treatments-xml/data/8D/73/87/8D7387F9FF9CFFF9BBC64942054FFAA2.xml
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<document id="5C2BDC9DC7DF311F4A9F953D321C62CA" ID-CLB-Dataset="22439" ID-DOI="10.5281/zenodo.3779118" ID-GBIF-Dataset="4a6a79da-9274-4617-8584-b5e45a85a1df" ID-ISSN="1942-1354" ID-Zenodo-Dep="3779118" ID-ZooBank="1A15BC98-F720-4063-ABFD-453124D3BE26" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.taxonomicNames_approvedBy="felipe" checkinTime="1588272917477" checkinUser="felipe" docAuthor="Minkina, Łukasz &amp; Kakizoe, Showtaro" docDate="2020" docId="8D7387F9FF9CFFF9BBC64942054FFAA2" docLanguage="en" docName="InsectaMundi.0762.1-6.pdf.imf" docOrigin="Insecta Mundi 762" docStyle="DocumentStyle:6D85C66E4B52897F25DA165649BC06DC.3:InsectaMundi.2009-2020.journal_article" docStyleId="6D85C66E4B52897F25DA165649BC06DC" docStyleName="InsectaMundi.2009-2020.journal_article" docStyleVersion="3" docTitle="Termitodiellus mindanaoensis Minkina and Kakizoe 2020, new species" docType="treatment" docVersion="6" lastPageNumber="3" masterDocId="714AFF81FF9FFFFDBB06485F0427FF90" masterDocTitle="Termitodiellus mindanaoensis, a new species of Rhyparini Schmidt, 1910 (Scarabaeidae: Aphodiinae) from the Philippines" masterLastPageNumber="6" masterPageNumber="1" pageNumber="2" updateTime="1698838241752" updateUser="ExternalLinkService" zenodo-license-document="CC-BY-NC-3.0">
<mods:mods id="8621F5CB5BC792F460BA7213364FBFEE" xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo id="32F65FF1D324BB8B77656E870C644327">
<mods:title id="5EC3E085CD1E163C283F6B6EAC7602BE">Termitodiellus mindanaoensis, a new species of Rhyparini Schmidt, 1910 (Scarabaeidae: Aphodiinae) from the Philippines</mods:title>
</mods:titleInfo>
<mods:name id="74B13390A559C6BECE368F7DD931B7CC" type="personal">
<mods:role id="E1DC1E8EE99384EEEB2220356B2EAD85">
<mods:roleTerm id="A319B676D619A0C28B92F067AC76FC82">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="A389C2D813724996C242D945FB217D4A">Minkina, Łukasz</mods:namePart>
<mods:affiliation id="E2FED00912EEB63449F39F2E017543A1">os. Polana Szaflarska 4 / 39 34 - 400 Nowy Targ, Poland klekel @ interia. eu</mods:affiliation>
</mods:name>
<mods:name id="64A0F2CEB30E553BB99BD7A890FB6258" type="personal">
<mods:role id="D76277EF47DDBFB77C31E23795592DA6">
<mods:roleTerm id="C31BBAB3BEB0C22AEA87CA0CA3043048">Author</mods:roleTerm>
</mods:role>
<mods:namePart id="C670252B57F772487AEFF2B25D048DEA">Kakizoe, Showtaro</mods:namePart>
<mods:affiliation id="1AD4EDC05CFCF21DF7A0C6374C669BA3">The Kyushu University Museum Hakozaki 6 - 10 - 1, Higashi-ku, Fukuoka, 812 - 8581, Japan</mods:affiliation>
</mods:name>
<mods:typeOfResource id="25457D7F4AD2337C12223778CB130D7A">text</mods:typeOfResource>
<mods:relatedItem id="539C8B0426C3C8CA772ACF865329363A" type="host">
<mods:titleInfo id="E78BDE0E13FC929FBBAD2963DB107D1E">
<mods:title id="99A0600FCD55E3CE191D3369E5E35B10">Insecta Mundi</mods:title>
</mods:titleInfo>
<mods:part id="2FEC9649596287B6F40C4A3596E5E77A">
<mods:date id="3D23E2572A3CF10A95B644E866203AF5">2020</mods:date>
<mods:detail id="3E54EE17DDA4B9665618D6FD0B6FB452" type="pubDate">
<mods:number id="368D786BFD708457B1A9402CD4757ABC">2020-04-24</mods:number>
</mods:detail>
<mods:detail id="B90C504773684ABF6AD91EFDE15A6DFD" type="issue">
<mods:number id="0BCFABE6DF000DA5E4DC5F09B34F141F">762</mods:number>
</mods:detail>
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<mods:start id="141AFD1F292CD07A70C0678D75756A3A">1</mods:start>
<mods:end id="E6C09213489958C6EB2D89CB5950557B">6</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:classification id="56E751ADDFF2FDDBF3333D96CD37C2B6">journal article</mods:classification>
<mods:identifier id="739523852C73084A7B68A79695E5278C" type="CLB-Dataset">22439</mods:identifier>
<mods:identifier id="8359369EDB185191C2FE4067715D9491" type="DOI">10.5281/zenodo.3779118</mods:identifier>
<mods:identifier id="7E6685C1DFEF5FF5E3E2B400E7FFB8C5" type="GBIF-Dataset">4a6a79da-9274-4617-8584-b5e45a85a1df</mods:identifier>
<mods:identifier id="BCEE6F7DB79CD3D8A2EA7D462CBE5D8D" type="ISSN">1942-1354</mods:identifier>
<mods:identifier id="CEBD1A059EA95A026EE1251268B0228C" type="Zenodo-Dep">3779118</mods:identifier>
<mods:identifier id="3A888B1955A3A06F985900174D94EF41" type="ZooBank">1A15BC98-F720-4063-ABFD-453124D3BE26</mods:identifier>
</mods:mods>
<treatment id="8D7387F9FF9CFFF9BBC64942054FFAA2" ID-DOI="http://doi.org/10.5281/zenodo.3795349" ID-GBIF-Taxon="163879523" ID-Zenodo-Dep="3795349" LSID="urn:lsid:plazi:treatment:8D7387F9FF9CFFF9BBC64942054FFAA2" httpUri="http://treatment.plazi.org/id/8D7387F9FF9CFFF9BBC64942054FFAA2" lastPageId="4" lastPageNumber="3" pageId="3" pageNumber="2">
<subSubSection id="4DC06564FF9CFFFEBBC649420066FEA5" box="[192,1089,285,309]" pageId="3" pageNumber="2" type="nomenclature">
<paragraph id="056536EFFF9CFFFEBBC649420066FEA5" blockId="3.[192,1089,285,342]" box="[192,1089,285,309]" pageId="3" pageNumber="2">
<heading id="5E2D8183FF9CFFFEBBC649420066FEA5" bold="true" box="[192,1089,285,309]" fontSize="10" level="2" pageId="3" pageNumber="2" reason="3">
<emphasis id="37AEEAFDFF9CFFFEBBC649420066FEA5" bold="true" box="[192,1089,285,309]" pageId="3" pageNumber="2">
<taxonomicName id="C2DA4D6CFF9CFFFEBBC6494207B7FEA5" authority="Minkina and Kakizoe" authorityName="Minkina and Kakizoe" authorityYear="2020" box="[192,912,285,309]" class="Insecta" family="Aphodiidae" genus="Termitodiellus" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="2" phylum="Arthropoda" rank="species" species="mindanaoensis" status="sp. nov.">
<emphasis id="37AEEAFDFF9CFFFEBBC649420646FEA5" bold="true" box="[192,609,285,309]" italics="true" pageId="3" pageNumber="2">Termitodiellus mindanaoensis</emphasis>
Minkina and Kakizoe
</taxonomicName>
,
<taxonomicNameLabel id="2C9D5786FF9CFFFEB89849420066FEA5" box="[926,1089,285,309]" pageId="3" pageNumber="2" rank="species">new species</taxonomicNameLabel>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="4DC06564FF9CFFFEBBC649610563FEC6" box="[192,324,318,342]" pageId="3" pageNumber="2" type="description">
<paragraph id="056536EFFF9CFFFEBBC649610563FEC6" blockId="3.[192,1089,285,342]" box="[192,324,318,342]" pageId="3" pageNumber="2">
(
<figureCitation id="9DE12A6AFF9CFFFEBBCF4961051DFEC6" box="[201,314,318,342]" captionStart-0="Figures 15" captionStart-1="Figures 611" captionStartId-0="6.[192,286,1879,1901]" captionStartId-1="7.[192,287,1877,1899]" captionTargetBox-0="[346,1278,235,1819]" captionTargetBox-1="[260,1308,216,1843]" captionTargetId-0="figure@6.[331,1300,215,1845]" captionTargetId-1="figure@7.[260,1328,213,1843]" captionTargetPageId-0="6" captionTargetPageId-1="7" captionText-0="Figures 15. Termitodiellus mindanaoensis, new species. 1) Dorsal view, holotype male. 2) Ventral view, paratype male. 3) Dorsal view, paratype female. 4) Ventral view, paratype female. 5) Dorso-lateral view, holotype male." captionText-1="Figures 611. Termitodiellus mindanaoensis, new species. 6) Head, holotype male. 7) Left mesotibia, paratype male. 8) Left mesotibia, paratype female. 9) Aedeagus, dorsal view. 10) Aedeagus, lateral view. 11) Epipharynx." figureDoi-0="http://doi.org/10.5281/zenodo.3779120" figureDoi-1="http://doi.org/10.5281/zenodo.3779122" httpUri-0="https://zenodo.org/record/3779120/files/figure.png" httpUri-1="https://zenodo.org/record/3779122/files/figure.png" pageId="3" pageNumber="2">Fig. 111</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="4DC06564FF9CFFFEBBC6493106D2FC86" pageId="3" pageNumber="2" type="materials_examined">
<paragraph id="056536EFFF9CFFFEBBC649310577FE36" blockId="3.[192,1440,365,422]" pageId="3" pageNumber="2">
<emphasis id="37AEEAFDFF9CFFFEBBC6493105ACFE15" bold="true" box="[192,395,365,390]" pageId="3" pageNumber="2">Type material.</emphasis>
<materialsCitation id="B5B23CB2FF9CFFFEBA9349320577FE36" ID-GBIF-Occurrence="2609529510" collectingDate="2019-01" collectionCode="LMC" country="Philippines" location="Alamada" municipality="Mindanao Island" pageId="3" pageNumber="2" specimenCount="1" specimenCount-male="1" stateProvince="North Cotabato" typeStatus="holotype">
<emphasis id="37AEEAFDFF9CFFFEBA9349320627FE15" box="[405,512,365,389]" italics="true" pageId="3" pageNumber="2">
<typeStatus id="DA61884DFF9CFFFEBA9349320627FE15" box="[405,512,365,389]" pageId="3" pageNumber="2" type="holotype">Holotype</typeStatus>
</emphasis>
<specimenCount id="13DCFD66FF9CFFFEB90F49310607FE15" box="[521,544,366,389]" pageId="3" pageNumber="2" type="male"></specimenCount>
.
<collectingCountry id="7DCD767FFF9CFFFEB9284931069BFE16" box="[558,700,366,390]" name="Philippines" pageId="3" pageNumber="2">Philippines</collectingCountry>
,
<collectingMunicipality id="E501AC95FF9CFFFEB9CC49310787FE16" box="[714,928,366,390]" pageId="3" pageNumber="2">Mindanao Island</collectingMunicipality>
,
<location id="00056034FF9CFFFEB8A849310038FE16" LSID="urn:lsid:plazi:treatment:8D7387F9FF9CFFF9BBC64942054FFAA2:00056034FF9CFFFEB8A849310038FE16" box="[942,1055,366,390]" country="Philippines" municipality="Mindanao Island" name="Alamada" pageId="3" pageNumber="2" stateProvince="North Cotabato">Alamada</location>
,
<collectingRegion id="C71EF80DFF9CFFFEBF28493100D5FE16" box="[1070,1266,366,390]" country="Philippines" name="North Cotabato" pageId="3" pageNumber="2">North Cotabato</collectingRegion>
,
<date id="7164102FFF9CFFFEBE064931017DFE16" box="[1280,1370,366,390]" pageId="3" pageNumber="2" value="2019-01">
<collectingDate id="6120E9C7FF9CFFFEBE064931017DFE16" box="[1280,1370,366,390]" pageId="3" pageNumber="2" value="2019-01">I. 2019</collectingDate>
</date>
, local coll. (
<collectionCode id="63CBAE2AFF9CFFFEBA0649D10564FE36" box="[256,323,398,422]" country="Mozambique" lsid="urn:lsid:biocol.org:col:13322" name="Instituto de Investigação Científica de Mozambique" pageId="3" pageNumber="2" type="Herbarium">LMC</collectionCode>
).
</materialsCitation>
</paragraph>
<paragraph id="056536EFFF9CFFFEBBC649E106D2FC86" blockId="3.[192,1441,446,790]" pageId="3" pageNumber="2">
<materialsCitation id="B5B23CB2FF9CFFFEBBC649E1016FFD66" ID-GBIF-Occurrence="2609529511" collectingDate="2017-03" collectingDateMax="2019-10" collectingDateMin="2017-03" collectionCode="CNC, SKC, MKC, LMC, MFC" collectorName="Local Collector &amp; Description &amp; Dorsum &amp; Body &amp; Dorsally &amp; Relatively" country="Philippines" county="South Catabato" location="Mindanao Island" municipality="Mindanao Island" pageId="3" pageNumber="2" specimenCount="38" specimenCount-female="24" specimenCount-male="14" stateProvince="Albay" typeStatus="paratype">
<emphasis id="37AEEAFDFF9CFFFEBBC649E10566FE45" box="[192,321,446,469]" italics="true" pageId="3" pageNumber="2">
<typeStatus id="DA61884DFF9CFFFEBBC649E1051AFE45" box="[192,317,446,469]" pageId="3" pageNumber="2" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<specimenCount id="13DCFD66FF9CFFFEBA4E49E1054BFE46" box="[328,364,446,470]" pageId="3" pageNumber="2" type="male">1♂</specimenCount>
,
<specimenCount id="13DCFD66FF9CFFFEBA7E49E105BAFE46" box="[376,413,446,470]" pageId="3" pageNumber="2" type="female">5♀</specimenCount>
, same data as holotype (
<collectionCode id="63CBAE2AFF9CFFFEB9C149E10720FE46" box="[711,775,446,470]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="3" pageNumber="2">CNC</collectionCode>
,
<collectionCode id="63CBAE2AFF9CFFFEB81449E10768FE46" box="[786,847,446,470]" pageId="3" pageNumber="2">SKC</collectionCode>
).
<specimenCount id="13DCFD66FF9CFFFEB86449E107A0FE46" box="[866,903,446,470]" pageId="3" pageNumber="2" type="male">4♂</specimenCount>
,
<specimenCount id="13DCFD66FF9CFFFEB89549E1079FFE46" box="[915,952,446,470]" pageId="3" pageNumber="2" type="female">4♀</specimenCount>
, same data as
<typeStatus id="DA61884DFF9CFFFEBF6D49E100F4FE46" box="[1131,1235,446,470]" pageId="3" pageNumber="2" type="holotype">holotype</typeStatus>
except,
<date id="7164102FFF9CFFFEBE3049E10187FE46" box="[1334,1440,446,470]" pageId="3" pageNumber="2" value="2019-03">
<collectingDate id="6120E9C7FF9CFFFEBE3049E10187FE46" box="[1334,1440,446,470]" pageId="3" pageNumber="2" value="2019-03">III. 2019</collectingDate>
</date>
(
<collectionCode id="63CBAE2AFF9CFFFEBBCE4981052AFE66" box="[200,269,478,502]" pageId="3" pageNumber="2">MKC</collectionCode>
,
<collectionCode id="63CBAE2AFF9CFFFEBA1C4981057EFE66" box="[282,345,478,502]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="3" pageNumber="2">CNC</collectionCode>
,
<collectionCode id="63CBAE2AFF9CFFFEBA614981058DFE66" box="[359,426,478,502]" country="Mozambique" lsid="urn:lsid:biocol.org:col:13322" name="Instituto de Investigação Científica de Mozambique" pageId="3" pageNumber="2" type="Herbarium">LMC</collectionCode>
).
<specimenCount id="13DCFD66FF9CFFFEBAB8498105C4FE66" box="[446,483,478,502]" pageId="3" pageNumber="2" type="female">2♀</specimenCount>
, same data as
<typeStatus id="DA61884DFF9CFFFEB99949810720FE66" box="[671,775,478,502]" pageId="3" pageNumber="2" type="holotype">holotype</typeStatus>
except,
<date id="7164102FFF9CFFFEB869498107C9FE66" box="[879,1006,478,502]" pageId="3" pageNumber="2" value="2019-08">
<collectingDate id="6120E9C7FF9CFFFEB869498107C9FE66" box="[879,1006,478,502]" pageId="3" pageNumber="2" value="2019-08">VIII. 2019</collectingDate>
</date>
(
<collectionCode id="63CBAE2AFF9CFFFEB8F949810065FE66" box="[1023,1090,478,502]" country="Mozambique" lsid="urn:lsid:biocol.org:col:13322" name="Instituto de Investigação Científica de Mozambique" pageId="3" pageNumber="2" type="Herbarium">LMC</collectionCode>
).
<specimenCount id="13DCFD66FF9CFFFEBF514981005DFE66" box="[1111,1146,478,502]" pageId="3" pageNumber="2" type="female">1♀</specimenCount>
, same data as
<typeStatus id="DA61884DFF9CFFFEBE3E49810187FE66" box="[1336,1440,478,502]" pageId="3" pageNumber="2" type="holotype">holotype</typeStatus>
except,
<collectingCounty id="EC044E63FF9CFFFEBA1A49A105F9FD86" box="[284,478,510,534]" pageId="3" pageNumber="2">South Catabato</collectingCounty>
,
<date id="7164102FFF9CFFFEBAEF49A1066BFD86" box="[489,588,510,534]" pageId="3" pageNumber="2" value="2019-03">
<collectingDate id="6120E9C7FF9CFFFEBAEF49A1066BFD86" box="[489,588,510,534]" pageId="3" pageNumber="2" value="2019-03">III.2019</collectingDate>
</date>
(
<collectionCode id="63CBAE2AFF9CFFFEB95D49A106B9FD86" box="[603,670,510,534]" country="Mozambique" lsid="urn:lsid:biocol.org:col:13322" name="Instituto de Investigação Científica de Mozambique" pageId="3" pageNumber="2" type="Herbarium">LMC</collectionCode>
).
<specimenCount id="13DCFD66FF9CFFFEB9B649A106F1FD86" box="[688,726,510,534]" pageId="3" pageNumber="2" type="male">2♂</specimenCount>
,
<specimenCount id="13DCFD66FF9CFFFEB9E749A10722FD86" box="[737,773,510,534]" pageId="3" pageNumber="2" type="female">3♀</specimenCount>
, same data as
<typeStatus id="DA61884DFF9CFFFEB8BE49A10007FD86" box="[952,1056,510,534]" pageId="3" pageNumber="2" type="holotype">holotype</typeStatus>
except,
<date id="7164102FFF9CFFFEBF8449A10130FD86" box="[1154,1303,510,534]" pageId="3" pageNumber="2" value="2019-03-10">
<collectingDate id="6120E9C7FF9CFFFEBF8449A10130FD86" box="[1154,1303,510,534]" pageId="3" pageNumber="2" value="2019-03-10">10. III. 2019</collectingDate>
</date>
(
<collectionCode id="63CBAE2AFF9CFFFEBE2049A10144FD86" box="[1318,1379,510,534]" pageId="3" pageNumber="2">SKC</collectionCode>
).
<specimenCount id="13DCFD66FF9CFFFEBE7049A101BCFD86" box="[1398,1435,510,534]" pageId="3" pageNumber="2" type="male">4♂</specimenCount>
,
<specimenCount id="13DCFD66FF9CFFFEBBC64A4104C2FDA6" box="[192,229,542,566]" pageId="3" pageNumber="2" type="female">4♀</specimenCount>
, same data as
<typeStatus id="DA61884DFF9CFFFEBA9E4A410627FDA6" box="[408,512,542,566]" pageId="3" pageNumber="2" type="holotype">holotype</typeStatus>
except,
<date id="7164102FFF9CFFFEB9654A4106EDFDA6" box="[611,714,542,566]" pageId="3" pageNumber="2" value="2019-04">
<collectingDate id="6120E9C7FF9CFFFEB9654A4106EDFDA6" box="[611,714,542,566]" pageId="3" pageNumber="2" value="2019-04">IV. 2019</collectingDate>
</date>
(
<collectionCode id="63CBAE2AFF9CFFFEB9DC4A410730FDA6" box="[730,791,542,566]" pageId="3" pageNumber="2">SKC</collectionCode>
). 1 ex, same data as
<typeStatus id="DA61884DFF9CFFFEBF164A41005FFDA6" box="[1040,1144,542,566]" pageId="3" pageNumber="2" type="holotype">holotype</typeStatus>
except,
<date id="7164102FFF9CFFFEBFDD4A410165FDA6" box="[1243,1346,542,566]" pageId="3" pageNumber="2" value="2019-06">
<collectingDate id="6120E9C7FF9CFFFEBFDD4A410165FDA6" box="[1243,1346,542,566]" pageId="3" pageNumber="2" value="2019-06">VI. 2019</collectingDate>
</date>
(
<collectionCode id="63CBAE2AFF9CFFFEBE574A4101B3FDA6" box="[1361,1428,542,566]" country="Mozambique" lsid="urn:lsid:biocol.org:col:13322" name="Instituto de Investigação Científica de Mozambique" pageId="3" pageNumber="2" type="Herbarium">LMC</collectionCode>
).
<specimenCount id="13DCFD66FF9CFFFEBBC64A6104C3FDC6" box="[192,228,574,598]" pageId="3" pageNumber="2" type="male">1♂</specimenCount>
, same data as
<typeStatus id="DA61884DFF9CFFFEBA9F4A610626FDC6" box="[409,513,574,598]" pageId="3" pageNumber="2" type="holotype">holotype</typeStatus>
except,
<date id="7164102FFF9CFFFEB9624A6006C6FDC6" box="[612,737,574,598]" pageId="3" pageNumber="2" value="2019-08">
<collectingDate id="6120E9C7FF9CFFFEB9624A6006C6FDC6" box="[612,737,574,598]" pageId="3" pageNumber="2" value="2019-08">VIII. 2019</collectingDate>
</date>
(
<collectionCode id="63CBAE2AFF9CFFFEB9F74A610709FDC6" box="[753,814,574,598]" pageId="3" pageNumber="2">SKC</collectionCode>
).
<specimenCount id="13DCFD66FF9CFFFEB8474A610743FDC6" box="[833,868,574,598]" pageId="3" pageNumber="2" type="female">1♀</specimenCount>
, same data as
<typeStatus id="DA61884DFF9CFFFEBF1C4A6100A5FDC6" box="[1050,1154,574,598]" pageId="3" pageNumber="2" type="holotype">holotype</typeStatus>
except,
<date id="7164102FFF9CFFFEBFE04A600165FDC6" box="[1254,1346,574,598]" pageId="3" pageNumber="2" value="2019-10">
<collectingDate id="6120E9C7FF9CFFFEBFE04A600165FDC6" box="[1254,1346,574,598]" pageId="3" pageNumber="2" value="2019-10">X. 2019</collectingDate>
</date>
(
<collectionCode id="63CBAE2AFF9CFFFEBE574A6101B3FDC6" box="[1361,1428,574,598]" country="Japan" httpUri="http://grbio.org/cool/fgye-365i" name="Matsushima Fungus Collection" pageId="3" pageNumber="2">MFC</collectionCode>
).
<specimenCount id="13DCFD66FF9CFFFEBBC64A0104C4FDE6" box="[192,227,606,630]" pageId="3" pageNumber="2" type="female">1♀</specimenCount>
,
<collectingCountry id="7DCD767FFF9CFFFEBBF54A0105A6FDE6" box="[243,385,606,630]" name="Philippines" pageId="3" pageNumber="2">Philippines</collectingCountry>
,
<collectingMunicipality id="E501AC95FF9CFFFEBA894A010644FDE6" box="[399,611,606,630]" pageId="3" pageNumber="2">Mindanao Island</collectingMunicipality>
, Composteia New
<collectingRegion id="C71EF80DFF9CFFFEB84C4A0107B3FDE6" box="[842,916,606,630]" country="Philippines" name="Albay" pageId="3" pageNumber="2">Albay</collectingRegion>
,
<date id="7164102FFF9CFFFEB8A44A000036FDE6" box="[930,1041,606,630]" pageId="3" pageNumber="2" value="2017-03">
<collectingDate id="6120E9C7FF9CFFFEB8A44A000036FDE6" box="[930,1041,606,630]" pageId="3" pageNumber="2" value="2017-03">III. 2017</collectingDate>
</date>
,
<collectorName id="A82F5339FF9CFFFEBF184A0100F1FDE6" box="[1054,1238,606,630]" pageId="3" pageNumber="2">Local Collector</collectorName>
leg. (
<collectionCode id="63CBAE2AFF9CFFFEBE1D4A010146FDE6" box="[1307,1377,606,630]" pageId="3" pageNumber="2">MKC</collectionCode>
).
<specimenCount id="13DCFD66FF9CFFFEBE704A0101BDFDE6" box="[1398,1434,606,630]" pageId="3" pageNumber="2" type="male">1♂</specimenCount>
,
<collectingCountry id="7DCD767FFF9CFFFEBBC64A210569FD06" box="[192,334,638,662]" name="Philippines" pageId="3" pageNumber="2">Philippines</collectingCountry>
,
<location id="00056034FF9CFFFEBA5C4A21060AFD06" LSID="urn:lsid:plazi:treatment:8D7387F9FF9CFFF9BBC64942054FFAA2:00056034FF9CFFFEBA5C4A21060AFD06" box="[346,557,638,662]" country="Philippines" county="South Catabato" municipality="Mindanao Island" name="Mindanao Island" pageId="3" pageNumber="2" stateProvince="Albay">Mindanao Island</location>
,
<collectingRegion id="C71EF80DFF9CFFFEB93F4A2006CFFD06" box="[569,744,638,662]" country="Philippines" name="Lanao del Sur" pageId="3" pageNumber="2">Lanao del Sur</collectingRegion>
, Amai Manabilang,
<location id="00056034FF9CFFFEB8D94A200091FD06" LSID="urn:lsid:plazi:treatment:8D7387F9FF9CFFF9BBC64942054FFAA2:00056034FF9CFFFEB8D94A200091FD06" box="[991,1206,638,662]" country="Philippines" county="South Catabato" municipality="Mindanao Island" name="Mt. Piapayungan" pageId="3" pageNumber="2" stateProvince="Albay">Mt. Piapayungan</location>
,
<date id="7164102FFF9CFFFEBFC44A20013DFD06" box="[1218,1306,638,662]" pageId="3" pageNumber="2" value="2019-01">
<collectingDate id="6120E9C7FF9CFFFEBFC44A20013DFD06" box="[1218,1306,638,662]" pageId="3" pageNumber="2" value="2019-01">I. 2019</collectingDate>
</date>
,
<collectorName id="A82F5339FF9CFFFEBE234A210522FD26" pageId="3" pageNumber="2">Local Collector</collectorName>
leg. (
<collectionCode id="63CBAE2AFF9CFFFEBA4A4AC105AEFD26" box="[332,393,670,694]" pageId="3" pageNumber="2">SKC</collectionCode>
).
<specimenCount id="13DCFD66FF9CFFFEBA994AC105E3FD26" box="[415,452,670,694]" pageId="3" pageNumber="2" type="male">1♂</specimenCount>
,
<collectingCountry id="7DCD767FFF9CFFFEBAD54AC10646FD26" box="[467,609,670,694]" name="Philippines" pageId="3" pageNumber="2">Philippines</collectingCountry>
, Mindanao, Wao, Lanao,
<date id="7164102FFF9CFFFEB8A54AC00037FD26" box="[931,1040,670,694]" pageId="3" pageNumber="2" value="2019-06">
<collectingDate id="6120E9C7FF9CFFFEB8A54AC00037FD26" box="[931,1040,670,694]" pageId="3" pageNumber="2" value="2019-06">VI. 2019</collectingDate>
</date>
,
<collectorName id="A82F5339FF9CFFFEBF184AC100F0FD26" box="[1054,1239,670,694]" pageId="3" pageNumber="2">Local Collector</collectorName>
leg. (
<collectionCode id="63CBAE2AFF9CFFFEBE194AC10178FD26" box="[1311,1375,670,694]" country="Canada" httpUri="http://grbio.org/cool/y2kv-9w3k" name="Canadian National Collection of Insects, Arachnids, and Nematodes" pageId="3" pageNumber="2">CNC</collectionCode>
),
<specimenCount id="13DCFD66FF9CFFFEBE704AC101BDFD26" box="[1398,1434,670,694]" pageId="3" pageNumber="2" type="female">3♀</specimenCount>
,
<collectingCountry id="7DCD767FFF9CFFFEBBC64AE10569FD46" box="[192,334,702,726]" name="Philippines" pageId="3" pageNumber="2">Philippines</collectingCountry>
,
<collectingRegion id="C71EF80DFF9CFFFEBA5E4AE10634FD46" box="[344,531,702,726]" country="Philippines" name="Agusan del Sur" pageId="3" pageNumber="2">Agusan del Sur</collectingRegion>
,
<date id="7164102FFF9CFFFEB91B4AE106A1FD46" box="[541,646,702,726]" pageId="3" pageNumber="2" value="2019-07">
<collectingDate id="6120E9C7FF9CFFFEB91B4AE106A1FD46" box="[541,646,702,726]" pageId="3" pageNumber="2" value="2019-07">vii. 2019</collectingDate>
</date>
,
<collectorName id="A82F5339FF9CFFFEB9964AE10763FD46" box="[656,836,702,726]" pageId="3" pageNumber="2">Local Collector</collectorName>
leg. (
<collectionCode id="63CBAE2AFF9CFFFEB8844AE107E2FD46" box="[898,965,702,726]" country="Mozambique" lsid="urn:lsid:biocol.org:col:13322" name="Instituto de Investigação Científica de Mozambique" pageId="3" pageNumber="2" type="Herbarium">LMC</collectionCode>
).
<collectorName id="A82F5339FF9CFFFEB8DD4AE1004FFD46" box="[987,1128,702,726]" pageId="3" pageNumber="2">Description</collectorName>
of
<typeStatus id="DA61884DFF9CFFFEBF8E4AE100D7FD46" box="[1160,1264,702,726]" pageId="3" pageNumber="2" type="holotype">holotype</typeStatus>
male.
<collectorName id="A82F5339FF9CFFFEBE384AE00187FD46" box="[1342,1440,703,726]" pageId="3" pageNumber="2">Dorsum</collectorName>
(
<figureCitation id="9DE12A6AFF9CFFFEBBCF4A81052AFD66" box="[201,269,734,758]" captionStart="Figures 15" captionStartId="6.[192,286,1879,1901]" captionTargetBox="[346,1278,235,1819]" captionTargetId="figure@6.[331,1300,215,1845]" captionTargetPageId="6" captionText="Figures 15. Termitodiellus mindanaoensis, new species. 1) Dorsal view, holotype male. 2) Ventral view, paratype male. 3) Dorsal view, paratype female. 4) Ventral view, paratype female. 5) Dorso-lateral view, holotype male." figureDoi="http://doi.org/10.5281/zenodo.3779120" httpUri="https://zenodo.org/record/3779120/files/figure.png" pageId="3" pageNumber="2">Fig. 1</figureCitation>
).
<collectorName id="A82F5339FF9CFFFEBA214A810542FD66" box="[295,357,734,758]" pageId="3" pageNumber="2">Body</collectorName>
length
<quantity id="C2229B0AFF9CFFFEBAC24A810604FD66" box="[452,547,734,758]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="3" pageNumber="2" unit="mm" value="3.9">3.9 mm</quantity>
, greatest width
<quantity id="C2229B0AFF9CFFFEB9ED4A81076DFD66" box="[747,842,734,758]" metricMagnitude="-3" metricUnit="m" metricValue="1.8" pageId="3" pageNumber="2" unit="mm" value="1.8">1.8 mm</quantity>
.
<collectorName id="A82F5339FF9CFFFEB8504A810799FD66" box="[854,958,734,758]" pageId="3" pageNumber="2">Dorsally</collectorName>
dark brownish.
<collectorName id="A82F5339FF9CFFFEBF8E4A810123FD66" box="[1160,1284,734,758]" pageId="3" pageNumber="2">Relatively</collectorName>
wide
</materialsCitation>
; shiny; all punctures bearing short, thick macrosetae.
</paragraph>
</subSubSection>
<subSubSection id="4DC06564FF9CFFF9BBC64B710147FDC2" lastPageId="4" lastPageNumber="3" pageId="3" pageNumber="2" type="description">
<paragraph id="056536EFFF9CFFFEBBC64B71012AFB37" blockId="3.[192,1440,814,1191]" pageId="3" pageNumber="2">
<emphasis id="37AEEAFDFF9CFFFEBBC64B710537FCD6" bold="true" box="[192,272,814,838]" pageId="3" pageNumber="2">Head.</emphasis>
Clypeus (
<figureCitation id="9DE12A6AFF9CFFFEBA8A4B7105F3FCD6" box="[396,468,814,838]" captionStart="Figures 611" captionStartId="7.[192,287,1877,1899]" captionTargetBox="[260,1308,216,1843]" captionTargetId="figure@7.[260,1328,213,1843]" captionTargetPageId="7" captionText="Figures 611. Termitodiellus mindanaoensis, new species. 6) Head, holotype male. 7) Left mesotibia, paratype male. 8) Left mesotibia, paratype female. 9) Aedeagus, dorsal view. 10) Aedeagus, lateral view. 11) Epipharynx." figureDoi="http://doi.org/10.5281/zenodo.3779122" httpUri="https://zenodo.org/record/3779122/files/figure.png" pageId="3" pageNumber="2">Fig. 6</figureCitation>
) anteriorly gently, widely rounded, with rounded angles at sides, considerable lateral emargination and a further widely rounded angle separated by a small emargination from the distinctly protruding, rounded genae. Clypeocentral disc distinctly convex, ringed by a shallow, pubescent peridiscal impression, with a pair of distinct, long, convergent costae, on sides with distinct, fine punctures bearing small setae. Frons with four short but distinct longitudinal costae (two frontodiscal and two frontolateral) of structure similar to costae on clypeal convexity; punctures of frons with setae similar to those on clypeus. Head covered by rather regularly spaced, dense, fine punctures bearing short, thick setae. Epipharynx (
<figureCitation id="9DE12A6AFF9CFFFEB9464C5006B4FBB7" box="[576,659,1039,1063]" captionStart="Figures 611" captionStartId="7.[192,287,1877,1899]" captionTargetBox="[260,1308,216,1843]" captionTargetId="figure@7.[260,1328,213,1843]" captionTargetPageId="7" captionText="Figures 611. Termitodiellus mindanaoensis, new species. 6) Head, holotype male. 7) Left mesotibia, paratype male. 8) Left mesotibia, paratype female. 9) Aedeagus, dorsal view. 10) Aedeagus, lateral view. 11) Epipharynx." figureDoi="http://doi.org/10.5281/zenodo.3779122" httpUri="https://zenodo.org/record/3779122/files/figure.png" pageId="3" pageNumber="2">Fig. 11</figureCitation>
) transverse, with lateral sides broadly rounded. Anterior border concave; corypha absent, on its place present very wide, weakly convex exalcation, with few small, short and sparsely distributed setae in middle, and much thinner, quite long, dense setae on sides; additionally on each side of that structure there are small teeth in shape of inverted scalene triangle, and few relatively long and thick setae on anterior part of pedia. Tormae short, regularly rounded.
</paragraph>
<paragraph id="056536EFFF9CFFFEBBC64CE005BCFA47" blockId="3.[192,1441,1215,1496]" pageId="3" pageNumber="2">
<emphasis id="37AEEAFDFF9CFFFEBBC64CE00571FB47" bold="true" box="[192,342,1215,1239]" pageId="3" pageNumber="2">Prothorax.</emphasis>
Pronotum shiny, with six distinct convex costae and seven longitudinal furrows, and two lateral lobes on each side. Anterior and posterior lobe of similar width, form widest part of pronotum. Paramedian and discolateral costae interrupted near middle; submarginal costa interrupted in apical median third; costae with small punctures bearing short, thick setae. Middle furrow with rather regularly spaced, medium sized, dense punctures nearly on whole surface, with small basal area lacking punctures; second furrow with regularly spaced, medium-sized, dense punctures over whole surface, except base and central part lacking punctures; third and fourth furrows with regularly spaced, medium-sized, dense punctures nearly on whole surface, except at level of costa interruption. All furrow punctures bearing short, thick setae.
</paragraph>
<paragraph id="056536EFFF9CFFFEBBC64DAF055EF958" blockId="3.[192,1441,1520,1928]" pageId="3" pageNumber="2">
<emphasis id="37AEEAFDFF9CFFFEBBC64DAF0557F998" bold="true" box="[192,368,1520,1544]" pageId="3" pageNumber="2">Pterothorax.</emphasis>
Each elytron with six elevated costae and five flat intervals. Almost entire surface of first pair of intervals, except just before apex, basal half of second intervals and basal part of remaining intervals, with distinct microreticulation, elytra here weakly shiny; remaining interval surface shiny, without microreticulation. Intervals irregularly punctured on microreticulate surface, all punctures with short, thick setae. Costae on sides with dense punctures bearing short, thick setae. Postdiscal bulb reduced. Caudal bulb nearly spherical, rounded when viewed from above, and elongately rounded in lateral view.
</paragraph>
<paragraph id="056536EFFF9CFFF9BBEE4E8E0037FE02" blockId="3.[192,1441,1520,1928]" lastBlockId="4.[192,1440,217,402]" lastPageId="4" lastPageNumber="3" pageId="3" pageNumber="2">
Venter (
<figureCitation id="9DE12A6AFF9CFFFEBA4F4E8F05AAF978" box="[329,397,1744,1768]" captionStart="Figures 15" captionStartId="6.[192,286,1879,1901]" captionTargetBox="[346,1278,235,1819]" captionTargetId="figure@6.[331,1300,215,1845]" captionTargetPageId="6" captionText="Figures 15. Termitodiellus mindanaoensis, new species. 1) Dorsal view, holotype male. 2) Ventral view, paratype male. 3) Dorsal view, paratype female. 4) Ventral view, paratype female. 5) Dorso-lateral view, holotype male." figureDoi="http://doi.org/10.5281/zenodo.3779120" httpUri="https://zenodo.org/record/3779120/files/figure.png" pageId="3" pageNumber="2">Fig. 2</figureCitation>
) moderately shiny, with distinct microreticulation on median impression and indistinct microreticulation on remaining surface. Metaventrite flattened in middle, with distinct wide, shallow median impression devoid of punctures; sides of median impression with flattened surface and rows of small punctures; anterolateral juxtacostal impressions large, wider than median impression, shallow; the rest of punctation of metaventrite dense, regularly spaced; all punctures bearing short, thick setae. Abdominal ventrites shiny, without microreticulation, with two transverse rows of punctures, laterally a small, rounded, shallow impression; anal ventrite with moderately large rounded and shallow impressions laterally, surface area with regular punctation; punctures of abdominal ventrites with short, thick setae. Pygidium in middle with longitudinal furrow on each side; punctation similar to punctation of abdominal ventrites. Meso- and metafemora lacking teeth on posterior margins; all femora moderately shiny, with distinct microreticulation; entire surface with dense, regular punctation bearing short, thick setae. Mesotibiae with large, flattened, inwardly hooked apical spine.
</paragraph>
<paragraph id="056536EFFF9BFFF9BBC649F500D9FE72" blockId="4.[192,1440,425,482]" pageId="4" pageNumber="3">
<emphasis id="37AEEAFDFF9BFFF9BBC649F505A9FE51" bold="true" box="[192,398,425,450]" pageId="4" pageNumber="3">Male genitalia.</emphasis>
Aedeagus (
<figureCitation id="9DE12A6AFF9BFFF9B92649F506B3FE52" box="[544,660,426,450]" captionStart="Figures 611" captionStartId="7.[192,287,1877,1899]" captionTargetBox="[260,1308,216,1843]" captionTargetId="figure@7.[260,1328,213,1843]" captionTargetPageId="7" captionText="Figures 611. Termitodiellus mindanaoensis, new species. 6) Head, holotype male. 7) Left mesotibia, paratype male. 8) Left mesotibia, paratype female. 9) Aedeagus, dorsal view. 10) Aedeagus, lateral view. 11) Epipharynx." figureDoi="http://doi.org/10.5281/zenodo.3779122" httpUri="https://zenodo.org/record/3779122/files/figure.png" pageId="4" pageNumber="3">Fig. 910</figureCitation>
) nearly rectangular in dorsal view, with phallobase noticeably longer than parameres, sinusoidal in lateral view, with two indistinct setae on each side.
</paragraph>
<paragraph id="056536EFFF9BFFF9BBC649A60147FDC2" blockId="4.[192,1440,505,594]" pageId="4" pageNumber="3">
<emphasis id="37AEEAFDFF9BFFF9BBC649A605F4FD81" bold="true" box="[192,467,505,529]" pageId="4" pageNumber="3">Sexual dimorphism.</emphasis>
Male mesotibia near apex gently sinuate along medial margin, with large, flattened, inwardly bent apical spine (
<figureCitation id="9DE12A6AFF9BFFF9B9724A4506FBFDA2" box="[628,732,538,562]" captionStart="Figures 15" captionStartId="6.[192,286,1879,1901]" captionTargetBox="[346,1278,235,1819]" captionTargetId="figure@6.[331,1300,215,1845]" captionTargetPageId="6" captionText="Figures 15. Termitodiellus mindanaoensis, new species. 1) Dorsal view, holotype male. 2) Ventral view, paratype male. 3) Dorsal view, paratype female. 4) Ventral view, paratype female. 5) Dorso-lateral view, holotype male." figureDoi="http://doi.org/10.5281/zenodo.3779120" httpUri="https://zenodo.org/record/3779120/files/figure.png" pageId="4" pageNumber="3">Fig. 12</figureCitation>
,
<figureCitation id="9DE12A6AFF9BFFF9B9EC4A4506DCFDA2" box="[746,763,538,562]" captionStart="Figures 611" captionStartId="7.[192,287,1877,1899]" captionTargetBox="[260,1308,216,1843]" captionTargetId="figure@7.[260,1328,213,1843]" captionTargetPageId="7" captionText="Figures 611. Termitodiellus mindanaoensis, new species. 6) Head, holotype male. 7) Left mesotibia, paratype male. 8) Left mesotibia, paratype female. 9) Aedeagus, dorsal view. 10) Aedeagus, lateral view. 11) Epipharynx." figureDoi="http://doi.org/10.5281/zenodo.3779122" httpUri="https://zenodo.org/record/3779122/files/figure.png" pageId="4" pageNumber="3">7</figureCitation>
). Female (
<figureCitation id="9DE12A6AFF9BFFF9B8874A4507CEFDA2" box="[897,1001,538,562]" captionStart="Figures 15" captionStartId="6.[192,286,1879,1901]" captionTargetBox="[346,1278,235,1819]" captionTargetId="figure@6.[331,1300,215,1845]" captionTargetPageId="6" captionText="Figures 15. Termitodiellus mindanaoensis, new species. 1) Dorsal view, holotype male. 2) Ventral view, paratype male. 3) Dorsal view, paratype female. 4) Ventral view, paratype female. 5) Dorso-lateral view, holotype male." figureDoi="http://doi.org/10.5281/zenodo.3779120" httpUri="https://zenodo.org/record/3779120/files/figure.png" pageId="4" pageNumber="3">Fig. 34</figureCitation>
,
<figureCitation id="9DE12A6AFF9BFFF9B8F14A450022FDA2" box="[1015,1029,538,562]" captionStart="Figures 611" captionStartId="7.[192,287,1877,1899]" captionTargetBox="[260,1308,216,1843]" captionTargetId="figure@7.[260,1328,213,1843]" captionTargetPageId="7" captionText="Figures 611. Termitodiellus mindanaoensis, new species. 6) Head, holotype male. 7) Left mesotibia, paratype male. 8) Left mesotibia, paratype female. 9) Aedeagus, dorsal view. 10) Aedeagus, lateral view. 11) Epipharynx." figureDoi="http://doi.org/10.5281/zenodo.3779122" httpUri="https://zenodo.org/record/3779122/files/figure.png" pageId="4" pageNumber="3">8</figureCitation>
) metatibia nearly straight along medial margin, near apex, with rather small, not so much flattened, nearly straight apical spine.
</paragraph>
</subSubSection>
<subSubSection id="4DC06564FF9BFFF9BBC64A3600B5FB72" pageId="4" pageNumber="3" type="discussion">
<paragraph id="056536EFFF9BFFF9BBC64A3600B5FB72" blockId="4.[192,1441,617,1250]" pageId="4" pageNumber="3">
<emphasis id="37AEEAFDFF9BFFF9BBC64A360508FD11" bold="true" box="[192,303,617,641]" pageId="4" pageNumber="3">Affinity.</emphasis>
The only species previously known from the
<collectingCountry id="7DCD767FFF9BFFF9B8434A3507F7FD12" box="[837,976,618,642]" name="Philippines" pageId="4" pageNumber="3">Philippines</collectingCountry>
is
<taxonomicName id="C2DA4D6CFF9BFFF9B8E94A360520FD31" authority="(Howden, 1995)" baseAuthorityName="Howden" baseAuthorityYear="1995" class="Insecta" family="Aphodiidae" genus="Termitodiellus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="luzonensis">
<emphasis id="37AEEAFDFF9BFFF9B8E94A360101FD11" box="[1007,1318,617,641]" italics="true" pageId="4" pageNumber="3">Termitodiellus luzonensis</emphasis>
(
<bibRefCitation id="614B4B1EFF9BFFF9BE354A3504D9FD31" author="Howden, H." pageId="4" pageNumber="3" pagination="23 - 27" refId="ref3211" refString="Howden, H. 1995. New Rhyparini from Fiji and the Philippines. The Coleopterists Bulletin 49 (1): 23 - 27." type="journal article" year="1995">Howden, 1995</bibRefCitation>
)
</taxonomicName>
, however it was collected only on Luzon and can be shown to notably differ from
<taxonomicName id="C2DA4D6CFF9BFFF9BFCE4AD50187FD31" authority="Minkina &amp; Kakizoe &amp; April, 2020" authorityName="Minkina &amp; Kakizoe &amp; April" authorityYear="2020" box="[1224,1440,649,673]" class="Insecta" family="Aphodiidae" genus="Termitodiellus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="mindanaoensis" status="sp. nov.">
<emphasis id="37AEEAFDFF9BFFF9BFCE4AD50187FD31" box="[1224,1440,649,673]" italics="true" pageId="4" pageNumber="3">T. mindanaoensis</emphasis>
</taxonomicName>
<emphasis id="37AEEAFDFF9BFFF9BBC64AF5052EFD52" bold="true" box="[192,265,682,706]" pageId="4" pageNumber="3">
<taxonomicNameLabel id="2C9D5786FF9BFFF9BBC64AF5052EFD52" box="[192,265,682,706]" pageId="4" pageNumber="3" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
by: the lack of dense punctures on the pronotum and elytral intervals, which additionally are always rounded; by much different, not spherical shape of caudal bulbs, which are incised in the middle; and relatively smaller body size (
<quantity id="C2229B0AFF9BFFF9B9524AB50694FC92" box="[596,691,746,770]" metricMagnitude="-3" metricUnit="m" metricValue="3.1" pageId="4" pageNumber="3" unit="mm" value="3.1">3.1 mm</quantity>
).
<bibRefCitation id="614B4B1EFF9BFFF9B9C04AB50759FC92" author="Howden, H." box="[710,894,746,770]" pageId="4" pageNumber="3" pagination="23 - 27" refId="ref3211" refString="Howden, H. 1995. New Rhyparini from Fiji and the Philippines. The Coleopterists Bulletin 49 (1): 23 - 27." type="journal article" year="1995">Howden (1995)</bibRefCitation>
mentioned that
<taxonomicName id="C2DA4D6CFF9BFFF9BF4F4AB500CEFC91" baseAuthorityName="Howden" baseAuthorityYear="1995" box="[1097,1257,745,769]" class="Insecta" family="Aphodiidae" genus="Termitodiellus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="luzonensis">
<emphasis id="37AEEAFDFF9BFFF9BF4F4AB500CEFC91" box="[1097,1257,745,769]" italics="true" pageId="4" pageNumber="3">T. luzonensis</emphasis>
</taxonomicName>
is most closely related to
<taxonomicName id="C2DA4D6CFF9BFFF9BA3E4B550779FCB2" authority="(Krikken and Huijbregts, 1987)" baseAuthorityName="Krikken and Huijbregts" baseAuthorityYear="1987" box="[312,862,778,802]" class="Insecta" family="Aphodiidae" genus="Termitodiellus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="interruptus">
<emphasis id="37AEEAFDFF9BFFF9BA3E4B5505C7FCB1" box="[312,480,778,801]" italics="true" pageId="4" pageNumber="3">T. interruptus</emphasis>
(
<bibRefCitation id="614B4B1EFF9BFFF9BAE84B550771FCB2" author="Krikken, J. &amp; J. Huijbregts" box="[494,854,778,802]" pageId="4" pageNumber="3" pagination="97 - 111" refId="ref3331" refString="Krikken, J., and J. Huijbregts. 1987. Southeast Asian Termitodius: a taxonomic review with descriptions of four new species. Zoologische Mededelingen Leyden 61 (7): 97 - 111." type="journal article" year="1987">Krikken and Huijbregts, 1987</bibRefCitation>
)
</taxonomicName>
without citing any arguments, however because of similar proportions of the body and discomedian costae not interrupted it seems to be more closely related to
<taxonomicName id="C2DA4D6CFF9BFFF9BA394B15063DFCF1" authority="Minkina &amp; Kakizoe &amp; April, 2020" authorityName="Minkina &amp; Kakizoe &amp; April" authorityYear="2020" box="[319,538,841,865]" class="Insecta" family="Aphodiidae" genus="Termitodiellus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="mindanaoensis" status="sp. nov.">
<emphasis id="37AEEAFDFF9BFFF9BA394B15063DFCF1" box="[319,538,841,865]" italics="true" pageId="4" pageNumber="3">T. mindanaoensis</emphasis>
</taxonomicName>
<emphasis id="37AEEAFDFF9BFFF9B9244B15064CFCF2" bold="true" box="[546,619,842,866]" pageId="4" pageNumber="3">
<taxonomicNameLabel id="2C9D5786FF9BFFF9B9244B15064CFCF2" box="[546,619,842,866]" pageId="4" pageNumber="3" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
than to
<taxonomicName id="C2DA4D6CFF9BFFF9B9D34B150136FCF2" authority="(Krikken and Huijbregts, 1987)" baseAuthorityName="Krikken and Huijbregts" baseAuthorityYear="1987" box="[725,1297,842,866]" class="Insecta" family="Aphodiidae" genus="Termitodiellus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="interruptus">
<emphasis id="37AEEAFDFF9BFFF9B9D34B1507A6FCF1" box="[725,897,842,865]" italics="true" pageId="4" pageNumber="3">T. interruptus</emphasis>
(
<bibRefCitation id="614B4B1EFF9BFFF9B8974B15012FFCF2" author="Krikken, J. &amp; J. Huijbregts" box="[913,1288,842,866]" pageId="4" pageNumber="3" pagination="97 - 111" refId="ref3331" refString="Krikken, J., and J. Huijbregts. 1987. Southeast Asian Termitodius: a taxonomic review with descriptions of four new species. Zoologische Mededelingen Leyden 61 (7): 97 - 111." type="journal article" year="1987">Krikken and Huijbregts, 1987</bibRefCitation>
)
</taxonomicName>
. Because of the dense punctation of pronotum and elytral intervals, and proportions of the body it seems to be most similar to
<taxonomicName id="C2DA4D6CFF9BFFF9BA314BD606D5FC32" authority="(Nomura, 1943)" baseAuthorityName="Nomura" baseAuthorityYear="1943" box="[311,754,905,930]" class="Insecta" family="Aphodiidae" genus="Termitodiellus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="esakii">
<emphasis id="37AEEAFDFF9BFFF9BA314BD60615FC31" box="[311,562,905,929]" italics="true" pageId="4" pageNumber="3">Termitodiellus esakii</emphasis>
(
<bibRefCitation id="614B4B1EFF9BFFF9B9394BD506CDFC32" author="Nomura, S." box="[575,746,906,930]" pageId="4" pageNumber="3" pagination="77 - 83" refId="ref3450" refString="Nomura, S. 1943. Zur Kenntnis der Aphodiiden aus Mikronesien. Mushi, Fukuoka 15: 77 - 83." type="journal article" year="1943">Nomura, 1943</bibRefCitation>
)
</taxonomicName>
and seems to be evolutionarily intermediate between that and the other known species. From
<taxonomicName id="C2DA4D6CFF9BFFF9B9774BF507B9FC52" authority="(Nomura, 1943)" baseAuthorityName="Nomura" baseAuthorityYear="1943" box="[625,926,937,962]" class="Insecta" family="Aphodiidae" genus="Termitodiellus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="esakii">
<emphasis id="37AEEAFDFF9BFFF9B9774BF506FFFC51" box="[625,728,937,961]" italics="true" pageId="4" pageNumber="3">T. esakii</emphasis>
(
<bibRefCitation id="614B4B1EFF9BFFF9B9E14BF507B2FC52" author="Nomura, S." box="[743,917,938,962]" pageId="4" pageNumber="3" pagination="77 - 83" refId="ref3450" refString="Nomura, S. 1943. Zur Kenntnis der Aphodiiden aus Mikronesien. Mushi, Fukuoka 15: 77 - 83." type="journal article" year="1943">Nomura, 1943</bibRefCitation>
)
</taxonomicName>
it can be easily distinguished by following characters: larger body size (
<quantity id="C2229B0AFF9BFFF9B9114B950656FC71" box="[535,625,970,994]" metricMagnitude="-3" metricUnit="m" metricValue="3.9" pageId="4" pageNumber="3" unit="mm" value="3.9">3.9 mm</quantity>
vs. 3.0 to
<quantity id="C2229B0AFF9BFFF9B9E34B950765FC72" box="[741,834,970,994]" metricMagnitude="-3" metricUnit="m" metricValue="3.3" pageId="4" pageNumber="3" unit="mm" value="3.3">3.3 mm</quantity>
); basal part of pronotal furrows without punctures (vs. whole surface of pronotal furrows with punctures); elytral intervals impunctate in apical part (vs. whole surface of elytral intervals with punctures); and different shape of caudal bulbs (
<bibRefCitation id="614B4B1EFF9BFFF9BFF64C5501BCFBB2" author="Nomura, S." box="[1264,1435,1034,1058]" pageId="4" pageNumber="3" pagination="77 - 83" refId="ref3450" refString="Nomura, S. 1943. Zur Kenntnis der Aphodiiden aus Mikronesien. Mushi, Fukuoka 15: 77 - 83." type="journal article" year="1943">Nomura 1943</bibRefCitation>
;
<bibRefCitation id="614B4B1EFF9BFFF9BBC64C75063AFBD2" author="Cartwright, O. L. &amp; R. D. Gordon" box="[192,541,1066,1090]" pageId="4" pageNumber="3" pagination="257 - 296" refId="ref3179" refString="Cartwright, O. L., and R. D. Gordon. 1971. Coleoptera: Scarabaeidae. Insects of Micronesia 17 (4): 257 - 296." type="journal article" year="1971">Cartwright and Gordon 1971</bibRefCitation>
;
<bibRefCitation id="614B4B1EFF9BFFF9B9214C750772FBD2" author="Howden, H. &amp; R. Storey" box="[551,853,1066,1090]" pageId="4" pageNumber="3" pagination="1810 - 1823" refId="ref3237" refString="Howden, H., and R. Storey. 1992. Phylogeny of the Rhyparini and the new tribe Stereomerini with descriptions of new genera and species. Canadian Journal of Zoology 70 (9): 1810 - 1823." type="journal article" year="1992">Howden and Storey 1992</bibRefCitation>
). From all other known species
<taxonomicName id="C2DA4D6CFF9BFFF9BFCC4C7501B8FBD1" authority="Minkina &amp; Kakizoe &amp; April, 2020" authorityName="Minkina &amp; Kakizoe &amp; April" authorityYear="2020" box="[1226,1439,1065,1089]" class="Insecta" family="Aphodiidae" genus="Termitodiellus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="mindanaoensis" status="sp. nov.">
<emphasis id="37AEEAFDFF9BFFF9BFCC4C7501B8FBD1" box="[1226,1439,1065,1089]" italics="true" pageId="4" pageNumber="3">T. mindanaoensis</emphasis>
</taxonomicName>
<emphasis id="37AEEAFDFF9BFFF9BBC64C15052EFBF2" bold="true" box="[192,265,1098,1122]" pageId="4" pageNumber="3">
<taxonomicNameLabel id="2C9D5786FF9BFFF9BBC64C15052EFBF2" box="[192,265,1098,1122]" pageId="4" pageNumber="3" rank="species">n. sp.</taxonomicNameLabel>
</emphasis>
can be easily distinguished by: punctures in elytral intervals irregularly spaced (vs. punctures in elytral intervals arranged in very distinct rows); proportions of the body. Only
<taxonomicName id="C2DA4D6CFF9BFFF9BFA54C350626FB32" authority="(Krikken and Huijbregts, 1987)" baseAuthorityName="Krikken and Huijbregts" baseAuthorityYear="1987" class="Insecta" family="Aphodiidae" genus="Termitodiellus" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="3" phylum="Arthropoda" rank="species" species="interruptus">
<emphasis id="37AEEAFDFF9BFFF9BFA54C350169FB11" box="[1187,1358,1130,1153]" italics="true" pageId="4" pageNumber="3">T. interruptus</emphasis>
(
<bibRefCitation id="614B4B1EFF9BFFF9BE584C3505DFFB32" author="Krikken, J. &amp; J. Huijbregts" pageId="4" pageNumber="3" pagination="97 - 111" refId="ref3331" refString="Krikken, J., and J. Huijbregts. 1987. Southeast Asian Termitodius: a taxonomic review with descriptions of four new species. Zoologische Mededelingen Leyden 61 (7): 97 - 111." type="journal article" year="1987">Krikken and Huijbregts, 1987</bibRefCitation>
)
</taxonomicName>
has similar proportions, but the new species can be easily distinguished by discomedian costae which are not interrupted, and dense and regularly spaced punctures on most of the pronotal surface (in other species a large part of pronotum lacks punctures).
</paragraph>
</subSubSection>
<subSubSection id="4DC06564FF9BFFF9BBC64CA5054FFAA2" pageId="4" pageNumber="3" type="etymology">
<paragraph id="056536EFFF9BFFF9BBC64CA5054FFAA2" blockId="4.[192,1440,1274,1330]" pageId="4" pageNumber="3">
<emphasis id="37AEEAFDFF9BFFF9BBC64CA5057CFA82" bold="true" box="[192,347,1274,1298]" pageId="4" pageNumber="3">Etymology.</emphasis>
Toponymic; an adjective derived from the name of Mindanao Island, where the new species was collected.
</paragraph>
</subSubSection>
</treatment>
</document>