageId="19" pageNumber="18">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
based on
|
||
<specimenCount id="9D2B51857943CB06FE6BFA2FEA0CFA62" box="[387,538,1426,1449]" pageId="19" pageNumber="18" type="generic">40 specimens</specimenCount>
|
||
from two separate localities,
|
||
<specimenCount id="9D2B51857943CB06FEF4FA0DE9A6FA0C" box="[284,432,1456,1479]" pageId="19" pageNumber="18" type="generic">15 specimens</specimenCount>
|
||
from Borama (
|
||
<typeStatus id="549624AE7943CB06FDB6FA0DEA98FA0C" box="[606,654,1456,1479]" pageId="19" pageNumber="18">type</typeStatus>
|
||
locality) and
|
||
<specimenCount id="9D2B51857943CB06FF28FA72E949FA2D" box="[192,351,1487,1510]" pageId="19" pageNumber="18" type="generic">25 specimens</specimenCount>
|
||
from
|
||
<collectingRegion id="49E954EE7943CB06FE57FA72E9E7FA2D" box="[447,497,1487,1510]" country="Tanzania" name="Mjini Magharibi" pageId="19" pageNumber="18">Laas</collectingRegion>
|
||
Gel. We noticed some variability between these two populations, with noticeable differences in coloration (
|
||
<figureCitation id="131686897943CB06FDF8F9B1EA63F9E8" box="[528,629,1548,1571]" captionStart="Figures 1–2" captionStartId="4.[192,278,1854,1877]" captionTargetBox="[209,1417,191,1825]" captionTargetId="figure-12@4.[210,1416,192,1824]" captionTargetPageId="4" captionText="Figures 1–2: Gint amoudensis sp. n., paratypes from locality No. 17SF (Laas Gel), in vivo habitus. Figure 1. Female and two males. Figure 2. Male." pageId="19" pageNumber="18">Figs. 1–2</figureCitation>
|
||
vs. 46), e.g. all species from the
|
||
<typeStatus id="549624AE7943CB06FE61F996E9A1F989" box="[393,439,1579,1602]" pageId="19" pageNumber="18">type</typeStatus>
|
||
locality are dark with partially fuscous chelicerae (
|
||
<figureCitation id="131686897943CB06FE4AF9F7EA3FF9AA" box="[418,553,1610,1633]" captionStart="Figures 11–20" captionStartId="7.[192,278,1254,1277]" captionTargetBox="[191,1437,217,1225]" captionTargetId="figure-6@7.[192,1437,218,1224]" captionTargetPageId="7" captionText="Figures 11–20: Gint amoudensis sp. n. Figures 11, 13, 17–20. Holotype male, chelicerae, carapace and tergites I–III (11), sternopectinal region and sternites III–V (13), distal segments of right legs I–IV, retrolateral views (17–20 respectively). Figures 12, 14. Paratype female from type locality, chelicerae, carapace and tergites I–III (12), sternopectinal region and sternites III–VI (14). Figure 15. Paratype male from Laas Gel, chelicerae, carapace and tergites I–III. Figure 16. Paratype female from Laas Gel, chelicerae, carapace and tergites I–III." pageId="19" pageNumber="18">Figs. 11–12</figureCitation>
|
||
) while all specimens from
|
||
<collectingRegion id="49E954EE7943CB06FF16F9D5E926F9B4" box="[254,304,1640,1663]" country="Tanzania" name="Mjini Magharibi" pageId="19" pageNumber="18">Laas</collectingRegion>
|
||
Gel are yellow with yellow chelicerae (
|
||
<figureCitation id="131686897943CB06FD1FF9D5E915F955" captionStart="Figures 11–20" captionStartId="7.[192,278,1254,1277]" captionTargetBox="[191,1437,217,1225]" captionTargetId="figure-6@7.[192,1437,218,1224]" captionTargetPageId="7" captionText="Figures 11–20: Gint amoudensis sp. n. Figures 11, 13, 17–20. Holotype male, chelicerae, carapace and tergites I–III (11), sternopectinal region and sternites III–V (13), distal segments of right legs I–IV, retrolateral views (17–20 respectively). Figures 12, 14. Paratype female from type locality, chelicerae, carapace and tergites I–III (12), sternopectinal region and sternites III–VI (14). Figure 15. Paratype male from Laas Gel, chelicerae, carapace and tergites I–III. Figure 16. Paratype female from Laas Gel, chelicerae, carapace and tergites I–III." pageId="19" pageNumber="18">Fig. 15–16</figureCitation>
|
||
). Other differences are listed in
|
||
<tableCitation id="C6AFAFB77943CB06FD9BF93AEAC4F955" box="[627,722,1671,1694]" captionStart="Table 1" captionStartId="12.[192,257,1044,1067]" captionText="Table 1: Comparative measurements of adults of Gint amoundensis sp. n. Abbreviations: length (L), width (W, in carapace it corresponds to" pageId="19" pageNumber="18">Tables 1</tableCitation>
|
||
and
|
||
<tableCitation id="C6AFAFB77943CB06FCE6F93AEB0DF955" box="[782,795,1671,1694]" captionStart="Table 3" captionStartId="29.[192,257,1072,1095]" captionText="Table 3: Comparison among Gint species (specimens), based upon selected morphometric ratios of adult males. Abbreviations: length (L), width (W), depth (D)." pageId="19" pageNumber="18">3</tableCitation>
|
||
. However, the DNA analysis (Just et al., in preparation) confirmed that these two populations form a monophyletic lineage and can be thus considered a single species.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="C337C9877943CB06FF28F8F8EC1DFC30" pageId="19" pageNumber="18" type="discussion">
|
||
<paragraph id="8B929A0C7943CB06FF28F8F8EB37F8B0" blockId="19.[192,802,1861,1915]" pageId="19" pageNumber="18">
|
||
<materialsCitation id="3B4590517943CB06FF28F8F8EB37F8B0" collectionCode="LOCALITIES, AND, LIFE, STRATEGY" pageId="19" pageNumber="18" specimenCount="1">
|
||
COMMENTS ON
|
||
<collectionCode id="ED3C02C97943CB06FE9DF8F5EAF4F890" box="[373,738,1864,1883]" pageId="19" pageNumber="18">LOCALITIES AND LIFE STRATEGY</collectionCode>
|
||
. The
|
||
<typeStatus id="549624AE7943CB06FF28F8D9E8F8F8B0" box="[192,238,1892,1915]" pageId="19" pageNumber="18">type</typeStatus>
|
||
locality, 17SR is a riverbed of an occasional river
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="8B929A0C7943CB06FCD6FF7AEB63FD85" blockId="19.[830,1441,199,1019]" pageId="19" pageNumber="18">
|
||
(
|
||
<figureCitation id="131686897943CB06FCAEFF7AEB82FF15" box="[838,916,199,222]" captionStart="Figures 46–47" captionStartId="10.[192,278,1843,1866]" captionTargetBox="[191,1440,191,1813]" captionTargetId="figure-12@10.[192,1439,192,1812]" captionTargetPageId="10" captionText="Figures 46–47: Gint amoudensis sp. n., paratype male from type locality, in vivo habitus (46) and the type locality (47)." pageId="19" pageNumber="18">Fig. 47</figureCitation>
|
||
and figs.
|
||
<quantity id="4CD537E97943CB06FC15FF7AEC4AFF15" box="[1021,1116,199,222]" metricMagnitude="0" metricUnit="m" metricValue="1.1811" metricValueMax="1.2192" metricValueMin="1.143" pageId="19" pageNumber="18" unit="in" value="46.5" valueMax="48.0" valueMin="45.0">45–48 in</quantity>
|
||
Kovařík et al., 2017: 18). The locality lies in the grounds of Amoud University Campus and is a study site for detailed research.
|
||
<taxonomicName id="4C2DE18F7943CB06FA6EFEB9EBAAFEF1" authority="Kovařík & Lowe & Just & Awale & Sh, 2018" authorityName="Kovařík & Lowe & Just & Awale & Sh" authorityYear="2018" class="Arachnida" family="Buthidae" genus="Gint" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="amoudensis" status="sp. nov.">
|
||
<emphasis id="B959461E7943CB06FA6EFEB9EBAAFEF1" italics="true" pageId="19" pageNumber="18">G. amoudensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26AFB657943CB06FC20FE9EEC1CFEF1" box="[968,1034,291,314]" pageId="19" pageNumber="18" rank="species">
|
||
<emphasis id="B959461E7943CB06FC20FE9EEBF7FEF1" bold="true" box="[968,993,291,314]" pageId="19" pageNumber="18">sp</emphasis>
|
||
.
|
||
<emphasis id="B959461E7943CB06FC1CFE9EEC15FEF1" bold="true" box="[1012,1027,291,314]" pageId="19" pageNumber="18">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
was recorded at night during ultraviolet (UV) light collecting together with
|
||
<taxonomicName id="4C2DE18F7943CB06FAEAFEFFED8FFE92" box="[1282,1433,322,345]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B959461E7943CB06FAEAFEFFED65FE92" box="[1282,1395,322,345]" italics="true" pageId="19" pageNumber="18">Neobuthus</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2DE18F7943CB06FCD7FEDDED70FEBC" authority="Pocock, 1901" authorityName="Pocock" authorityYear="1901" box="[831,1382,352,375]" class="Arachnida" family="Buthidae" genus="Parabuthus" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="abyssinicus">
|
||
<emphasis id="B959461E7943CB06FCD7FEDDEC55FEBC" box="[831,1091,352,375]" italics="true" pageId="19" pageNumber="18">Parabuthus abyssinicus</emphasis>
|
||
Pocock, 1901 (Buthidae)
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C2DE18F7943CB06FCD6FEC2ED63FE5D" authority="(Pocock, 1900)" baseAuthorityName="Pocock" baseAuthorityYear="1900" box="[830,1397,383,406]" class="Arachnida" family="Scorpionidae" genus="Pandinops" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="pugilator">
|
||
<emphasis id="B959461E7943CB06FCD6FEC2EC0BFE5D" box="[830,1053,383,406]" italics="true" pageId="19" pageNumber="18">Pandinops pugilator</emphasis>
|
||
(Pocock, 1900) (Scorpionidae)
|
||
</taxonomicName>
|
||
. At this locality, the first author recorded maximum daytime temperatures of 29.1 ºC (
|
||
<date id="FF93BCCC7943CB06FBA4FE01ED25FE18" box="[1100,1331,444,467]" pageId="19" pageNumber="18" value="2017-09-10">10th September 2017</date>
|
||
) and 31.8 ºC (
|
||
<date id="FF93BCCC7943CB06FC86FE66EC4BFE39" box="[878,1117,475,498]" pageId="19" pageNumber="18" value="2017-09-12">12th September 2017</date>
|
||
), and a minimum nighttime temperature of 19.6 ºC. The recorded humidity was between 31% (minimum at night) and 79% (maximum at day).
|
||
</paragraph>
|
||
<paragraph id="8B929A0C7943CB06FC86FDEBEC1DFC30" blockId="19.[830,1441,199,1019]" pageId="19" pageNumber="18">
|
||
The second locality 17SF (
|
||
<figureCitation id="131686897943CB06FB7FFDEBECF0FDA6" box="[1175,1254,598,621]" captionStart="Figure 48" captionStartId="11.[192,268,1329,1352]" captionTargetBox="[191,1917,227,1297]" captionTargetId="figure-6@11.[192,1916,228,1296]" captionTargetPageId="11" captionText="Figure 48: Gint amoudensis sp. n., locality No. 17SF, Laas Gel, 50 km NE Hargeisa." pageId="19" pageNumber="18">Fig. 48</figureCitation>
|
||
,
|
||
<collectingRegion id="49E954EE7943CB06FB1EFDEBED3EFDA6" box="[1270,1320,598,621]" country="Tanzania" name="Mjini Magharibi" pageId="19" pageNumber="18">Laas</collectingRegion>
|
||
Gel), is in rocky semi-desert terrain with the riverbed of an occasional river in the center.
|
||
<taxonomicName id="4C2DE18F7943CB06FB62FD2EED3EFD61" authority="Kovařík & Lowe & Just & Awale & Sh, 2018" authorityName="Kovařík & Lowe & Just & Awale & Sh" authorityYear="2018" box="[1162,1320,659,682]" class="Arachnida" family="Buthidae" genus="Gint" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="amoudensis" status="sp. nov.">
|
||
<emphasis id="B959461E7943CB06FB62FD2EED3EFD61" box="[1162,1320,659,682]" italics="true" pageId="19" pageNumber="18">G. amoudensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="A26AFB657943CB06FAD8FD2EED79FD61" box="[1328,1391,659,682]" pageId="19" pageNumber="18" rank="species">
|
||
<emphasis id="B959461E7943CB06FAD8FD2EED5FFD61" bold="true" box="[1328,1353,659,682]" pageId="19" pageNumber="18">sp</emphasis>
|
||
.
|
||
<emphasis id="B959461E7943CB06FAB1FD2EED7EFD61" bold="true" box="[1369,1384,659,682]" pageId="19" pageNumber="18">n</emphasis>
|
||
.
|
||
</taxonomicNameLabel>
|
||
was recorded at night during UV light collecting together with
|
||
<taxonomicName id="4C2DE18F7943CB06FC96FD6DEC09FD2C" box="[894,1055,720,743]" class="Arachnida" family="Buthidae" genus="Neobuthus" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="undetermined">
|
||
<emphasis id="B959461E7943CB06FC96FD6DEBE6FD2C" box="[894,1008,720,743]" italics="true" pageId="19" pageNumber="18">Neobuthus</emphasis>
|
||
sp.
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2DE18F7943CB06FBDEFD6DEB68FCCD" authority="(Pocock, 1896)" baseAuthorityName="Pocock" baseAuthorityYear="1896" class="Arachnida" family="Buthidae" genus="Hottentotta" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="polystictus">
|
||
<emphasis id="B959461E7943CB06FBDEFD6DED24FD2C" box="[1078,1330,720,743]" italics="true" pageId="19" pageNumber="18">Hottentotta polystictus</emphasis>
|
||
(Pocock, 1896)
|
||
</taxonomicName>
|
||
,
|
||
<taxonomicName id="4C2DE18F7943CB06FC66FD52ED8CFCCD" authority="Pocock, 1897" authorityName="Pocock" authorityYear="1897" box="[910,1434,751,774]" class="Arachnida" family="Buthidae" genus="Parabuthus" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="heterurus">
|
||
<emphasis id="B959461E7943CB06FC66FD52EC6AFCCD" box="[910,1148,751,774]" italics="true" pageId="19" pageNumber="18">Parabuthus heterurus</emphasis>
|
||
Pocock, 1897 (Buthidae)
|
||
</taxonomicName>
|
||
, and
|
||
<taxonomicName id="4C2DE18F7943CB06FC85FCB3EBB0FC88" authority="(Kovarik, 2012)" baseAuthorityName="Kovarik" baseAuthorityYear="2012" class="Arachnida" family="Scorpionidae" genus="Pandiborellius" kingdom="Animalia" order="Scorpiones" pageId="19" pageNumber="18" phylum="Arthropoda" rank="species" species="somalilandus">
|
||
<emphasis id="B959461E7943CB06FC85FCB3ECB4FCEE" box="[877,1186,782,805]" italics="true" pageId="19" pageNumber="18">Pandiborellius somalilandus</emphasis>
|
||
(Kovařík, 2012) (Scorpionidae)
|
||
</taxonomicName>
|
||
. At this locality, the first author recorded maximum daytime temperatures of 33.8 ºC (
|
||
<date id="FF93BCCC7943CB06FAF5FCF6EB62FC4A" pageId="19" pageNumber="18" value="2017-08-28">28th August 2017</date>
|
||
) and 34.7 ºC (
|
||
<date id="FF93BCCC7943CB06FBFFFCD7ECCDFC4A" box="[1047,1243,874,897]" pageId="19" pageNumber="18" value="2017-08-29">29th August 2017</date>
|
||
), and a minimum nighttime temperature of 22.8 ºC (
|
||
<date id="FF93BCCC7943CB06FB22FC35ED81FC54" box="[1226,1431,904,927]" pageId="19" pageNumber="18" value="2017-08-29">29th August 2017</date>
|
||
) and 23.2 ºC (
|
||
<date id="FF93BCCC7943CB06FC27FC1AEC85FC75" box="[975,1171,935,958]" pageId="19" pageNumber="18" value="2017-08-30">30th August 2017</date>
|
||
). The recorded humidity was between 26% (minimum at night) and 54% (maximum at day).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |