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<document id="363ED24FFF605E3B4D7838C9147E1765" ID-CLB-Dataset="4591" ID-DOI="10.5281/zenodo.5353931" ID-GBIF-Dataset="71f0b51e-9437-4b21-9e8c-8dfef3442df0" ID-ISSN="1942-1354" ID-Zenodo-Dep="5353931" ID-ZooBank="6B2D5CD0-9036-44FD-B80F-4E1F10530ED4" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630212937847" checkinUser="felipe" docAuthor="Keller, Oliver &amp; Skelley, Paul E." docDate="2020" docId="C33D40133255FFEBFF39F990DE73FE38" docLanguage="en" docName="0839.pdf" docOrigin="Insecta Mundi 2020 (839)" docStyle="DocumentStyle:F9976E9B7F8A094D890004515A7A0592.3:InsectaMundi.2020-.journal_article" docStyleId="F9976E9B7F8A094D890004515A7A0592" docStyleName="InsectaMundi.2020-.journal_article" docStyleVersion="3" docTitle="Glaresis franki Keller and Skelley 2020, new species" docType="treatment" docVersion="7" lastPageNumber="4" masterDocId="3F04386B3256FFEEFFF9FFA1DB5EFFD3" masterDocTitle="New family record for the West Indies and two new species of Glaresis Erichson (Coleoptera: Scarabaeoidea: Glaresidae) from Hispaniola" masterLastPageNumber="6" masterPageNumber="1" pageNumber="2" updateTime="1699211413939" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-3.0">
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<mods:title id="E29DEF8A740F9E53346BFCFEB9EF42B6">New family record for the West Indies and two new species of Glaresis Erichson (Coleoptera: Scarabaeoidea: Glaresidae) from Hispaniola</mods:title>
</mods:titleInfo>
<mods:name id="0828A6360C08AD2DCABFD125AF39C7B5" type="personal">
<mods:role id="9DEF8620F6FECAA43A3AAA9C412628EB">
<mods:roleTerm id="09D6B6F31C4C22D36CB90D4BFB53CDB2">Author</mods:roleTerm>
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<mods:namePart id="BC0CCD177222DC697E8DC5881EB8C8EC">Keller, Oliver</mods:namePart>
<mods:affiliation id="0CE595FA352BF8FDF62F35F7F84889C9">University of Florida Department of Entomology and Nematology 1881 Natural Area Drive Gainesville, FL 32611</mods:affiliation>
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<mods:name id="EEC2636B02A0ED692F89B50C2018ECEF" type="personal">
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<mods:namePart id="041A0342FC814A1492A0611B440EAE5C">Skelley, Paul E.</mods:namePart>
<mods:affiliation id="A481D52C00420A6CDB6D20B136E27B00">Florida State Collection of Arthropods Florida Department of Agriculture and Consumer Services P. O. Box 147100 Gainesville, FL 32614 - 7100</mods:affiliation>
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<mods:title id="6CFDD380466369B265E603F14514F8DF">Insecta Mundi</mods:title>
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<mods:part id="9DBC88C2C2A65086C844B6D3D45B5550">
<mods:date id="F7B69A6C3CDE23E3BC095983167DC1C7">2020</mods:date>
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<mods:number id="2B82CCB671933A1FE7D946B04AFC0991">2020-12-25</mods:number>
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<mods:number id="50E66EFACCF1D8286353C4F1711833DB">2020</mods:number>
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<treatment id="C33D40133255FFEBFF39F990DE73FE38" ID-DOI="http://doi.org/10.5281/zenodo.5452241" ID-GBIF-Taxon="185260859" ID-Zenodo-Dep="5452241" LSID="urn:lsid:plazi:treatment:C33D40133255FFEBFF39F990DE73FE38" httpUri="http://treatment.plazi.org/id/C33D40133255FFEBFF39F990DE73FE38" lastPageId="5" lastPageNumber="4" pageId="3" pageNumber="2">
<subSubSection id="038EA28E3255FFEDFF39F990D85BF99D" box="[192,773,1585,1614]" pageId="3" pageNumber="2" type="nomenclature">
<paragraph id="4B2BF1053255FFEDFF39F990D85BF99D" blockId="3.[192,773,1585,1647]" box="[192,773,1585,1614]" pageId="3" pageNumber="2">
<heading id="106346693255FFEDFF39F990D85BF99D" box="[192,773,1585,1614]" fontSize="12" level="2" pageId="3" pageNumber="2" reason="2">
<emphasis id="79E02D173255FFEDFF39F990D85BF99D" bold="true" box="[192,773,1585,1614]" pageId="3" pageNumber="2">
<taxonomicName id="8C948A863255FFEDFF39F990D93AF99D" ID-CoL="7WBQM" authority="Keller and Skelley" authorityName="Keller and Skelley" authorityYear="2020" box="[192,612,1585,1614]" class="Insecta" family="Glaresidae" genus="Glaresis" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="franki" status="sp. nov.">
<emphasis id="79E02D173255FFEDFF39F990DA26F99D" bold="true" box="[192,376,1585,1614]" italics="true" pageId="3" pageNumber="2">Glaresis franki</emphasis>
Keller and Skelley
</taxonomicName>
,
<taxonomicNameLabel id="62D3906C3255FFEDFD89F990D85BF99D" box="[624,773,1585,1614]" pageId="3" pageNumber="2" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="038EA28E3255FFEBFF39F9F4D913FEE7" lastPageId="5" lastPageNumber="4" pageId="3" pageNumber="2" type="description">
<paragraph id="4B2BF1053255FFEDFF39F9F4DA1DF9BC" blockId="3.[192,773,1585,1647]" box="[192,323,1621,1647]" pageId="3" pageNumber="2">
<figureCitation id="D3AFED803255FFEDFF39F9F4DA1DF9BC" box="[192,323,1621,1647]" captionStart="Figures 16" captionStartId="4.[192,279,1310,1336]" captionTargetBox="[192,1440,213,1279]" captionTargetId="figure-60@4.[192,1440,213,1279]" captionTargetPageId="4" captionText="Figures 16. Glaresis franki Keller and Skelley, holotype male. 1) Dorsal habitus. 2) Ventral habitus. 3) Middle and hind legs. 4) Clypeus and head surface. 5) Pronotal surface, lateral view. 6) Male genitalia. Figures 12 scale bar = 1.0 mm, Figures 36 scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5353933" httpUri="https://zenodo.org/record/5353933/files/figure.png" pageId="3" pageNumber="2">Figures 16</figureCitation>
</paragraph>
<paragraph id="4B2BF1053255FFEBFF39F9DFD913FEE7" blockId="3.[192,1441,1662,1927]" lastBlockId="5.[192,1440,214,308]" lastPageId="5" lastPageNumber="4" pageId="3" pageNumber="2">
<emphasis id="79E02D173255FFEDFF39F9DFD956F94A" bold="true" box="[192,520,1662,1689]" pageId="3" pageNumber="2">
Description. Male
<typeStatus id="942F4FA73255FFEDFE63F9DED95DF94A" box="[410,515,1663,1689]" pageId="3" pageNumber="2" type="holotype">holotype</typeStatus>
.
</emphasis>
Length
<quantity id="8C6C5CE03255FFEDFD9CF9DED9E0F94A" box="[613,702,1663,1689]" metricMagnitude="-3" metricUnit="m" metricValue="3.8" pageId="3" pageNumber="2" unit="mm" value="3.8">3.8 mm</quantity>
, width 2.0 mm; body form elongate, slightly widened from elytral base to apical 1/3 (
<figureCitation id="D3AFED803255FFEDFE7DF900DABDF968" box="[388,483,1697,1723]" captionStart="Figures 16" captionStartId="4.[192,279,1310,1336]" captionTargetBox="[192,1440,213,1279]" captionTargetId="figure-60@4.[192,1440,213,1279]" captionTargetPageId="4" captionText="Figures 16. Glaresis franki Keller and Skelley, holotype male. 1) Dorsal habitus. 2) Ventral habitus. 3) Middle and hind legs. 4) Clypeus and head surface. 5) Pronotal surface, lateral view. 6) Male genitalia. Figures 12 scale bar = 1.0 mm, Figures 36 scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5353933" httpUri="https://zenodo.org/record/5353933/files/figure.png" pageId="3" pageNumber="2">Fig. 12</figureCitation>
). Color dark brown, nearly black. Head surface densely microreticulate, strongly dull with shiny tubercles each bearing a single seta, separated by 23 diameters, tubercle diameters equal 12 ocular facets; setae short, barely visible medially, longer and distinct laterally and basally. Vertex with basal groove and carina complete from side to side, groove continuing laterally above eyes. Clypeal apex weakly emarginate medially, somewhat sinuate, with small, dense, evenly spaced tubercles, appearing serrate, without large tubercle on each side of middle, lateral angles sharply oblique, pronounced, lacking tooth (
<figureCitation id="D3AFED803255FFEDFBA8F8EBDFEEF8B6" box="[1105,1200,1866,1893]" captionStart="Figures 16" captionStartId="4.[192,279,1310,1336]" captionTargetBox="[192,1440,213,1279]" captionTargetId="figure-60@4.[192,1440,213,1279]" captionTargetPageId="4" captionText="Figures 16. Glaresis franki Keller and Skelley, holotype male. 1) Dorsal habitus. 2) Ventral habitus. 3) Middle and hind legs. 4) Clypeus and head surface. 5) Pronotal surface, lateral view. 6) Male genitalia. Figures 12 scale bar = 1.0 mm, Figures 36 scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5353933" httpUri="https://zenodo.org/record/5353933/files/figure.png" pageId="3" pageNumber="2">Fig. 45</figureCitation>
). Mandible pair symmetrical; mesal tooth strong; lateral prominence weak; outer margin abruptly rounded. Pronotum with all foveae weakly impressed except fovea on each side medially near lateral margin strongly impressed; surface with straight setae-bearing carinae short, weakly raised; surface between carinae rugose and strongly microreticulate, dulled (
<figureCitation id="D3AFED803252FFEAFF30FA54DA52F9C3" box="[201,268,1525,1552]" captionStart="Figures 16" captionStartId="4.[192,279,1310,1336]" captionTargetBox="[192,1440,213,1279]" captionTargetId="figure-60@4.[192,1440,213,1279]" captionTargetPageId="4" captionText="Figures 16. Glaresis franki Keller and Skelley, holotype male. 1) Dorsal habitus. 2) Ventral habitus. 3) Middle and hind legs. 4) Clypeus and head surface. 5) Pronotal surface, lateral view. 6) Male genitalia. Figures 12 scale bar = 1.0 mm, Figures 36 scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5353933" httpUri="https://zenodo.org/record/5353933/files/figure.png" pageId="4" pageNumber="3">Fig. 5</figureCitation>
); carinae weakly raised, indistinct above lateral foveae. Elytra with surface dull, densely microreticulate; striae convex, feebly carinate, carinal segments widely separated, each with short seta; intervals with deep, circular to nearly rectangular punctures. Metasternum long, feebly shiny, densely microreticulate and rugose, surface medially flat with weak median carina on posterior 1/3, surface either side of glabrous middle with short seta bearing ridges; metasternal groove nearly invisible. Lateral protibial teeth unevenly spaced, basal 2 teeth close together. Mesotibia with 8 spines laterally, evenly spaced in posterolateral emargination terminating at base of strong apical projection (
<figureCitation id="D3AFED803252FFEAFE18F960D97AF90F" box="[481,548,1729,1756]" captionStart="Figures 16" captionStartId="4.[192,279,1310,1336]" captionTargetBox="[192,1440,213,1279]" captionTargetId="figure-60@4.[192,1440,213,1279]" captionTargetPageId="4" captionText="Figures 16. Glaresis franki Keller and Skelley, holotype male. 1) Dorsal habitus. 2) Ventral habitus. 3) Middle and hind legs. 4) Clypeus and head surface. 5) Pronotal surface, lateral view. 6) Male genitalia. Figures 12 scale bar = 1.0 mm, Figures 36 scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5353933" httpUri="https://zenodo.org/record/5353933/files/figure.png" pageId="4" pageNumber="3">Fig. 3</figureCitation>
). Posterior metatrochanteral margin strongly crenate, almost toothed (
<figureCitation id="D3AFED803252FFEAFAA9F960DECAF90F" box="[1360,1428,1729,1756]" captionStart="Figures 16" captionStartId="4.[192,279,1310,1336]" captionTargetBox="[192,1440,213,1279]" captionTargetId="figure-60@4.[192,1440,213,1279]" captionTargetPageId="4" captionText="Figures 16. Glaresis franki Keller and Skelley, holotype male. 1) Dorsal habitus. 2) Ventral habitus. 3) Middle and hind legs. 4) Clypeus and head surface. 5) Pronotal surface, lateral view. 6) Male genitalia. Figures 12 scale bar = 1.0 mm, Figures 36 scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5353933" httpUri="https://zenodo.org/record/5353933/files/figure.png" pageId="4" pageNumber="3">Fig. 3</figureCitation>
); posterosuperior surface of metatrochanter with tooth. Metafemoral surface with widely scattered, elongate, setaebearing tubercles, strongly microreticulate; width to length ratio 1.0:1.5, with narrow flange on anterior margin; small blunt tooth at angle near trochanter; posterosuperior margin without teeth (
<figureCitation id="D3AFED803252FFEAFBAEF889DFC7F891" box="[1111,1177,1832,1858]" captionStart="Figures 16" captionStartId="4.[192,279,1310,1336]" captionTargetBox="[192,1440,213,1279]" captionTargetId="figure-60@4.[192,1440,213,1279]" captionTargetPageId="4" captionText="Figures 16. Glaresis franki Keller and Skelley, holotype male. 1) Dorsal habitus. 2) Ventral habitus. 3) Middle and hind legs. 4) Clypeus and head surface. 5) Pronotal surface, lateral view. 6) Male genitalia. Figures 12 scale bar = 1.0 mm, Figures 36 scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5353933" httpUri="https://zenodo.org/record/5353933/files/figure.png" pageId="4" pageNumber="3">Fig. 3</figureCitation>
). Metatibia broadly triangular, surface rugose-microreticulate, outer margin evenly strongly crenate, lacking large teeth, inner margin smooth, pubescent (
<figureCitation id="D3AFED803252FFEAFE5FF8CDDAB6F855" box="[422,488,1900,1926]" captionStart="Figures 16" captionStartId="4.[192,279,1310,1336]" captionTargetBox="[192,1440,213,1279]" captionTargetId="figure-60@4.[192,1440,213,1279]" captionTargetPageId="4" captionText="Figures 16. Glaresis franki Keller and Skelley, holotype male. 1) Dorsal habitus. 2) Ventral habitus. 3) Middle and hind legs. 4) Clypeus and head surface. 5) Pronotal surface, lateral view. 6) Male genitalia. Figures 12 scale bar = 1.0 mm, Figures 36 scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5353933" httpUri="https://zenodo.org/record/5353933/files/figure.png" pageId="4" pageNumber="3">Fig. 3</figureCitation>
). Apex of 5th abdominal ventrite truncate. Genitalia with basal piece slightly longer than parameres, proximal end curved; median lobe shorter than parameres, tapered from base to rounded apex, broad, about 1.5× width of paramere at middle, curved upward before apex; paramere weakly curved in lateral view, apex bluntly rounded (
<figureCitation id="D3AFED803253FFEBFE04FEB8D961FEE7" box="[509,575,281,308]" captionStart="Figures 16" captionStartId="4.[192,279,1310,1336]" captionTargetBox="[192,1440,213,1279]" captionTargetId="figure-60@4.[192,1440,213,1279]" captionTargetPageId="4" captionText="Figures 16. Glaresis franki Keller and Skelley, holotype male. 1) Dorsal habitus. 2) Ventral habitus. 3) Middle and hind legs. 4) Clypeus and head surface. 5) Pronotal surface, lateral view. 6) Male genitalia. Figures 12 scale bar = 1.0 mm, Figures 36 scale bar = 0.5 mm." figureDoi="http://doi.org/10.5281/zenodo.5353933" httpUri="https://zenodo.org/record/5353933/files/figure.png" pageId="5" pageNumber="4">Fig. 6</figureCitation>
).
</paragraph>
</subSubSection>
<caption id="1FEBA18D3252FFEAFF39FABFD999FAAF" ID-DOI="http://doi.org/10.5281/zenodo.5353933" ID-Zenodo-Dep="5353933" httpUri="https://zenodo.org/record/5353933/files/figure.png" pageId="4" pageNumber="3" startId="4.[192,279,1310,1336]" targetBox="[192,1440,213,1279]" targetPageId="4">
<paragraph id="4B2BF1053252FFEAFF39FABFD999FAAF" blockId="4.[192,1440,1310,1404]" pageId="4" pageNumber="3">
<emphasis id="79E02D173252FFEAFF39FABFDA17FAEB" bold="true" box="[192,329,1310,1336]" pageId="4" pageNumber="3">Figures 16</emphasis>
.
<taxonomicName id="8C948A863252FFEAFEAEFABFD997FAEB" authority="Keller and Skelley" authorityName="Keller and Skelley" authorityYear="2020" box="[343,713,1310,1336]" class="Insecta" family="Glaresidae" genus="Glaresis" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="franki">
<emphasis id="79E02D173252FFEAFEAEFABFDAA9FAEB" box="[343,503,1310,1336]" italics="true" pageId="4" pageNumber="3">Glaresis franki</emphasis>
Keller and Skelley
</taxonomicName>
, holotype male.
<emphasis id="79E02D173252FFEAFC7CFABFD8C3FAEB" bold="true" box="[901,925,1310,1336]" pageId="4" pageNumber="3">1)</emphasis>
Dorsal habitus.
<emphasis id="79E02D173252FFEAFBAEFABFDF2EFAEB" bold="true" box="[1111,1136,1310,1336]" pageId="4" pageNumber="3">2)</emphasis>
Ventral habitus.
<emphasis id="79E02D173252FFEAFAC8FABFDE17FAEB" bold="true" box="[1329,1353,1310,1336]" pageId="4" pageNumber="3">3)</emphasis>
Middle and hind legs.
<emphasis id="79E02D173252FFEAFE9AFA9EDA25FA8A" bold="true" box="[355,379,1343,1369]" pageId="4" pageNumber="3">4)</emphasis>
Clypeus and head surface.
<emphasis id="79E02D173252FFEAFD55FA9ED99AFA8A" bold="true" box="[684,708,1343,1369]" pageId="4" pageNumber="3">5)</emphasis>
Pronotal surface, lateral view.
<emphasis id="79E02D173252FFEAFBE0FA9EDF6FFA8A" bold="true" box="[1049,1073,1343,1369]" pageId="4" pageNumber="3">6)</emphasis>
Male genitalia. Figures 12 scale bar = 1.0 mm, Figures 36 scale bar = 0.5 mm.
</paragraph>
</caption>
<subSubSection id="038EA28E3253FFEBFF39FEE2DA73FEAC" pageId="5" pageNumber="4" type="materials_examined">
<paragraph id="4B2BF1053253FFEBFF39FEE2DA73FEAC" blockId="5.[192,1440,322,383]" pageId="5" pageNumber="4">
<emphasis id="79E02D173253FFEBFF39FEE2DA37FE8F" bold="true" box="[192,361,322,349]" pageId="5" pageNumber="4">Type material.</emphasis>
<materialsCitation id="FBFCFB583253FFEBFE89FEE3DA73FEAC" ID-GBIF-Occurrence="3340071301" collectingDate="1970-11-26" collectionCode="FSCA" collectorName="J. H. Frank" country="Haiti" location="Ganthier" pageId="5" pageNumber="4" specimenCount="1" typeStatus="holotype">
<typeStatus id="942F4FA73253FFEBFE89FEE3DA86FE8F" box="[368,472,322,348]" pageId="5" pageNumber="4" type="holotype">Holotype</typeStatus>
male (
<collectionCode id="2D8569C03253FFEBFDDCFEE2D930FE8E" box="[549,622,323,349]" country="USA" httpUri="http://grbio.org/cool/jekt-x82m" name="Florida State Collection of Arthropods, The Museum of Entomology" pageId="5" pageNumber="4" type="Museum">FSCA</collectionCode>
):
<emphasis id="79E02D173253FFEBFD78FEE3D9E2FE8F" bold="true" box="[641,700,322,348]" pageId="5" pageNumber="4">
<collectingCountry id="3383B1953253FFEBFD78FEE3D9E2FE8F" box="[641,700,322,348]" name="Haiti" pageId="5" pageNumber="4">Haiti</collectingCountry>
</emphasis>
:
<quantity id="8C6C5CE03253FFEBFD30FEE3D85DFE8F" box="[713,771,322,349]" metricMagnitude="3" metricUnit="m" metricValue="8.04672" pageId="5" pageNumber="4" unit="mi" value="5.0">5 mi.</quantity>
W. of /
<location id="4E4BA7DE3253FFEBFCA5FEE3D89EFE8F" LSID="urn:lsid:plazi:treatment:C33D40133255FFEBFF39F990DE73FE38:4E4BA7DE3253FFEBFCA5FEE3D89EFE8F" box="[860,960,322,348]" country="Haiti" name="Ganthier" pageId="5" pageNumber="4">Ganthier</location>
by road /
<date id="3F2AD7C53253FFEBFBCCFEE3DFE9FE8F" box="[1077,1207,322,348]" pageId="5" pageNumber="4" value="1970-11-26">
<collectingDate id="2F6E2E2D3253FFEBFBCCFEE3DFE9FE8F" box="[1077,1207,322,348]" pageId="5" pageNumber="4" value="1970-11-26">26-XI-1970</collectingDate>
</date>
,
<collectorName id="E66194D33253FFEBFB3AFEE2DE61FE8F" box="[1219,1343,322,349]" pageId="5" pageNumber="4">J. H. Frank</collectorName>
, /
<collectingMethod id="92D589123253FFEBFAA2FEE3DA77FEAC" pageId="5" pageNumber="4">blacklight trap</collectingMethod>
.
</materialsCitation>
</paragraph>
</subSubSection>
<subSubSection id="038EA28E3253FFEBFF39FE2FDE73FE38" pageId="5" pageNumber="4" type="etymology">
<paragraph id="4B2BF1053253FFEBFF39FE2FDE73FE38" blockId="5.[191,1440,398,491]" pageId="5" pageNumber="4">
<emphasis id="79E02D173253FFEBFF39FE2FDA1AFE7B" bold="true" box="[192,324,398,424]" pageId="5" pageNumber="4">Etymology.</emphasis>
This species is named for the collector of the species, J. Howard Frank, a close friend and colleague of Mike Thomas. They were both long-term members of the Editorial Board for
<taxonomicName id="8C948A863253FFEBFBD8FE11DF9AFE1A" authority="Mundi" authorityName="Mundi" box="[1057,1220,431,458]" class="Insecta" kingdom="Animalia" pageId="5" pageNumber="4" phylum="Arthropoda" rank="class">Insecta Mundi</taxonomicName>
from its early years until recently. Naming the two
<taxonomicName id="8C948A863253FFEBFDE3FE70D998FE38" box="[538,710,465,491]" class="Insecta" family="Glaresidae" genus="Glaresis" kingdom="Animalia" order="Coleoptera" pageId="5" pageNumber="4" phylum="Arthropoda" rank="species" species="undetermined">
<emphasis id="79E02D173253FFEBFDE3FE70D92EFE38" box="[538,624,465,491]" italics="true" pageId="5" pageNumber="4">Glaresis</emphasis>
species
</taxonomicName>
on Hispaniola for these two friends seems appropriate.
</paragraph>
</subSubSection>
</treatment>
</document>