417 lines
50 KiB
XML
417 lines
50 KiB
XML
<document ID-DOI="http://dx.doi.org/10.3897/zookeys.1109.83807" ID-GBIF-Dataset="178820d5-d720-47fd-a85e-1ac28fca6c90" ID-Pensoft-Pub="1313-2970-1109-67" ID-Pensoft-UUID="534F47F344D858A49331F110D3D21111" ID-ZooBank="9C8BF86D194A46EF9D49072D09BF9E48" ModsDocID="1313-2970-1109-67" checkinTime="1656747431702" checkinUser="pensoft" docAuthor="Suzuki, Yuya, Hiramatsu, Takehisa & Tatsuta, Haruki" docDate="2022" docId="ECB883B6802053219478AC0800388169" docLanguage="en" docName="ZooKeys 1109: 67-101" docOrigin="ZooKeys 1109" docPubDate="2022-07-01" docSource="http://dx.doi.org/10.3897/zookeys.1109.83807" docTitle="Zoma dibaiyin Miller, Griswold & Yin 2009" docType="treatment" docVersion="2" id="534F47F344D858A49331F110D3D21111" lastPageNumber="67" masterDocId="534F47F344D858A49331F110D3D21111" masterDocTitle="Two new species and a new genus of ray spiders (Araneae, Theridiosomatidae) from the Ryukyu Islands, southwest Japan, with notes on their natural history" masterLastPageNumber="101" masterPageNumber="67" pageNumber="67" updateTime="1656748190795" updateUser="ExternalLinkService">
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<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
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<mods:titleInfo>
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<mods:title>Two new species and a new genus of ray spiders (Araneae, Theridiosomatidae) from the Ryukyu Islands, southwest Japan, with notes on their natural history</mods:title>
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</mods:titleInfo>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Suzuki, Yuya</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-6523-9272</mods:nameIdentifier>
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<mods:affiliation>The United Graduate School of Agricultural Sciences, Kagoshima University, 1 - 21 - 24, Korimoto, Kagoshima-shi, Kagoshima, 890 - 0065, Japan & Graduate School of Systems Life Sciences, Kyushu University, 744 Motooka, Nishi-ku, Fukuoka, 819 - 0395 Japan</mods:affiliation>
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<mods:nameIdentifier type="email">sasaganiya1206@gmail.com</mods:nameIdentifier>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Hiramatsu, Takehisa</mods:namePart>
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<mods:affiliation>Fregrance-Uwado 203, Uwado, Kawagoe-shi, Saitama, 350 - 0816, Japan</mods:affiliation>
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</mods:name>
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<mods:name type="personal">
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<mods:role>
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<mods:roleTerm>Author</mods:roleTerm>
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</mods:role>
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<mods:namePart>Tatsuta, Haruki</mods:namePart>
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<mods:nameIdentifier type="ORCID">https://orcid.org/0000-0001-8847-8874</mods:nameIdentifier>
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<mods:affiliation>Graduate School of Systems Life Sciences, Kyushu University, 744 Motooka, Nishi-ku, Fukuoka, 819 - 0395 Japan</mods:affiliation>
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</mods:name>
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<mods:typeOfResource>text</mods:typeOfResource>
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<mods:title>ZooKeys</mods:title>
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<mods:part>
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<mods:date>2022</mods:date>
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<mods:detail type="pubDate">
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<mods:number>2022-07-01</mods:number>
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</mods:detail>
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<mods:detail type="volume">
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<mods:number>1109</mods:number>
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</mods:detail>
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<mods:extent unit="page">
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<mods:start>67</mods:start>
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<mods:end>101</mods:end>
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</mods:extent>
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</mods:part>
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</mods:relatedItem>
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<mods:location>
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<mods:url>http://dx.doi.org/10.3897/zookeys.1109.83807</mods:url>
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</mods:location>
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<mods:classification>journal article</mods:classification>
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<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.1109.83807</mods:identifier>
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<mods:identifier type="Pensoft-Pub">1313-2970-1109-67</mods:identifier>
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<treatment LSID="urn:lsid:plazi:treatment:ECB883B6802053219478AC0800388169" httpUri="http://treatment.plazi.org/id/ECB883B6802053219478AC0800388169" lastPageNumber="67" pageId="0" pageNumber="67">
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<subSubSection pageId="0" pageNumber="67" type="nomenclature">
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<paragraph pageId="0" pageNumber="67">
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<taxonomicName LSID="ECB883B6-8020-5321-9478-AC0800388169" authority="Miller, Griswold & Yin, 2009" authorityName="Miller, Griswold & Yin" authorityYear="2009" class="Arachnida" family="Theridiosomatidae" genus="Zoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zoma dibaiyin" order="Araneae" pageId="0" pageNumber="67" phylum="Arthropoda" rank="species" species="dibaiyin">Zoma dibaiyin Miller, Griswold & Yin, 2009</taxonomicName>
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</paragraph>
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</subSubSection>
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<subSubSection pageId="0" pageNumber="67" type="description">
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<paragraph pageId="0" pageNumber="67">
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<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Zoma dibaiyin Miller, Griswold & Yin, 2009, male habitus and genitalia (NSMT-Ar 21720 A-H) and female habitus and genitalia (NSMT-Ar 21721 I-K) A habitus, dorsal view B habitus, lateral view C palp, retrolateral view D palp, ventral view E palp, posterior-ventral view F palp, prolateral view G embolic division, prolateral view H embolic division, posterior-ventral view I habitus, dorsal view J epigyne, ventral view K vulva, dorsal view. Abbreviations: C conductor CD copulatory ducts E embolus EA embolic apophysis FD fertilization ducts MA median apophysis PC paracymbium PCP posterior conductor projection RCP retrolateral conductor projection S spermatheca ST subtegulum T tegulum. Arrows in E, F, H indicate a cornered margin of posterior membrane of conductor. Arrow in J indicates a sclerotized median pit. Scale bars: 1.0 mm (A, B, I); 0.1 mm (C-H, J, K)." figureDoi="10.3897/zookeys.1109.83807.figure4" httpUri="https://binary.pensoft.net/fig/708570" pageId="0" pageNumber="67">Figs 4</figureCitation>
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||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Zoma dibaiyin Miller, Griswold & Yin, 2009, male genitalia (NSMT-Ar 21720) A ventral view B posterior-ventral view C prolateral view D conductor and embolic division, retrolateral view E conductor and embolic division, posterior-ventral view. Abbreviations: C conductor CD copulatory ducts E embolus EA embolic apophysis MA median apophysis PC paracymbium PCP posterior conductor projection RCP retrolateral conductor projection ST subtegulum T tegulum. Arrows in B-E indicate a cornered margin of posterior membrane of conductor. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1109.83807.figure5" httpUri="https://binary.pensoft.net/fig/708571" pageId="0" pageNumber="67">, 5</figureCitation>
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<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Distribution of theridiosomatid species in the Ryukyu Islands, Japan." figureDoi="10.3897/zookeys.1109.83807.figure11" httpUri="https://binary.pensoft.net/fig/708577" pageId="0" pageNumber="67">, 11</figureCitation>
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<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Habitats of theridiosomatid species in Ryukyu Islands A streamside in dim forest at Iriomote Island B forest floor of secondary forest at Kume Island C grassland at Kume Island D grassland at Yonaguni Island E crevices on limestone rocky wall at Iriomote Island." figureDoi="10.3897/zookeys.1109.83807.figure12" httpUri="https://binary.pensoft.net/fig/708578" pageId="0" pageNumber="67">, 12B</figureCitation>
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<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Web of Theridiosoma and Zoma spiders at Ryukyu Islands, Japan A-C web of Theridiosoma nigrivirgatum sp. nov. D-E web of Theridiosoma alboannulatum F-H web of Zoma dibaiyin. Abbreviations: RA radial anastomosis SS sticky spirals TL tension line." figureDoi="10.3897/zookeys.1109.83807.figure13" httpUri="https://binary.pensoft.net/fig/708579" pageId="0" pageNumber="67">, 13F-H</figureCitation>
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</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="0" pageNumber="67" type="reference_group">
|
||
<paragraph pageId="0" pageNumber="67">
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||
<taxonomicName authorityName="Miller, Griswold & Yin" authorityYear="2009" class="Arachnida" family="Theridiosomatidae" genus="Zoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zoma dibaiyin" order="Araneae" pageId="0" pageNumber="67" phylum="Arthropoda" rank="species" species="dibaiyin">Zoma dibaiyin</taxonomicName>
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Miller, Griswold & Yin, 2009: 27, figs 10A-F, 11A-B, 13A-D (holotype male and paratypes from China; not examined);
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<bibRefCitation author="Ono, H" journalOrPublisher="Tokai University Press, Kanagawa" pageId="0" pageNumber="67" refId="B15" refString="Ono, H, Ogata, K, 2018. Spiders of Japan: their natural history and diversity. Tokai University Press, Kanagawa" title="Spiders of Japan: their natural history and diversity." year="2018">Ono and Ogata 2018</bibRefCitation>
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: 120, 504, 505;
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<bibRefCitation author="Suzuki, Y" journalOrPublisher="Kishidaia" pageId="0" pageNumber="67" pagination="230 - 236" refId="B27" refString="Suzuki, Y, Serita, R, 2021. . Kishidaia 118: 230 - 236" title="." volume="118" year="2021">Suzuki and Serita 2021</bibRefCitation>
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a: 231, figs 1-3.
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</paragraph>
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||
</subSubSection>
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||
<subSubSection pageId="0" pageNumber="67" type="materials_examined">
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||
<paragraph pageId="0" pageNumber="67">Material examined.</paragraph>
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<paragraph pageId="0" pageNumber="67">
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||
<materialsCitation collectingDate="2021-07-04" collectorName="Y. Suzuki" country="Japan" county="Oshima District" elevation="354" latitude="28.209362" location="Katsuura" longLatPrecision="1" longitude="129.33167" municipality="Setouchi Town" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Kagoshima Prefecture">
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||
<emphasis bold="true" pageId="0" pageNumber="67">
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||
<collectingCountry name="Japan">Japan</collectingCountry>
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, Amami Is. (
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<collectingRegion country="Japan" name="Kagoshima">Kagoshima Prefecture</collectingRegion>
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||
)
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||
</emphasis>
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||
:
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<specimenCount type="male">1♂</specimenCount>
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<specimenCount type="female">1 ♀</specimenCount>
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(NSMT-Ar. 21720, 21721),
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<collectingCounty>
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||
<normalizedToken originalValue="Ôshima">Oshima</normalizedToken>
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District
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</collectingCounty>
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||
,
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||
<collectingMunicipality>Setouchi Town</collectingMunicipality>
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||
,
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||
<location LSID="urn:lsid:plazi:treatment:ECB883B6802053219478AC0800388169:1E5278D606399CE1183C086E6DDCE518" country="Japan" county="Oshima District" latitude="28.209362" longLatPrecision="1" longitude="129.33167" municipality="Setouchi Town" name="Katsuura" stateProvince="Kagoshima Prefecture">Katsuura</location>
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||
(
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||
<geoCoordinate degrees="28" direction="north" minutes="12" orientation="latitude" precision="1" seconds="33.7" value="28.209362">28°12'33.7"N</geoCoordinate>
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||
,
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||
<geoCoordinate degrees="129" direction="east" minutes="19" orientation="longitude" precision="1" seconds="54.0" value="129.33167">129°19'54.0"E</geoCoordinate>
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||
, alt.
|
||
<quantity metricMagnitude="2" metricUnit="m" metricValue="3.54" unit="m" value="354.0">
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||
<elevation metricMagnitude="2" metricUnit="m" metricValue="3.54" unit="m" value="354.0">354 m</elevation>
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||
</quantity>
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||
),
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||
<collectingDate value="2021-07-04">4 Jul. 2021</collectingDate>
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||
,
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||
<collectorName>Y. Suzuki</collectorName>
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||
leg.
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||
</materialsCitation>
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||
;
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||
<materialsCitation collectingDate="2021-07-04" collectorName="Y. Suzuki" country="Japan" elevation="111" latitude="28.220945" location="Amurogama" longLatPrecision="1" longitude="129.31648" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Amami Is.">
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||
<specimenCount type="male">1 ♂</specimenCount>
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||
<specimenCount type="female">1 ♀</specimenCount>
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||
,
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||
<location LSID="urn:lsid:plazi:treatment:ECB883B6802053219478AC0800388169:B7E7B14FA04FD74B3960BF9F295C2E4F" country="Japan" latitude="28.220945" longLatPrecision="1" longitude="129.31648" name="Amurogama" stateProvince="Amami Is.">Amurogama</location>
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||
(
|
||
<geoCoordinate degrees="28" direction="north" minutes="13" orientation="latitude" precision="1" seconds="15.4" value="28.220945">28°13'15.4"N</geoCoordinate>
|
||
,
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||
<geoCoordinate degrees="129" direction="east" minutes="18" orientation="longitude" precision="1" seconds="59.3" value="129.31648">129°18'59.3"E</geoCoordinate>
|
||
, alt.
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||
<quantity metricMagnitude="2" metricUnit="m" metricValue="1.1099999999999999" unit="m" value="111.0">
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||
<elevation metricMagnitude="2" metricUnit="m" metricValue="1.1099999999999999" unit="m" value="111.0">111 m</elevation>
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||
</quantity>
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||
),
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<collectingDate value="2021-07-04">4 Jul. 2021</collectingDate>
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||
,
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||
<collectorName>Y. Suzuki</collectorName>
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||
leg.
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||
</materialsCitation>
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||
;
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||
<materialsCitation collectingDate="2021-05-06" collectorName="Y. Suzuki" country="Japan" county="Amami City" elevation="141" latitude="28.263527" location="Nishinakama" longLatPrecision="1" longitude="129.41917" municipality="Sumiyo Town" specimenCount="1" specimenCount-male="1" stateProvince="Amami Is.">
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||
<specimenCount type="male">1 ♂</specimenCount>
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||
,
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||
<collectingCounty>Amami City</collectingCounty>
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||
,
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||
<collectingMunicipality>Sumiyo Town</collectingMunicipality>
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||
,
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||
<location LSID="urn:lsid:plazi:treatment:ECB883B6802053219478AC0800388169:8675D922B743C678B590A7C35E9CF299" country="Japan" county="Amami City" latitude="28.263527" longLatPrecision="1" longitude="129.41917" municipality="Sumiyo Town" name="Nishinakama" stateProvince="Amami Is.">Nishinakama</location>
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||
,
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||
<location LSID="urn:lsid:plazi:treatment:ECB883B6802053219478AC0800388169:421466A17CA0AD355044872CD82F79A3" country="Japan" county="Amami City" latitude="28.263527" longLatPrecision="1" longitude="129.41917" municipality="Sumiyo Town" name="Santarou-toge Pass" stateProvince="Amami Is.">Santarou-toge Pass</location>
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||
(
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||
<geoCoordinate degrees="28" direction="north" minutes="15" orientation="latitude" precision="1" seconds="48.7" value="28.263527">28°15'48.7"N</geoCoordinate>
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||
,
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||
<geoCoordinate degrees="129" direction="east" minutes="25" orientation="longitude" precision="1" seconds="09.0" value="129.41917">129°25'09.0"E</geoCoordinate>
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||
, alt.
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||
<quantity metricMagnitude="2" metricUnit="m" metricValue="1.41" unit="m" value="141.0">
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||
<elevation metricMagnitude="2" metricUnit="m" metricValue="1.41" unit="m" value="141.0">141 m</elevation>
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||
</quantity>
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||
),
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<collectingDate value="2021-05-06">6 May 2021</collectingDate>
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||
,
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||
<collectorName>Y. Suzuki</collectorName>
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||
leg.
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||
</materialsCitation>
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||
;
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||
<materialsCitation collectingDate="2021-04-14" collectorName="Y. Suzuki" country="Japan" county="Kunigami District" elevation="128" latitude="26.680471" location="Nerome" longLatPrecision="1" longitude="128.13364" municipality="Ogimi Village" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Okinawa">
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||
<emphasis bold="true" pageId="0" pageNumber="67">
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||
<collectingRegion country="Japan" name="Okinawa">Okinawa</collectingRegion>
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Is. (
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<collectingRegion country="Japan" name="Okinawa">Okinawa Prefecture</collectingRegion>
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||
)
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</emphasis>
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||
:
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||
<specimenCount type="male">1 ♂</specimenCount>
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||
<specimenCount type="female">1 ♀</specimenCount>
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||
,
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||
<collectingCounty>Kunigami District</collectingCounty>
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||
,
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||
<collectingMunicipality>
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||
<normalizedToken originalValue="Ôgimi">Ogimi</normalizedToken>
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||
Village
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||
</collectingMunicipality>
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||
,
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||
<location LSID="urn:lsid:plazi:treatment:ECB883B6802053219478AC0800388169:09929AD5B4841721B811B8093B6094F4" country="Japan" county="Kunigami District" latitude="26.680471" longLatPrecision="1" longitude="128.13364" municipality="Ogimi Village" name="Nerome" stateProvince="Okinawa">Nerome</location>
|
||
(
|
||
<geoCoordinate degrees="26" direction="north" minutes="40" orientation="latitude" precision="1" seconds="49.7" value="26.680471">26°40'49.7"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="128" direction="east" minutes="08" orientation="longitude" precision="1" seconds="01.1" value="128.13364">128°08'01.1"E</geoCoordinate>
|
||
, alt.
|
||
<quantity metricMagnitude="2" metricUnit="m" metricValue="1.28" unit="m" value="128.0">
|
||
<elevation metricMagnitude="2" metricUnit="m" metricValue="1.28" unit="m" value="128.0">128 m</elevation>
|
||
</quantity>
|
||
),
|
||
<collectingDate value="2021-04-14">14 Apr. 2021</collectingDate>
|
||
,
|
||
<collectorName>Y. Suzuki</collectorName>
|
||
leg.
|
||
</materialsCitation>
|
||
;
|
||
<materialsCitation collectingDate="2021-09-10" collectorName="Y. Suzuki" country="Japan" county="Shimajiri District" elevation="149" latitude="26.361917" location="Uezu" longLatPrecision="1" longitude="126.7597" municipality="Kumejima Town" specimenCount="2" specimenCount-female="1" specimenCount-male="1" stateProvince="Okinawa Prefecture">
|
||
<emphasis bold="true" pageId="0" pageNumber="67">
|
||
Kume Is. (
|
||
<collectingRegion country="Japan" name="Okinawa">Okinawa Prefecture</collectingRegion>
|
||
)
|
||
</emphasis>
|
||
:
|
||
<specimenCount type="male">1 ♂</specimenCount>
|
||
<specimenCount type="female">1 ♀</specimenCount>
|
||
,
|
||
<collectingCounty>Shimajiri District</collectingCounty>
|
||
,
|
||
<collectingMunicipality>Kumejima Town</collectingMunicipality>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:ECB883B6802053219478AC0800388169:F188BE2C8DE8B1A9482097C5F10525B9" country="Japan" county="Shimajiri District" latitude="26.361917" longLatPrecision="1" longitude="126.7597" municipality="Kumejima Town" name="Uezu" stateProvince="Okinawa Prefecture">Uezu</location>
|
||
,
|
||
<location LSID="urn:lsid:plazi:treatment:ECB883B6802053219478AC0800388169:736C8895DBC61629AFBE4D942EF5B1CD" country="Japan" county="Shimajiri District" latitude="26.361917" longLatPrecision="1" longitude="126.7597" municipality="Kumejima Town" name="Mt. Daruma-yama" stateProvince="Okinawa Prefecture">Mt. Daruma-yama</location>
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||
(
|
||
<geoCoordinate degrees="26" direction="north" minutes="21" orientation="latitude" precision="1" seconds="42.9" value="26.361917">26°21'42.9"N</geoCoordinate>
|
||
,
|
||
<geoCoordinate degrees="126" direction="east" minutes="45" orientation="longitude" precision="1" seconds="34.9" value="126.7597">126°45'34.9"E</geoCoordinate>
|
||
, alt.
|
||
<quantity metricMagnitude="2" metricUnit="m" metricValue="1.49" unit="m" value="149.0">
|
||
<elevation metricMagnitude="2" metricUnit="m" metricValue="1.49" unit="m" value="149.0">149 m</elevation>
|
||
</quantity>
|
||
),
|
||
<collectingDate value="2021-09-10">10 Sep. 2021</collectingDate>
|
||
,
|
||
<collectorName>Y. Suzuki</collectorName>
|
||
leg.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="67" type="diagnosis">
|
||
<paragraph pageId="0" pageNumber="67">Diagnosis.</paragraph>
|
||
<paragraph pageId="0" pageNumber="67">
|
||
Males of this species can be distinguished from congeners by the embolic apophysis with a curved tip running along the sclerotized surface of the ventral tegulum, and females by a nearly transverse posterior margin of the epigynal plate (more convex in
|
||
<taxonomicName genus="Z." lsidName="Z. zoma" pageId="0" pageNumber="67" rank="species" species="zoma">
|
||
<emphasis italics="true" pageId="0" pageNumber="67">Z. zoma</emphasis>
|
||
</taxonomicName>
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||
and rounded in
|
||
<taxonomicName genus="Z." lsidName="Z. taiwanica" pageId="0" pageNumber="67" rank="species" species="taiwanica">
|
||
<emphasis italics="true" pageId="0" pageNumber="67">Z. taiwanica</emphasis>
|
||
</taxonomicName>
|
||
) and lower position of the spermathecae (higher in
|
||
<taxonomicName genus="Z." lsidName="Z. taiwanica" pageId="0" pageNumber="67" rank="species" species="taiwanica">
|
||
<emphasis italics="true" pageId="0" pageNumber="67">Z. taiwanica</emphasis>
|
||
</taxonomicName>
|
||
) (
|
||
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.11.160" author="Miller, JA" journalOrPublisher="ZooKeys" pageId="0" pageNumber="67" pagination="9 - 195" refId="B13" refString="Miller, JA, Griswold, CE, Yin, CM, 2009. The symphytognathoid spiders of the Gaoligongshan, Yunnan, China (Araneae, Araneoidea): Systematics and diversity of micro-orbweavers. ZooKeys 11: 9 - 195, DOI: https://doi.org/10.3897/zookeys.11.160" title="The symphytognathoid spiders of the Gaoligongshan, Yunnan, China (Araneae, Araneoidea): Systematics and diversity of micro-orbweavers." url="https://doi.org/10.3897/zookeys.11.160" volume="11" year="2009">Miller et al. 2009</bibRefCitation>
|
||
;
|
||
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4927.2.2" author="Ballarin, F" journalOrPublisher="Zootaxa" pageId="0" pageNumber="67" pagination="197 - 208" refId="B2" refString="Ballarin, F, Yamasaki, T, Su, YC, 2021. A survey on poorly known rainforest litter-dwelling spiders of Orchid Island (Lanyu, Taiwan) with the description of a new species (Araneae: Linyphiidae, Tetrablemmidae, and Theridiosomatidae). Zootaxa 4927 (2): 197 - 208, DOI: https://doi.org/10.11646/zootaxa.4927.2.2" title="A survey on poorly known rainforest litter-dwelling spiders of Orchid Island (Lanyu, Taiwan) with the description of a new species (Araneae: Linyphiidae, Tetrablemmidae, and Theridiosomatidae)." url="https://doi.org/10.11646/zootaxa.4927.2.2" volume="4927" year="2021">Ballarin et al. 2021</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="67" type="description">
|
||
<paragraph pageId="0" pageNumber="67">Description.</paragraph>
|
||
<paragraph pageId="0" pageNumber="67">
|
||
<emphasis bold="true" pageId="0" pageNumber="67">Male</emphasis>
|
||
(NSMT-Ar 21720). Measurements. Body 1.62 long. Carapace 0.74 long, 0.68 wide, 0.60 high. Eye size and interdistances: AME 0.090, ALE 0.076, PME 0.093, PLE 0.065, AME-AME 0.015, AME-ALE 0.032, PME-PME 0.008, PLE-PLE 0.070. Leg length: leg I 0.80 + 0.23 + 0.57 + 0.47 + 0.28 = 2.35; leg II 0.50 + 0.21 + 0.46 + 0.31 + 0.28 = 1.76; leg III 0.40 + 0.19 + 0.23 + 0.24 + 0.24 = 1.30; leg IV 0.44 + 0.14 + 0.34 + 0.27 + 0.27 = 1.46. Abdomen 0.88 long, 0.95 wide, 1.01 high.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="67">Carapace oval, longer than wide (CaL/CaW 1.34). Chelicerae with three teeth on promargin. Abdomen oval, wider than long (AL/AW 0.93).</paragraph>
|
||
<paragraph pageId="0" pageNumber="67">
|
||
Coloration and markings (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Zoma dibaiyin Miller, Griswold & Yin, 2009, male habitus and genitalia (NSMT-Ar 21720 A-H) and female habitus and genitalia (NSMT-Ar 21721 I-K) A habitus, dorsal view B habitus, lateral view C palp, retrolateral view D palp, ventral view E palp, posterior-ventral view F palp, prolateral view G embolic division, prolateral view H embolic division, posterior-ventral view I habitus, dorsal view J epigyne, ventral view K vulva, dorsal view. Abbreviations: C conductor CD copulatory ducts E embolus EA embolic apophysis FD fertilization ducts MA median apophysis PC paracymbium PCP posterior conductor projection RCP retrolateral conductor projection S spermatheca ST subtegulum T tegulum. Arrows in E, F, H indicate a cornered margin of posterior membrane of conductor. Arrow in J indicates a sclerotized median pit. Scale bars: 1.0 mm (A, B, I); 0.1 mm (C-H, J, K)." figureDoi="10.3897/zookeys.1109.83807.figure4" httpUri="https://binary.pensoft.net/fig/708570" pageId="0" pageNumber="67">4A, B</figureCitation>
|
||
). Carapace, chelicerae, maxillae, labium, sternum, and legs yellowish brown. Eyes on the dark bases. Cephalic groove stained with dark spots. Legs lacking annulation. Abdomen dark brown encircled dorsolaterally with a whitish silver band.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="67">
|
||
Palp (Figs
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Zoma dibaiyin Miller, Griswold & Yin, 2009, male habitus and genitalia (NSMT-Ar 21720 A-H) and female habitus and genitalia (NSMT-Ar 21721 I-K) A habitus, dorsal view B habitus, lateral view C palp, retrolateral view D palp, ventral view E palp, posterior-ventral view F palp, prolateral view G embolic division, prolateral view H embolic division, posterior-ventral view I habitus, dorsal view J epigyne, ventral view K vulva, dorsal view. Abbreviations: C conductor CD copulatory ducts E embolus EA embolic apophysis FD fertilization ducts MA median apophysis PC paracymbium PCP posterior conductor projection RCP retrolateral conductor projection S spermatheca ST subtegulum T tegulum. Arrows in E, F, H indicate a cornered margin of posterior membrane of conductor. Arrow in J indicates a sclerotized median pit. Scale bars: 1.0 mm (A, B, I); 0.1 mm (C-H, J, K)." figureDoi="10.3897/zookeys.1109.83807.figure4" httpUri="https://binary.pensoft.net/fig/708570" pageId="0" pageNumber="67">4C-H</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Zoma dibaiyin Miller, Griswold & Yin, 2009, male genitalia (NSMT-Ar 21720) A ventral view B posterior-ventral view C prolateral view D conductor and embolic division, retrolateral view E conductor and embolic division, posterior-ventral view. Abbreviations: C conductor CD copulatory ducts E embolus EA embolic apophysis MA median apophysis PC paracymbium PCP posterior conductor projection RCP retrolateral conductor projection ST subtegulum T tegulum. Arrows in B-E indicate a cornered margin of posterior membrane of conductor. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1109.83807.figure5" httpUri="https://binary.pensoft.net/fig/708571" pageId="0" pageNumber="67">5</figureCitation>
|
||
). Paracymbium with sharp tip. Tegulum bulbous. Median apophysis weakly sclerotized, wider than long. Embolic division branched into a few bristle-like apophyses, embolus short and tubular, and a long and filiform embolic apophysis emerging beneath from a translucent conductor. Tip of embolic apophysis curved and running along sclerotized surface of the tegulum beneath the conductor. Conductor having two projections: posterior conductor projection strongly sclerotized and triangular with a blunt tip; retrolateral conductor projection strongly sclerotized and weakly curved anteriorly with a triangular posterior tip. Posterior margin of conductor with a sharp tip.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="67">
|
||
<emphasis bold="true" pageId="0" pageNumber="67">Female</emphasis>
|
||
(NSMT-Ar 21721). Measurements. Body 2.04 long. Carapace 0.82 long, 0.76 wide, 0.67 high. Eye size and interdistances: AME 0.089, ALE 0.082, PME 084, PLE 0.076, AME-AME 0.023, AME-ALE 0.034, PME-PME 0.009, PLE-PLE 0.076. Leg length: leg I 0.66 + 0.32 + 0.42 + 0.33 + 0.22 = 1.95; leg II 0.64 + 0.30 + 0.41 + 0.33 + 0.21 = 1.89; leg III 0.36 + 0.18 + 0.20 + 0.25 + 0.18 = 1.17; leg IV 0.53 + 0.21 + 0.34 + 0.27 + 0.22 = 1.57. Abdomen 1.21 long, 1.32 wide, 1.16 high.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="67">Carapace, mouthparts, and abdomen as in male (CaL/CaW 1.08; AL/AW 0.92).</paragraph>
|
||
<paragraph pageId="0" pageNumber="67">
|
||
Coloration and markings (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Zoma dibaiyin Miller, Griswold & Yin, 2009, male habitus and genitalia (NSMT-Ar 21720 A-H) and female habitus and genitalia (NSMT-Ar 21721 I-K) A habitus, dorsal view B habitus, lateral view C palp, retrolateral view D palp, ventral view E palp, posterior-ventral view F palp, prolateral view G embolic division, prolateral view H embolic division, posterior-ventral view I habitus, dorsal view J epigyne, ventral view K vulva, dorsal view. Abbreviations: C conductor CD copulatory ducts E embolus EA embolic apophysis FD fertilization ducts MA median apophysis PC paracymbium PCP posterior conductor projection RCP retrolateral conductor projection S spermatheca ST subtegulum T tegulum. Arrows in E, F, H indicate a cornered margin of posterior membrane of conductor. Arrow in J indicates a sclerotized median pit. Scale bars: 1.0 mm (A, B, I); 0.1 mm (C-H, J, K)." figureDoi="10.3897/zookeys.1109.83807.figure4" httpUri="https://binary.pensoft.net/fig/708570" pageId="0" pageNumber="67">4I</figureCitation>
|
||
) similar to male.
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="67">
|
||
Genitalia (Fig.
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Zoma dibaiyin Miller, Griswold & Yin, 2009, male habitus and genitalia (NSMT-Ar 21720 A-H) and female habitus and genitalia (NSMT-Ar 21721 I-K) A habitus, dorsal view B habitus, lateral view C palp, retrolateral view D palp, ventral view E palp, posterior-ventral view F palp, prolateral view G embolic division, prolateral view H embolic division, posterior-ventral view I habitus, dorsal view J epigyne, ventral view K vulva, dorsal view. Abbreviations: C conductor CD copulatory ducts E embolus EA embolic apophysis FD fertilization ducts MA median apophysis PC paracymbium PCP posterior conductor projection RCP retrolateral conductor projection S spermatheca ST subtegulum T tegulum. Arrows in E, F, H indicate a cornered margin of posterior membrane of conductor. Arrow in J indicates a sclerotized median pit. Scale bars: 1.0 mm (A, B, I); 0.1 mm (C-H, J, K)." figureDoi="10.3897/zookeys.1109.83807.figure4" httpUri="https://binary.pensoft.net/fig/708570" pageId="0" pageNumber="67">4J, K</figureCitation>
|
||
). Epigynal plate flat and wider than long with a sclerotized posterior margin and a median pit. Spermathecae touching each other, copulatory ducts wide at their openings, and the course of the ducts simple. See
|
||
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.11.160" author="Miller, JA" journalOrPublisher="ZooKeys" pageId="0" pageNumber="67" pagination="9 - 195" refId="B13" refString="Miller, JA, Griswold, CE, Yin, CM, 2009. The symphytognathoid spiders of the Gaoligongshan, Yunnan, China (Araneae, Araneoidea): Systematics and diversity of micro-orbweavers. ZooKeys 11: 9 - 195, DOI: https://doi.org/10.3897/zookeys.11.160" title="The symphytognathoid spiders of the Gaoligongshan, Yunnan, China (Araneae, Araneoidea): Systematics and diversity of micro-orbweavers." url="https://doi.org/10.3897/zookeys.11.160" volume="11" year="2009">Miller et al. (2009)</bibRefCitation>
|
||
for further details.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="67" type="remarks">
|
||
<paragraph pageId="0" pageNumber="67">Remarks.</paragraph>
|
||
<paragraph pageId="0" pageNumber="67">
|
||
A strongly sclerotized triangular projection with a rounded tip (Figs
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Zoma dibaiyin Miller, Griswold & Yin, 2009, male habitus and genitalia (NSMT-Ar 21720 A-H) and female habitus and genitalia (NSMT-Ar 21721 I-K) A habitus, dorsal view B habitus, lateral view C palp, retrolateral view D palp, ventral view E palp, posterior-ventral view F palp, prolateral view G embolic division, prolateral view H embolic division, posterior-ventral view I habitus, dorsal view J epigyne, ventral view K vulva, dorsal view. Abbreviations: C conductor CD copulatory ducts E embolus EA embolic apophysis FD fertilization ducts MA median apophysis PC paracymbium PCP posterior conductor projection RCP retrolateral conductor projection S spermatheca ST subtegulum T tegulum. Arrows in E, F, H indicate a cornered margin of posterior membrane of conductor. Arrow in J indicates a sclerotized median pit. Scale bars: 1.0 mm (A, B, I); 0.1 mm (C-H, J, K)." figureDoi="10.3897/zookeys.1109.83807.figure4" httpUri="https://binary.pensoft.net/fig/708570" pageId="0" pageNumber="67">4E</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Zoma dibaiyin Miller, Griswold & Yin, 2009, male genitalia (NSMT-Ar 21720) A ventral view B posterior-ventral view C prolateral view D conductor and embolic division, retrolateral view E conductor and embolic division, posterior-ventral view. Abbreviations: C conductor CD copulatory ducts E embolus EA embolic apophysis MA median apophysis PC paracymbium PCP posterior conductor projection RCP retrolateral conductor projection ST subtegulum T tegulum. Arrows in B-E indicate a cornered margin of posterior membrane of conductor. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1109.83807.figure5" httpUri="https://binary.pensoft.net/fig/708571" pageId="0" pageNumber="67">5B</figureCitation>
|
||
) and a cornered margin of the posterior membrane of the conductor (Figs
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Zoma dibaiyin Miller, Griswold & Yin, 2009, male habitus and genitalia (NSMT-Ar 21720 A-H) and female habitus and genitalia (NSMT-Ar 21721 I-K) A habitus, dorsal view B habitus, lateral view C palp, retrolateral view D palp, ventral view E palp, posterior-ventral view F palp, prolateral view G embolic division, prolateral view H embolic division, posterior-ventral view I habitus, dorsal view J epigyne, ventral view K vulva, dorsal view. Abbreviations: C conductor CD copulatory ducts E embolus EA embolic apophysis FD fertilization ducts MA median apophysis PC paracymbium PCP posterior conductor projection RCP retrolateral conductor projection S spermatheca ST subtegulum T tegulum. Arrows in E, F, H indicate a cornered margin of posterior membrane of conductor. Arrow in J indicates a sclerotized median pit. Scale bars: 1.0 mm (A, B, I); 0.1 mm (C-H, J, K)." figureDoi="10.3897/zookeys.1109.83807.figure4" httpUri="https://binary.pensoft.net/fig/708570" pageId="0" pageNumber="67">4E</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Zoma dibaiyin Miller, Griswold & Yin, 2009, male genitalia (NSMT-Ar 21720) A ventral view B posterior-ventral view C prolateral view D conductor and embolic division, retrolateral view E conductor and embolic division, posterior-ventral view. Abbreviations: C conductor CD copulatory ducts E embolus EA embolic apophysis MA median apophysis PC paracymbium PCP posterior conductor projection RCP retrolateral conductor projection ST subtegulum T tegulum. Arrows in B-E indicate a cornered margin of posterior membrane of conductor. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1109.83807.figure5" httpUri="https://binary.pensoft.net/fig/708571" pageId="0" pageNumber="67">5B</figureCitation>
|
||
, arrowed) were visible in the ventro-posterior view of the male palp. The triangular projection does not seem to be homologous to conductor projection in
|
||
<taxonomicName authorityName="O. Pickard-Cambridge" authorityYear="1879" class="Arachnida" family="Theridiosomatidae" genus="Theridiosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theridiosoma" order="Araneae" pageId="0" pageNumber="67" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="67">Theridiosoma</emphasis>
|
||
</taxonomicName>
|
||
, as the former protrudes from the posterior margin of the conductor, while the latter was positioned on the surface of the conductor. Herein, we define it as posterior conductor projection (PCP).
|
||
</paragraph>
|
||
<paragraph pageId="0" pageNumber="67">
|
||
The embolus on the male palp was not determined in
|
||
<taxonomicName genus="Z." lsidName="Z. dibaiyin" pageId="0" pageNumber="67" rank="species" species="dibaiyin">
|
||
<emphasis italics="true" pageId="0" pageNumber="67">Z. dibaiyin</emphasis>
|
||
</taxonomicName>
|
||
and
|
||
<taxonomicName genus="Z." lsidName="Z. fascia" pageId="0" pageNumber="67" rank="species" species="fascia">
|
||
<emphasis italics="true" pageId="0" pageNumber="67">Z. fascia</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.11.160" author="Miller, JA" journalOrPublisher="ZooKeys" pageId="0" pageNumber="67" pagination="9 - 195" refId="B13" refString="Miller, JA, Griswold, CE, Yin, CM, 2009. The symphytognathoid spiders of the Gaoligongshan, Yunnan, China (Araneae, Araneoidea): Systematics and diversity of micro-orbweavers. ZooKeys 11: 9 - 195, DOI: https://doi.org/10.3897/zookeys.11.160" title="The symphytognathoid spiders of the Gaoligongshan, Yunnan, China (Araneae, Araneoidea): Systematics and diversity of micro-orbweavers." url="https://doi.org/10.3897/zookeys.11.160" volume="11" year="2009">Miller et al. 2009</bibRefCitation>
|
||
;
|
||
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.255.3272" author="Zhao, QY" journalOrPublisher="ZooKeys" pageId="0" pageNumber="67" pagination="1 - 48" refId="B35" refString="Zhao, QY, Li, SQ, 2012. Eleven new species of theridiosomatid spiders from southern China (Araneae, Theridiosomatidae). ZooKeys 255: 1 - 48, DOI: https://doi.org/10.3897/zookeys.255.3272" title="Eleven new species of theridiosomatid spiders from southern China (Araneae, Theridiosomatidae)." url="https://doi.org/10.3897/zookeys.255.3272" volume="255" year="2012">Zhao and Li 2012</bibRefCitation>
|
||
). In
|
||
<taxonomicName genus="Z." lsidName="Z. taiwanica" pageId="0" pageNumber="67" rank="species" species="taiwanica">
|
||
<emphasis italics="true" pageId="0" pageNumber="67">Z. taiwanica</emphasis>
|
||
</taxonomicName>
|
||
, the embolus is described as a 'short and tubular
|
||
<normalizedToken originalValue="structure’">structure'</normalizedToken>
|
||
but lacks explanations in illustrations (
|
||
<bibRefCitation author="Zhang, JX" journalOrPublisher="Bulletin - British Arachnological Society" pageId="0" pageNumber="67" pagination="265 - 266" refId="B34" refString="Zhang, JX, Zhu, MS, Tso, IM, 2006. First record of the family Theridiosomatidae from Taiwan, with description of a new species (Arachnida: Araneae). Bulletin - British Arachnological Society 13: 265 - 266" title="First record of the family Theridiosomatidae from Taiwan, with description of a new species (Arachnida: Araneae)." volume="13" year="2006">Zhang et al. 2006</bibRefCitation>
|
||
).
|
||
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4927.2.2" author="Ballarin, F" journalOrPublisher="Zootaxa" pageId="0" pageNumber="67" pagination="197 - 208" refId="B2" refString="Ballarin, F, Yamasaki, T, Su, YC, 2021. A survey on poorly known rainforest litter-dwelling spiders of Orchid Island (Lanyu, Taiwan) with the description of a new species (Araneae: Linyphiidae, Tetrablemmidae, and Theridiosomatidae). Zootaxa 4927 (2): 197 - 208, DOI: https://doi.org/10.11646/zootaxa.4927.2.2" title="A survey on poorly known rainforest litter-dwelling spiders of Orchid Island (Lanyu, Taiwan) with the description of a new species (Araneae: Linyphiidae, Tetrablemmidae, and Theridiosomatidae)." url="https://doi.org/10.11646/zootaxa.4927.2.2" volume="4927" year="2021">Ballarin et al. (2021)</bibRefCitation>
|
||
determined the embolus as a strongly sclerotized, thin, stick-like apophysis located on the retrolateral side of the embolic division (
|
||
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4927.2.2" author="Ballarin, F" journalOrPublisher="Zootaxa" pageId="0" pageNumber="67" pagination="197 - 208" refId="B2" refString="Ballarin, F, Yamasaki, T, Su, YC, 2021. A survey on poorly known rainforest litter-dwelling spiders of Orchid Island (Lanyu, Taiwan) with the description of a new species (Araneae: Linyphiidae, Tetrablemmidae, and Theridiosomatidae). Zootaxa 4927 (2): 197 - 208, DOI: https://doi.org/10.11646/zootaxa.4927.2.2" title="A survey on poorly known rainforest litter-dwelling spiders of Orchid Island (Lanyu, Taiwan) with the description of a new species (Araneae: Linyphiidae, Tetrablemmidae, and Theridiosomatidae)." url="https://doi.org/10.11646/zootaxa.4927.2.2" volume="4927" year="2021">Ballarin et al. 2021</bibRefCitation>
|
||
). Considering the shape of
|
||
<taxonomicName authorityName="O. Pickard-Cambridge" authorityYear="1879" class="Arachnida" family="Theridiosomatidae" genus="Theridiosoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Theridiosoma" order="Araneae" pageId="0" pageNumber="67" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="67">Theridiosoma</emphasis>
|
||
</taxonomicName>
|
||
's embolus, which is short, tubular, and hidden under conductor (see
|
||
<bibRefCitation DOI="https://doi.org/10.5479/si.00810282.422" author="Coddington, JA" journalOrPublisher="Smithsonian Contributions to Zoology" pageId="0" pageNumber="67" pagination="1 - 96" refId="B4" refString="Coddington, JA, 1986a. The genera of the spider family Theridiosomatidae. Smithsonian Contributions to Zoology 422 (422): 1 - 96, DOI: https://doi.org/10.5479/si.00810282.422" title="The genera of the spider family Theridiosomatidae." url="https://doi.org/10.5479/si.00810282.422" volume="422" year="1986 a">Coddington 1986a</bibRefCitation>
|
||
: fig. 131), we suppose the
|
||
<normalizedToken originalValue="‘embolus’">'embolus'</normalizedToken>
|
||
of
|
||
<taxonomicName authorityName="Saaristo" authorityYear="1996" class="Arachnida" family="Theridiosomatidae" genus="Zoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zoma" order="Araneae" pageId="0" pageNumber="67" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="67">Zoma</emphasis>
|
||
</taxonomicName>
|
||
species determined in
|
||
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4927.2.2" author="Ballarin, F" journalOrPublisher="Zootaxa" pageId="0" pageNumber="67" pagination="197 - 208" refId="B2" refString="Ballarin, F, Yamasaki, T, Su, YC, 2021. A survey on poorly known rainforest litter-dwelling spiders of Orchid Island (Lanyu, Taiwan) with the description of a new species (Araneae: Linyphiidae, Tetrablemmidae, and Theridiosomatidae). Zootaxa 4927 (2): 197 - 208, DOI: https://doi.org/10.11646/zootaxa.4927.2.2" title="A survey on poorly known rainforest litter-dwelling spiders of Orchid Island (Lanyu, Taiwan) with the description of a new species (Araneae: Linyphiidae, Tetrablemmidae, and Theridiosomatidae)." url="https://doi.org/10.11646/zootaxa.4927.2.2" volume="4927" year="2021">Ballarin et al. (2021)</bibRefCitation>
|
||
is not an embolus. The true embolus of
|
||
<taxonomicName genus="Z." lsidName="Z. dibaiyin" pageId="0" pageNumber="67" rank="species" species="dibaiyin">
|
||
<emphasis italics="true" pageId="0" pageNumber="67">Z. dibaiyin</emphasis>
|
||
</taxonomicName>
|
||
is found beneath basal part of embolic apophyses (Fig.
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Zoma dibaiyin Miller, Griswold & Yin, 2009, male genitalia (NSMT-Ar 21720) A ventral view B posterior-ventral view C prolateral view D conductor and embolic division, retrolateral view E conductor and embolic division, posterior-ventral view. Abbreviations: C conductor CD copulatory ducts E embolus EA embolic apophysis MA median apophysis PC paracymbium PCP posterior conductor projection RCP retrolateral conductor projection ST subtegulum T tegulum. Arrows in B-E indicate a cornered margin of posterior membrane of conductor. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1109.83807.figure5" httpUri="https://binary.pensoft.net/fig/708571" pageId="0" pageNumber="67">5D</figureCitation>
|
||
). Hereafter we defined the sclerotized stick-like structure as a retrolateral conductor projection (RCP), and distinguishable from an embolus. The shape of RCP is useful as a taxonomic character for differentiating species within the genus
|
||
<taxonomicName authorityName="Saaristo" authorityYear="1996" class="Arachnida" family="Theridiosomatidae" genus="Zoma" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Zoma" order="Araneae" pageId="0" pageNumber="67" phylum="Arthropoda" rank="genus">
|
||
<emphasis italics="true" pageId="0" pageNumber="67">Zoma</emphasis>
|
||
</taxonomicName>
|
||
: S-shaped with pointed tip in
|
||
<taxonomicName genus="Z." lsidName="Z. fascia" pageId="0" pageNumber="67" rank="species" species="fascia">
|
||
<emphasis italics="true" pageId="0" pageNumber="67">Z. fascia</emphasis>
|
||
</taxonomicName>
|
||
, claviform with rounded tip in
|
||
<taxonomicName genus="Z." lsidName="Z. taiwanica" pageId="0" pageNumber="67" rank="species" species="taiwanica">
|
||
<emphasis italics="true" pageId="0" pageNumber="67">Z. taiwanica</emphasis>
|
||
</taxonomicName>
|
||
(
|
||
<bibRefCitation DOI="https://doi.org/10.3897/zookeys.255.3272" author="Zhao, QY" journalOrPublisher="ZooKeys" pageId="0" pageNumber="67" pagination="1 - 48" refId="B35" refString="Zhao, QY, Li, SQ, 2012. Eleven new species of theridiosomatid spiders from southern China (Araneae, Theridiosomatidae). ZooKeys 255: 1 - 48, DOI: https://doi.org/10.3897/zookeys.255.3272" title="Eleven new species of theridiosomatid spiders from southern China (Araneae, Theridiosomatidae)." url="https://doi.org/10.3897/zookeys.255.3272" volume="255" year="2012">Zhao and Li 2012</bibRefCitation>
|
||
: figs 28, 30;
|
||
<bibRefCitation author="Zhang, JX" journalOrPublisher="Bulletin - British Arachnological Society" pageId="0" pageNumber="67" pagination="265 - 266" refId="B34" refString="Zhang, JX, Zhu, MS, Tso, IM, 2006. First record of the family Theridiosomatidae from Taiwan, with description of a new species (Arachnida: Araneae). Bulletin - British Arachnological Society 13: 265 - 266" title="First record of the family Theridiosomatidae from Taiwan, with description of a new species (Arachnida: Araneae)." volume="13" year="2006">Zhang et al. 2006</bibRefCitation>
|
||
: figs 4-6;
|
||
<bibRefCitation DOI="https://doi.org/10.11646/zootaxa.4927.2.2" author="Ballarin, F" journalOrPublisher="Zootaxa" pageId="0" pageNumber="67" pagination="197 - 208" refId="B2" refString="Ballarin, F, Yamasaki, T, Su, YC, 2021. A survey on poorly known rainforest litter-dwelling spiders of Orchid Island (Lanyu, Taiwan) with the description of a new species (Araneae: Linyphiidae, Tetrablemmidae, and Theridiosomatidae). Zootaxa 4927 (2): 197 - 208, DOI: https://doi.org/10.11646/zootaxa.4927.2.2" title="A survey on poorly known rainforest litter-dwelling spiders of Orchid Island (Lanyu, Taiwan) with the description of a new species (Araneae: Linyphiidae, Tetrablemmidae, and Theridiosomatidae)." url="https://doi.org/10.11646/zootaxa.4927.2.2" volume="4927" year="2021">Ballarin et al. 2021</bibRefCitation>
|
||
: fig. 5B, C), and weakly curved anteriorly with a triangular posterior tip in
|
||
<taxonomicName genus="Z." lsidName="Z. dibaiyin" pageId="0" pageNumber="67" rank="species" species="dibaiyin">
|
||
<emphasis italics="true" pageId="0" pageNumber="67">Z. dibaiyin</emphasis>
|
||
</taxonomicName>
|
||
(Figs
|
||
<figureCitation captionStart="Figure 4" captionStartId="F4" captionText="Figure 4. Zoma dibaiyin Miller, Griswold & Yin, 2009, male habitus and genitalia (NSMT-Ar 21720 A-H) and female habitus and genitalia (NSMT-Ar 21721 I-K) A habitus, dorsal view B habitus, lateral view C palp, retrolateral view D palp, ventral view E palp, posterior-ventral view F palp, prolateral view G embolic division, prolateral view H embolic division, posterior-ventral view I habitus, dorsal view J epigyne, ventral view K vulva, dorsal view. Abbreviations: C conductor CD copulatory ducts E embolus EA embolic apophysis FD fertilization ducts MA median apophysis PC paracymbium PCP posterior conductor projection RCP retrolateral conductor projection S spermatheca ST subtegulum T tegulum. Arrows in E, F, H indicate a cornered margin of posterior membrane of conductor. Arrow in J indicates a sclerotized median pit. Scale bars: 1.0 mm (A, B, I); 0.1 mm (C-H, J, K)." figureDoi="10.3897/zookeys.1109.83807.figure4" httpUri="https://binary.pensoft.net/fig/708570" pageId="0" pageNumber="67">4D</figureCitation>
|
||
,
|
||
<figureCitation captionStart="Figure 5" captionStartId="F5" captionText="Figure 5. Zoma dibaiyin Miller, Griswold & Yin, 2009, male genitalia (NSMT-Ar 21720) A ventral view B posterior-ventral view C prolateral view D conductor and embolic division, retrolateral view E conductor and embolic division, posterior-ventral view. Abbreviations: C conductor CD copulatory ducts E embolus EA embolic apophysis MA median apophysis PC paracymbium PCP posterior conductor projection RCP retrolateral conductor projection ST subtegulum T tegulum. Arrows in B-E indicate a cornered margin of posterior membrane of conductor. Scale bars: 0.1 mm." figureDoi="10.3897/zookeys.1109.83807.figure5" httpUri="https://binary.pensoft.net/fig/708571" pageId="0" pageNumber="67">5D</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="67" type="distribution">
|
||
<paragraph pageId="0" pageNumber="67">Distribution.</paragraph>
|
||
<paragraph pageId="0" pageNumber="67">
|
||
China (Yunnan), Japan (Honshu to the Ryukyu Islands; Fig.
|
||
<figureCitation captionStart="Figure 11" captionStartId="F11" captionText="Figure 11. Distribution of theridiosomatid species in the Ryukyu Islands, Japan." figureDoi="10.3897/zookeys.1109.83807.figure11" httpUri="https://binary.pensoft.net/fig/708577" pageId="0" pageNumber="67">11</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="67" type="habitat">
|
||
<paragraph pageId="0" pageNumber="67">Habitat.</paragraph>
|
||
<paragraph pageId="0" pageNumber="67">
|
||
This species inhabits the forest floor and streamside in dim and wet forests (Fig.
|
||
<figureCitation captionStart="Figure 12" captionStartId="F12" captionText="Figure 12. Habitats of theridiosomatid species in Ryukyu Islands A streamside in dim forest at Iriomote Island B forest floor of secondary forest at Kume Island C grassland at Kume Island D grassland at Yonaguni Island E crevices on limestone rocky wall at Iriomote Island." figureDoi="10.3897/zookeys.1109.83807.figure12" httpUri="https://binary.pensoft.net/fig/708578" pageId="0" pageNumber="67">12B</figureCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="67" type="web morphology">
|
||
<paragraph pageId="0" pageNumber="67">Web morphology.</paragraph>
|
||
<paragraph pageId="0" pageNumber="67">
|
||
This species weaves a concave orb web with radial anastomosis above the ground (Fig.
|
||
<figureCitation captionStart="Figure 13" captionStartId="F13" captionText="Figure 13. Web of Theridiosoma and Zoma spiders at Ryukyu Islands, Japan A-C web of Theridiosoma nigrivirgatum sp. nov. D-E web of Theridiosoma alboannulatum F-H web of Zoma dibaiyin. Abbreviations: RA radial anastomosis SS sticky spirals TL tension line." figureDoi="10.3897/zookeys.1109.83807.figure13" httpUri="https://binary.pensoft.net/fig/708579" pageId="0" pageNumber="67">13F-H</figureCitation>
|
||
). The orb web is almost horizontal. A tension line stretched from the center of the web and attached to substrates such as rocks and dead leaves. The mesh of sticky spirals tends to be fine (occasionally, the number of sticky lines is> 30). For details of the web morphology, see
|
||
<bibRefCitation author="Hiramatsu, T" journalOrPublisher="The University of Chicago Press, Chicago, USA" pageId="0" pageNumber="67" refId="B11" refString="Hiramatsu, T, 2021. Kishidaia 118: 14-23. [Note on the natural history of Zoma dibaiyin (Araneae: Theridiosomatidae)] [In Japanese]" title="Kishidaia 118: 14 - 23. [Note on the natural history of Zoma dibaiyin (Araneae: Theridiosomatidae)] [In Japanese]" year="2021">Hiramatsu (2021)</bibRefCitation>
|
||
.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection pageId="0" pageNumber="67" type="egg sac">
|
||
<paragraph pageId="0" pageNumber="67">Egg sac.</paragraph>
|
||
<paragraph pageId="0" pageNumber="67">
|
||
light brown with a distinct circular suture at the upper end. The sac was suspended from a long horizontal line with a short stalk (see
|
||
<bibRefCitation author="Hiramatsu, T" journalOrPublisher="The University of Chicago Press, Chicago, USA" pageId="0" pageNumber="67" refId="B11" refString="Hiramatsu, T, 2021. Kishidaia 118: 14-23. [Note on the natural history of Zoma dibaiyin (Araneae: Theridiosomatidae)] [In Japanese]" title="Kishidaia 118: 14 - 23. [Note on the natural history of Zoma dibaiyin (Araneae: Theridiosomatidae)] [In Japanese]" year="2021">Hiramatsu 2021</bibRefCitation>
|
||
).
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |