287 lines
28 KiB
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287 lines
28 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.905.39592" ID-GBIF-Dataset="ee5a0817-7cfc-4c21-ba7d-3beabb06cccd" ID-PMC="PMC7004242" ID-Pensoft-Pub="1313-2970-905-1" ID-Pensoft-UUID="4C7E0CB428DF56BDB15B7AC8D6707961" ID-PubMed="32063726" ID-ZooBank="F4C766E4633A41039FFDE952718F41FB" ModsDocID="1313-2970-905-1" checkinTime="1579539902851" checkinUser="pensoft" docAuthor="Salata, Sebastian & Fisher, Brian L." docDate="2020" docId="FE9EE26351DF4A7FD85F6AB8757CD313" docLanguage="en" docName="ZooKeys 905: 1-235" docOrigin="ZooKeys 905" docSource="http://dx.doi.org/10.3897/zookeys.905.39592" docTitle="Pheidole ankerana Salata & Fisher 2020, sp. nov." docType="treatment" docUuid="5845FFBC-72D4-43A6-AF1D-11C57445217F" docUuidSource="ZooBank" docVersion="5" id="4C7E0CB428DF56BDB15B7AC8D6707961" lastPageNumber="1" masterDocId="4C7E0CB428DF56BDB15B7AC8D6707961" masterDocTitle="Pheidole Westwood, 1839 (Hymenoptera, Formicidae) of Madagascar - an introduction and a taxonomic revision of eleven species groups" masterLastPageNumber="235" masterPageNumber="1" pageNumber="1" updateTime="1668168337901" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Pheidole Westwood, 1839 (Hymenoptera, Formicidae) of Madagascar - an introduction and a taxonomic revision of eleven species groups</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Salata, Sebastian</mods:namePart>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Fisher, Brian L.</mods:namePart>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:relatedItem type="host">
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<mods:titleInfo>
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<mods:title>ZooKeys</mods:title>
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</mods:titleInfo>
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<mods:part>
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<mods:date>2020</mods:date>
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<mods:detail type="volume">
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<mods:number>905</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>1</mods:start>
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<mods:end>235</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.905.39592</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.905.39592</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-905-1</mods:identifier>
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<mods:identifier type="ZooBank">F4C766E4633A41039FFDE952718F41FB</mods:identifier>
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<mods:identifier type="Pensoft-UUID">4C7E0CB428DF56BDB15B7AC8D6707961</mods:identifier>
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</mods:mods>
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<treatment ID-GBIF-Taxon="161776991" LSID="urn:lsid:zoobank.org:act:5845FFBC-72D4-43A6-AF1D-11C57445217F" httpUri="http://treatment.plazi.org/id/FE9EE26351DF4A7FD85F6AB8757CD313" lastPageNumber="1" pageId="0" pageNumber="1">
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<subSubSection pageId="0" pageNumber="1" type="nomenclature">
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<paragraph pageId="0" pageNumber="1">
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<taxonomicName LSID="http://zoobank.org/5845FFBC-72D4-43A6-AF1D-11C57445217F" authority="Salata & Fisher, 2020" authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole ankerana" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ankerana" status="sp. nov.">Pheidole ankerana</taxonomicName>
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<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
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<figureCitation captionStart="Figure 71" captionStartId="F71" captionText="Figure 71. Pheidole ankerana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0274003) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923171)." figureDoi="10.3897/zookeys.905.39592.figure71" httpUri="https://binary.pensoft.net/fig/373118" pageId="0" pageNumber="1">Figs 71A-F</figureCitation>
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<figureCitation captionStart="Figure 84" captionStartId="F84" captionText="Figure 84. Hypostomal teeth. Pheidole aelloea sp. nov. (A). P. ala sp. nov. (B). P. andapa sp. nov. (C). P. ankerana sp. nov. (D). P. annemariae Forel (E). P. avaratra sp. nov. (F). P. bemarahaensis sp. nov. (G). P. bemarivoensis sp. nov. (H). P. binara sp. nov. (I). P. boribora sp. nov. (J). P. brevipilosa sp. nov. (K). P. curvistriata sp. nov. (L). P. diakritos sp. nov. (M). P. ehazoara sp. nov. (N). P. ensifera Forel (O). P. ferruginea sp. nov. (P). P. fisaka sp. nov. (R). P. fitarata sp. nov. (S). P. glabra sp. nov. (T). P. goavana sp. nov. (U). P. lamperos sp. nov. (W). P. longipilosa sp. nov. (X). P. longispinosa Forel (Y). P. lutea sp. nov. (Z)." figureDoi="10.3897/zookeys.905.39592.figure84" httpUri="https://binary.pensoft.net/fig/373131" pageId="0" pageNumber="1">, 84D</figureCitation>
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<figureCitation captionStart="Figure 86" captionStartId="F86" captionText="Figure 86. Distribution. Pheidole aelloea sp. nov. (A). P. ala sp. nov. (B). P. andapa sp. nov. (C). P. ankerana sp. nov. (D). P. annemariae Forel (E). P. avaratra sp. nov. (F). P. bemarahaensis sp. nov. (G). P. bemarivoensis sp. nov. (H). P. binara sp. nov. (I). P. boribora sp. nov. (J). P. brevipilosa sp. nov. (K). P. curvistriata sp. nov. (L). P. diakritos sp. nov. (M). P. ehazoara sp. nov. (N). P. ensifera Forel (O). P. ferruginea sp. nov. (P). P. fisaka sp. nov. (R). P. fitarata sp. nov. (S). P. glabra sp. nov. (T). P. goavana sp. nov. (U)." figureDoi="10.3897/zookeys.905.39592.figure86" httpUri="https://binary.pensoft.net/fig/373133" pageId="0" pageNumber="1">, 86D</figureCitation>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="type material">
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<paragraph pageId="0" pageNumber="1">Type material.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
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.
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</emphasis>
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Madagascar. •1 major worker; Toamasina; Ankerana;
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<geoCoordinate degrees="18.4104" direction="south" orientation="latitude" precision="5" value="-18.4104">-18.4104</geoCoordinate>
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,
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<geoCoordinate degrees="48.8189" direction="east" orientation="longitude" precision="5" value="48.8189">48.8189</geoCoordinate>
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; alt. 855 m; 27 Jan 2012; B.L. Fisher et al. leg.; BLF28148, CASENT0923171 (CASC).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
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.
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</emphasis>
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Madagascar. •2w., 1q.; same data as for holotype; CASENT0274002, CASENT0274003, CASENT0872185 (CASC).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="materials_examined">
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<paragraph pageId="0" pageNumber="1">Other material.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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Madagascar. -
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<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Toamasina</emphasis>
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: •3w.; Ankerana;
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<geoCoordinate degrees="18.4061" direction="south" orientation="latitude" precision="5" value="-18.4061">-18.4061</geoCoordinate>
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,
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<geoCoordinate degrees="48.82029" direction="east" orientation="longitude" precision="1" value="48.82029">48.82029</geoCoordinate>
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; alt. 725 m; 20 Jan 2012; B.L. Fisher et al. leg.; CASENT0274747, CASENT0274762 (CASC). •3w., 1m.; Ankerana;
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<geoCoordinate degrees="18.40829" direction="south" orientation="latitude" precision="1" value="-18.40829">-18.40829</geoCoordinate>
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,
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<geoCoordinate degrees="48.82107" direction="east" orientation="longitude" precision="1" value="48.82107">48.82107</geoCoordinate>
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; alt. 750 m; 21 Jan 2012; B.L. Fisher et al. leg.; CASENT0275260, CASENT0275261 (CASC). •2w.; Ankerana;
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<geoCoordinate degrees="18.4104" direction="south" orientation="latitude" precision="5" value="-18.4104">-18.4104</geoCoordinate>
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,
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<geoCoordinate degrees="48.8189" direction="east" orientation="longitude" precision="5" value="48.8189">48.8189</geoCoordinate>
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; alt. 855 m; 22 Jan 2012; B.L. Fisher et al. leg.; CASENT0274408, CASENT0275713 (CASC). •2w.; Ankerana;
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<geoCoordinate degrees="18.40062" direction="south" orientation="latitude" precision="1" value="-18.40062">-18.40062</geoCoordinate>
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,
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<geoCoordinate degrees="48.81311" direction="east" orientation="longitude" precision="1" value="48.81311">48.81311</geoCoordinate>
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; alt. 865 m; 17 Jan 2012; B.L. Fisher et al. leg.; CASENT0274909 (CASC). •1w., 1s.; Parc National de Zahamena, Besaky River;
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<geoCoordinate degrees="17.75244" direction="south" orientation="latitude" precision="1" value="-17.75244">-17.75244</geoCoordinate>
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,
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<geoCoordinate degrees="48.85321" direction="east" orientation="longitude" precision="1" value="48.85321">48.85321</geoCoordinate>
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; alt. 760 m; 22 Feb 2009; B.L. Fisher et al. leg.; CASENT0149769 (CASC).
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="diagnosis">
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<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
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.
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</emphasis>
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Head in full-face view rectangular, with lateral sides relatively straight, only posteriormost part slightly convex; sides of the head with dense, long, suberect pilosity; occipital lobes and genae shiny, with thick, sparse, irregular rugae; frons with sparse, thick, and longitudinal rugae, interspaces smooth; inner hypostomal teeth distinct, small, and low, closely spaced, lobe-like, with rounded apex and wide base; outer hypostomal teeth bigger and wider than inner hypostomal teeth, triangular; inner and outer teeth closely spaced and connected by concavity; propodeal spines moderately long, triangular, with acute apex; gaster smooth; body reddish brown.
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
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.
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</emphasis>
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Head foveolate with additional thin, sparse rugae on the whole surface; promesonotum low, short, slightly convex, with relatively steep posterior declivity; propodeal spines minute, triangular; mesosoma foveolate with additional thin, sparse rugae, katepisternum smooth.
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="1" type="description">
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<paragraph pageId="0" pageNumber="1">Description.</paragraph>
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Major workers.</emphasis>
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Measurements (
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<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
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= 5): HL: 1.42-1.52 (1.48); HW: 1.34-1.43 (1.38); SL: 0.61-0.65 (0.63); EL: 0.16-0.2 (0.18); WL: 1.07-1.16 (1.12); PSL: 0.19-0.21 (0.2); MTL: 0.65-0.68 (0.66); PNW: 0.62-0.67 (0.64); PTW: 0.15-0.17 (0.16); PPW: 0.47-0.51 (0.49); CI: 92.1-94.1 (93.3); SI: 44.7-46.2 (45.5); PSLI: 12.1-14.6 (13.4); PPI: 28.9-32.7 (31.6); PNI: 45.7-47.5 (46.6); MTI: 47.4-48.8 (48.3).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Head</emphasis>
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.
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</emphasis>
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In full-face view rectangular, with lateral sides relatively straight, only posteriormost part slightly convex (Fig.
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<figureCitation captionStart="Figure 71" captionStartId="F71" captionText="Figure 71. Pheidole ankerana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0274003) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923171)." figureDoi="10.3897/zookeys.905.39592.figure71" httpUri="https://binary.pensoft.net/fig/373118" pageId="0" pageNumber="1">71B</figureCitation>
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). In lateral view sub-rectangular; ventral and dorsal faces finely convex; dorsal face finely depressed posteriorly, forming shallow transverse depression between frons and occipital lobes; inner hypostomal teeth invisible. Sides of the head with dense, long, suberect pilosity; whole head with dense, moderately long, suberect to erect pilosity. Antennal scrobes indistinct and not delimited by carinulae; scrobe surface shiny, with dense, thick, and irregular rugoreticulation. Occipital lobes and genae shiny, with thick, sparse, irregular rugae; frons with sparse, thick, and longitudinal rugae, interspaces smooth; malar area with dense, thick, and longitudinal rugae, interspaces smooth to finely rugulose. Centre of clypeus smooth and shiny, lateral sides with longitudinal rugae; median notch present, narrow and shallow to moderate; median longitudinal carina present; lateral longitudinal carinae absent. Scape, when laid back, slightly exceeding the midlength of head; pilosity decumbent to erect (Fig.
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<figureCitation captionStart="Figure 71" captionStartId="F71" captionText="Figure 71. Pheidole ankerana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0274003) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923171)." figureDoi="10.3897/zookeys.905.39592.figure71" httpUri="https://binary.pensoft.net/fig/373118" pageId="0" pageNumber="1">71B, D</figureCitation>
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). Inner hypostomal teeth distinct, small and low, closely spaced, lobe-like, with rounded apex and wide base; outer hypostomal teeth bigger and wider than inner hypostomal teeth, triangular; inner and outer teeth closely spaced and connected by concavity (Fig.
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<figureCitation captionStart="Figure 84" captionStartId="F84" captionText="Figure 84. Hypostomal teeth. Pheidole aelloea sp. nov. (A). P. ala sp. nov. (B). P. andapa sp. nov. (C). P. ankerana sp. nov. (D). P. annemariae Forel (E). P. avaratra sp. nov. (F). P. bemarahaensis sp. nov. (G). P. bemarivoensis sp. nov. (H). P. binara sp. nov. (I). P. boribora sp. nov. (J). P. brevipilosa sp. nov. (K). P. curvistriata sp. nov. (L). P. diakritos sp. nov. (M). P. ehazoara sp. nov. (N). P. ensifera Forel (O). P. ferruginea sp. nov. (P). P. fisaka sp. nov. (R). P. fitarata sp. nov. (S). P. glabra sp. nov. (T). P. goavana sp. nov. (U). P. lamperos sp. nov. (W). P. longipilosa sp. nov. (X). P. longispinosa Forel (Y). P. lutea sp. nov. (Z)." figureDoi="10.3897/zookeys.905.39592.figure84" httpUri="https://binary.pensoft.net/fig/373131" pageId="0" pageNumber="1">84D</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
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.
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</emphasis>
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In lateral view, promesonotum short, angular, and low, posterior mesonotum relatively steep, with indistinct tubercle-like projection; promesonotal groove absent; metanotal groove absent; propodeal spines moderately long, triangular, with acute apex; humeral area laterally weakly produced (Fig.
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<figureCitation captionStart="Figure 71" captionStartId="F71" captionText="Figure 71. Pheidole ankerana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0274003) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923171)." figureDoi="10.3897/zookeys.905.39592.figure71" httpUri="https://binary.pensoft.net/fig/373118" pageId="0" pageNumber="1">71D</figureCitation>
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). Surface shiny, with fine to thin, dense rugoreticulation, sculpture weakening on dorsum, katepisternum smooth. Pilosity moderately sparse, long, and erect (Fig.
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<figureCitation captionStart="Figure 71" captionStartId="F71" captionText="Figure 71. Pheidole ankerana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0274003) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923171)." figureDoi="10.3897/zookeys.905.39592.figure71" httpUri="https://binary.pensoft.net/fig/373118" pageId="0" pageNumber="1">71D, F</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
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.
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</emphasis>
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Shiny; peduncle shagreened, short, without horizontal lobes on its basal part; node smooth, relatively high and thin, triangular, with rounded apex, in rear view node straight; pilosity moderately sparse and erect (Fig.
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<figureCitation captionStart="Figure 71" captionStartId="F71" captionText="Figure 71. Pheidole ankerana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0274003) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923171)." figureDoi="10.3897/zookeys.905.39592.figure71" httpUri="https://binary.pensoft.net/fig/373118" pageId="0" pageNumber="1">71D, F</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
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.
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</emphasis>
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Finely shagreened, shiny; in dorsal view sides with moderately short, acute, and horn-like projections; pilosity long, moderately sparse and erect (Fig.
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<figureCitation captionStart="Figure 71" captionStartId="F71" captionText="Figure 71. Pheidole ankerana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0274003) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923171)." figureDoi="10.3897/zookeys.905.39592.figure71" httpUri="https://binary.pensoft.net/fig/373118" pageId="0" pageNumber="1">71D, F</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
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.
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</emphasis>
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Smooth; pilosity moderately dense, long, and erect (Fig.
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<figureCitation captionStart="Figure 71" captionStartId="F71" captionText="Figure 71. Pheidole ankerana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0274003) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923171)." figureDoi="10.3897/zookeys.905.39592.figure71" httpUri="https://binary.pensoft.net/fig/373118" pageId="0" pageNumber="1">71D, F</figureCitation>
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).
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<emphasis bold="true" pageId="0" pageNumber="1">
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<emphasis italics="true" pageId="0" pageNumber="1">Colour</emphasis>
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.
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</emphasis>
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Unicolourous, reddish brown; legs dark yellow (Fig.
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<figureCitation captionStart="Figure 71" captionStartId="F71" captionText="Figure 71. Pheidole ankerana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0274003) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923171)." figureDoi="10.3897/zookeys.905.39592.figure71" httpUri="https://binary.pensoft.net/fig/373118" pageId="0" pageNumber="1">71D, F</figureCitation>
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).
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</paragraph>
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<caption doi="10.3897/zookeys.905.39592.figure71" httpUri="https://binary.pensoft.net/fig/373118" pageId="0" pageNumber="1" start="Figure 71" startId="F71">
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<paragraph pageId="0" pageNumber="1">
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<emphasis bold="true" pageId="0" pageNumber="1">Figure 71.</emphasis>
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<taxonomicName authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole ankerana" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ankerana">
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<emphasis italics="true" pageId="0" pageNumber="1">Pheidole ankerana</emphasis>
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</taxonomicName>
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sp. nov., full-face view (
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<emphasis bold="true" pageId="0" pageNumber="1">A</emphasis>
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), profile (
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<emphasis bold="true" pageId="0" pageNumber="1">C</emphasis>
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), and dorsal view (
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<emphasis bold="true" pageId="0" pageNumber="1">E</emphasis>
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) of paratype minor worker (CASENT0274003) and full-face view (
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<emphasis bold="true" pageId="0" pageNumber="1">B</emphasis>
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), profile (
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<emphasis bold="true" pageId="0" pageNumber="1">D</emphasis>
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), and dorsal view (
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<emphasis bold="true" pageId="0" pageNumber="1">F</emphasis>
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|
) of holotype major worker (CASENT0923171).
|
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</paragraph>
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</caption>
|
|
<paragraph pageId="0" pageNumber="1">
|
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<emphasis bold="true" pageId="0" pageNumber="1">Minor workers.</emphasis>
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Measurements (
|
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<emphasis italics="true" pageId="0" pageNumber="1">N</emphasis>
|
|
= 5): HL: 0.61-0.69 (0.66); HW: 0.55-0.63 (0.59); SL: 0.58-0.64 (0.61); EL: 0.11-0.13 (0.12); WL: 0.75-0.85 (0.8); PSL: 0.09-0.11 (0.1); MTL: 0.48-0.55 (0.51); PNW: 0.37-0.43 (0.4); PTW: 0.06-0.08 (0.07); PPW: 0.13-0.2 (0.16); CI: 88.5-93.8 (90.3); SI: 99.2-105.1 (102.4); PSLI: 14.4-17.3 (15.5); PPI: 38.2-52.7 (45.5); PNI: 66.1-68.6 (67.6); MTI: 79.5-90.1 (86.9).
|
|
<emphasis bold="true" pageId="0" pageNumber="1">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Head</emphasis>
|
|
.
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|
</emphasis>
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|
Occipital margin straight or indistinctly concave; occipital carina indistinct, weakly developed (Fig.
|
|
<figureCitation captionStart="Figure 71" captionStartId="F71" captionText="Figure 71. Pheidole ankerana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0274003) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923171)." figureDoi="10.3897/zookeys.905.39592.figure71" httpUri="https://binary.pensoft.net/fig/373118" pageId="0" pageNumber="1">71A</figureCitation>
|
|
). Pilosity moderately dense, long, erect. Whole head foveolate with additional thin, sparse rugae on the whole surface. Clypeus shiny, with sparse, irregular rugae; median longitudinal carina absent; two lateral longitudinal carinae absent. Scape, when laid back, surpassing the posterior head margin by one-fifth of its length; pilosity erect (Fig.
|
|
<figureCitation captionStart="Figure 71" captionStartId="F71" captionText="Figure 71. Pheidole ankerana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0274003) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923171)." figureDoi="10.3897/zookeys.905.39592.figure71" httpUri="https://binary.pensoft.net/fig/373118" pageId="0" pageNumber="1">71A, C</figureCitation>
|
|
).
|
|
<emphasis bold="true" pageId="0" pageNumber="1">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Mesosoma</emphasis>
|
|
.
|
|
</emphasis>
|
|
In lateral view, promesonotum low, short, slightly convex, with relatively steep posterior declivity; promesonotal groove absent; metanotal groove absent; propodeal spines minute, triangular, apex acute (Fig.
|
|
<figureCitation captionStart="Figure 71" captionStartId="F71" captionText="Figure 71. Pheidole ankerana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0274003) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923171)." figureDoi="10.3897/zookeys.905.39592.figure71" httpUri="https://binary.pensoft.net/fig/373118" pageId="0" pageNumber="1">71C</figureCitation>
|
|
). Sculpture foveolate with additional thin, sparse rugae on the whole surface, katepisternum smooth. Pilosity sparse, long, and erect (Fig.
|
|
<figureCitation captionStart="Figure 71" captionStartId="F71" captionText="Figure 71. Pheidole ankerana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0274003) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923171)." figureDoi="10.3897/zookeys.905.39592.figure71" httpUri="https://binary.pensoft.net/fig/373118" pageId="0" pageNumber="1">71C, E</figureCitation>
|
|
).
|
|
<emphasis bold="true" pageId="0" pageNumber="1">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
|
.
|
|
</emphasis>
|
|
Peduncle short and thin; node moderately high, triangular and small; with few long, erect setae (Fig.
|
|
<figureCitation captionStart="Figure 71" captionStartId="F71" captionText="Figure 71. Pheidole ankerana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0274003) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923171)." figureDoi="10.3897/zookeys.905.39592.figure71" httpUri="https://binary.pensoft.net/fig/373118" pageId="0" pageNumber="1">71C, E</figureCitation>
|
|
).
|
|
<emphasis bold="true" pageId="0" pageNumber="1">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Postpetiole</emphasis>
|
|
.
|
|
</emphasis>
|
|
Short, low, and slightly convex; with few short, erect setae (Fig.
|
|
<figureCitation captionStart="Figure 71" captionStartId="F71" captionText="Figure 71. Pheidole ankerana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0274003) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923171)." figureDoi="10.3897/zookeys.905.39592.figure71" httpUri="https://binary.pensoft.net/fig/373118" pageId="0" pageNumber="1">71C, E</figureCitation>
|
|
).
|
|
<emphasis bold="true" pageId="0" pageNumber="1">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Petiole</emphasis>
|
|
.
|
|
</emphasis>
|
|
With moderately sparse, erect pilosity (Fig.
|
|
<figureCitation captionStart="Figure 71" captionStartId="F71" captionText="Figure 71. Pheidole ankerana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0274003) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923171)." figureDoi="10.3897/zookeys.905.39592.figure71" httpUri="https://binary.pensoft.net/fig/373118" pageId="0" pageNumber="1">71C, E</figureCitation>
|
|
).
|
|
<emphasis bold="true" pageId="0" pageNumber="1">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Colour</emphasis>
|
|
.
|
|
</emphasis>
|
|
Unicolourous, yellow (Fig.
|
|
<figureCitation captionStart="Figure 71" captionStartId="F71" captionText="Figure 71. Pheidole ankerana sp. nov., full-face view (A), profile (C), and dorsal view (E) of paratype minor worker (CASENT 0274003) and full-face view (B), profile (D), and dorsal view (F) of holotype major worker (CASENT 0923171)." figureDoi="10.3897/zookeys.905.39592.figure71" httpUri="https://binary.pensoft.net/fig/373118" pageId="0" pageNumber="1">71C, E</figureCitation>
|
|
).
|
|
</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="1" type="etymology">
|
|
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">From the type locality.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="1" type="biology_ecology">
|
|
<paragraph pageId="0" pageNumber="1">Biology.</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">The species was collected between 725-865 m in elevation, in rainforest. Nests were located in rotten logs.</paragraph>
|
|
</subSubSection>
|
|
<subSubSection pageId="0" pageNumber="1" type="comments">
|
|
<paragraph pageId="0" pageNumber="1">Comments.</paragraph>
|
|
<paragraph pageId="0" pageNumber="1">
|
|
<emphasis bold="true" pageId="0" pageNumber="1">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Major workers</emphasis>
|
|
.
|
|
</emphasis>
|
|
<taxonomicName authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole ankerana" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ankerana">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole ankerana</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. is most similar to
|
|
<taxonomicName lsidName="P. vatovavensis" pageId="0" pageNumber="1" rank="species" species="vatovavensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. vatovavensis</emphasis>
|
|
</taxonomicName>
|
|
. It differs from
|
|
<taxonomicName lsidName="P. vatovavensis" pageId="0" pageNumber="1" rank="species" species="vatovavensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. vatovavensis</emphasis>
|
|
</taxonomicName>
|
|
in denser and longer pilosity on the sides of head, smooth sculpture of interspaces on frons, sculpture of scrobes never foveolate, lower promesonotum and lobe-like shape of inner hypostomal teeth.
|
|
<emphasis bold="true" pageId="0" pageNumber="1">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Minor workers</emphasis>
|
|
.
|
|
</emphasis>
|
|
<taxonomicName authorityName="Salata & Fisher" authorityYear="2020" class="Insecta" family="Formicidae" genus="Pheidole" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pheidole ankerana" order="Hymenoptera" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="ankerana">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">Pheidole ankerana</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. differs from
|
|
<taxonomicName lsidName="P. vatovavensis" pageId="0" pageNumber="1" rank="species" species="vatovavensis">
|
|
<emphasis italics="true" pageId="0" pageNumber="1">P. vatovavensis</emphasis>
|
|
</taxonomicName>
|
|
sp. nov. by presence of additional thin, sparse rugae on the whole surface of head and mesosoma and minute size of propodeal spines.
|
|
</paragraph>
|
|
</subSubSection>
|
|
</treatment>
|
|
</document> |