treatments-xml/data/03/87/47/038747324C45C6611EB72DBAFED12521.xml

<document id="E955ED8330107951BB1D6C8BDBBC0EF7" ID-CLB-Dataset="38930" ID-DOI="10.11646/zootaxa.4118.1.1" ID-GBIF-Dataset="a4e408c4-2575-4b5c-878b-e1b9010ec619" ID-ISSN="1175-5326" ID-Zenodo-Dep="264597" ID-ZooBank="CFA586DA-10EE-468B-80EE-35351E3845FD" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1465242077321" checkinUser="plazi" docAuthor="C, Lees David" docDate="2016" docId="038747324C45C6611EB72DBAFED12521" docLanguage="en" docName="zootaxa.4118.pdf.imf" docOrigin="Zootaxa 4118 (1)" docStyle="DocumentStyle:5EBBA59367AD13919D70D935FA04F6A3.14:Zootaxa.2013-.monograph" docStyleId="5EBBA59367AD13919D70D935FA04F6A3" docStyleName="Zootaxa.2013-.monograph" docStyleVersion="14" docTitle="Heteropsis avaratra Lees &amp; Kremen, sp. nov." docType="treatment" docVersion="9" lastPageNumber="54" masterDocId="FFBE3F4A4C76C6571E202D2DFFD32174" masterDocTitle="Heteropsis (Nymphalidae: Satyrinae: Satyrini: Mycalesina): 19 new species from Madagascar and interim revision" masterLastPageNumber="97" masterPageNumber="1" pageNumber="52" updateTime="1698662715872" updateUser="ExternalLinkService">
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<mods:title id="3D698A548A1BBB00BBA7FA55DF1F1385">Heteropsis (Nymphalidae: Satyrinae: Satyrini: Mycalesina): 19 new species from Madagascar and interim revision</mods:title>
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<mods:namePart id="E31A264526F90CA99452C487D18AF373">C, Lees David</mods:namePart>
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<mods:date id="6B27F3CFACDA3165066CDD4654F87147">2016</mods:date>
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<treatment id="038747324C45C6611EB72DBAFED12521" ID-DOI="http://doi.org/10.5281/zenodo.6086434" ID-GBIF-Taxon="120817127" ID-Zenodo-Dep="6086434" LSID="urn:lsid:plazi:treatment:038747324C45C6611EB72DBAFED12521" httpUri="http://treatment.plazi.org/id/038747324C45C6611EB72DBAFED12521" lastPageId="54" lastPageNumber="54" pageId="51" pageNumber="52">
<subSubSection id="C334A5AF4C45C6641EB72DBAFBC021A7" pageId="51" pageNumber="52" type="nomenclature">
<paragraph id="8B91F6244C45C6641EB72DBAFD6C21C6" blockId="51.[151,1043,151,211]" box="[151,703,151,178]" pageId="51" pageNumber="52">
<heading id="D0D941484C45C6641EB72DBAFD6C21C6" bold="true" box="[151,703,151,178]" fontSize="11" level="1" pageId="51" pageNumber="52" reason="1">
<emphasis id="B95A2A364C45C6641EB72DBAFD6C21C6" bold="true" box="[151,703,151,178]" pageId="51" pageNumber="52">
<taxonomicName id="4C2E8DA74C45C6641EB72DBAFD8721C6" authority="Lees &amp; Kremen" authorityName="Lees &amp; Kremen" box="[151,596,151,178]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="avaratra" status="sp. nov.">
<emphasis id="B95A2A364C45C6641EB72DBAFE5621C5" bold="true" box="[151,389,151,177]" italics="true" pageId="51" pageNumber="52">Heteropsis avaratra</emphasis>
Lees &amp; Kremen
</taxonomicName>
, 
<taxonomicNameLabel id="A269974D4C45C6641C412DB5FD6C21C6" box="[609,703,152,178]" pageId="51" pageNumber="52" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
<paragraph id="8B91F6244C45C6641EB72D96FBC021A7" blockId="51.[151,1043,151,211]" box="[151,1043,186,211]" pageId="51" pageNumber="52">LSID: urn:lsid:zoobank.org:act:17C4A71B-C04D-4AC1-9407-4D52B343F803</paragraph>
</subSubSection>
<subSubSection id="C334A5AF4C45C6641EB72DDAFF012040" pageId="51" pageNumber="52" type="description">
<paragraph id="8B91F6244C45C6641EB72DDAFF012040" blockId="51.[151,1437,247,2013]" pageId="51" pageNumber="52">
<emphasis id="B95A2A364C45C6641EB72DDAFEB22064" bold="true" box="[151,353,247,272]" pageId="51" pageNumber="52">Prior references:</emphasis>
sp. 25 (
<bibRefCitation id="EFBF8BD54C45C6641F9C2DD5FDE52064" author="Lees" box="[444,566,248,272]" pageId="51" pageNumber="52" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees 1997</bibRefCitation>
: 64, 
<bibRefCitation id="EFBF8BD54C45C6641C4C2DD5FCED2064" author="Torres" box="[620,830,247,272]" pageId="51" pageNumber="52" refString="Torres, E., Lees, D. C., Vane-Wright, R. I., Kremen, C., Leonard, J. A. &amp; Wayne, R. K. (2001) Examining monophyly in a large radiation of Madagascan butterflies (Lepidoptera: Satyrinae: Mycalesina) based on mitochondrial DNA data. Molecular Phylogenetics and Evolution, 20, 460 - 473. http: // dx. doi. org / 10.1006 / mpev. 2001.0984" type="journal article" year="2001">
Torres 
<emphasis id="B95A2A364C45C6641C9B2DD4FD292064" box="[699,762,247,272]" italics="true" pageId="51" pageNumber="52">et al.,</emphasis>
2001
</bibRefCitation>
: 462); ‘ 
<taxonomicName id="4C2E8DA74C45C6641DB62DD4FBFA2064" box="[918,1065,247,272]" class="Insecta" family="Nymphalidae" genus="Pseudonympha" kingdom="Animalia" order="Lepidoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="subsimilis">
<emphasis id="B95A2A364C45C6641DB62DD4FBFA2064" box="[918,1065,247,272]" italics="true" pageId="51" pageNumber="52">H. subsimilis</emphasis>
</taxonomicName>
’ ♂, (
<bibRefCitation id="EFBF8BD54C45C6641A4D2DDAFAC42064" author="d'Abrera" box="[1133,1303,247,272]" pageId="51" pageNumber="52" refString="d'Abrera, B. L. (1980) Butterflies of the Afrotropical Region. Lansdowne Editions, Melbourne, 593 pp." type="book" year="1980">d’Abrera 1980</bibRefCitation>
: 182; 1997: 221).
</paragraph>
</subSubSection>
<subSubSection id="C334A5AF4C45C6641EE72C6FFC0B2381" pageId="51" pageNumber="52" type="materials_examined">
<paragraph id="8B91F6244C45C6641EE72C6FFE9A20D1" blockId="51.[151,1437,247,2013]" pageId="51" pageNumber="52">
<emphasis id="B95A2A364C45C6641EE72C6FFD35202F" bold="true" box="[199,742,322,347]" pageId="51" pageNumber="52">
<typeStatus id="549548864C45C6641EE72C6FFED1202F" box="[199,258,322,347]" pageId="51" pageNumber="52">Type</typeStatus>
material., Deposition 
<collectionCode id="ED3F6EE14C45C6641C292C6FFDB7202F" box="[521,612,322,347]" pageId="51" pageNumber="52">BMNH</collectionCode>
: 
<typeStatus id="549548864C45C6641C502C6FFD32202F" box="[624,737,322,347]" pageId="51" pageNumber="52" type="holotype">Holotype</typeStatus>
:
</emphasis>
♂ (
<figureCitation id="1315EAA14C45C6641D312C6FFCB3202E" box="[785,864,322,347]" captionStart="FIGURE 13" captionStartId="47.[151,250,1920,1942]" captionTargetBox="[243,1342,223,1890]" captionTargetId="figure@47.[226,1359,193,1906]" captionTargetPageId="47" captionText="FIGURE 13. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis kremenae, Lees, sp. nov. (Ambavaony R., Masoala; NHMUK 010289135). B: PT ♀ of Ht. kremenae (Masoala; NHMUK 010289139). C: HT ♂ of Heteropsis avaratra Lees &amp; Kremen, sp. nov. (Montagne d’Ambre; NHMUK 010289141). D: PT ♀ of Ht. avaratra (Montagne d’Ambre, Les Roussettes; DCL-DB- 3385, MNHN)." httpUri="https://zenodo.org/record/264610/files/figure.png" pageId="51" pageNumber="52">Fig. 13</figureCitation>
C), 
<collectingCountry id="F339B6B44C45C6641DAB2C6FFBC1202F" box="[907,1042,322,347]" name="Madagascar" pageId="51" pageNumber="52">Madagascar</collectingCountry>
N, Montagne d'Ambre, 
<geoCoordinate id="EE1A90E34C45C6641B032C6FFA45202E" box="[1315,1430,318,347]" direction="south" orientation="latitude" pageId="51" pageNumber="52" precision="5" value="-12.5289">12.5289o S</geoCoordinate>
, 
<geoCoordinate id="EE1A90E34C45C6641EB72C44FED920F4" box="[151,266,357,385]" direction="east" orientation="longitude" pageId="51" pageNumber="52" precision="5" value="49.1727">49.1727o E</geoCoordinate>
+/- 
<quantity id="4CD65BC14C45C6641F1C2C45FE4E20F5" box="[316,413,360,385]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="51" pageNumber="52" unit="km" value="0.15">0.15 km</quantity>
, 
<quantity id="4CD65BC14C45C6641F892C45FD8320F4" box="[425,592,360,385]" metricMagnitude="3" metricUnit="m" metricValue="1.057" metricValueMax="1.082" metricValueMin="1.032" pageId="51" pageNumber="52" unit="m" value="1057.0" valueMax="1082.0" valueMin="1032.0">1057 +/- 25 m</quantity>
, 
<date id="FF90D0E44C45C6641C7E2C44FD0C20F5" box="[606,735,361,385]" pageId="51" pageNumber="52" value="2004-11-18">18/11/2004</date>
, fruit trap MD3, D.C. Lees: DL-05-169, 
<collectionCode id="ED3F6EE14C45C6641A942C44FACC20F5" box="[1204,1311,361,385]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="51" pageNumber="52">NHMUK</collectionCode>
010289141 [
<collectingCountry id="F339B6B44C45C6641EBD2CA0FF0B20D1" box="[157,216,397,421]" name="Qatar" pageId="51" pageNumber="52">QTR</collectingCountry>
barcode].
</paragraph>
<paragraph id="8B91F6244C45C6641EE72C9EFC0B2381" blockId="51.[151,1437,247,2013]" pageId="51" pageNumber="52">
<typeStatus id="549548864C45C6641EE72C9EFEEC20B8" box="[199,319,435,460]" pageId="51" pageNumber="52" type="paratype">
<emphasis id="B95A2A364C45C6641EE72C9EFEEC20B8" bold="true" box="[199,319,435,460]" pageId="51" pageNumber="52">Paratypes</emphasis>
</typeStatus>
: 
<emphasis id="B95A2A364C45C6641F6D2C9EFDF820B8" bold="true" box="[333,555,435,460]" pageId="51" pageNumber="52">
Deposition 
<collectionCode id="ED3F6EE14C45C6641FF42C9EFDF820B8" box="[468,555,435,460]" pageId="51" pageNumber="52">BMNH</collectionCode>
</emphasis>
: ♂, 
<collectingCountry id="F339B6B44C45C6641C7C2C9EFD3020B8" box="[604,739,435,460]" name="Madagascar" pageId="51" pageNumber="52">Madagascar</collectingCountry>
N, Montagne d'Ambre, 
<geoCoordinate id="EE1A90E34C45C6641DD42C9EFBB420BF" box="[1012,1127,431,459]" direction="south" orientation="latitude" pageId="51" pageNumber="52" precision="5" value="-12.4984">12.4984o S</geoCoordinate>
, 
<geoCoordinate id="EE1A90E34C45C6641A542C9EFB3420BF" box="[1140,1255,431,460]" direction="east" orientation="longitude" pageId="51" pageNumber="52" precision="5" value="49.1754">49.1754o E</geoCoordinate>
+/- 
<quantity id="4CD65BC14C45C6641B362C9EFAB520B8" box="[1302,1382,435,460]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="51" pageNumber="52" unit="km" value="0.5">0.5 km</quantity>
, 
<quantity id="4CD65BC14C45C6641B522C9EFF39209B" metricMagnitude="2" metricUnit="m" metricValue="8.12" metricValueMax="8.17" metricValueMin="8.07" pageId="51" pageNumber="52" unit="m" value="812.0" valueMax="817.0" valueMin="807.0">812 +/- 5 m</quantity>
<date id="FF90D0E44C45C6641ED02CFAFEA2209B" box="[240,369,471,495]" pageId="51" pageNumber="52" value="2004-11-17">17/11/2004</date>
, D.C. Lees: DL-05-69, 
<collectionCode id="ED3F6EE14C45C6641C5A2CFAFD02209B" box="[634,721,471,495]" pageId="51" pageNumber="52">BMNH</collectionCode>
(E) #676332; IA82 [isotope voucher], CCDB-02225-B04 [
<collectionCode id="ED3F6EE14C45C6641B7C2CFAFA4F209B" box="[1372,1436,471,495]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="51" pageNumber="52">DNA</collectionCode>
barcode voucher]; ♂, N, Montagne d'Ambre, 
<geoCoordinate id="EE1A90E34C45C6641CB72CD0FCD92362" box="[663,778,506,534]" direction="south" orientation="latitude" pageId="51" pageNumber="52" precision="5" value="-12.5289">12.5289o S</geoCoordinate>
, 
<geoCoordinate id="EE1A90E34C45C6641D372CD3FC5E2361" box="[791,909,506,534]" direction="east" orientation="longitude" pageId="51" pageNumber="52" precision="5" value="49.1727">49.1727o E</geoCoordinate>
, +/- 
<quantity id="4CD65BC14C45C6641DE12CD0FBF22362" box="[961,1057,509,534]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="51" pageNumber="52" unit="km" value="0.15">0.15 km</quantity>
, 
<quantity id="4CD65BC14C45C6641A0C2CD0FB1D2361" box="[1068,1230,509,534]" metricMagnitude="3" metricUnit="m" metricValue="1.057" metricValueMax="1.082" metricValueMin="1.032" pageId="51" pageNumber="52" unit="m" value="1057.0" valueMax="1082.0" valueMin="1032.0">1057 +/- 25 m</quantity>
, 
<date id="FF90D0E44C45C6641AFA2CD0FA882362" box="[1242,1371,509,534]" pageId="51" pageNumber="52" value="2004-11-16">16/11/2004</date>
, D.C. Lees: DL-05-68, 
<collectionCode id="ED3F6EE14C45C6641F412F0CFE6B234D" box="[353,440,545,569]" pageId="51" pageNumber="52">BMNH</collectionCode>
(E) #676331, 2133 (= 
<collectionCode id="ED3F6EE14C45C6641C942F0CFD0F234D" box="[692,732,545,569]" pageId="51" pageNumber="52">DL</collectionCode>
2133; 
<collectionCode id="ED3F6EE14C45C6641D062F0CFCB0234D" box="[806,867,545,569]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="51" pageNumber="52">DNA</collectionCode>
extract number), IA91 [isotope voucher]; ♀, N, Montagne d'Ambre, 
<quantity id="4CD65BC14C45C6641FA12F65FE09232B" box="[385,474,584,608]" metricMagnitude="3" metricUnit="m" metricValue="1.2" pageId="51" pageNumber="52" unit="m" value="1200.0">1200 m</quantity>
, 
<geoCoordinate id="EE1A90E34C45C6641FC72F6AFD882314" box="[487,603,580,608]" direction="south" orientation="latitude" pageId="51" pageNumber="52" precision="5" value="-12.5405">12.5405o S</geoCoordinate>
, 
<geoCoordinate id="EE1A90E34C45C6641C492F6AFD0E232B" box="[617,733,580,608]" direction="east" orientation="longitude" pageId="51" pageNumber="52" precision="5" value="49.1682">49.1682o E</geoCoordinate>
+/- 
<quantity id="4CD65BC14C45C6641D2E2F65FC9F2314" box="[782,844,583,608]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="51" pageNumber="52" unit="km" value="1.0">1 km</quantity>
, 
<quantity id="4CD65BC14C45C6641D782F65FC2C232B" box="[856,1023,583,608]" metricMagnitude="3" metricUnit="m" metricValue="1.2" metricValueMax="1.25" metricValueMin="1.15" pageId="51" pageNumber="52" unit="m" value="1200.0" valueMax="1250.0" valueMin="1150.0">1200 +/- 50 m</quantity>
, 
<date id="FF90D0E44C45C6641A2C2F6AFB532314" box="[1036,1152,583,608]" pageId="51" pageNumber="52" value="1995-01-13">13/1/1995</date>
, D.C. Lees: 
<collectionCode id="ED3F6EE14C45C6641B302F6AFAA92314" box="[1296,1402,583,608]" pageId="51" pageNumber="52">DLMDA</collectionCode>
95- 0 0 0 1, 
<collectionCode id="ED3F6EE14C45C6641EFE2F43FE9B23F2" box="[222,328,622,646]" country="United Kingdom" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34665" name="Natural History Museum, London" pageId="51" pageNumber="52">NHMUK</collectionCode>
010289142 [
<collectingCountry id="F339B6B44C45C6641FF42F43FDDC23F2" box="[468,527,622,646]" name="Qatar" pageId="51" pageNumber="52">QTR</collectingCountry>
barcode]; ♂, N, Montagne d'Ambre, 
<quantity id="4CD65BC14C45C6641D9A2F43FBD723F2" box="[954,1028,622,646]" metricMagnitude="2" metricUnit="m" metricValue="9.0" pageId="51" pageNumber="52" unit="m" value="900.0">900 m</quantity>
, 
<geoCoordinate id="EE1A90E34C45C6641A312F40FBA523F2" box="[1041,1142,618,646]" direction="south" orientation="latitude" pageId="51" pageNumber="52" precision="55" value="-12.525">12.525o S</geoCoordinate>
, 
<geoCoordinate id="EE1A90E34C45C6641AA42F43FB2B23F2" box="[1156,1272,618,646]" direction="east" orientation="longitude" pageId="51" pageNumber="52" precision="5" value="49.1722">49.1722o E</geoCoordinate>
+/- 
<quantity id="4CD65BC14C45C6641B092F40FAB523F2" box="[1321,1382,621,646]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="51" pageNumber="52" unit="km" value="5.0">5 km</quantity>
, 
<quantity id="4CD65BC14C45C6641B522F43FF2B23DD" metricMagnitude="2" metricUnit="m" metricValue="9.0" metricValueMax="9.05" metricValueMin="8.95" pageId="51" pageNumber="52" unit="m" value="900.0" valueMax="905.0" valueMin="895.0">900 +/- 5 m</quantity>
, 
<date id="FF90D0E44C45C6641F2B2FBCFEAD23DE" box="[267,382,657,682]" pageId="51" pageNumber="52" value="1995-01-16">16/1/1995</date>
, 0 945 [= 
<collectionCode id="ED3F6EE14C45C6641FCC2FBCFDC623DE" box="[492,533,657,682]" pageId="51" pageNumber="52">DL</collectionCode>
0945], 
<collectionCode id="ED3F6EE14C45C6641C4A2FBFFD1223DE" box="[618,705,658,682]" pageId="51" pageNumber="52">BMNH</collectionCode>
(E) #672310 [
<collectionCode id="ED3F6EE14C45C6641D442FBFFC7723DE" box="[868,932,658,682]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="51" pageNumber="52">DNA</collectionCode>
voucher], 
<accessionNumber id="947D6BC74C45C6641A172FBFFB7B23DE" box="[1079,1192,658,682]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/FJ819380" pageId="51" pageNumber="52">
<collectionCode id="ED3F6EE14C45C6641A172FBFFB8123DE" box="[1079,1106,658,682]" pageId="51" pageNumber="52">FJ</collectionCode>
819380
</accessionNumber>
, cob 357 bp; ♂, N, Montagne d'Ambre, c. 
<quantity id="4CD65BC14C45C6641F802F94FE2923A4" box="[416,506,697,721]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="51" pageNumber="52" unit="m" value="1000.0">1000 m</quantity>
, 
<geoCoordinate id="EE1A90E34C45C6641C282F95FDBD23A5" box="[520,622,693,721]" direction="south" orientation="latitude" pageId="51" pageNumber="52" precision="55" value="-12.525">12.525o S</geoCoordinate>
, 
<geoCoordinate id="EE1A90E34C45C6641C5D2F94FD2223A4" box="[637,753,693,721]" direction="east" orientation="longitude" pageId="51" pageNumber="52" precision="5" value="49.1722">49.1722o E</geoCoordinate>
+/- 
<quantity id="4CD65BC14C45C6641D052F95FCB123A5" box="[805,866,696,721]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="51" pageNumber="52" unit="km" value="5.0">5 km</quantity>
, 
<quantity id="4CD65BC14C45C6641D512F94FC2F23A4" box="[881,1020,696,721]" metricMagnitude="2" metricUnit="m" metricValue="9.0" metricValueMax="9.05" metricValueMin="8.95" pageId="51" pageNumber="52" unit="m" value="900.0" valueMax="905.0" valueMin="895.0">900 +/- 5 m</quantity>
<date id="FF90D0E44C45C6641A262F95FBA923A5" box="[1030,1146,696,721]" pageId="51" pageNumber="52" value="1995-01-16">16/1/1995</date>
, D.C. Lees, DCLW-0153 [wing prep.]; 
<collectionCode id="ED3F6EE14C45C6641F102FF1FE542380" box="[304,391,732,756]" pageId="51" pageNumber="52">BMNH</collectionCode>
(E) #672438 [
<collectionCode id="ED3F6EE14C45C6641C3E2FF1FD8D2380" box="[542,606,732,756]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="51" pageNumber="52">DNA</collectionCode>
voucher; 
<accessionNumber id="947D6BC74C45C6641CEE2FF1FCEC2381" box="[718,831,732,757]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/FJ819358" pageId="51" pageNumber="52">
<collectionCode id="ED3F6EE14C45C6641CEE2FF1FD3A2381" box="[718,745,732,757]" pageId="51" pageNumber="52">FJ</collectionCode>
819358
</accessionNumber>
, cob 357 bp];
</paragraph>
</subSubSection>
<subSubSection id="C334A5AF4C45C6641EE72FD2FAF82285" pageId="51" pageNumber="52" type="description">
<paragraph id="8B91F6244C45C6641EE72FD2FB9C22F0" blockId="51.[151,1437,247,2013]" pageId="51" pageNumber="52">
<emphasis id="B95A2A364C45C6641EE72FD2FE7A226C" bold="true" box="[199,425,767,792]" pageId="51" pageNumber="52">Deposition MNHN</emphasis>
: ♂ [MNHN], 
<collectingCountry id="F339B6B44C45C6641C712E2DFCCB226C" box="[593,792,768,792]" name="Madagascar" pageId="51" pageNumber="52">MADAGASCAR</collectingCountry>
NORD Montagne d’Ambre Les Roussettes alt. 
<quantity id="4CD65BC14C45C6641B652E2DFA4F2263" box="[1349,1436,768,792]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="51" pageNumber="52" unit="m" value="1000.0">1000 m</quantity>
<date id="FF90D0E44C45C6641EB72E0EFEC42248" box="[151,279,803,828]" pageId="51" pageNumber="52" value="1958-11-27">27.XI.1958</date>
P. Viette|DCL-DB-3364; ♂ [MNHN], 
<collectingCountry id="F339B6B44C45C6641CD62E09FC6E2248" box="[758,957,804,828]" name="Madagascar" pageId="51" pageNumber="52">MADAGASCAR</collectingCountry>
NORD MONTAGNE D’AMBRE LES ROUSSETTES 
<quantity id="4CD65BC14C45C6641F712E6AFE78222B" box="[337,427,839,863]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="51" pageNumber="52" unit="m" value="1000.0">1000 m</quantity>
<date id="FF90D0E44C45C6641F972E6AFDE32214" box="[439,560,839,864]" pageId="51" pageNumber="52" value="1970-05-25">25-V-1970</date>
P. GRIVEAUD|DCL-DB-3381; ♀ [MNHN] (
<figureCitation id="1315EAA14C45C6641A6E2E6AFB77222B" box="[1102,1188,839,864]" captionStart="FIGURE 13" captionStartId="47.[151,250,1920,1942]" captionTargetBox="[243,1342,223,1890]" captionTargetId="figure@47.[226,1359,193,1906]" captionTargetPageId="47" captionText="FIGURE 13. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis kremenae, Lees, sp. nov. (Ambavaony R., Masoala; NHMUK 010289135). B: PT ♀ of Ht. kremenae (Masoala; NHMUK 010289139). C: HT ♂ of Heteropsis avaratra Lees &amp; Kremen, sp. nov. (Montagne d’Ambre; NHMUK 010289141). D: PT ♀ of Ht. avaratra (Montagne d’Ambre, Les Roussettes; DCL-DB- 3385, MNHN)." httpUri="https://zenodo.org/record/264610/files/figure.png" pageId="51" pageNumber="52">Fig. 13</figureCitation>
D), 
<collectingCountry id="F339B6B44C45C6641AF52E6AFA4F222B" box="[1237,1436,839,863]" name="Madagascar" pageId="51" pageNumber="52">MADAGASCAR</collectingCountry>
NORD Montagne d’Ambre Les Roussettes alt. 
<quantity id="4CD65BC14C45C6641C8A2E46FD2C22F7" box="[682,767,875,899]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="51" pageNumber="52" unit="m" value="1000.0">1000 m</quantity>
<date id="FF90D0E44C45C6641D262E46FC4B22F0" box="[774,920,875,900]" pageId="51" pageNumber="52" value="1958-12-06" valueMax="1958-12-07" valueMin="1958-12-06">6-7 XII 1958</date>
|DCL-DB-3385;
</paragraph>
<paragraph id="8B91F6244C45C6641EE72EBCFDB722BA" blockId="51.[151,1437,247,2013]" pageId="51" pageNumber="52">
<emphasis id="B95A2A364C45C6641EE72EBCFE4722DE" bold="true" box="[199,404,913,938]" pageId="51" pageNumber="52">Deposition ABRI</emphasis>
: ♂, 
<collectingCountry id="F339B6B44C45C6641FE72EBCFD9D22DE" box="[455,590,913,938]" name="Madagascar" pageId="51" pageNumber="52">Madagascar</collectingCountry>
N, Montagne d'Ambre, 
<geoCoordinate id="EE1A90E34C45C6641D422EBCFC0522DE" box="[866,982,910,938]" direction="south" orientation="latitude" pageId="51" pageNumber="52" precision="5" value="-12.5289">12.5289o S</geoCoordinate>
, 
<geoCoordinate id="EE1A90E34C45C6641DC32EBFFB8422DD" box="[995,1111,910,938]" direction="east" orientation="longitude" pageId="51" pageNumber="52" precision="5" value="49.1727">49.1727o E</geoCoordinate>
+/- 
<quantity id="4CD65BC14C45C6641AA82EBCFB3422DE" box="[1160,1255,913,938]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="51" pageNumber="52" unit="km" value="0.15">0.15 km</quantity>
, 
<quantity id="4CD65BC14C45C6641AD32EBCFA4422DD" box="[1267,1431,913,938]" metricMagnitude="3" metricUnit="m" metricValue="1.057" metricValueMax="1.082" metricValueMin="1.032" pageId="51" pageNumber="52" unit="m" value="1057.0" valueMax="1082.0" valueMin="1032.0">1057 +/- 25 m</quantity>
, 
<date id="FF90D0E44C45C6641EB72E9BFECB22BA" box="[151,280,950,974]" pageId="51" pageNumber="52" value="2004-11-19">19/11/2004</date>
, R. Ranaivosolo: DL-05-181.
</paragraph>
<paragraph id="8B91F6244C45C6641EE72EF4FAF82285" blockId="51.[151,1437,247,2013]" box="[199,1323,985,1010]" pageId="51" pageNumber="52">
<emphasis id="B95A2A364C45C6641EE72EF4FE1B2286" bold="true" box="[199,456,985,1010]" pageId="51" pageNumber="52">Deposition summary:</emphasis>
BMNH (HT♂, 4 PT♂♂, 1 PT♀)), MNHN (2 PT♂♂, 1 PT ♀), ABRI (PT♂).
</paragraph>
</subSubSection>
<subSubSection id="C334A5AF4C45C6641EE72ED2FAE42563" box="[199,1335,1020,1048]" pageId="51" pageNumber="52" type="materials_examined">
<paragraph id="8B91F6244C45C6641EE72ED2FAE42563" blockId="51.[151,1437,247,2013]" box="[199,1335,1020,1048]" pageId="51" pageNumber="52">
<emphasis id="B95A2A364C45C6641EE72ED2FEB6256C" bold="true" box="[199,357,1023,1048]" pageId="51" pageNumber="52">
<typeStatus id="549548864C45C6641EE72ED2FED1256C" box="[199,258,1023,1048]" pageId="51" pageNumber="52">Type</typeStatus>
locality.
</emphasis>
<collectingCountry id="F339B6B44C45C6641F4C2ED2FE20256C" box="[364,499,1023,1048]" name="Madagascar" pageId="51" pageNumber="52">Madagascar</collectingCountry>
N, Montagne d'Ambre, 
<geoCoordinate id="EE1A90E34C45C6641D242ED2FCA4256C" box="[772,887,1020,1048]" direction="south" orientation="latitude" pageId="51" pageNumber="52" precision="5" value="-12.5289">12.5289o S</geoCoordinate>
, 
<geoCoordinate id="EE1A90E34C45C6641DA4292DFC2B2563" box="[900,1016,1020,1048]" direction="east" orientation="longitude" pageId="51" pageNumber="52" precision="5" value="49.1727">49.1727o E</geoCoordinate>
+/- 
<quantity id="4CD65BC14C45C6641A072ED2FB56256C" box="[1063,1157,1023,1048]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="51" pageNumber="52" unit="km" value="0.15">0.15 km</quantity>
, 
<quantity id="4CD65BC14C45C6641AB02ED2FAE02563" box="[1168,1331,1023,1048]" metricMagnitude="3" metricUnit="m" metricValue="1.057" metricValueMax="1.082" metricValueMin="1.032" pageId="51" pageNumber="52" unit="m" value="1057.0" valueMax="1082.0" valueMin="1032.0">
1057 +/- 
<specimenCount id="9D283DAD4C45C6641AD72ED2FAE02563" box="[1271,1331,1023,1048]" pageId="51" pageNumber="52" type="generic">25 m</specimenCount>
</quantity>
.
</paragraph>
</subSubSection>
<subSubSection id="C334A5AF4C45C6641EE7290EFE832441" pageId="51" pageNumber="52" type="diagnosis">
<paragraph id="8B91F6244C45C6641EE7290EFE832441" blockId="51.[151,1437,247,2013]" pageId="51" pageNumber="52">
<emphasis id="B95A2A364C45C6641EE7290EFE922548" bold="true" box="[199,321,1059,1084]" pageId="51" pageNumber="52">Diagnosis.</emphasis>
compared with 
<emphasis id="B95A2A364C45C6641FDD2909FCEB2548" box="[509,824,1059,1084]" italics="true" pageId="51" pageNumber="52">
<taxonomicName id="4C2E8DA74C45C6641FDD2909FD5C254F" box="[509,655,1060,1083]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="avaratra">Ht. avaratra</taxonomicName>
, 
<taxonomicName id="4C2E8DA74C45C6641CBC2909FCEB2548" box="[668,824,1059,1084]" class="Insecta" family="Nymphalidae" genus="Pseudonympha" kingdom="Animalia" order="Lepidoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="subsimilis">Ht. subsimilis</taxonomicName>
</emphasis>
has the Rs ocellus on HWV usually quite prominent and 
<taxonomicName id="4C2E8DA74C45C6641EE82965FEE2252B" box="[200,305,1096,1119]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="sogai">
<emphasis id="B95A2A364C45C6641EE82965FEE2252B" box="[200,305,1096,1119]" italics="true" pageId="51" pageNumber="52">Ht. sogai</emphasis>
</taxonomicName>
has the Mb and PMb strongly dentate between M1 and CuA2. Populations of 
<taxonomicName id="4C2E8DA74C45C6641AE32965FA82252B" box="[1219,1361,1096,1119]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="avaratra">
<emphasis id="B95A2A364C45C6641AE32965FA82252B" box="[1219,1361,1096,1119]" italics="true" pageId="51" pageNumber="52">Ht. avaratra</emphasis>
</taxonomicName>
in the northern reaches of the main rainforest belt can be hard to distinguish by wing pattern from the sympatric 
<taxonomicName id="4C2E8DA74C45C6641B592941FED425D3" class="Insecta" family="Nymphalidae" genus="Pseudonympha" kingdom="Animalia" order="Lepidoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="subsimilis">
<emphasis id="B95A2A364C45C6641B592941FED425D3" italics="true" pageId="51" pageNumber="52">Ht. subsimilis</emphasis>
</taxonomicName>
though easily discriminated by DNA barcoding, but 
<taxonomicName id="4C2E8DA74C45C6641D4E29A2FC2F25D2" box="[878,1020,1167,1190]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="avaratra">
<emphasis id="B95A2A364C45C6641D4E29A2FC2F25D2" box="[878,1020,1167,1190]" italics="true" pageId="51" pageNumber="52">Ht. avaratra</emphasis>
</taxonomicName>
lacks an inflated M1-M2 vein (
<figureCitation id="1315EAA14C45C6641B4F29A3FF6725BE" captionStart="FIGURE 12" captionStartId="44.[151,250,1007,1029]" captionTargetBox="[242,1351,208,982]" captionTargetId="figure@44.[212,1370,177,1004]" captionTargetPageId="44" captionText="FIGURE 12. Diagrams of ♂ HW inflated vein system in the Ht. subsimilis group (modified after Lees, 1997: 96); wings in DV and R vein component on the anterior border of the cell in VV (flipped). A: Ht. pauper Oberthür, 1916 (Ankazomivady; “ H. 23 ” in Torres et al., 2001). B: Ht. pauper (Anjanaharibe Sud). C: Ht. kremenae Lees, sp. nov. (Masoala). D: Ht. subsimilis (Butler, 1878) (Ambohitantely), showing M 1 + M 2 inflation and asymmetric swelling above cell. E: Ht. avaratra Lees &amp; Kremen, sp. nov. (Montagne d’Ambre). No wing preparation was available for Ht. sogai." httpUri="https://zenodo.org/record/264609/files/figure.png" pageId="51" pageNumber="52">Fig. 12</figureCitation>
E), and the inflated vein below the Sdb basad of the fork of Rs+M1 is more symmetric (more rounded on the anteriad edge) in 
<taxonomicName id="4C2E8DA74C45C6641F4029FAFE2F259A" box="[352,508,1237,1262]" class="Insecta" family="Nymphalidae" genus="Pseudonympha" kingdom="Animalia" order="Lepidoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="subsimilis">
<emphasis id="B95A2A364C45C6641F4029FAFE2F259A" box="[352,508,1237,1262]" italics="true" pageId="51" pageNumber="52">Ht. subsimilis</emphasis>
</taxonomicName>
(
<figureCitation id="1315EAA14C45C6641C2E29F8FDB2259A" box="[526,609,1237,1262]" captionStart="FIGURE 12" captionStartId="44.[151,250,1007,1029]" captionTargetBox="[242,1351,208,982]" captionTargetId="figure@44.[212,1370,177,1004]" captionTargetPageId="44" captionText="FIGURE 12. Diagrams of ♂ HW inflated vein system in the Ht. subsimilis group (modified after Lees, 1997: 96); wings in DV and R vein component on the anterior border of the cell in VV (flipped). A: Ht. pauper Oberthür, 1916 (Ankazomivady; “ H. 23 ” in Torres et al., 2001). B: Ht. pauper (Anjanaharibe Sud). C: Ht. kremenae Lees, sp. nov. (Masoala). D: Ht. subsimilis (Butler, 1878) (Ambohitantely), showing M 1 + M 2 inflation and asymmetric swelling above cell. E: Ht. avaratra Lees &amp; Kremen, sp. nov. (Montagne d’Ambre). No wing preparation was available for Ht. sogai." httpUri="https://zenodo.org/record/264609/files/figure.png" pageId="51" pageNumber="52">Fig. 12</figureCitation>
D). 
<taxonomicName id="4C2E8DA74C45C6641CAD29FAFCC8259A" box="[653,795,1239,1262]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="avaratra">
<emphasis id="B95A2A364C45C6641CAD29FAFCC8259A" box="[653,795,1239,1262]" italics="true" pageId="51" pageNumber="52">Ht. avaratra</emphasis>
</taxonomicName>
, especially from the topotypical population, has a more prominently ventral yellow cast than in other species including 
<taxonomicName id="4C2E8DA74C45C6641D4029D6FC292466" box="[864,1018,1273,1298]" class="Insecta" family="Nymphalidae" genus="Pseudonympha" kingdom="Animalia" order="Lepidoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="subsimilis">
<emphasis id="B95A2A364C45C6641D4029D6FC292466" box="[864,1018,1273,1298]" italics="true" pageId="51" pageNumber="52">Ht. subsimilis</emphasis>
</taxonomicName>
, and is not variegated with orange as in 
<taxonomicName id="4C2E8DA74C45C6641E942833FE9A2442" box="[180,329,1309,1334]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="kremenae">
<emphasis id="B95A2A364C45C6641E942833FE9A2442" box="[180,329,1309,1334]" italics="true" pageId="51" pageNumber="52">Ht. kremenae</emphasis>
</taxonomicName>
.
</paragraph>
</subSubSection>
<subSubSection id="C334A5AF4C45C6631EE7286CFC752398" lastPageId="52" lastPageNumber="53" pageId="51" pageNumber="52" type="description">
<paragraph id="8B91F6244C45C6631EE7286CFD3420DF" blockId="51.[151,1437,247,2013]" lastBlockId="52.[151,1437,151,2035]" lastPageId="52" lastPageNumber="53" pageId="51" pageNumber="52">
<emphasis id="B95A2A364C45C6641EE7286CFE7E242E" bold="true" box="[199,429,1345,1370]" pageId="51" pageNumber="52">Description. Wings</emphasis>
: Dorsal surface homogeneous mid-brown as in others of the 
<taxonomicName id="4C2E8DA74C45C6641A49286FFAD7242E" box="[1129,1284,1345,1370]" class="Insecta" family="Nymphalidae" genus="Pseudonympha" kingdom="Animalia" order="Lepidoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="subsimilis">
<emphasis id="B95A2A364C45C6641A49286FFAD7242E" box="[1129,1284,1345,1370]" italics="true" pageId="51" pageNumber="52">Ht. subsimilis</emphasis>
</taxonomicName>
group. FWD space-CuA1 ocellus with black iris at least spanning CuA1-CuA2, with a fairly wide pale orange roundsubhexagonal ring, not perfectly concentric, often protruding a bit proximad within space-CuA1. FWD space-M1 ocellus usually small but conspicuous and subelliptic, with narrow orange ring. HWD space-CuA1 ocellus subelliptic and a little larger than that of FWD space-M1, usually spanning at least ¾ of HW veins CuA1-CuA2, including the fairly narrow concentric orange ring. Other ocelli not expressed on HWD. Ventral surface a more strongly ochreous background colour, especially in Montagne d’Ambre populations, than in others of the 
<taxonomicName id="4C2E8DA74C45C6641B592B35FED42720" class="Insecta" family="Nymphalidae" genus="Pseudonympha" kingdom="Animalia" order="Lepidoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="subsimilis">
<emphasis id="B95A2A364C45C6641B592B35FED42720" italics="true" pageId="51" pageNumber="52">Ht. subsimilis</emphasis>
</taxonomicName>
group, and the area between Mb and PMb is finely irrorated in brown to a greater extent than other parts of the wing. FWV space-CuA1 ocellus with black iris same size as upperside and ring fairly subhexagonal and mainly yellow with gradation to orange towards tergal edge. The ring is slightly more pointed towards proximad edge where it hugs a strongly concave rufous brown Mb before it bends back to mid costa and Ore as a yellow hook like the small tail of a comma, representing the end of a spiral arm. FWV space-M1 ocellus as on upperside, slightly more conspicuous with a yellowish ring of background colour. On HWV, space-CuA1 ocellus is the largest as usual and subelliptic and including the concentric yellow ring spanning about 4/5 of veins CuA1-CuA2 (generally smaller than in 
<taxonomicName id="4C2E8DA74C45C6641F9F2A1AFD85263A" box="[447,598,1845,1870]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="51" pageNumber="52" phylum="Arthropoda" rank="species" species="kremenae">
<emphasis id="B95A2A364C45C6641F9F2A1AFD85263A" box="[447,598,1845,1870]" italics="true" pageId="51" pageNumber="52">Ht. kremenae</emphasis>
</taxonomicName>
). HWV space-CuA2 ocellus not evident in the HT ♂. HWV space Rs-M3 ocellus reduced to white point often with narrow black iris and Orng blending with background. The FWV cell area has two unequal brown edged more yellow transverse Cbs, the distad one wider. There is no distinctly orange shoulder to the basal FWV costa, which is strigulated evenly with black. The HWV is highlighted with coronal-like yellow-ochreous flares distad of the Mb, particularly in M2, where the Mb is toothed outwards slightly at vein intersections M2, M3 and CuA1, inflexed proximad of ocellus space-CuA1 and bends back towards 1A. 
<emphasis id="B95A2A364C42C6631B042DBAFA4F21C4" bold="true" box="[1316,1436,151,176]" pageId="52" pageNumber="53">Variation.</emphasis>
♀♀ similar but slightly larger. Referred material from the main rainforest block is sometimes harder to distinguish by wing pattern from 
<taxonomicName id="4C2E8DA74C42C6631FB72DCDFDE7218C" box="[407,564,223,248]" class="Insecta" family="Nymphalidae" genus="Pseudonympha" kingdom="Animalia" order="Lepidoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="subsimilis">
<emphasis id="B95A2A364C42C6631FB72DCDFDE7218C" box="[407,564,223,248]" italics="true" pageId="52" pageNumber="53">Ht. subsimilis</emphasis>
</taxonomicName>
, but 
<taxonomicName id="4C2E8DA74C42C6631C532DCDFCD22183" box="[627,769,224,247]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="avaratra">
<emphasis id="B95A2A364C42C6631C532DCDFCD22183" box="[627,769,224,247]" italics="true" pageId="52" pageNumber="53">Ht. avaratra</emphasis>
</taxonomicName>
tends to be a larger species (see also 
<figureCitation id="1315EAA14C42C6631A9F2DF2FAC1218C" box="[1215,1298,223,248]" captionStart="FIGURE 16" captionStartId="56.[151,250,1770,1792]" captionTargetBox="[242,1343,191,1739]" captionTargetId="figure@56.[234,1354,177,1752]" captionTargetPageId="56" captionText="FIGURE 16. Adults in DV (left), VV (right) for mounted specimens and adults in natural posture (photos: D. C. Lees). A: HT ♂ of Heteropsis sogai, sp. nov. (Tsaratanana; DCL-DB- 3386). B: PT ♀ of Ht. sogai (Tsaratanana; DCL-DB- 3387). Scale bar: 30 mm. C: ♀ of Ht. kremenae Lees, sp. nov. (Vohitaly, Makira; 30 / 12 / 2002). D: ♂ of Ht. pauper (Vohitaly, Makira; 30 / 12 / 2002). E: ♀ of Ht. subsimilis (Ambohitantely; 6 / 12 / 2006); F: ♀ of Ht. avaratra Lees &amp; Kremen, sp. nov. (Anjanaharibe Sud; 23 / 11 / 2014)." httpUri="https://zenodo.org/record/264613/files/figure.png" pageId="52" pageNumber="53">Fig. 16</figureCitation>
F). Mb well marked and sometimes more jagged-slightly inflexed than in HT, with yellow Mf in space-M2 distad of the Mb sometimes rectangular, although this character is not distinct in the HT specimen. HWV space-CuA2 ocellus pointlike in some specimens, or subelliptic with a pale yellow ring. HWV space-Rs-M3 ocelli usually but not always point-like with that of HWV space-Rs sometimes slightly more evident, as may be that of space-CuA2. FWV Cbs sometimes delineate indistinctly a third yellow band.
</paragraph>
<paragraph id="8B91F6244C42C6631EE72C98FC052362" blockId="52.[151,1437,151,2035]" pageId="52" pageNumber="53">
<emphasis id="B95A2A364C42C6631EE72C98FE8720BA" bold="true" box="[199,340,437,462]" pageId="52" pageNumber="53">Androconia</emphasis>
: Sdb dark brown to russet brown, underlying Sdp varying from ‘bullet’-shaped, pointed to flattopped. M3-A2 veins, but not M2, linearly inflated in the HWD (1A from 1/6 and widest at about 40%, tapering thence to margin; 3A from base to margin +/- parallel from ca. 1/8 to 7/8).
</paragraph>
<paragraph id="8B91F6244C42C6631EE72F0CFE7623F5" blockId="52.[151,1437,151,2035]" pageId="52" pageNumber="53">
<emphasis id="B95A2A364C42C6631EE72F0CFEBF234E" bold="true" box="[199,364,545,570]" pageId="52" pageNumber="53">Wingspan/fwl</emphasis>
: range 32.1–38/ 
<quantity id="4CD65BC14C42C6631C042F0CFD63234D" box="[548,688,545,569]" metricMagnitude="-2" metricUnit="m" metricValue="1.935" metricValueMax="2.1" metricValueMin="1.77" pageId="52" pageNumber="53" unit="mm" value="19.35" valueMax="21.0" valueMin="17.7">17.7–21 mm</quantity>
(n=7 ♂♂); mean = 35.8 +/- 2.3 SD/19.7 +/- 0.5 SD mm (n=7 ♂♂), including HT ♂ 33.2/ 
<quantity id="4CD65BC14C42C6631FAA2F69FE3D2328" box="[394,494,580,605]" metricMagnitude="-2" metricUnit="m" metricValue="1.95" pageId="52" pageNumber="53" unit="mm" value="19.5">19.5 mm</quantity>
. Range 35.8–37.0 (n=2 ♀♀)/ 
<quantity id="4CD65BC14C42C6631D092F68FC042328" box="[809,983,581,605]" metricMagnitude="-2" metricUnit="m" metricValue="2.0549999999999997" metricValueMax="2.07" metricValueMin="2.04" pageId="52" pageNumber="53" unit="mm" value="20.549999999999997" valueMax="20.7" valueMin="20.4">20.4–20.7 mm</quantity>
(n=4 ♀♀); mean 36.4 (n=2 ♀♀)/20.5 +/ - 0.1 SD mm (n=4 ♀♀).
</paragraph>
<paragraph id="8B91F6244C42C6631EE72FA1FC752398" blockId="52.[151,1437,151,2035]" pageId="52" pageNumber="53">
<emphasis id="B95A2A364C42C6631EE72FA1FEDB23D1" bold="true" box="[199,264,652,677]" pageId="52" pageNumber="53">Palps</emphasis>
: penultimate segment on outside face cream ochreous towards compound eye, dark brown away from eye with a few light coloured scales within this dark border (more apparent in the ♀); fringed dark brown where not contacting eye. Face of palp away from compound eye pale ochreous.
</paragraph>
</subSubSection>
<subSubSection id="C334A5AF4C42C6631EE72FDAFB6D25C9" pageId="52" pageNumber="53" type="etymology">
<paragraph id="8B91F6244C42C6631EE72FDAFA952264" blockId="52.[151,1437,151,2035]" box="[199,1350,759,784]" pageId="52" pageNumber="53">
<emphasis id="B95A2A364C42C6631EE72FDAFE9D2264" bold="true" box="[199,334,759,784]" pageId="52" pageNumber="53">Etymology.</emphasis>
‘avaratra’ means ‘north’ in 
<collectingCountry id="F339B6B44C42C6631CA82FDAFD252264" box="[648,758,759,784]" name="Madagascar" pageId="52" pageNumber="53">Malagasy</collectingCountry>
, referring to its distribution, as it is currently known.
</paragraph>
<paragraph id="8B91F6244C42C6631EE72E36FB6D25C9" blockId="52.[151,1437,151,2035]" pageId="52" pageNumber="53">
<emphasis id="B95A2A364C42C6631EE72E36FDBD2240" bold="true" box="[199,622,795,820]" pageId="52" pageNumber="53">Additional information. ♂ genitalia</emphasis>
: 315DL (referred specimen, Joffreville); 
<bibRefCitation id="EFBF8BD54C42C6631A602E31FED52223" author="Lees" pageId="52" pageNumber="53" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997" yearSuffix="f">
Lees (1997: 106, 
<figureCitation id="1315EAA14C42C6631B252E36FA942240" box="[1285,1351,795,820]" captionStart="FIGURE 7" captionStartId="25.[151,250,1840,1862]" captionTargetBox="[267,1325,216,1803]" captionTargetId="figure@25.[244,1342,193,1819]" captionTargetPageId="25" captionText="FIGURE 7. Adults in DV (left), VV (right) or in natural postures. A: HT ♂ of Heteropsis harveyi Lees &amp; Kremen, sp. nov. (NHMUK 010289119). B: PT ♀ of Ht. harveyi (NHMUK 010289121). C: HT ♂ of Henotesia andravahana ab. marmorata Aurivillius, 1911 (NHRS-TOBI 000000050). D: pair of Ht. harveyi in copula (Maromizaha, photo courtesy Lance Featherstone). E: ♂ of Ht. imerina (Ankazomivady, 23 / 03 / 2003, photo: D. C. Lees), for comparison, showing relatively produced ‘ tail’ on HW vein M 3." httpUri="https://zenodo.org/record/264604/files/figure.png" pageId="52" pageNumber="53">Fig. 7</figureCitation>
f; “25”; 
<figureCitation id="1315EAA14C42C6631EB72E13FF3B2223" box="[151,232,830,855]" captionStart="FIGURE 15" captionStartId="49.[151,250,1351,1373]" captionTargetBox="[386,1205,210,1306]" captionTargetId="figure@49.[363,1224,193,1330]" captionTargetPageId="49" captionText="FIGURE 15. ♂ genitalia in LV (left), DV (right) unless otherwise stated. A: Heteropsis kremenae Lees, sp. nov. (Masoala, Ambanivony; 19 DL). B: Ht. avaratra Lees &amp; Kremen, sp. nov. (315 DL, referred specimen, Joffreville). C: Heteropsis sogai Lees, sp. nov. PT (Tsaratanana; 96 DL, MNHN), LV, SV. Scale bar: 3 mm." httpUri="https://zenodo.org/record/264612/files/figure.png" pageId="52" pageNumber="53">Fig. 15</figureCitation>
B)
</bibRefCitation>
. Miniaturised, about 
<quantity id="4CD65BC14C42C6631FD42E13FD982223" box="[500,587,830,855]" metricMagnitude="-3" metricUnit="m" metricValue="1.6" pageId="52" pageNumber="53" unit="mm" value="1.6">1.6 mm</quantity>
long, and with configuration typical of the 
<taxonomicName id="4C2E8DA74C42C6631A142E6DFB1C2223" box="[1076,1231,830,855]" class="Insecta" family="Nymphalidae" genus="Pseudonympha" kingdom="Animalia" order="Lepidoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="subsimilis">
<emphasis id="B95A2A364C42C6631A142E6DFB1C2223" box="[1076,1231,830,855]" italics="true" pageId="52" pageNumber="53">Ht. subsimilis</emphasis>
</taxonomicName>
group; there is no obvious difference in shapes of the genitalic components from other members (
<bibRefCitation id="EFBF8BD54C42C6631A432E4EFAE7220F" author="Lees" box="[1123,1332,866,891]" pageId="52" pageNumber="53" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees (1997: 106)</bibRefCitation>
and the description below might apply to almost any of the 
<taxonomicName id="4C2E8DA74C42C6631CDF2EAAFC4E22EB" box="[767,925,902,927]" class="Insecta" family="Nymphalidae" genus="Pseudonympha" kingdom="Animalia" order="Lepidoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="subsimilis">
<emphasis id="B95A2A364C42C6631CDF2EAAFC4E22EB" box="[767,925,902,927]" italics="true" pageId="52" pageNumber="53">Ht. subsimilis</emphasis>
</taxonomicName>
group; from LV, the valve bases are rather symmetrically-‘skittle’-shaped (
<bibRefCitation id="EFBF8BD54C42C6631FDD2E87FDA422B6" author="Lees" box="[509,631,938,962]" pageId="52" pageNumber="53" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees 1997</bibRefCitation>
). Uncus is slightly longer than tegumen (which is shallowly and widely notched proximad from DV) and quite inflated dorsoventrally before the slightly pointed downturned tip (distinctly inflated from DV before tip). Gnathos tapered from small base, fairly straight, not reaching tip of uncus (from DV, the appearance in the examined specimen is quite quadrate, not particularly sinuate or bull’s horn-like). Valve with prominent rounded dorsal ‘shoulder’, valve arm tapering to distinct ‘beak’ that points to uncus base (strongly incurved from DV) with limited serration (more obvious distad from DV) and not much indication of a club at valve end. Saccus relatively small, slightly bulbous and juxta also not prominent proximad. Aedeagus about same length as valve, shallowly recurved distad of ostium, which is slightly bulbous at proximad tip.
</paragraph>
</subSubSection>
<subSubSection id="C334A5AF4C42C6631EE729EAFAE82668" pageId="52" pageNumber="53" type="description">
<paragraph id="8B91F6244C42C6631EE729EAFBEB241B" blockId="52.[151,1437,151,2035]" pageId="52" pageNumber="53">
<emphasis id="B95A2A364C42C6631EE729EAFE482594" bold="true" box="[199,411,1223,1248]" pageId="52" pageNumber="53">DNA divergences</emphasis>
: the COI-5P barcode (cluster number BOLD:AAD0195, exemplar BMAD016-09 from Montagne d’Ambre) is 3.46% pairwise divergent to 
<taxonomicName id="4C2E8DA74C42C6631CD829C0FC452470" box="[760,918,1259,1284]" class="Insecta" family="Nymphalidae" genus="Pseudonympha" kingdom="Animalia" order="Lepidoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="subsimilis">
<emphasis id="B95A2A364C42C6631CD829C0FC452470" box="[760,918,1259,1284]" italics="true" pageId="52" pageNumber="53">Ht. subsimilis</emphasis>
</taxonomicName>
(cluster number BOLD:AAB4493, currently with 27 members on the BOLD database; e.g. exemplar 
<accessionNumber id="947D6BC74C42C6631D0A2822FC48245C" box="[810,923,1295,1320]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/FJ666680" pageId="52" pageNumber="53">FJ666680</accessionNumber>
, 
<bibRefCitation id="EFBF8BD54C42C6631D8A2822FB5F245C" author="Linares" box="[938,1164,1295,1320]" pageId="52" pageNumber="53" refString="Linares, M. C., Soto-Calderon, I. D., Lees, D. C. &amp; Anthony, N. M. (2009) High mitochondrial diversity in geographically widespread butterflies of Madagascar: a test of the DNA barcoding approach. Molecular Phylogenetics and Evolution, 50, 485 - 495. http: // dx. doi. org / 10.1016 / j. ympev. 2008.11.008" type="journal article" year="2009">
Linares 
<emphasis id="B95A2A364C42C6631A28283DFB99245C" box="[1032,1098,1295,1320]" italics="true" pageId="52" pageNumber="53">et al.,</emphasis>
2009
</bibRefCitation>
dataset) whereas about 6.55% divergent to that of 
<taxonomicName id="4C2E8DA74C42C6631FE62819FD93243F" box="[454,576,1332,1355]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="pauper">
<emphasis id="B95A2A364C42C6631FE62819FD93243F" box="[454,576,1332,1355]" italics="true" pageId="52" pageNumber="53">Ht. pauper</emphasis>
</taxonomicName>
(BOLD:AAA6758). The COII dataset of 
<bibRefCitation id="EFBF8BD54C42C6631A3C2819FB2D243F" box="[1052,1278,1331,1356]" pageId="52" pageNumber="53" refString="Torres, E., Lees, D. C., Vane-Wright, R. I., Kremen, C., Leonard, J. A. &amp; Wayne, R. K. (2001) Examining monophyly in a large radiation of Madagascan butterflies (Lepidoptera: Satyrinae: Mycalesina) based on mitochondrial DNA data. Molecular Phylogenetics and Evolution, 20, 460 - 473. http: // dx. doi. org / 10.1006 / mpev. 2001.0984" type="journal article">
Torres 
<emphasis id="B95A2A364C42C6631A4C2819FB782438" box="[1132,1195,1331,1356]" italics="true" pageId="52" pageNumber="53">et al.,</emphasis>
(2001)
</bibRefCitation>
shows 5.27% pairwise divergence (398 bp compared) of 
<taxonomicName id="4C2E8DA74C42C6631C582875FCD7241B" box="[632,772,1368,1391]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="avaratra">
<emphasis id="B95A2A364C42C6631C582875FCD7241B" box="[632,772,1368,1391]" italics="true" pageId="52" pageNumber="53">Ht. avaratra</emphasis>
</taxonomicName>
(‘sp. 25’) to 
<taxonomicName id="4C2E8DA74C42C6631DB72875FBE2241B" box="[919,1073,1366,1391]" class="Insecta" family="Nymphalidae" genus="Pseudonympha" kingdom="Animalia" order="Lepidoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="subsimilis">
<emphasis id="B95A2A364C42C6631DB72875FBE2241B" box="[919,1073,1366,1391]" italics="true" pageId="52" pageNumber="53">Ht. subsimilis</emphasis>
</taxonomicName>
.
</paragraph>
<paragraph id="8B91F6244C42C6631EE72857FAE82668" blockId="52.[151,1437,151,2035]" pageId="52" pageNumber="53">
<emphasis id="B95A2A364C42C6631EE72857FE3524E7" bold="true" box="[199,486,1402,1427]" pageId="52" pageNumber="53">Phylogeny/sister species</emphasis>
: in the morphological analysis of 
<bibRefCitation id="EFBF8BD54C42C6631D532856FC3F24E7" author="Lees" box="[883,1004,1403,1427]" pageId="52" pageNumber="53" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees 1997</bibRefCitation>
, this taxon (“TWNFV”) connected to 
<taxonomicName id="4C2E8DA74C42C6631EB728B2FE8424C2" box="[151,343,1439,1462]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="sogai">
<emphasis id="B95A2A364C42C6631EB728B2FE8424C2" box="[151,343,1439,1462]" italics="true" pageId="52" pageNumber="53">Heteropsis sogai</emphasis>
</taxonomicName>
(“THRNN”) but in only one (jackknife) tree with support (
<bibRefCitation id="EFBF8BD54C42C6631A3B28B2FAFE24C3" author="Lees" box="[1051,1325,1438,1463]" pageId="52" pageNumber="53" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">
Lees 1997: 157, 
<figureCitation id="1315EAA14C42C6631AC528B3FAFE24C3" box="[1253,1325,1438,1463]" captionStart="FIGURE 2" captionStartId="11.[151,250,1868,1890]" captionTargetBox="[266,1318,198,1839]" captionTargetId="figure@11.[254,1332,193,1847]" captionTargetPageId="11" captionText="FIGURE 2. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis mimetica Lees &amp; Kremen, sp. nov. (Anjanaharibe Sud; BMNH (E) 1717105), B: PT ♀ of Ht. mimetica (Anjanaharibe Sud; BMNH (E) 1717113). C: HT ♂ of Pseudonympha cowani Butler, 1880 (‘ Fianarantsoa’). D: LT ♂ of Culapa houlbertiana Oberthür, 1923 (‘ Tamatave’, BMNH (E) # 674789). F. LT ♂ of Mycalesis antahala Ward, 1872 ([Madagascar], BMNH (E) 672567). Scale bar: 30 mm. The figure shows apparent contrasting mimicry on the two surfaces across three different Heteropsis clades." httpUri="https://zenodo.org/record/264599/files/figure.png" pageId="52" pageNumber="53">Fig. 2</figureCitation>
</bibRefCitation>
). In their analysis of COII sequences (where 
<taxonomicName id="4C2E8DA74C42C6631C0828EEFD5C24AE" box="[552,655,1475,1498]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="sogai">
<emphasis id="B95A2A364C42C6631C0828EEFD5C24AE" box="[552,655,1475,1498]" italics="true" pageId="52" pageNumber="53">Ht. sogai</emphasis>
</taxonomicName>
was the only member of the 
<taxonomicName id="4C2E8DA74C42C6631DFA28EEFBA624AF" box="[986,1141,1474,1499]" class="Insecta" family="Nymphalidae" genus="Pseudonympha" kingdom="Animalia" order="Lepidoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="subsimilis">
<emphasis id="B95A2A364C42C6631DFA28EEFBA624AF" box="[986,1141,1474,1499]" italics="true" pageId="52" pageNumber="53">Ht. subsimilis</emphasis>
</taxonomicName>
group on 
<collectingCountry id="F339B6B44C42C6631AC928EFFAA324AF" box="[1257,1392,1474,1499]" name="Madagascar" pageId="52" pageNumber="53">Madagascar</collectingCountry>
not included), 
<bibRefCitation id="EFBF8BD54C42C6631F2F28CAFE39248A" author="Torres" box="[271,490,1509,1534]" pageId="52" pageNumber="53" refString="Torres, E., Lees, D. C., Vane-Wright, R. I., Kremen, C., Leonard, J. A. &amp; Wayne, R. K. (2001) Examining monophyly in a large radiation of Madagascan butterflies (Lepidoptera: Satyrinae: Mycalesina) based on mitochondrial DNA data. Molecular Phylogenetics and Evolution, 20, 460 - 473. http: // dx. doi. org / 10.1006 / mpev. 2001.0984" type="journal article" year="2001">
Torres 
<emphasis id="B95A2A364C42C6631F7D28CAFE4E248A" box="[349,413,1509,1534]" italics="true" pageId="52" pageNumber="53">et al.,</emphasis>
(2001
</bibRefCitation>
: 366) had a meager 64% bootstrap support for a sister relation of 
<taxonomicName id="4C2E8DA74C42C6631AEE28CAFA89248A" box="[1230,1370,1511,1534]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="avaratra">
<emphasis id="B95A2A364C42C6631AEE28CAFA89248A" box="[1230,1370,1511,1534]" italics="true" pageId="52" pageNumber="53">Ht. avaratra</emphasis>
</taxonomicName>
, their “ 
<emphasis id="B95A2A364C42C6631E832B27FF012755" box="[163,210,1546,1569]" italics="true" pageId="52" pageNumber="53">Hen</emphasis>
. sp. 25”, 
<accessionNumber id="947D6BC74C42C6631F1E2B24FE6B2756" box="[318,440,1545,1570]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/AY040146" pageId="52" pageNumber="53">
AY 
<date id="FF90D0E44C42C6631F452B24FE6B2756" box="[357,440,1545,1570]" pageId="52" pageNumber="53" value="46-04-01">040146</date>
</accessionNumber>
based on one individual from the 
<typeStatus id="549548864C42C6631D192B27FCBA2756" box="[825,873,1546,1570]" pageId="52" pageNumber="53">type</typeStatus>
locality) with 
<taxonomicName id="4C2E8DA74C42C6631A2D2B27FB742756" box="[1037,1191,1545,1570]" class="Insecta" family="Nymphalidae" genus="Pseudonympha" kingdom="Animalia" order="Lepidoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="subsimilis">
<emphasis id="B95A2A364C42C6631A2D2B27FB742756" box="[1037,1191,1545,1570]" italics="true" pageId="52" pageNumber="53">Ht. subsimilis</emphasis>
</taxonomicName>
(
<accessionNumber id="947D6BC74C42C6631A962B24FAE12756" box="[1206,1330,1545,1570]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/AY040145" pageId="52" pageNumber="53">
AY 
<date id="FF90D0E44C42C6631AFE2B24FAE12756" box="[1246,1330,1545,1570]" pageId="52" pageNumber="53" value="45-04-01">040145</date>
</accessionNumber>
based on one individual from Ranomafana National Park). 
<taxonomicName id="4C2E8DA74C42C6631CF32B03FCB12731" box="[723,866,1582,1605]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="avaratra">
<emphasis id="B95A2A364C42C6631CF32B03FCB12731" box="[723,866,1582,1605]" italics="true" pageId="52" pageNumber="53">Ht. avaratra</emphasis>
</taxonomicName>
(as “sp. 25”) was however strongly supported as sister to 
<taxonomicName id="4C2E8DA74C42C6631ED72B7FFE41271E" box="[247,402,1617,1642]" class="Insecta" family="Nymphalidae" genus="Pseudonympha" kingdom="Animalia" order="Lepidoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="subsimilis">
<emphasis id="B95A2A364C42C6631ED72B7FFE41271E" box="[247,402,1617,1642]" italics="true" pageId="52" pageNumber="53">Ht. subsimilis</emphasis>
</taxonomicName>
in the phylogeographic analysis of 
<bibRefCitation id="EFBF8BD54C42C6631D052B7CFBD8271D" author="Linares" box="[805,1035,1617,1642]" pageId="52" pageNumber="53" refString="Linares, M. C., Soto-Calderon, I. D., Lees, D. C. &amp; Anthony, N. M. (2009) High mitochondrial diversity in geographically widespread butterflies of Madagascar: a test of the DNA barcoding approach. Molecular Phylogenetics and Evolution, 50, 485 - 495. http: // dx. doi. org / 10.1016 / j. ympev. 2008.11.008" type="journal article" year="2009">
Linares 
<emphasis id="B95A2A364C42C6631DA02B7FFC13271E" box="[896,960,1617,1642]" italics="true" pageId="52" pageNumber="53">et al.,</emphasis>
(2009
</bibRefCitation>
, p. 488, their 
<figureCitation id="1315EAA14C42C6631A872B7CFB38271D" box="[1191,1259,1617,1642]" captionStart="FIGURE 2" captionStartId="11.[151,250,1868,1890]" captionTargetBox="[266,1318,198,1839]" captionTargetId="figure@11.[254,1332,193,1847]" captionTargetPageId="11" captionText="FIGURE 2. Adults in DV (left), VV (right). A: HT ♂ of Heteropsis mimetica Lees &amp; Kremen, sp. nov. (Anjanaharibe Sud; BMNH (E) 1717105), B: PT ♀ of Ht. mimetica (Anjanaharibe Sud; BMNH (E) 1717113). C: HT ♂ of Pseudonympha cowani Butler, 1880 (‘ Fianarantsoa’). D: LT ♂ of Culapa houlbertiana Oberthür, 1923 (‘ Tamatave’, BMNH (E) # 674789). F. LT ♂ of Mycalesis antahala Ward, 1872 ([Madagascar], BMNH (E) 672567). Scale bar: 30 mm. The figure shows apparent contrasting mimicry on the two surfaces across three different Heteropsis clades." httpUri="https://zenodo.org/record/264599/files/figure.png" pageId="52" pageNumber="53">Fig. 2</figureCitation>
) including COI or also EF-1a and wingless (p. 490, their 
<figureCitation id="1315EAA14C42C6631C4D2B59FD6127F9" box="[621,690,1652,1677]" captionStart="FIGURE 4" captionStartId="18.[151,250,1903,1925]" captionTargetBox="[266,1320,200,1853]" captionTargetId="figure@18.[253,1332,193,1866]" captionTargetPageId="18" captionText="FIGURE 4. Adults in DV (left), VV (right) unless otherwise stated. A: HT ♂ of Heteropsis hazovola Lees &amp; Raharitsimba, sp. nov. (Masoala; NHMUK 010289116). B: PT ♀ of Ht. hazovola (Masoala; NHMUK 010289118). C: Abdomen of HT ♂ of Ht. hazovola (VV, left). D. Abdomen of Mycalesis ankoma LT ♂ (VV, right). E: LT ♂ of Mycalesis ankoma Mabille, 1878 (Madagascar; BMHH (E) 672615). F: PLT ♀ of M. ankoma (BMNH (E) 672618). Scale bar: 30 mm (C – D with separate scale)." httpUri="https://zenodo.org/record/264601/files/figure.png" pageId="52" pageNumber="53">Fig. 4</figureCitation>
). However, neither the latter study nor that of 
<bibRefCitation id="EFBF8BD54C42C6631AE02B59FEDC27C5" pageId="52" pageNumber="53" refString="Kodandaramaiah, U., Lees, D. C., Muller, C. J., Torres, E., Karanth, C. P. &amp; Wahlberg, N. (2010) Phylogenetics and biogeography of a spectacular Old World radiation of butterflies: the subtribe Mycalesina (Lepidoptera: Nymphalidae: Satyrini). BMC Evolutionary Biology, 10, 172. http: // dx. doi. org / 10.1186 / 1471 - 2148 - 10 - 172" type="journal article">
Kodandaramaiah 
<emphasis id="B95A2A364C42C6631BA82B5BFF6827C5" italics="true" pageId="52" pageNumber="53">et al.,</emphasis>
(2010)
</bibRefCitation>
, which found the same relationship, included 
<taxonomicName id="4C2E8DA74C42C6631D302BB4FC7527C5" box="[784,934,1688,1713]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="kremenae">
<emphasis id="B95A2A364C42C6631D302BB4FC7527C5" box="[784,934,1688,1713]" italics="true" pageId="52" pageNumber="53">Ht. kremenae</emphasis>
</taxonomicName>
nor 
<taxonomicName id="4C2E8DA74C42C6631DF92BB4FB9327C4" box="[985,1088,1689,1712]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="sogai">
<emphasis id="B95A2A364C42C6631DF92BB4FB9327C4" box="[985,1088,1689,1712]" italics="true" pageId="52" pageNumber="53">Ht. sogai</emphasis>
</taxonomicName>
(but that of 
<bibRefCitation id="EFBF8BD54C42C6631AE92BB4FA4B27C5" author="Torres" box="[1225,1432,1688,1713]" pageId="52" pageNumber="53" refString="Torres, E., Lees, D. C., Vane-Wright, R. I., Kremen, C., Leonard, J. A. &amp; Wayne, R. K. (2001) Examining monophyly in a large radiation of Madagascan butterflies (Lepidoptera: Satyrinae: Mycalesina) based on mitochondrial DNA data. Molecular Phylogenetics and Evolution, 20, 460 - 473. http: // dx. doi. org / 10.1006 / mpev. 2001.0984" type="journal article" year="2001">
Torres 
<emphasis id="B95A2A364C42C6631B372BB4FA8527C5" box="[1303,1366,1688,1713]" italics="true" pageId="52" pageNumber="53">et al.,</emphasis>
2001
</bibRefCitation>
, who found the same with COII, did include 
<taxonomicName id="4C2E8DA74C42C6631CA02B90FCC627A1" box="[640,789,1724,1749]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="kremenae">
<emphasis id="B95A2A364C42C6631CA02B90FCC627A1" box="[640,789,1724,1749]" italics="true" pageId="52" pageNumber="53">Ht. kremenae</emphasis>
</taxonomicName>
as their ‘ 
<emphasis id="B95A2A364C42C6631D5C2B90FC7827A0" box="[892,939,1725,1748]" italics="true" pageId="52" pageNumber="53">Hen</emphasis>
. sp. 14A’), in fact only otherwise 
<taxonomicName id="4C2E8DA74C42C6631B032B90FA4827A0" box="[1315,1435,1725,1748]" class="Insecta" family="Nymphalidae" genus="Culapa" higherTaxonomySource="CoL" kingdom="Animalia" order="Lepidoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="pauper">
<emphasis id="B95A2A364C42C6631B032B90FA4827A0" box="[1315,1435,1725,1748]" italics="true" pageId="52" pageNumber="53">Ht. pauper</emphasis>
</taxonomicName>
was included among the 
<taxonomicName id="4C2E8DA74C42C6631F902BCCFD99278C" box="[432,586,1759,1784]" class="Insecta" family="Nymphalidae" genus="Pseudonympha" kingdom="Animalia" order="Lepidoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="subsimilis">
<emphasis id="B95A2A364C42C6631F902BCCFD99278C" box="[432,586,1759,1784]" italics="true" pageId="52" pageNumber="53">Ht. subsimilis</emphasis>
</taxonomicName>
group. 
<taxonomicName id="4C2E8DA74C42C6631C822BCCFCEB278C" box="[674,824,1759,1784]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="kremenae">
<emphasis id="B95A2A364C42C6631C822BCCFCEB278C" box="[674,824,1759,1784]" italics="true" pageId="52" pageNumber="53">Ht. kremenae</emphasis>
</taxonomicName>
is more distant to 
<taxonomicName id="4C2E8DA74C42C6631A2B2BCCFB44278C" box="[1035,1175,1761,1784]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="avaratra">
<emphasis id="B95A2A364C42C6631A2B2BCCFB44278C" box="[1035,1175,1761,1784]" italics="true" pageId="52" pageNumber="53">Ht. avaratra</emphasis>
</taxonomicName>
by COII data (see also description above). 
<taxonomicName id="4C2E8DA74C42C6631F542A28FE092668" box="[372,474,1797,1820]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="sogai">
<emphasis id="B95A2A364C42C6631F542A28FE092668" box="[372,474,1797,1820]" italics="true" pageId="52" pageNumber="53">Ht. sogai</emphasis>
</taxonomicName>
might be its sister (as in combined tree in 
<bibRefCitation id="EFBF8BD54C42C6631D972A2EFB1A2668" author="Aduse-Poku" box="[951,1225,1795,1820]" pageId="52" pageNumber="53" refString="Aduse-Poku, K., Lees, D. C., Brattstrom, O., Kodandaramaiah, U., Collins, S. &amp; Brakefield, P. M., (2016) Molecular phylogeny and generic-level taxonomy of the widespread palaeotropical ' Heteropsis clade' (Nymphalidae: Satyrinae: Mycalesina). Systematic Entomology, in press. http: // dx. doi. org / syen. 12183" type="book" year="2016">
Aduse-Poku 
<emphasis id="B95A2A364C42C6631A682A28FB522668" box="[1096,1153,1795,1820]" italics="true" pageId="52" pageNumber="53">et al.</emphasis>
, 2016
</bibRefCitation>
, in press).
</paragraph>
</subSubSection>
<subSubSection id="C334A5AF4C42C6631EE72A0AFE252687" pageId="52" pageNumber="53" type="discussion">
<paragraph id="8B91F6244C42C6631EE72A0AFBC326BB" blockId="52.[151,1437,151,2035]" pageId="52" pageNumber="53">
<emphasis id="B95A2A364C42C6631EE72A0AFE9F2634" bold="true" box="[199,332,1831,1856]" pageId="52" pageNumber="53">Discussion.</emphasis>
This species was first recognized in the field as ‘sp. 25’ by Claire Kremen on 
<date id="FF90D0E44C42C6631AFA2A05FAAD2634" box="[1242,1406,1832,1856]" pageId="52" pageNumber="53" value="1992-02-19" valueMax="1992-02-20" valueMin="1992-02-19">19-20/02/1992</date>
at Montagne d’Ambre, but there were sparse earlier collections in BMNH (Meade-Waldo, 1906) and at MNHN. It is understandable however that these specimens were not described earlier owing to their superficial similarity with 
<taxonomicName id="4C2E8DA74C42C6631EB72AB9FEE026DF" box="[151,307,1938,1963]" class="Insecta" family="Nymphalidae" genus="Pseudonympha" kingdom="Animalia" order="Lepidoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="subsimilis">
<emphasis id="B95A2A364C42C6631EB72AB9FEE026DF" box="[151,307,1938,1963]" italics="true" pageId="52" pageNumber="53">Ht. subsimilis</emphasis>
</taxonomicName>
. All other available 
<typeStatus id="549548864C42C6631C022ABEFD8E26DF" box="[546,605,1939,1963]" pageId="52" pageNumber="53">types</typeStatus>
of the 
<taxonomicName id="4C2E8DA74C42C6631C932AB9FC9C26DF" box="[691,847,1938,1963]" class="Insecta" family="Nymphalidae" genus="Pseudonympha" kingdom="Animalia" order="Lepidoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="subsimilis">
<emphasis id="B95A2A364C42C6631C932AB9FC9C26DF" box="[691,847,1938,1963]" italics="true" pageId="52" pageNumber="53">Ht. subsimilis</emphasis>
</taxonomicName>
group have been examined and are clearly distinct with their 
<typeStatus id="549548864C42C6631F2A2A9AFEE926BB" box="[266,314,1975,1999]" pageId="52" pageNumber="53">type</typeStatus>
localities further south in 
<collectingCountry id="F339B6B44C42C6631C412A9BFD3B26BB" box="[609,744,1974,1999]" name="Madagascar" pageId="52" pageNumber="53">Madagascar</collectingCountry>
(see under 
<taxonomicName id="4C2E8DA74C42C6631D4A2A9AFBD326BB" box="[874,1024,1974,1999]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="52" pageNumber="53" phylum="Arthropoda" rank="species" species="kremenae">
<emphasis id="B95A2A364C42C6631D4A2A9AFBD326BB" box="[874,1024,1974,1999]" italics="true" pageId="52" pageNumber="53">Ht. kremenae</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8B91F6244C42C6631EE72AF7FE252687" blockId="52.[151,1437,151,2035]" box="[199,502,2010,2035]" pageId="52" pageNumber="53">
<emphasis id="B95A2A364C42C6631EE72AF7FE252687" bold="true" box="[199,502,2010,2035]" pageId="52" pageNumber="53">Ecology and distribution.</emphasis>
</paragraph>
</subSubSection>
<subSubSection id="C334A5AF4C43C6621EE72DBAFE042017" pageId="53" pageNumber="54" type="description">
<paragraph id="8B91F6244C43C6621EE72DBAFC6421C4" blockId="53.[151,1437,151,2031]" box="[199,951,151,176]" pageId="53" pageNumber="54">
<emphasis id="B95A2A364C43C6621EE72DBAFEF021C4" bold="true" box="[199,291,151,176]" pageId="53" pageNumber="54">Habitat</emphasis>
: montane primary rainforest, particularly on ridges/slopes.
</paragraph>
<paragraph id="8B91F6244C43C6621EE72D96FA562068" blockId="53.[151,1437,151,2031]" pageId="53" pageNumber="54">
<emphasis id="B95A2A364C43C6621EE72D96FE9021A0" bold="true" box="[199,323,187,212]" pageId="53" pageNumber="54">Behaviour</emphasis>
: unusually for 
<taxonomicName id="4C2E8DA74C43C6621FD82D90FDA321A0" box="[504,624,189,212]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="genus">
<emphasis id="B95A2A364C43C6621FD82D90FDA321A0" box="[504,624,189,212]" italics="true" pageId="53" pageNumber="54">Heteropsis</emphasis>
</taxonomicName>
, adults have been seen visiting 
<taxonomicName id="4C2E8DA74C43C6621DC92D90FACE21A0" box="[1001,1309,187,212]" class="Magnoliopsida" family="Balsaminaceae" genus="Impatiens" kingdom="Plantae" order="Ericales" pageId="53" pageNumber="54" phylum="Tracheophyta" rank="genus">
<emphasis id="B95A2A364C43C6621DC92D90FB8421A0" box="[1001,1111,189,212]" italics="true" pageId="53" pageNumber="54">Impatiens</emphasis>
(Balsaminaceae)
</taxonomicName>
flowers in Montagne d’Ambre (pers. obs.). Adults fly low and would usually feed on fruits on the ground, but also feed on sap runs. In Anjanaharibe Sud, however, adults were attracted to banana bait and no flower visitation was witnessed.
</paragraph>
<paragraph id="8B91F6244C43C6621EE72C0BFD652034" blockId="53.[151,1437,151,2031]" box="[199,694,294,320]" pageId="53" pageNumber="54">
<emphasis id="B95A2A364C43C6621EE72C0BFEEE204B" bold="true" box="[199,317,294,319]" pageId="53" pageNumber="54">Hostplant</emphasis>
: unknown, probably low grasses.
</paragraph>
<paragraph id="8B91F6244C43C6621EE72C67FE042017" blockId="53.[151,1437,151,2031]" box="[199,471,330,355]" pageId="53" pageNumber="54">
<emphasis id="B95A2A364C43C6621EE72C67FE8B2017" bold="true" box="[199,344,330,355]" pageId="53" pageNumber="54">Early stages</emphasis>
: unknown.
</paragraph>
</subSubSection>
<subSubSection id="C334A5AF4C43C6621EE72C43FB7D2398" pageId="53" pageNumber="54" type="distribution">
<paragraph id="8B91F6244C43C6621EE72C43FD8723BC" blockId="53.[151,1437,151,2031]" pageId="53" pageNumber="54">
<emphasis id="B95A2A364C43C6621EE72C43FE8A20F3" bold="true" box="[199,345,366,391]" pageId="53" pageNumber="54">Distribution</emphasis>
: Montagne d’Ambre is an important locality but the species occurs in rainforest also in the main rainforest block from Tsaratanana to Makira (
<figureCitation id="1315EAA14C43C6621CBC2CBFFD2320DE" box="[668,752,402,427]" captionStart="FIGURE 30" captionStartId="92.[151,250,1763,1785]" captionTargetBox="[325,1224,254,1731]" captionTargetId="figure@92.[316,1247,230,1746]" captionTargetPageId="92" captionText="FIGURE 30. Distribution maps. A. Purple: Heteropsis vanewrighti Lees, sp. nov. Green: Ht. pauliani Lees, sp. nov. Yellow: Ht. mimetica Lees &amp; Kremen, sp. nov. Light blue: Ht. hazovola Lees &amp; Raharitsimba, sp. nov. Turquoise: Ht. imerina Lees, sp. nov. [northern populations overlapping with Ht. vanewrighti and Ht. pauliani are not shown]. Pink: Ht. westwoodi Lees, sp. nov. B. Red: Ht. avaratra Lees &amp; Kremen, sp. nov. Light blue: Ht. kremenae Lees, sp. nov. Fawn: Ht. vertigo Lees &amp; Raharitsimba, sp. nov. C. Ochre: Ht. roussettae Lees &amp; Kremen, sp. nov. Slate-blue: Ht. menamenoides Lees, sp. nov. Olive: Ht. lanyvary Lees, sp. nov. Dark green: Ht. borgo Lees, sp. nov. Light green: Ht. barbarae Lees &amp; Kremen, sp. nov. Brown: Ht. oberthueri Lees, sp. nov. Dark pink: Ht. tianae Lees &amp; Kremen, sp. nov. D. Mid green: Ht. sogai Lees, sp. nov. Dark brown: Ht. tornado Lees, Allaoui &amp; Aduse-Poku, sp. nov. Lilac: Ht. harveyi Lees &amp; Kremen, sp. nov. Light brown: centraleastern populations of Ht. harveyi complex. Scale bar: 500 km." httpUri="https://zenodo.org/record/264627/files/figure.png" pageId="53" pageNumber="54">Fig. 30</figureCitation>
B, red dots). If 
<emphasis id="B95A2A364C43C6621DBD2CBEFBD820DE" box="[925,1035,403,426]" italics="true" pageId="53" pageNumber="54">
<taxonomicName id="4C2E8DA74C43C6621DBD2CBEFBD420DE" box="[925,1031,403,426]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="sogai">Ht. sogai</taxonomicName>
,
</emphasis>
not sequenced, is not more closely related, 
<taxonomicName id="4C2E8DA74C43C6621ED52C9AFE5720BA" box="[245,388,439,462]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="avaratra">
<emphasis id="B95A2A364C43C6621ED52C9AFE5720BA" box="[245,388,439,462]" italics="true" pageId="53" pageNumber="54">Ht. avaratra</emphasis>
</taxonomicName>
might represent 
<taxonomicName id="4C2E8DA74C43C6621C6C2C9AFD3920BA" box="[588,746,437,462]" class="Insecta" family="Nymphalidae" genus="Pseudonympha" kingdom="Animalia" order="Lepidoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="subsimilis">
<emphasis id="B95A2A364C43C6621C6C2C9AFD3920BA" box="[588,746,437,462]" italics="true" pageId="53" pageNumber="54">Ht. subsimilis</emphasis>
</taxonomicName>
’s replacement in Montagne d’Ambre, and in that case, 
<taxonomicName id="4C2E8DA74C43C6621B592C9AFF2A2085" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="avaratra">
<emphasis id="B95A2A364C43C6621B592C9AFF2A2085" italics="true" pageId="53" pageNumber="54">Ht. avaratra</emphasis>
</taxonomicName>
has achieved sympatry and secondary contact with 
<taxonomicName id="4C2E8DA74C43C6621D672CF7FC312086" box="[839,994,473,498]" class="Insecta" family="Nymphalidae" genus="Pseudonympha" kingdom="Animalia" order="Lepidoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="subsimilis">
<emphasis id="B95A2A364C43C6621D672CF7FC312086" box="[839,994,473,498]" italics="true" pageId="53" pageNumber="54">Ht. subsimilis</emphasis>
</taxonomicName>
in northwestern and northeastern parts of the main rainforest belt. Due to a formatting error, the locality was incorrectly given in Table 1 of 
<bibRefCitation id="EFBF8BD54C43C6621B2D2CD3FF1C234D" author="Torres" pageId="53" pageNumber="54" refString="Torres, E., Lees, D. C., Vane-Wright, R. I., Kremen, C., Leonard, J. A. &amp; Wayne, R. K. (2001) Examining monophyly in a large radiation of Madagascan butterflies (Lepidoptera: Satyrinae: Mycalesina) based on mitochondrial DNA data. Molecular Phylogenetics and Evolution, 20, 460 - 473. http: // dx. doi. org / 10.1006 / mpev. 2001.0984" type="journal article" year="2001">
Torres 
<emphasis id="B95A2A364C43C6621B7C2CD3FA462362" box="[1372,1429,509,534]" italics="true" pageId="53" pageNumber="54">et al.</emphasis>
, 2001
</bibRefCitation>
as “Ankazomivady, 
<quantity id="4CD65BC14C43C6621F982F0CFDD3234E" box="[440,512,545,570]" metricMagnitude="4" metricUnit="m" metricValue="3.2" pageId="53" pageNumber="54" unit="km" value="32.0">32 km</quantity>
S. Ambositra” instead of “Montagne d’Ambre”. 
<bibRefCitation id="EFBF8BD54C43C6621A082F0FFB09234E" author="Lees" box="[1064,1242,545,570]" pageId="53" pageNumber="54" refString="Lees, D. C. (1997) Systematics and biogeography of Madagascan mycalesine butterflies (Lepidoptera: Satyrinae). Unpubl. PhD thesis, University of London, London, 411 pp." type="book" year="1997">Lees (1997: 66)</bibRefCitation>
assigned a series of specimens in BMNH from Mahassabe [= Mahasoabe], forêt SE Fianarantsoa (“sp. 55”) to this species, but this needs re-examination as they might easily be referable to 
<taxonomicName id="4C2E8DA74C43C6621D632F44FC3123F5" box="[835,994,616,641]" class="Insecta" family="Nymphalidae" genus="Pseudonympha" kingdom="Animalia" order="Lepidoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="subsimilis">
<emphasis id="B95A2A364C43C6621D632F44FC3123F5" box="[835,994,616,641]" italics="true" pageId="53" pageNumber="54">Ht. subsimilis</emphasis>
</taxonomicName>
or another species (plus this location includes a range of material that seems more characteristic of northeastern localities, for example 
<taxonomicName id="4C2E8DA74C43C6621B392FA0FE8523BC" authority="Oberthur, 1916" authorityName="Oberthur" authorityYear="1916" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="erebina">
<emphasis id="B95A2A364C43C6621B392FA0FA4F23D1" box="[1305,1436,652,677]" italics="true" pageId="53" pageNumber="54">Ht. erebina</emphasis>
(
<bibRefCitation id="EFBF8BD54C43C6621EBE2F82FE9D23BC" author="Oberthur" box="[158,334,687,712]" pageId="53" pageNumber="54" refString="Oberthur, C. (1916) Observations sur une centurie d'Especes de Lepidopteres Rhopaloceres malagaches. Etudes de Lepidopterologie Comparee, 11, 177 - 244." type="journal article" year="1916">Oberthür, 1916</bibRefCitation>
)
</taxonomicName>
; see also 
<taxonomicName id="4C2E8DA74C43C6621FE12F9CFD9723BC" box="[449,580,687,712]" class="Insecta" family="Curculionidae" genus="Heteropsis" kingdom="Animalia" order="Coleoptera" pageId="53" pageNumber="54" phylum="Arthropoda" rank="species" species="tornado">
<emphasis id="B95A2A364C43C6621FE12F9CFD9723BC" box="[449,580,687,712]" italics="true" pageId="53" pageNumber="54">Ht. tornado</emphasis>
</taxonomicName>
).
</paragraph>
<paragraph id="8B91F6244C43C6621EE72FFEFB7D2398" blockId="53.[151,1437,151,2031]" box="[199,1198,723,748]" pageId="53" pageNumber="54">
<emphasis id="B95A2A364C43C6621EE72FFEFE4A2398" bold="true" box="[199,409,723,748]" pageId="53" pageNumber="54">Elevational range</emphasis>
: 
<quantity id="4CD65BC14C43C6621F882FFEFDEB239F" box="[424,568,723,748]" metricMagnitude="3" metricUnit="m" metricValue="1.08" metricValueMax="1.63" metricValueMin="0.53" pageId="53" pageNumber="54" unit="m" value="1080.0" valueMax="1630.0" valueMin="530.0">530–1630 m</quantity>
. (n=64, including referred specimens and observations).
</paragraph>
</subSubSection>
<subSubSection id="C334A5AF4C43C6611EE72FD7FED12521" lastPageId="54" lastPageNumber="55" pageId="53" pageNumber="54" type="materials_examined">
<paragraph id="8B91F6244C43C6611EE72FD7FED12521" blockId="53.[151,1437,151,2031]" lastBlockId="54.[151,1437,151,1109]" lastPageId="54" lastPageNumber="55" pageId="53" pageNumber="54">
<emphasis id="B95A2A364C43C6621EE72FD7FE682267" bold="true" box="[199,443,762,787]" pageId="53" pageNumber="54">Referred specimens.</emphasis>
♂, 
<collectingCountry id="F339B6B44C43C6621FC92FD7FDA32267" box="[489,624,762,787]" name="Madagascar" pageId="53" pageNumber="54">Madagascar</collectingCountry>
N, Montagne d'Ambre, path to Petit Cascade, 
<geoCoordinate id="EE1A90E34C43C6621AAA2FD7FB232266" box="[1162,1264,758,787]" direction="south" orientation="latitude" pageId="53" pageNumber="54" precision="55" value="-12.526">12.526o S</geoCoordinate>
, 
<geoCoordinate id="EE1A90E34C43C6621ADE2FD7FAA12266" box="[1278,1394,758,786]" direction="east" orientation="longitude" pageId="53" pageNumber="54" precision="5" value="49.1749">49.1749o E</geoCoordinate>
+/- 
<quantity id="4CD65BC14C43C6621EB72E33FF3A2242" box="[151,233,797,822]" metricMagnitude="2" metricUnit="m" metricValue="2.0" pageId="53" pageNumber="54" unit="km" value="0.2">0.2 km</quantity>
, 
<quantity id="4CD65BC14C43C6621ED52E33FE4A2242" box="[245,409,797,822]" metricMagnitude="3" metricUnit="m" metricValue="1.084" metricValueMax="1.159" metricValueMin="1.009" pageId="53" pageNumber="54" unit="m" value="1084.0" valueMax="1159.0" valueMin="1009.0">1084 +/- 75 m</quantity>
, 
<date id="FF90D0E44C43C6621F862E30FDF42242" box="[422,551,797,822]" pageId="53" pageNumber="54" value="2004-11-16">16/11/2004</date>
, D.C. Lees: DL-05-71, 
<collectionCode id="ED3F6EE14C43C6621D142E33FC582242" box="[820,907,798,822]" pageId="53" pageNumber="54">BMNH</collectionCode>
(E) #675489 [
<collectionCode id="ED3F6EE14C43C6621A042E33FBB72242" box="[1060,1124,798,822]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="53" pageNumber="54">DNA</collectionCode>
voucher; 
<accessionNumber id="947D6BC74C43C6621AF52E30FA992242" box="[1237,1354,797,822]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/FJ819359" pageId="53" pageNumber="54">
<collectionCode id="ED3F6EE14C43C6621AF52E30FB222242" box="[1237,1265,797,822]" pageId="53" pageNumber="54">FJ</collectionCode>
819359
</accessionNumber>
]; ♂, N, Montagne d'Ambre, 
<geoCoordinate id="EE1A90E34C43C6621F5F2E69FE212229" box="[383,498,833,861]" direction="south" orientation="latitude" pageId="53" pageNumber="54" precision="5" value="-12.5289">12.5289o S</geoCoordinate>
, 
<geoCoordinate id="EE1A90E34C43C6621FDF2E68FDA02228" box="[511,627,833,861]" direction="east" orientation="longitude" pageId="53" pageNumber="54" precision="5" value="49.1727">49.1727o E</geoCoordinate>
+/- 
<quantity id="4CD65BC14C43C6621C832E69FCD22229" box="[675,769,836,861]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="53" pageNumber="54" unit="km" value="0.15">0.15 km</quantity>
, 
<quantity id="4CD65BC14C43C6621D2D2E69FC632228" box="[781,944,836,861]" metricMagnitude="3" metricUnit="m" metricValue="1.057" metricValueMax="1.082" metricValueMin="1.032" pageId="53" pageNumber="54" unit="m" value="1057.0" valueMax="1082.0" valueMin="1032.0">1057 +/- 25 m</quantity>
, D.C. Lees: DL-05-66, 
<collectionCode id="ED3F6EE14C43C6621A9B2E68FAC12229" box="[1211,1298,837,861]" pageId="53" pageNumber="54">BMNH</collectionCode>
(E) #676329 [
<collectionCode id="ED3F6EE14C43C6621EBD2E44FF0E22F5" box="[157,221,873,897]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="53" pageNumber="54">DNA</collectionCode>
voucher; 
<accessionNumber id="947D6BC74C43C6621F702E45FE1622F5" box="[336,453,872,897]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/FJ819360" pageId="53" pageNumber="54">
<collectionCode id="ED3F6EE14C43C6621F702E45FEBF22F5" box="[336,364,872,897]" pageId="53" pageNumber="54">FJ</collectionCode>
819360
</accessionNumber>
]; ♂, data as above but: DL-05-67, 
<collectionCode id="ED3F6EE14C43C6621D742E45FC7822F4" box="[852,939,872,896]" pageId="53" pageNumber="54">BMNH</collectionCode>
(E) #676330 [
<collectionCode id="ED3F6EE14C43C6621A652E44FB5622F5" box="[1093,1157,873,897]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="53" pageNumber="54">DNA</collectionCode>
voucher; 
<accessionNumber id="947D6BC74C43C6621AD82E45FABE22F5" box="[1272,1389,872,897]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/FJ819361" pageId="53" pageNumber="54">
<collectionCode id="ED3F6EE14C43C6621AD82E45FAC622F5" box="[1272,1301,872,897]" pageId="53" pageNumber="54">FJ</collectionCode>
819361
</accessionNumber>
]; ♂, N, Montagne d'Ambre, 
<geoCoordinate id="EE1A90E34C43C6621F812EA3FDF122D3" box="[417,546,907,935]" direction="south" orientation="latitude" pageId="53" pageNumber="54" precision="1" value="-12.51405">12.51405o S</geoCoordinate>
, 
<geoCoordinate id="EE1A90E34C43C6621C0F2EA3FD6222D3" box="[559,689,907,935]" direction="east" orientation="longitude" pageId="53" pageNumber="54" precision="1" value="49.17385">49.17385o E</geoCoordinate>
+/- 
<quantity id="4CD65BC14C43C6621CC02EA3FCE222D3" box="[736,817,910,935]" metricMagnitude="3" metricUnit="m" metricValue="1.6" pageId="53" pageNumber="54" unit="km" value="1.6">1.6 km</quantity>
, 
<quantity id="4CD65BC14C43C6621D1C2EA3FC0D22D3" box="[828,990,910,935]" metricMagnitude="2" metricUnit="m" metricValue="9.85" metricValueMax="11.1" metricValueMin="8.6" pageId="53" pageNumber="54" unit="m" value="985.0" valueMax="1110.0" valueMin="860.0">985 +/- 125 m</quantity>
, R. Ranaivosolo: DL-05-77, 
<collectionCode id="ED3F6EE14C43C6621B042EA2FAA822D3" box="[1316,1403,911,935]" pageId="53" pageNumber="54">BMNH</collectionCode>
(E) #676336 [
<collectionCode id="ED3F6EE14C43C6621F262E9BFE9B22BA" box="[262,328,950,974]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="53" pageNumber="54">DNA</collectionCode>
, 
<accessionNumber id="947D6BC74C43C6621F732E98FE1422BA" box="[339,455,949,974]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/FJ819364" pageId="53" pageNumber="54">
<collectionCode id="ED3F6EE14C43C6621F732E98FEBC22BA" box="[339,367,949,974]" pageId="53" pageNumber="54">FJ</collectionCode>
819364
</accessionNumber>
], IA92 [isotope voucher]; ♂, N, Montagne d'Ambre, piste vers petit cascade, 
<geoCoordinate id="EE1A90E34C43C6621B032E98FA4522BA" box="[1315,1430,946,974]" direction="south" orientation="latitude" pageId="53" pageNumber="54" precision="5" value="-12.5264">12.5264o S</geoCoordinate>
, 
<geoCoordinate id="EE1A90E34C43C6621EB72EF6FED92287" box="[151,266,984,1012]" direction="east" orientation="longitude" pageId="53" pageNumber="54" precision="5" value="49.1761">49.1761o E</geoCoordinate>
+/- 
<quantity id="4CD65BC14C43C6621F6B2EF6FE602280" box="[331,435,987,1012]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="53" pageNumber="54" unit="km" value="0.25">0.25 km</quantity>
, 
<quantity id="4CD65BC14C43C6621FE62EF6FD512287" box="[454,642,987,1012]" metricMagnitude="3" metricUnit="m" metricValue="1.064" metricValueMax="1.089" metricValueMin="1.039" pageId="53" pageNumber="54" unit="m" value="1064.0" valueMax="1089.0" valueMin="1039.0">1064 +/- 25 m</quantity>
, R. Ranaivosolo: DL-05-99, 
<collectionCode id="ED3F6EE14C43C6621DCB2EF1FB912280" box="[1003,1090,988,1012]" pageId="53" pageNumber="54">BMNH</collectionCode>
(E) #676342 [
<collectionCode id="ED3F6EE14C43C6621ACA2EF1FAF92280" box="[1258,1322,988,1012]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="53" pageNumber="54">DNA</collectionCode>
voucher; 
<accessionNumber id="947D6BC74C43C6621EB72ED2FEDF256C" box="[151,268,1023,1048]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/FJ819365" pageId="53" pageNumber="54">
<collectionCode id="ED3F6EE14C43C6621EB72ED2FF60256C" box="[151,179,1023,1048]" pageId="53" pageNumber="54">FJ</collectionCode>
819365
</accessionNumber>
]; specimen, N, Montagne d’Ambre, 
<date id="FF90D0E44C43C6621C862ED2FCFB256C" box="[678,808,1023,1048]" pageId="53" pageNumber="54" value="1995-01-14">14/01/1995</date>
, WG33 [wing imaged], IA138 [isotope voucher]; ♂, N, Montagne d'Ambre, piste petit cascade, 
<geoCoordinate id="EE1A90E34C43C6621C76290BFD19254B" box="[598,714,1059,1087]" direction="south" orientation="latitude" pageId="53" pageNumber="54" precision="5" value="-12.5264">12.5264o S</geoCoordinate>
, 
<geoCoordinate id="EE1A90E34C43C6621CF6290BFC99254A" box="[726,842,1059,1087]" direction="east" orientation="longitude" pageId="53" pageNumber="54" precision="5" value="49.1761">49.1761o E</geoCoordinate>
+/- 
<quantity id="4CD65BC14C43C6621D58290BFC05254B" box="[888,982,1062,1087]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="53" pageNumber="54" unit="km" value="0.25">0.25 km</quantity>
, 
<quantity id="4CD65BC14C43C6621DC1290BFB51254A" box="[993,1154,1062,1087]" metricMagnitude="3" metricUnit="m" metricValue="1.064" metricValueMax="1.089" metricValueMin="1.039" pageId="53" pageNumber="54" unit="m" value="1064.0" valueMax="1089.0" valueMin="1039.0">1064 +/- 25 m</quantity>
, R. Ranaivosolo: DL-05- 100, 
<collectionCode id="ED3F6EE14C43C6621EF72960FEFD2511" box="[215,302,1101,1125]" pageId="53" pageNumber="54">BMNH</collectionCode>
(E) #676343 [
<collectionCode id="ED3F6EE14C43C6621FF62960FDC52511" box="[470,534,1101,1125]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="53" pageNumber="54">DNA</collectionCode>
voucher; 
<accessionNumber id="947D6BC74C43C6621CB72961FCDF2511" box="[663,780,1100,1125]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/FJ819366" pageId="53" pageNumber="54">
<collectionCode id="ED3F6EE14C43C6621CB72961FD602511" box="[663,691,1100,1125]" pageId="53" pageNumber="54">FJ</collectionCode>
819366
</accessionNumber>
]; ♂, N, Montagne d'Ambre, petit cascade, 
<geoCoordinate id="EE1A90E34C43C6621B032961FA452511" box="[1315,1430,1097,1125]" direction="south" orientation="latitude" pageId="53" pageNumber="54" precision="5" value="-12.5264">12.5264o S</geoCoordinate>
, 
<geoCoordinate id="EE1A90E34C43C6621EB7295EFED925FF" box="[151,266,1135,1164]" direction="east" orientation="longitude" pageId="53" pageNumber="54" precision="5" value="49.1761">49.1761o E</geoCoordinate>
+/- 
<quantity id="4CD65BC14C43C6621F69295EFE7C25F8" box="[329,431,1139,1164]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="53" pageNumber="54" unit="km" value="0.25">0.25 km</quantity>
, 
<quantity id="4CD65BC14C43C6621FE2295EFDA825FF" box="[450,635,1139,1164]" metricMagnitude="3" metricUnit="m" metricValue="1.064" metricValueMax="1.089" metricValueMin="1.039" pageId="53" pageNumber="54" unit="m" value="1064.0" valueMax="1089.0" valueMin="1039.0">1064 +/- 25 m</quantity>
, R. Ranaivosolo: DL-05-101, 
<collectionCode id="ED3F6EE14C43C6621DCE295EFB9625FF" box="[1006,1093,1139,1163]" pageId="53" pageNumber="54">BMNH</collectionCode>
(E) #676344 [
<collectionCode id="ED3F6EE14C43C6621ACB295EFAF825FF" box="[1259,1323,1139,1163]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="53" pageNumber="54">DNA</collectionCode>
voucher; 
<accessionNumber id="947D6BC74C43C6621EB729B4FEDF25C6" box="[151,268,1177,1202]" httpUri="https://www.ebi.ac.uk/ena/browser/api/embl/FJ819367" pageId="53" pageNumber="54">
<collectionCode id="ED3F6EE14C43C6621EB729B4FF6025C6" box="[151,179,1177,1202]" pageId="53" pageNumber="54">FJ</collectionCode>
819367
</accessionNumber>
]; ♂, N, Montagne d'Ambre, 
<quantity id="4CD65BC14C43C6621C6D29B7FD7025C5" box="[589,675,1178,1202]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="53" pageNumber="54" unit="m" value="1100.0">1100 m</quantity>
, 
<geoCoordinate id="EE1A90E34C43C6621C9029B4FCC725C6" box="[688,788,1174,1202]" direction="south" orientation="latitude" pageId="53" pageNumber="54" precision="55" value="-12.511">12.511o S</geoCoordinate>
, 
<geoCoordinate id="EE1A90E34C43C6621D0129B4FC5425C5" box="[801,903,1174,1202]" direction="east" orientation="longitude" pageId="53" pageNumber="54" precision="55" value="49.161">49.161o E</geoCoordinate>
+/- 
<quantity id="4CD65BC14C43C6621D9729B4FBC525C6" box="[951,1046,1177,1202]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="53" pageNumber="54" unit="km" value="0.75">0.75 km</quantity>
, 
<quantity id="4CD65BC14C43C6621A0229B7FBAB25C5" box="[1058,1144,1178,1202]" metricMagnitude="3" metricUnit="m" metricValue="1.1" pageId="53" pageNumber="54" unit="m" value="1100.0">1100 m</quantity>
, 
<date id="FF90D0E44C43C6621AA529B7FB2A25C6" box="[1157,1273,1178,1202]" pageId="53" pageNumber="54" value="1992-02-19">19/2/1992</date>
, C. Kremen et al.: Genitalia 146; ♂, N, Montagne d'Ambre, 
<geoCoordinate id="EE1A90E34C43C6621C8A29EDFCCE25AC" box="[682,797,1213,1241]" direction="south" orientation="latitude" pageId="53" pageNumber="54" precision="5" value="-12.5253">12.5253o S</geoCoordinate>
, 
<geoCoordinate id="EE1A90E34C43C6621D0C29EDFC7325AC" box="[812,928,1213,1240]" direction="east" orientation="longitude" pageId="53" pageNumber="54" precision="5" value="49.1483">49.1483o E</geoCoordinate>
+/- 
<quantity id="4CD65BC14C43C6621DF529EDFBEB25AD" box="[981,1080,1216,1241]" metricMagnitude="3" metricUnit="m" metricValue="2.47" pageId="53" pageNumber="54" unit="km" value="2.47">2.47 km</quantity>
, 
<quantity id="4CD65BC14C43C6621A6529EDFB2225AC" box="[1093,1265,1216,1241]" metricMagnitude="2" metricUnit="m" metricValue="9.5" metricValueMax="13.0" metricValueMin="6.0" pageId="53" pageNumber="54" unit="m" value="950.0" valueMax="1300.0" valueMin="600.0">950 +/- 350 m</quantity>
, 
<date id="FF90D0E44C43C6621B2029EDFAA725AC" box="[1280,1396,1216,1240]" pageId="53" pageNumber="54" value="1992-02-20">20/2/1992</date>
, C. Kremen et al.: Genitalia 147; ♂, data as above but: Genitalia 149; ♂, N, Montagne d'Ambre, 
<geoCoordinate id="EE1A90E34C43C6621A8829CBFACF258B" box="[1192,1308,1251,1279]" direction="south" orientation="latitude" pageId="53" pageNumber="54" precision="5" value="-12.5253">12.5253o S</geoCoordinate>
, 
<geoCoordinate id="EE1A90E34C43C6621B0829CAFA4F258B" box="[1320,1436,1251,1279]" direction="east" orientation="longitude" pageId="53" pageNumber="54" precision="5" value="49.1483">49.1483o E</geoCoordinate>
+/- 
<quantity id="4CD65BC14C43C6621EE22823FEF72452" box="[194,292,1293,1318]" metricMagnitude="3" metricUnit="m" metricValue="2.47" pageId="53" pageNumber="54" unit="km" value="2.47">2.47 km</quantity>
, 
<quantity id="4CD65BC14C43C6621F122820FE3A2451" box="[306,489,1293,1318]" metricMagnitude="3" metricUnit="m" metricValue="1.185" metricValueMax="1.3" metricValueMin="1.0699999999999998" pageId="53" pageNumber="54" unit="m" value="1185.0" valueMax="1300.0" valueMin="1070.0">1185 +/- 115 m</quantity>
, 
<date id="FF90D0E44C43C6621FD82823FDBF2452" box="[504,620,1294,1318]" pageId="53" pageNumber="54" value="1992-02-20">20/2/1992</date>
, C. Kremen: CK841; ♂, N, Montagne d'Ambre, site MD1, 
<geoCoordinate id="EE1A90E34C43C6621B032820FA452452" box="[1315,1430,1290,1318]" direction="south" orientation="latitude" pageId="53" pageNumber="54" precision="5" value="-12.5289">12.5289o S</geoCoordinate>
, 
<geoCoordinate id="EE1A90E34C43C6621EB72819FED9243F" box="[151,266,1328,1356]" direction="east" orientation="longitude" pageId="53" pageNumber="54" precision="5" value="49.1727">49.1727o E</geoCoordinate>
+/- 
<quantity id="4CD65BC14C43C6621F1F281EFE762438" box="[319,421,1331,1356]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="53" pageNumber="54" unit="km" value="0.15">0.15 km</quantity>
., 
<quantity id="4CD65BC14C43C6621F97281EFDB1243F" box="[439,610,1331,1356]" metricMagnitude="3" metricUnit="m" metricValue="1.057" metricValueMax="1.082" metricValueMin="1.032" pageId="53" pageNumber="54" unit="m" value="1057.0" valueMax="1082.0" valueMin="1032.0">1057 +/- 25 m</quantity>
, D.C. Lees: DL-05-176, 
<date id="FF90D0E44C43C6621DA62819FBD42438" box="[902,1031,1332,1356]" pageId="53" pageNumber="54" value="2004-11-19">19/11/2004</date>
, KA521 [=KA-P521; 
<collectionCode id="ED3F6EE14C43C6621B292819FA952438" box="[1289,1350,1332,1356]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="53" pageNumber="54">DNA</collectionCode>
extract voucher]; ♀, NE, Lac Amparihibe, Makira, 
<geoCoordinate id="EE1A90E34C43C6621CA42877FD2B2407" box="[644,760,1367,1395]" direction="south" orientation="latitude" pageId="53" pageNumber="54" precision="5" value="-15.0349">15.0349o S</geoCoordinate>
, 
<geoCoordinate id="EE1A90E34C43C6621D242877FCAB2406" box="[772,888,1367,1395]" direction="east" orientation="longitude" pageId="53" pageNumber="54" precision="5" value="49.5836">49.5836o E</geoCoordinate>
+/- 
<quantity id="4CD65BC14C43C6621D872877FC2B2407" box="[935,1016,1370,1395]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="53" pageNumber="54" unit="km" value="0.5">0.5 km</quantity>
, 
<quantity id="4CD65BC14C43C6621A242876FB4B2406" box="[1028,1176,1370,1395]" metricMagnitude="2" metricUnit="m" metricValue="8.49" metricValueMax="8.99" metricValueMin="7.99" pageId="53" pageNumber="54" unit="m" value="849.0" valueMax="899.0" valueMin="799.0">849 +/- 50 m</quantity>
, 
<date id="FF90D0E44C43C6621A842876FACB2407" box="[1188,1304,1371,1395]" pageId="53" pageNumber="54" value="2003-02-31">31/2/2003</date>
, D.C. Lees, 
<collectionCode id="ED3F6EE14C43C6621EB728ACFF3D24ED" box="[151,238,1409,1433]" pageId="53" pageNumber="54">BMNH</collectionCode>
(E) #697130 [
<collectionCode id="ED3F6EE14C43C6621FA628ACFE1524ED" box="[390,454,1409,1433]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="53" pageNumber="54">DNA</collectionCode>
voucher]; ♀, NE, Lac Amparihibe, Makira, 
<geoCoordinate id="EE1A90E34C43C6621D9F28ADFBF624ED" box="[959,1061,1405,1433]" direction="south" orientation="latitude" pageId="53" pageNumber="54" precision="55" value="-15.039">15.039o S</geoCoordinate>
, 
<geoCoordinate id="EE1A90E34C43C6621A1228ADFB4424EC" box="[1074,1175,1405,1433]" direction="east" orientation="longitude" pageId="53" pageNumber="54" precision="55" value="49.572">49.572o E</geoCoordinate>
+/- 
<quantity id="4CD65BC14C43C6621AE828ADFAC924ED" box="[1224,1306,1408,1433]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="53" pageNumber="54" unit="km" value="0.5">0.5 km</quantity>
, 
<quantity id="4CD65BC14C43C6621B0528ACFF7C24C8" metricMagnitude="2" metricUnit="m" metricValue="9.0" metricValueMax="9.5" metricValueMin="8.5" pageId="53" pageNumber="54" unit="m" value="900.0" valueMax="950.0" valueMin="850.0">900 +/- 50 m</quantity>
, 
<date id="FF90D0E44C43C6621E9A2888FEFD24C9" box="[186,302,1445,1469]" pageId="53" pageNumber="54" value="2003-02-31">31/2/2003</date>
, D.C. Lees, 
<collectionCode id="ED3F6EE14C43C6621F982888FDDC24C9" box="[440,527,1445,1469]" pageId="53" pageNumber="54">BMNH</collectionCode>
(E) #697131 [
<collectionCode id="ED3F6EE14C43C6621C852888FD3624C9" box="[677,741,1445,1469]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="53" pageNumber="54">DNA</collectionCode>
voucher], KA528 [=KA-P538; 
<collectionCode id="ED3F6EE14C43C6621A692888FB5524C9" box="[1097,1158,1445,1469]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="53" pageNumber="54">DNA</collectionCode>
extract number]; ♂, NE, Lac Amparihibe-Ankilandy, walkout, 
<geoCoordinate id="EE1A90E34C43C6621C6528E6FD4E2497" box="[581,669,1479,1508]" direction="south" orientation="latitude" pageId="53" pageNumber="54" precision="555" value="-15.07">15.07o S</geoCoordinate>
, 
<geoCoordinate id="EE1A90E34C43C6621C8B28E6FCD12497" box="[683,770,1479,1508]" direction="east" orientation="longitude" pageId="53" pageNumber="54" precision="555" value="49.58">49.58o E</geoCoordinate>
+/- 
<quantity id="4CD65BC14C43C6621D1428E6FCA22490" box="[820,881,1483,1508]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="53" pageNumber="54" unit="km" value="5.0">5 km</quantity>
, 
<quantity id="4CD65BC14C43C6621D5D28E6FBF02497" box="[893,1059,1483,1507]" metricMagnitude="2" metricUnit="m" metricValue="7.0" metricValueMax="8.0" metricValueMin="6.0" pageId="53" pageNumber="54" unit="m" value="700.0" valueMax="800.0" valueMin="600.0">700 +/- 100 m</quantity>
, 
<date id="FF90D0E44C43C6621A1028E6FB452497" box="[1072,1174,1483,1507]" pageId="53" pageNumber="54" value="2003-03-01">1/3/2003</date>
, D.C. Lees, 
<collectionCode id="ED3F6EE14C43C6621B0428E6FAA82497" box="[1316,1403,1483,1507]" pageId="53" pageNumber="54">BMNH</collectionCode>
(E) #697942 [
<collectionCode id="ED3F6EE14C43C6621F2728DFFE94277E" box="[263,327,1522,1546]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="53" pageNumber="54">DNA</collectionCode>
voucher]; ♀, NE, Sahantaha, 
<geoCoordinate id="EE1A90E34C43C6621CBA28DCFD21277E" box="[666,754,1518,1546]" direction="south" orientation="latitude" pageId="53" pageNumber="54" precision="555" value="-15.23">15.23o S</geoCoordinate>
, 
<geoCoordinate id="EE1A90E34C43C6621CDF28DCFC84277D" box="[767,855,1518,1546]" direction="east" orientation="longitude" pageId="53" pageNumber="54" precision="555" value="49.53">49.53o E</geoCoordinate>
+/- 
<quantity id="4CD65BC14C43C6621DA728DCFC34277E" box="[903,999,1521,1546]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="53" pageNumber="54" unit="km" value="0.15">0.15 km</quantity>
, 
<quantity id="4CD65BC14C43C6621DD228DCFB45277D" box="[1010,1174,1521,1546]" metricMagnitude="2" metricUnit="m" metricValue="5.5" metricValueMax="6.5" metricValueMin="4.5" pageId="53" pageNumber="54" unit="m" value="550.0" valueMax="650.0" valueMin="450.0">550 +/- 100 m</quantity>
, 
<date id="FF90D0E44C43C6621A8328DFFAC4277E" box="[1187,1303,1522,1546]" pageId="53" pageNumber="54" value="2003-01-28">28/1/2003</date>
, D.C. Lees: 1645 [= 
<collectionCode id="ED3F6EE14C43C6621ECF2B35FECB2745" box="[239,280,1560,1585]" pageId="53" pageNumber="54">DL</collectionCode>
1645; 
<collectionCode id="ED3F6EE14C43C6621F402B34FE4E2745" box="[352,413,1561,1585]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="53" pageNumber="54">DNA</collectionCode>
extract number], 
<collectionCode id="ED3F6EE14C43C6621C4C2B35FD102744" box="[620,707,1560,1584]" pageId="53" pageNumber="54">BMNH</collectionCode>
(E) #697949; ♀, NE, Sahantaha, 
<geoCoordinate id="EE1A90E34C43C6621A602B35FB672744" box="[1088,1204,1557,1585]" direction="south" orientation="latitude" pageId="53" pageNumber="54" precision="5" value="-15.2406">15.2406o S</geoCoordinate>
, 
<geoCoordinate id="EE1A90E34C43C6621AE32B35FAE52744" box="[1219,1334,1557,1585]" direction="east" orientation="longitude" pageId="53" pageNumber="54" precision="5" value="49.5436">49.5436o E</geoCoordinate>
+/- 
<quantity id="4CD65BC14C43C6621B4B2B35FF6D2721" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="53" pageNumber="54" unit="km" value="0.15">0.15 km</quantity>
, 
<quantity id="4CD65BC14C43C6621EEA2B11FEA22720" box="[202,369,1596,1621]" metricMagnitude="2" metricUnit="m" metricValue="6.970000000000001" metricValueMax="7.970000000000001" metricValueMin="5.970000000000001" pageId="53" pageNumber="54" unit="m" value="697.0" valueMax="797.0" valueMin="597.0">697 +/- 100 m</quantity>
, 
<date id="FF90D0E44C43C6621F5E2B11FE212720" box="[382,498,1596,1620]" pageId="53" pageNumber="54" value="2003-01-18">18/1/2003</date>
, D.C. Lees: 1649 [= 
<collectionCode id="ED3F6EE14C43C6621CF92B11FCD12721" box="[729,770,1596,1621]" pageId="53" pageNumber="54">DL</collectionCode>
1649; 
<collectionCode id="ED3F6EE14C43C6621D692B11FC552720" box="[841,902,1596,1620]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="53" pageNumber="54">DNA</collectionCode>
extract number], 
<collectionCode id="ED3F6EE14C43C6621A722B11FB7A2720" box="[1106,1193,1596,1620]" pageId="53" pageNumber="54">BMNH</collectionCode>
(E) #697953; ♀, NW, Tsaratanana, descent, 
<geoCoordinate id="EE1A90E34C43C6621FB62B4EFDDA270F" box="[406,521,1631,1659]" direction="south" orientation="latitude" pageId="53" pageNumber="54" precision="5" value="-14.1898">14.1898o S</geoCoordinate>
, 
<geoCoordinate id="EE1A90E34C43C6621C3B2B4FFD5D270E" box="[539,654,1631,1659]" direction="east" orientation="longitude" pageId="53" pageNumber="54" precision="5" value="48.9452">48.9452o E</geoCoordinate>
+/- 
<quantity id="4CD65BC14C43C6621CE72B4EFCCE270F" box="[711,797,1634,1659]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="53" pageNumber="54" unit="km" value="0.8">0.8 km</quantity>
, 
<quantity id="4CD65BC14C43C6621D0D2B4FFC38270E" box="[813,1003,1634,1659]" metricMagnitude="3" metricUnit="m" metricValue="1.63" metricValueMax="1.73" metricValueMin="1.53" pageId="53" pageNumber="54" unit="m" value="1630.0" valueMax="1730.0" valueMin="1530.0">1630 +/- 100 m</quantity>
, 
<date id="FF90D0E44C43C6621DDC2B4EFBAD270F" box="[1020,1150,1635,1659]" pageId="53" pageNumber="54" value="2004-12-24">24/12/2004</date>
, D.C. Lees: DL-05-855, 
<collectionCode id="ED3F6EE14C43C6621EB72BAAFF3D27EB" box="[151,238,1671,1695]" pageId="53" pageNumber="54">BMNH</collectionCode>
(E) #675490 [
<collectionCode id="ED3F6EE14C43C6621FA92BAAFE1A27EB" box="[393,457,1671,1695]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="53" pageNumber="54">DNA</collectionCode>
voucher]; ♂, data as above but: DL-05-853, 
<collectionCode id="ED3F6EE14C43C6621DF12BAAFBFB27EB" box="[977,1064,1671,1695]" pageId="53" pageNumber="54">BMNH</collectionCode>
(E) #675491 [
<collectionCode id="ED3F6EE14C43C6621AE32BAAFAD027EB" box="[1219,1283,1671,1695]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="53" pageNumber="54">DNA</collectionCode>
voucher]; ♂, NW, Tsaratanana, descent, near forest margin, 
<geoCoordinate id="EE1A90E34C43C6621C802B80FCC727B1" box="[672,788,1705,1733]" direction="south" orientation="latitude" pageId="53" pageNumber="54" precision="5" value="-14.2034">14.2034o S</geoCoordinate>
, 
<geoCoordinate id="EE1A90E34C43C6621D002B81FC4727B0" box="[800,916,1705,1733]" direction="east" orientation="longitude" pageId="53" pageNumber="54" precision="5" value="48.9532">48.9532o E</geoCoordinate>
+/- 
<quantity id="4CD65BC14C43C6621DE32B81FBF127B1" box="[963,1058,1708,1733]" metricMagnitude="2" metricUnit="m" metricValue="2.5" pageId="53" pageNumber="54" unit="km" value="0.25">0.25 km</quantity>
, 
<quantity id="4CD65BC14C43C6621A0D2B80FB1D27B0" box="[1069,1230,1708,1733]" metricMagnitude="3" metricUnit="m" metricValue="1.441" metricValueMax="1.491" metricValueMin="1.391" pageId="53" pageNumber="54" unit="m" value="1441.0" valueMax="1491.0" valueMin="1391.0">1441 +/- 50 m</quantity>
, 
<date id="FF90D0E44C43C6621AFA2B80FA8F27B1" box="[1242,1372,1709,1733]" pageId="53" pageNumber="54" value="2004-12-24">24/12/2004</date>
, D.C. Lees: DL-05-844, 
<collectionCode id="ED3F6EE14C43C6621F4D2BFCFE17279D" box="[365,452,1745,1769]" pageId="53" pageNumber="54">BMNH</collectionCode>
(E) #675492 [
<collectionCode id="ED3F6EE14C43C6621C432BFCFD70279D" box="[611,675,1745,1769]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="53" pageNumber="54">DNA</collectionCode>
voucher]; ♂, data as above but: DL-05-843, 
<collectionCode id="ED3F6EE14C43C6621A982BFCFADC279D" box="[1208,1295,1745,1769]" pageId="53" pageNumber="54">BMNH</collectionCode>
(E) #675493 [
<collectionCode id="ED3F6EE14C43C6621EBD2BD8FF0E2679" box="[157,221,1781,1805]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="53" pageNumber="54">DNA</collectionCode>
voucher]; ♂, data as above but: DL-05-842, 
<collectionCode id="ED3F6EE14C43C6621CFC2BD8FCE02679" box="[732,819,1781,1805]" pageId="53" pageNumber="54">BMNH</collectionCode>
(E) #675494 [
<collectionCode id="ED3F6EE14C43C6621DEB2BD8FBD82679" box="[971,1035,1781,1805]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="53" pageNumber="54">DNA</collectionCode>
voucher]; ♀, NE, Lac Amparihibe, 
<geoCoordinate id="EE1A90E34C43C6621EB72A37FED92647" box="[151,266,1815,1843]" direction="south" orientation="latitude" pageId="53" pageNumber="54" precision="5" value="-15.0351">15.0351o S</geoCoordinate>
, 
<geoCoordinate id="EE1A90E34C43C6621F382A37FE5F2646" box="[280,396,1815,1843]" direction="east" orientation="longitude" pageId="53" pageNumber="54" precision="5" value="49.5862">49.5862o E</geoCoordinate>
+/- 
<quantity id="4CD65BC14C43C6621F9E2A37FDCD2647" box="[446,542,1818,1843]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="53" pageNumber="54" unit="km" value="0.15">0.15 km</quantity>
, 
<quantity id="4CD65BC14C43C6621C0B2A36FD102646" box="[555,707,1818,1843]" metricMagnitude="2" metricUnit="m" metricValue="8.7" metricValueMax="9.2" metricValueMin="8.2" pageId="53" pageNumber="54" unit="m" value="870.0" valueMax="920.0" valueMin="820.0">870 +/- 50 m</quantity>
, 
<date id="FF90D0E44C43C6621CF02A36FC962647" box="[720,837,1819,1843]" pageId="53" pageNumber="54" value="2002-02-23">23/2/2002</date>
, D.C. Lees, 
<collectionCode id="ED3F6EE14C43C6621DF32A36FBF92647" box="[979,1066,1819,1843]" pageId="53" pageNumber="54">BMNH</collectionCode>
(E) #675516 [
<collectionCode id="ED3F6EE14C43C6621AE42A36FAD72647" box="[1220,1284,1819,1843]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="53" pageNumber="54">DNA</collectionCode>
voucher]; ♀, NE, Lac Amparihibe, 
<geoCoordinate id="EE1A90E34C43C6621FB72A6CFE2E262E" box="[407,509,1854,1882]" direction="south" orientation="latitude" pageId="53" pageNumber="54" precision="55" value="-15.039">15.039o S</geoCoordinate>
, 
<geoCoordinate id="EE1A90E34C43C6621C2C2A6CFDA2262D" box="[524,625,1854,1882]" direction="east" orientation="longitude" pageId="53" pageNumber="54" precision="55" value="49.572">49.572o E</geoCoordinate>
+/- 
<quantity id="4CD65BC14C43C6621C862A6CFD2A262E" box="[678,761,1857,1882]" metricMagnitude="2" metricUnit="m" metricValue="5.0" pageId="53" pageNumber="54" unit="km" value="0.5">0.5 km</quantity>
, 
<quantity id="4CD65BC14C43C6621D272A6FFC70262D" box="[775,931,1857,1882]" metricMagnitude="2" metricUnit="m" metricValue="8.14" metricValueMax="8.64" metricValueMin="7.64" pageId="53" pageNumber="54" unit="m" value="814.0" valueMax="864.0" valueMin="764.0">814 +/- 50 m</quantity>
, 
<date id="FF90D0E44C43C6621D912A6FFBF6262E" box="[945,1061,1858,1882]" pageId="53" pageNumber="54" value="2003-02-19">19/2/2003</date>
, D.C. Lees; 
<collectionCode id="ED3F6EE14C43C6621A982A6FFADC262E" box="[1208,1295,1858,1882]" pageId="53" pageNumber="54">BMNH</collectionCode>
(E) #675520 [
<collectionCode id="ED3F6EE14C43C6621EBD2A48FF0E2609" box="[157,221,1893,1917]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="53" pageNumber="54">DNA</collectionCode>
voucher]; specimen, NW, Tsaratanana, Antetykalambazaha, below camp, “fougeres”, fruit low trap, 
<geoCoordinate id="EE1A90E34C43C6621EB72AA1FED926D0" box="[151,266,1928,1956]" direction="south" orientation="latitude" pageId="53" pageNumber="54" precision="5" value="-14.1898">14.1898o S</geoCoordinate>
, 
<geoCoordinate id="EE1A90E34C43C6621F3B2AA6FE5C26D7" box="[283,399,1928,1956]" direction="east" orientation="longitude" pageId="53" pageNumber="54" precision="5" value="48.9452">48.9452o E</geoCoordinate>
+/- 
<quantity id="4CD65BC14C43C6621FE72AA1FDCF26D0" box="[455,540,1931,1956]" metricMagnitude="2" metricUnit="m" metricValue="1.0" pageId="53" pageNumber="54" unit="km" value="0.1">0.1 km</quantity>
, 
<quantity id="4CD65BC14C43C6621C0C2AA6FD0826D7" box="[556,731,1931,1956]" metricMagnitude="3" metricUnit="m" metricValue="1.63" metricValueMax="1.655" metricValueMin="1.605" pageId="53" pageNumber="54" unit="m" value="1630.0" valueMax="1655.0" valueMin="1605.0">1630 +/- 25 m</quantity>
, R. R. Ranaivosolo: DL-05-829, 
<collectionCode id="ED3F6EE14C43C6621A452AA1FB6F26D0" box="[1125,1212,1932,1956]" pageId="53" pageNumber="54">BMNH</collectionCode>
(E) #676598 [
<collectionCode id="ED3F6EE14C43C6621B7C2AA1FA4F26D0" box="[1372,1436,1932,1956]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="53" pageNumber="54">DNA</collectionCode>
voucher]; ♀, NW, Tsaratanana, Ampidiranala, 
<geoCoordinate id="EE1A90E34C43C6621C812A9FFCF126BE" box="[673,802,1966,1995]" direction="south" orientation="latitude" pageId="53" pageNumber="54" precision="1" value="-14.20025">14.20025o S</geoCoordinate>
, 
<geoCoordinate id="EE1A90E34C43C6621D0E2A9FFC6326BE" box="[814,944,1966,1995]" direction="east" orientation="longitude" pageId="53" pageNumber="54" precision="1" value="48.95175">48.95175o E</geoCoordinate>
+/- 
<quantity id="4CD65BC14C43C6621DFF2A9FFBEE26BF" box="[991,1085,1970,1995]" metricMagnitude="2" metricUnit="m" metricValue="5.4" pageId="53" pageNumber="54" unit="km" value="0.54">0.54 km</quantity>
, 
<quantity id="4CD65BC14C43C6621A682A9FFB3A26BE" box="[1096,1257,1970,1995]" metricMagnitude="3" metricUnit="m" metricValue="1.5419999999999998" metricValueMax="1.627" metricValueMin="1.4569999999999999" pageId="53" pageNumber="54" unit="m" value="1542.0" valueMax="1627.0" valueMin="1457.0">1542 +/- 85 m</quantity>
, 
<date id="FF90D0E44C43C6621AD52A9FFAA426BE" box="[1269,1399,1970,1994]" pageId="53" pageNumber="54" value="2004-12-20">20/12/2004</date>
, R. Ranaivosolo: DL-05-757, 2487 [= 
<collectionCode id="ED3F6EE14C43C6621C332AFBFDE8269A" box="[531,571,2006,2030]" pageId="53" pageNumber="54">DL</collectionCode>
2487; 
<collectionCode id="ED3F6EE14C43C6621CA22AFBFD6C269A" box="[642,703,2006,2030]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="53" pageNumber="54">DNA</collectionCode>
extract number], 
<collectionCode id="ED3F6EE14C43C6621DA62AFBFC0E269A" box="[902,989,2006,2030]" pageId="53" pageNumber="54">BMNH</collectionCode>
(E) #676687; ♀, N, Montagne d'Ambre, 
<geoCoordinate id="EE1A90E34C40C6611EB72DB7FED921C7" box="[151,266,151,179]" direction="south" orientation="latitude" pageId="54" pageNumber="55" precision="5" value="-12.5289">12.5289o S</geoCoordinate>
, 
<geoCoordinate id="EE1A90E34C40C6611F382DB6FE5F21C6" box="[280,396,151,179]" direction="east" orientation="longitude" pageId="54" pageNumber="55" precision="5" value="49.1727">49.1727o E</geoCoordinate>
+/- 
<quantity id="4CD65BC14C40C6611F9E2DB7FDCD21C7" box="[446,542,154,179]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="54" pageNumber="55" unit="km" value="0.15">0.15 km</quantity>
, 
<quantity id="4CD65BC14C40C6611C0B2DB7FD0221C6" box="[555,721,154,179]" metricMagnitude="3" metricUnit="m" metricValue="1.057" metricValueMax="1.082" metricValueMin="1.032" pageId="54" pageNumber="55" unit="m" value="1057.0" valueMax="1082.0" valueMin="1032.0">1057 +/- 25 m</quantity>
, 
<date id="FF90D0E44C40C6611CFE2DB6FCB221C7" box="[734,865,155,179]" pageId="54" pageNumber="55" value="2004-11-19">19/11/2004</date>
; D.C. Lees: DL-05-182: IA204 [isotope voucher]; ♀, N, Montagne d'Ambre, 
<geoCoordinate id="EE1A90E34C40C6611FE72DECFDE921AD" box="[455,570,190,218]" direction="south" orientation="latitude" pageId="54" pageNumber="55" precision="5" value="-12.5289">12.5289o S</geoCoordinate>
, 
<geoCoordinate id="EE1A90E34C40C6611C672DECFD6821AD" box="[583,699,190,217]" direction="east" orientation="longitude" pageId="54" pageNumber="55" precision="5" value="49.1727">49.1727o E</geoCoordinate>
+/- 
<quantity id="4CD65BC14C40C6611CCB2DECFC9821AE" box="[747,843,193,218]" metricMagnitude="2" metricUnit="m" metricValue="1.5" pageId="54" pageNumber="55" unit="km" value="0.15">0.15 km</quantity>
, 
<quantity id="4CD65BC14C40C6611D772DECFC2821AD" box="[855,1019,193,218]" metricMagnitude="3" metricUnit="m" metricValue="1.057" metricValueMax="1.082" metricValueMin="1.032" pageId="54" pageNumber="55" unit="m" value="1057.0" valueMax="1082.0" valueMin="1032.0">1057 +/- 25 m</quantity>
, 
<date id="FF90D0E44C40C6611A272DECFB5921AD" box="[1031,1162,193,217]" pageId="54" pageNumber="55" value="2004-11-19">19/11/2004</date>
; D.C. Lees: DL-05-183, IA205 [isotope voucher]; ♂ [
<collectionCode id="ED3F6EE14C40C6611FC52DC5FDED218B" box="[485,574,232,255]" pageId="54" pageNumber="55">BMNH</collectionCode>
] (
<figureCitation id="1315EAA14C40C6611C762DCAFD792074" box="[598,682,231,256]" captionStart="FIGURE 15" captionStartId="49.[151,250,1351,1373]" captionTargetBox="[386,1205,210,1306]" captionTargetId="figure@49.[363,1224,193,1330]" captionTargetPageId="49" captionText="FIGURE 15. ♂ genitalia in LV (left), DV (right) unless otherwise stated. A: Heteropsis kremenae Lees, sp. nov. (Masoala, Ambanivony; 19 DL). B: Ht. avaratra Lees &amp; Kremen, sp. nov. (315 DL, referred specimen, Joffreville). C: Heteropsis sogai Lees, sp. nov. PT (Tsaratanana; 96 DL, MNHN), LV, SV. Scale bar: 3 mm." httpUri="https://zenodo.org/record/264612/files/figure.png" pageId="54" pageNumber="55">Fig. 15</figureCitation>
B), N, Joffreville [ca. 
<geoCoordinate id="EE1A90E34C40C6611D882DC5FBDE2074" box="[936,1037,228,256]" direction="south" orientation="latitude" pageId="54" pageNumber="55" precision="55" value="-12.483">12.483o S</geoCoordinate>
, 
<geoCoordinate id="EE1A90E34C40C6611A3C2DC5FBB5218B" box="[1052,1126,228,256]" direction="east" orientation="longitude" pageId="54" pageNumber="55" precision="5555" value="49.2">49.2o E</geoCoordinate>
+/- 
<quantity id="4CD65BC14C40C6611ABA2DCAFB362074" box="[1178,1253,231,256]" metricMagnitude="4" metricUnit="m" metricValue="1.5" pageId="54" pageNumber="55" unit="km" value="15.0">15 km</quantity>
, 
<quantity id="4CD65BC14C40C6611AD32DC5FA43218B" box="[1267,1424,231,256]" metricMagnitude="2" metricUnit="m" metricValue="7.17" metricValueMax="7.67" metricValueMin="6.67" pageId="54" pageNumber="55" unit="m" value="717.0" valueMax="767.0" valueMin="667.0">717 +/- 50 m</quantity>
], 1921, E. Brouhard|315 
<collectionCode id="ED3F6EE14C40C6611FBB2C26FE122050" box="[411,449,267,292]" pageId="54" pageNumber="55">DL</collectionCode>
[genitalia]; specimen [malaise], N, Parc National Montagne d'Ambre [1st campsite], 
<geoCoordinate id="EE1A90E34C40C6611EB72C1FFED9203E" box="[151,266,302,331]" direction="south" orientation="latitude" pageId="54" pageNumber="55" precision="5" value="-12.5144">12.5144o S</geoCoordinate>
, 
<geoCoordinate id="EE1A90E34C40C6611F3A2C1FFE43203E" box="[282,400,302,330]" direction="east" orientation="longitude" pageId="54" pageNumber="55" precision="5" value="49.1814">49.1814o E</geoCoordinate>
, 
<quantity id="4CD65BC14C40C6611FBF2C1FFE38203E" box="[415,491,306,331]" metricMagnitude="2" metricUnit="m" metricValue="9.6" pageId="54" pageNumber="55" unit="m" value="960.0">960 m</quantity>
, Irwin, Schlinger, Harin'Hala: MA-01-01A-02; specimen [malaise], data as above but: MA-01-01A-06; specimen [malaise], N, Parc National Montagne d'Ambre [Petit Lac road], 
<geoCoordinate id="EE1A90E34C40C6611B032C75FA452005" box="[1315,1430,341,369]" direction="south" orientation="latitude" pageId="54" pageNumber="55" precision="5" value="-12.5203">12.5203o S</geoCoordinate>
, 
<geoCoordinate id="EE1A90E34C40C6611EB72C52FEDF20E3" box="[151,268,380,407]" direction="east" orientation="longitude" pageId="54" pageNumber="55" precision="5" value="49.1792">49.1792o E</geoCoordinate>
, 
<quantity id="4CD65BC14C40C6611F3E2C52FEA920E3" box="[286,378,383,408]" metricMagnitude="3" metricUnit="m" metricValue="1.125" pageId="54" pageNumber="55" unit="m" value="1125.0">1125 m</quantity>
, R. Harin’Hala: MA-01-01D-05; specimen [malaise], data as above but: MA-01-01D-06; specimen [malaise], data as above but: MA-01-01D-07; specimen [malaise], data as above but: MA-01-01D-08; series DCL-DL-3369 to DCL-DB-3385 [
<collectionCode id="ED3F6EE14C40C6611CA22CEAFD3320AA" box="[642,736,455,478]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="54" pageNumber="55">MNHN</collectionCode>
], including: ♂ [
<collectionCode id="ED3F6EE14C40C6611D872CEAFBD620AA" box="[935,1029,455,478]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="54" pageNumber="55">MNHN</collectionCode>
], Nord, Les Rousettes, Montagne d'Ambre, 
<quantity id="4CD65BC14C40C6611F282CC3FEB12371" box="[264,354,494,518]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="54" pageNumber="55" unit="m" value="1000.0">1000 m</quantity>
, [ca. 
<geoCoordinate id="EE1A90E34C40C6611F822CC0FDDB2372" box="[418,520,490,518]" direction="south" orientation="latitude" pageId="54" pageNumber="55" precision="55" value="-12.525">12.525o S</geoCoordinate>
, 
<geoCoordinate id="EE1A90E34C40C6611C372CC3FD582371" box="[535,651,490,518]" direction="east" orientation="longitude" pageId="54" pageNumber="55" precision="5" value="49.1722">49.1722o E</geoCoordinate>
+/- 
<quantity id="4CD65BC14C40C6611C9E2CC0FD2C2372" box="[702,767,493,518]" metricMagnitude="3" metricUnit="m" metricValue="5.0" pageId="54" pageNumber="55" unit="km" value="5.0">5 km</quantity>
], 
<date id="FF90D0E44C40C6611D342CC0FC542372" box="[788,903,493,518]" pageId="54" pageNumber="55" value="1958-11-03">3/11/1958</date>
; ♂ [
<collectionCode id="ED3F6EE14C40C6611D9A2CC3FBCB2371" box="[954,1048,494,517]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="54" pageNumber="55">MNHN</collectionCode>
], data as above but: 
<date id="FF90D0E44C40C6611B272CC0FAA82372" box="[1287,1403,493,518]" pageId="54" pageNumber="55" value="1958-12-06">6/12/1958</date>
, P. Viette; 3 ♂♂ [
<collectionCode id="ED3F6EE14C40C6611F1D2F3CFE48235C" box="[317,411,529,552]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="54" pageNumber="55">MNHN</collectionCode>
], data as above but: 
<date id="FF90D0E44C40C6611CAA2F3DFD2D235D" box="[650,766,528,553]" pageId="54" pageNumber="55" value="1970-05-25">25/5/1970</date>
, P. Griveaud; specimen, N, Petit Lac, Montagne d'Ambre, 
<quantity id="4CD65BC14C40C6611EB72F1AFF3D233B" box="[151,238,567,592]" metricMagnitude="3" metricUnit="m" metricValue="1.115" pageId="54" pageNumber="55" unit="m" value="1115.0">1115 m</quantity>
, [
<geoCoordinate id="EE1A90E34C40C6611F222F1AFEAB2324" box="[258,376,564,592]" direction="south" orientation="latitude" pageId="54" pageNumber="55" precision="5" value="-12.5333">12.5333o S</geoCoordinate>
, 
<geoCoordinate id="EE1A90E34C40C6611FA72F1AFE28233B" box="[391,507,564,592]" direction="east" orientation="longitude" pageId="54" pageNumber="55" precision="5" value="49.1667">49.1667o E</geoCoordinate>
+/- 
<quantity id="4CD65BC14C40C6611C0E2F15FD5D2324" box="[558,654,567,592]" metricMagnitude="2" metricUnit="m" metricValue="9.1" pageId="54" pageNumber="55" unit="km" value="0.91">0.91 km</quantity>
, 
<quantity id="4CD65BC14C40C6611CBB2F1AFD27233B" box="[667,756,567,592]" metricMagnitude="3" metricUnit="m" metricValue="1.115" pageId="54" pageNumber="55" unit="m" value="1115.0">1115 m</quantity>
], 
<date id="FF90D0E44C40C6611D292F1AFCAD2324" box="[777,894,567,592]" pageId="54" pageNumber="55" value="1970-05-26">26/5/1970</date>
, P. Griveaud; ♂ [
<collectionCode id="ED3F6EE14C40C6611A652F15FB70233B" box="[1093,1187,568,591]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="54" pageNumber="55">MNHN</collectionCode>
], data as above but ♂ [
<collectionCode id="ED3F6EE14C40C6611EBD2F71FF2F2307" box="[157,252,604,627]" country="France" httpUri="http://biocol.org/urn:lsid:biocol.org:col:34988" name="Museum National d'Histoire Naturelle" pageId="54" pageNumber="55">MNHN</collectionCode>
]: 
<date id="FF90D0E44C40C6611F2E2F76FE7F2300" box="[270,428,603,628]" pageId="54" pageNumber="55" value="1970-05-26" valueMax="1970-05-27" valueMin="1970-05-26">26–27/5/1970</date>
, P. Griveaud; ♂, N, Montagne d'Ambre, vers premier petit sommet, route GCASC-&gt;Gite, 
<geoCoordinate id="EE1A90E34C40C6611EB72FACFECB23EE" box="[151,280,638,666]" direction="south" orientation="latitude" pageId="54" pageNumber="55" precision="1" value="-12.59435">12.59435o S</geoCoordinate>
, 
<geoCoordinate id="EE1A90E34C40C6611F082FACFE4823ED" box="[296,411,638,666]" direction="east" orientation="longitude" pageId="54" pageNumber="55" precision="5" value="49.1583">49.1583o E</geoCoordinate>
+/- 
<quantity id="4CD65BC14C40C6611FF02FAFFDDD23EE" box="[464,526,641,666]" metricMagnitude="3" metricUnit="m" metricValue="1.0" pageId="54" pageNumber="55" unit="km" value="1.0">1 km</quantity>
, 
<quantity id="4CD65BC14C40C6611C3C2FAFFD6B23ED" box="[540,696,641,666]" metricMagnitude="2" metricUnit="m" metricValue="8.72" metricValueMax="8.97" metricValueMin="8.47" pageId="54" pageNumber="55" unit="m" value="872.0" valueMax="897.0" valueMin="847.0">872 +/- 25 m</quantity>
, 
<date id="FF90D0E44C40C6611CE62FAFFC9423EE" box="[710,839,642,666]" pageId="54" pageNumber="55" value="2004-11-17">17/11/2004</date>
, R. Ranaivosolo: DL-05-121; ♂ [
<collectionCode id="ED3F6EE14C40C6611AEB2FAFFAF423ED" box="[1227,1319,642,665]" pageId="54" pageNumber="55">BMNH</collectionCode>
], N, Forêt d'Ambre, 
<quantity id="4CD65BC14C40C6611F292F85FE8C23B5" box="[265,351,680,705]" metricMagnitude="2" metricUnit="m" metricValue="9.144" pageId="54" pageNumber="55" unit="ft" value="3000.0">3000 ft</quantity>
, [ca. 
<geoCoordinate id="EE1A90E34C40C6611FB12F85FDDA23B4" box="[401,521,677,705]" direction="south" orientation="latitude" pageId="54" pageNumber="55" precision="5" value="-12.6282">12.6282o S</geoCoordinate>
, 
<geoCoordinate id="EE1A90E34C40C6611C392F85FD5E23B4" box="[537,653,677,704]" direction="east" orientation="longitude" pageId="54" pageNumber="55" precision="5" value="49.1448">49.1448o E</geoCoordinate>
+/- 
<quantity id="4CD65BC14C40C6611CE12F85FCF023B5" box="[705,803,680,705]" metricMagnitude="3" metricUnit="m" metricValue="8.51" pageId="54" pageNumber="55" unit="km" value="8.51">8.51 km</quantity>
, 
<quantity id="4CD65BC14C40C6611D112F85FCAC23B4" box="[817,895,680,704]" metricMagnitude="2" metricUnit="m" metricValue="9.14" pageId="54" pageNumber="55" unit="m" value="914.0">914 m</quantity>
], 
<date id="FF90D0E44C40C6611DB52F85FC2823B5" box="[917,1019,680,705]" pageId="54" pageNumber="55" value="1906-03-07">7/3/1906</date>
, Meade-Waldo| Photographed by B. D’Abrera 77/78| 
<collectionCode id="ED3F6EE14C40C6611F6C2FE2FE762393" box="[332,421,719,743]" pageId="54" pageNumber="55">BMNH</collectionCode>
(E) 674887; ♂, E, Anjanaharibe Sud, ~ 
<quantity id="4CD65BC14C40C6611D752FE3FC4C2393" box="[853,927,718,743]" metricMagnitude="2" metricUnit="m" metricValue="9.5" pageId="54" pageNumber="55" unit="m" value="950.0">950 m</quantity>
, 
<geoCoordinate id="EE1A90E34C40C6611D8B2FE3FBCD2393" box="[939,1054,715,743]" direction="south" orientation="latitude" pageId="54" pageNumber="55" precision="5" value="-14.7502">14.7502o S</geoCoordinate>
, 
<geoCoordinate id="EE1A90E34C40C6611A0B2FE2FB4D2393" box="[1067,1182,715,743]" direction="east" orientation="longitude" pageId="54" pageNumber="55" precision="5" value="49.4998">49.4998o E</geoCoordinate>
+/- 
<quantity id="4CD65BC14C40C6611AEC2FE2FACE2393" box="[1228,1309,718,743]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="54" pageNumber="55" unit="km" value="0.8">0.8 km</quantity>
, 
<quantity id="4CD65BC14C40C6611B082FE3FF7C227E" metricMagnitude="2" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="54" pageNumber="55" unit="m" value="950.0" valueMax="1000.0" valueMin="900.0">950 +/- 50 m</quantity>
, 
<date id="FF90D0E44C40C6611EE12FDFFE92227F" box="[193,321,754,779]" pageId="54" pageNumber="55" value="1995-11-25">25/11/1995</date>
, 11:00–13:15, C. Kremen: CK484, IA336 [isotope voucher]; specimen, NE, Lac Amparihibe, Makira, 
<geoCoordinate id="EE1A90E34C40C6611ED62E34FEB92245" box="[246,362,790,818]" direction="south" orientation="latitude" pageId="54" pageNumber="55" precision="5" value="-15.0301">15.0301o S</geoCoordinate>
, 
<geoCoordinate id="EE1A90E34C40C6611F572E34FE392245" box="[375,490,790,818]" direction="east" orientation="longitude" pageId="54" pageNumber="55" precision="5" value="49.5827">49.5827o E</geoCoordinate>
+/- 
<quantity id="4CD65BC14C40C6611C3A2E34FDAA2246" box="[538,633,793,818]" metricMagnitude="2" metricUnit="m" metricValue="7.5" pageId="54" pageNumber="55" unit="km" value="0.75">0.75 km</quantity>
, 
<quantity id="4CD65BC14C40C6611CA42E34FCC92245" box="[644,794,793,818]" metricMagnitude="2" metricUnit="m" metricValue="9.2" metricValueMax="9.95" metricValueMin="8.45" pageId="54" pageNumber="55" unit="m" value="920.0" valueMax="995.0" valueMin="845.0">920 +/- 75 m</quantity>
, 
<date id="FF90D0E44C40C6611D062E34FC492245" box="[806,922,793,817]" pageId="54" pageNumber="55" value="2003-02-19">19/2/2003</date>
, R. Ranaivosolo: 
<collectionCode id="ED3F6EE14C40C6611A422E34FB5A2245" box="[1122,1161,793,817]" pageId="54" pageNumber="55">DL</collectionCode>
03-0002, IA337 [isotope voucher]; ♂, NE, Lohan i' Sahantaha, site SD1, Makira, 
<geoCoordinate id="EE1A90E34C40C6611D2B2E12FCAD222C" box="[779,894,828,856]" direction="south" orientation="latitude" pageId="54" pageNumber="55" precision="5" value="-15.2287">15.2287o S</geoCoordinate>
, 
<geoCoordinate id="EE1A90E34C40C6611DAB2E12FC2D2223" box="[907,1022,828,856]" direction="east" orientation="longitude" pageId="54" pageNumber="55" precision="5" value="49.532">49.5320o E</geoCoordinate>
+/- 
<quantity id="4CD65BC14C40C6611A0C2E12FBAF2223" box="[1068,1148,831,856]" metricMagnitude="-1" metricUnit="m" metricValue="1.5" pageId="54" pageNumber="55" unit="m" value="0.15">0.15 m</quantity>
, 
<quantity id="4CD65BC14C40C6611AA72E6DFACA2223" box="[1159,1305,831,856]" metricMagnitude="2" metricUnit="m" metricValue="4.0" metricValueMax="4.5" metricValueMin="3.5" pageId="54" pageNumber="55" unit="m" value="400.0" valueMax="450.0" valueMin="350.0">400 +/- 50 m</quantity>
, D.C. Lees, 
<date id="FF90D0E44C40C6611EB72E49FEC92208" box="[151,282,868,892]" pageId="54" pageNumber="55" value="2003-01-24">24/01/2003</date>
: 10:00, 
<collectionCode id="ED3F6EE14C40C6611FA02E49FE042208" box="[384,471,868,892]" pageId="54" pageNumber="55">BMNH</collectionCode>
(E) #697350, 0 234 [= 
<collectionCode id="ED3F6EE14C40C6611CF82E49FCD32208" box="[728,768,868,892]" pageId="54" pageNumber="55">DL</collectionCode>
0234, 
<collectionCode id="ED3F6EE14C40C6611D6E2E49FC582208" box="[846,907,868,892]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="54" pageNumber="55">DNA</collectionCode>
extract number], KA538 [=KA-P538, 
<collectionCode id="ED3F6EE14C40C6611B402E49FA4E2208" box="[1376,1437,868,892]" country="Australia" httpUri="http://biocol.org/urn:lsid:biocol.org:col:15536" name="Department of Natural Resources, Environment, The Arts and Sport" pageId="54" pageNumber="55">DNA</collectionCode>
extract number]; ♂, E, Anjanaharibe Sud, 
<geoCoordinate id="EE1A90E34C40C6611CA82EA4FD2F22D6" box="[648,764,902,930]" direction="south" orientation="latitude" pageId="54" pageNumber="55" precision="5" value="-14.7502">14.7502o S</geoCoordinate>
, 
<geoCoordinate id="EE1A90E34C40C6611D2C2EA7FC5322D5" box="[780,896,902,930]" direction="east" orientation="longitude" pageId="54" pageNumber="55" precision="5" value="49.4998">49.4998o E</geoCoordinate>
+/- 
<quantity id="4CD65BC14C40C6611D8B2EA7FBD222D6" box="[939,1025,905,930]" metricMagnitude="2" metricUnit="m" metricValue="8.0" pageId="54" pageNumber="55" unit="km" value="0.8">0.8 km</quantity>
, 
<quantity id="4CD65BC14C40C6611A302EA4FB7C22D5" box="[1040,1199,905,930]" metricMagnitude="2" metricUnit="m" metricValue="9.5" metricValueMax="10.0" metricValueMin="9.0" pageId="54" pageNumber="55" unit="m" value="950.0" valueMax="1000.0" valueMin="900.0">950 +/- 50 m</quantity>
, 
<date id="FF90D0E44C40C6611A9F2EA4FAEC22D6" box="[1215,1343,905,930]" pageId="54" pageNumber="55" value="1995-11-25">25/11/1995</date>
, 11:00– 13:15, C. Kremen: CK485, IA180 [isotope voucher]; ♂, NE, Anjanaharibe Sud, 11/2014, D.C. Lees: DL-14A- 0 0 83, IA495 [isotope voucher]; ♂, NE, Anjanaharibe Sud, 11/2014; D.C. Lees: DL-14A-0059, IA496 [isotope voucher]; ♀, NE, Anjanaharibe Sud, Takhtajania trail, 
<date id="FF90D0E44C40C6611D0D2ED8FC7D2579" box="[813,942,1013,1037]" pageId="54" pageNumber="55" value="2014-11-23">23/11/2014</date>
, D.C.Lees: 
<collectionCode id="ED3F6EE14C40C6611A602ED8FBBB257A" box="[1088,1128,1013,1038]" pageId="54" pageNumber="55">DL</collectionCode>
14A-0365, IA561 [isotope voucher]; ♂, NE, Anjanaharibe Sud, summit trail, 
<date id="FF90D0E44C40C6611CC82934FCB82545" box="[744,875,1049,1073]" pageId="54" pageNumber="55" value="2014-11-19">19/11/2014</date>
: 05:51, D.C.Lees: 
<collectionCode id="ED3F6EE14C40C6611A672934FBBD2545" box="[1095,1134,1049,1073]" pageId="54" pageNumber="55">DL</collectionCode>
14A-0170, IA563 [isotope voucher].
</paragraph>
</subSubSection>
</treatment>
</document>