treatments-xml/data/6F/3E/CF/6F3ECF1F406AFFD65786B95F70F0FFDE.xml

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<mods:title id="BEA4C34BD21645478175F07C2775042A">On the type species of the genus Dendarus Dejean, 1821 (Coleoptera: Tenebrionidae)</mods:title>
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<mods:namePart id="040B4C09AECDC6ECB3DEE68E836D3257">Leo, Piero</mods:namePart>
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Status of 
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<emphasis id="D5E3A21B406AFFD6561CB95F73A0FD72" bold="true" box="[269,428,1051,1078]" italics="true" pageId="1" pageNumber="576">Helops tristis</emphasis>
<bibRefCitation id="830603F8406AFFD656A2B95F7033FD72" author="Rossi, P." box="[435,575,1051,1077]" pageId="1" pageNumber="576" refId="ref4418" refString="Rossi, P. (1790) Fauna Etrusca sistens insecta quae in provinciis Florentina et Pisana praesertim collegit. Tomus primus. Thomae Masi &amp; Sociorum, Liburni, xxii + [1] + 272 pp. https: // doi. org / 10.5962 / bhl. title. 15771" type="book" year="1790">Rossi, 1790</bibRefCitation>
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<bibRefCitation id="830603F8406AFFD65786B9277326FD3A" author="Rossi, P." box="[151,298,1123,1149]" pageId="1" pageNumber="576" refId="ref4418" refString="Rossi, P. (1790) Fauna Etrusca sistens insecta quae in provinciis Florentina et Pisana praesertim collegit. Tomus primus. Thomae Masi &amp; Sociorum, Liburni, xxii + [1] + 272 pp. https: // doi. org / 10.5962 / bhl. title. 15771" type="book" year="1790">Rossi (1790)</bibRefCitation>
described and illustrated 
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<emphasis id="D5E3A21B406AFFD65547B92770FCFD3A" box="[598,752,1123,1149]" italics="true" pageId="1" pageNumber="576">Helops tristis</emphasis>
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from the Central Italian region of Etruria. Type specimens from Rossi’s collection have been considered lost (
<bibRefCitation id="830603F8406AFFD655DEB9C37163FDE6" author="Seidlitz, G. von" box="[719,879,1159,1185]" pageId="1" pageNumber="576" pagination="201 - 400" refId="ref4471" refString="Seidlitz, G. von (1893) Tenebrionidae. Pp. 201 - 400. In: Kiesenwetter, H. von &amp; Seidlitz, G. von: Naturgeschichte der Insecten Deutschlands. Erste Abtheilung. Coleoptera. Funfter Band. Erste Halfte. Nicolaische Verlags-Buchhandlung, Berlin, xxviii + 877 pp." type="book chapter" year="1893">Seidlitz, 1893</bibRefCitation>
), although 
<bibRefCitation id="830603F8406AFFD654FDB9CC76FCFDE6" author="Horn, W. &amp; Kahle, I." box="[1004,1264,1159,1186]" pageId="1" pageNumber="576" pagination="38" refId="ref3615" refString="Horn, W. &amp; Kahle, I. (1937) &quot; Uber entomologische Sammlungen, Entomologen und Entomo-Museologie &quot;. Issued in 3 pts. as: Sonderdruck von Entomologische Beihefte, Bd. 2 - 4. Dec. 1935, Oct. 1936, Aug. 1937. Dahlem, Berlin, 536 + 12 + 38 b / w pls." type="journal article" year="1937">Horn and Kahle (1937)</bibRefCitation>
and 
<bibRefCitation id="830603F8406AFFD65237B9CC72EEFD82" author="Horn, W. &amp; Kahle, I. &amp; Friese, G. &amp; Gaedike, R." pageId="1" pageNumber="576" refId="ref3681" refString="Horn, W., Kahle, I., Friese, G. &amp; Gaedike, R. (1990) Collectiones Entomologicae. Ein Kompendium uber den Verbleib entomologischer Sammlungen der Welt bis 1960. Teil 1. A bis K. Akademie der Landwirtschaftswissenschaften der Deutschen Demokratischen Republik, Berlin, 573 pp." type="book" year="1990">Horn et al. (1990)</bibRefCitation>
indicated that some of his material went to the Museum für Naturkunde (ZMB - Berlin). Based on contacts with Bernd Jaeger of the ZMB, two historical specimens from 
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under the species name 
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<emphasis id="D5E3A21B406AFFD65397B98B7717FDAE" box="[1158,1307,1231,1257]" italics="true" pageId="1" pageNumber="576">Helops tristis</emphasis>
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were located in the Rossi material. As 
<bibRefCitation id="830603F8406AFFD656C0B9B7706CFC4A" author="Rossi, P." box="[465,608,1267,1293]" pageId="1" pageNumber="576" refId="ref4418" refString="Rossi, P. (1790) Fauna Etrusca sistens insecta quae in provinciis Florentina et Pisana praesertim collegit. Tomus primus. Thomae Masi &amp; Sociorum, Liburni, xxii + [1] + 272 pp. https: // doi. org / 10.5962 / bhl. title. 15771" type="book" year="1790">Rossi (1790)</bibRefCitation>
did not specify the number of studied specimens nor designate a 
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while describing 
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<emphasis id="D5E3A21B406AFFD65649B85373E2FC76" box="[344,494,1303,1329]" italics="true" pageId="1" pageNumber="576">Helops tristis</emphasis>
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a 
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designation is needed in order to fix the taxonomic status of the species. The male specimen presented on the 
<figureCitation id="7FAC628C406AFFD65527B87F7085FC12" box="[566,649,1339,1365]" captionStart="FIGURE 1" captionStartId="2.[151,250,1073,1097]" captionTargetBox="[151,1436,181,1049]" captionTargetId="figure-20@2.[151,1436,181,1049]" captionTargetPageId="2" captionText="FIGURE 1. Habitus photographs of selected Dendarus species. Lectotype (A) and paralectotype (B) of Helops tristis, lectotype (C) of Pandarus coarcticollis (photos by MNHN/Christophe Rivier)." figureDoi="http://doi.org/10.5281/zenodo.5556809" httpUri="https://zenodo.org/record/5556809/files/figure.png" pageId="1" pageNumber="576">Fig. 1A</figureCitation>
is hereby designated as the 
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. The second specimen, a female, is accordingly treated as a 
<typeStatus id="382CC0AB406AFFD656D0B81B7050FC3E" box="[449,604,1375,1401]" pageId="1" pageNumber="576" type="paralectotype">paralectotype</typeStatus>
. Labels indicating the status of these 
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were added.
</paragraph>
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Up to now, the following five species of 
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<emphasis id="D5E3A21B406AFFD65587B8C77108FCDA" box="[662,772,1411,1437]" italics="true" pageId="1" pageNumber="576">Dendarus</emphasis>
</taxonomicName>
have been reported from 
<collectingCountry id="9F803E99406AFFD6533BB8C77651FCDA" box="[1066,1117,1411,1437]" name="Italy" pageId="1" pageNumber="576">Italy</collectingCountry>
(
<bibRefCitation id="830603F8406AFFD6537DB8C77732FCDA" author="Aliquo, V. &amp; Rastelli, M. &amp; Rastelli, S. &amp; Soldati, F." box="[1132,1342,1411,1438]" pageId="1" pageNumber="576" refId="ref2750" refString="Aliquo, V., Rastelli, M., Rastelli, S. &amp; Soldati, F. (2007) Coleotteri Tenebrionidi d'Italia - Darkling Beetles of Italy. Piccole Faune II. Museo Civico di Storia Naturale di Carmagnola, Torino, Associazione Naturalistica Piemontese, Torino and Progetto Biodiversita, Comitato Parchi, Roma. [CD-ROM]" type="book" year="2007">Aliquo et al. 2007</bibRefCitation>
, 
<bibRefCitation id="830603F8406AFFD65258B8C772DEFC86" author="Trichas, A." pageId="1" pageNumber="576" pagination="417 - 462" refId="ref4580" refString="Trichas, A. (2008) The genus Dendarus Latreille, 1829 (Coleoptera, Tenebrionidae: Dendarini) in Greece. (A systematic account of the genus with description of a new species and four new systematic combinations.). In: Makarov, S. E. &amp; Dimitrijevic, R. N. (Eds.), Advances in arachnology and developmental biology. Papers dedicated to Prof. Dr. Bozidar Curcic. Institute of Zoology, Belgrade, BAS, Sofia, Faculty of Life Science, Vienna, SASA, Belgrade and UNESCO MAB Serbia, Belgrade, pp. 417 - 462." type="book chapter" year="2008">Trichas 2008</bibRefCitation>
, 
<bibRefCitation id="830603F8406AFFD657CFB8E37363FC86" author="Soldati, F." box="[222,367,1447,1473]" pageId="1" pageNumber="576" pagination="52 - 59" refId="ref4527" refString="Soldati, F. (2012) A new species of the genus Dendarus Dejean, 1821 from Greece. Une nouvelle espece du genre Dendarus Dejean, 1821 decrite de Grece (Coleoptera, Tenebrionidae). R. A. R. E., 21, 52 - 59." type="journal article" year="2012">Soldati 2012</bibRefCitation>
, 
<bibRefCitation id="830603F8406AFFD6566AB8EC703BFC86" author="Iwan, D. &amp; Lobl, I. &amp; Bouchard, P. &amp; Bousquet, Y. &amp; Kaminski, M. &amp; Merkl, O &amp; Ando, K. &amp; Schawaller, W." box="[379,567,1447,1474]" pageId="1" pageNumber="576" pagination="1 - 969" refId="ref3773" refString="Iwan, D., Lobl, I., Bouchard, P., Bousquet, Y., Kaminski, M., Merkl, O, Ando, K. &amp; Schawaller, W. (2020) family Tenebrionidae Latreille, 1802. In: Iwan, D. &amp; Lobl, I. (Eds.), Catalogue of Palaearctic Coleoptera. Vol. 5. Tenebrionoidea. Revised and Updated 2 nd Edition. I. Brill NV, Leiden, pp. 1 - 969. https: // doi. org / 10.1163 / 9789004434998" type="book chapter" year="2020">Iwan et al. 2020</bibRefCitation>
): 
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<emphasis id="D5E3A21B406AFFD65558B8EC7069FC86" box="[585,613,1448,1473]" italics="true" pageId="1" pageNumber="576">D.</emphasis>
(
<emphasis id="D5E3A21B406AFFD65564B8E370EFFC86" box="[629,739,1447,1473]" italics="true" pageId="1" pageNumber="576">Dendarus</emphasis>
) 
<emphasis id="D5E3A21B406AFFD655E5B8EC7152FC86" box="[756,862,1448,1473]" italics="true" pageId="1" pageNumber="576">carinatus</emphasis>
Mulsant &amp; Rey, 1854
</taxonomicName>
, 
<taxonomicName id="2097058A406AFFD65377B8EC7330FCA1" authority="Mulsant, 1854" authorityName="Mulsant. The" authorityYear="1854" class="Hexanauplia" family="Pandaridae" genus="Pandarus" kingdom="Animalia" order="Siphonostomatoida" pageId="1" pageNumber="576" phylum="Arthropoda" rank="species" species="coarcticollis" subGenus="Dendarus">
<emphasis id="D5E3A21B406AFFD65377B8EC7790FC86" box="[1126,1436,1447,1473]" italics="true" pageId="1" pageNumber="576">D. (Dendarus) coarcticollis</emphasis>
Mulsant, 1854
</taxonomicName>
, 
<taxonomicName id="2097058A406AFFD65656B8887134FCA2" authority="(Germar, 1823)" baseAuthorityName="Germar" baseAuthorityYear="1823" box="[327,824,1483,1510]" class="Insecta" family="Tenebrionidae" genus="Dendarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="576" phylum="Arthropoda" rank="species" species="dalmatinus" subGenus="Pandarinus">
<emphasis id="D5E3A21B406AFFD65656B888736FFCA2" box="[327,355,1484,1509]" italics="true" pageId="1" pageNumber="576">D.</emphasis>
(
<emphasis id="D5E3A21B406AFFD65662B88F73FAFCA2" box="[371,502,1483,1509]" italics="true" pageId="1" pageNumber="576">Pandarinus</emphasis>
) 
<emphasis id="D5E3A21B406AFFD65517B88F708EFCA2" box="[518,642,1483,1509]" italics="true" pageId="1" pageNumber="576">dalmatinus</emphasis>
(Germar, 1823)
</taxonomicName>
, 
<taxonomicName id="2097058A406AFFD65455B88876FAFCA2" authority="Soldati, 2012" authorityName="Soldati" authorityYear="2012" box="[836,1270,1483,1510]" class="Insecta" family="Tenebrionidae" genus="Dendarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="576" phylum="Arthropoda" rank="species" species="peslieri" subGenus="Pandarinus">
<emphasis id="D5E3A21B406AFFD65455B888716CFCA2" box="[836,864,1484,1509]" italics="true" pageId="1" pageNumber="576">D.</emphasis>
(
<emphasis id="D5E3A21B406AFFD65461B88F71FFFCA2" box="[880,1011,1483,1509]" italics="true" pageId="1" pageNumber="576">Pandarinus</emphasis>
) 
<emphasis id="D5E3A21B406AFFD65312B88F765BFCA2" box="[1027,1111,1483,1509]" italics="true" pageId="1" pageNumber="576">peslieri</emphasis>
<bibRefCitation id="830603F8406AFFD6534FB88F76FAFCA2" author="Soldati, F." box="[1118,1270,1483,1509]" pageId="1" pageNumber="576" pagination="52 - 59" refId="ref4527" refString="Soldati, F. (2012) A new species of the genus Dendarus Dejean, 1821 from Greece. Une nouvelle espece du genre Dendarus Dejean, 1821 decrite de Grece (Coleoptera, Tenebrionidae). R. A. R. E., 21, 52 - 59." type="journal article" year="2012">Soldati, 2012</bibRefCitation>
</taxonomicName>
, and 
<taxonomicName id="2097058A406AFFD65221B8887049FF4D" authority="(Mulsant &amp; Rey, 1854)" baseAuthorityName="Mulsant &amp; Rey" baseAuthorityYear="1854" class="Insecta" family="Tenebrionidae" genus="Dendarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="576" phylum="Arthropoda" rank="species" species="lugens" subGenus="Paroderus">
<emphasis id="D5E3A21B406AFFD65221B8887740FCA2" box="[1328,1356,1484,1509]" italics="true" pageId="1" pageNumber="576">D.</emphasis>
(
<emphasis id="D5E3A21B406AFFD6524DB88872D9FF4E" italics="true" pageId="1" pageNumber="576">Paroderus</emphasis>
) 
<emphasis id="D5E3A21B406AFFD657F6B8AB733CFF4E" box="[231,304,1519,1545]" italics="true" pageId="1" pageNumber="576">lugens</emphasis>
(Mulsant &amp; Rey, 1854)
</taxonomicName>
. Italian records of 
<taxonomicName id="2097058A406AFFD6540DB8B476DEFF4E" authority="Brulle, 1832" authorityName="Brulle" authorityYear="1832" box="[796,1234,1519,1546]" class="Insecta" family="Tenebrionidae" genus="Dendarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="576" phylum="Arthropoda" rank="species" species="coelatus" subGenus="Pandarinus">
<emphasis id="D5E3A21B406AFFD6540DB8B47134FF4E" box="[796,824,1520,1545]" italics="true" pageId="1" pageNumber="576">D.</emphasis>
(
<emphasis id="D5E3A21B406AFFD6545BB8AB71C1FF4E" box="[842,973,1519,1545]" italics="true" pageId="1" pageNumber="576">Pandarinus</emphasis>
) 
<emphasis id="D5E3A21B406AFFD654CFB8AB7637FF4E" box="[990,1083,1519,1545]" italics="true" pageId="1" pageNumber="576">coelatus</emphasis>
Brullé, 1832
</taxonomicName>
are artificial, and due to misinterpretation of this species with 
<taxonomicName id="2097058A406AFFD65587BB50763EFF6A" authority="(Aliquo &amp; Soldati 2014)" baseAuthorityName="Aliquo &amp; Soldati" baseAuthorityYear="2014" box="[662,1074,1555,1581]" class="Insecta" family="Tenebrionidae" genus="Dendarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="576" phylum="Arthropoda" rank="species" species="peslieri">
<emphasis id="D5E3A21B406AFFD65587BB507102FF6A" box="[662,782,1555,1581]" italics="true" pageId="1" pageNumber="576">D. peslieri</emphasis>
(
<bibRefCitation id="830603F8406AFFD6540FBB577627FF6A" author="Aliquo, V. &amp; Soldati, F." box="[798,1067,1555,1581]" pageId="1" pageNumber="576" pagination="429 - 442" refId="ref2812" refString="Aliquo, V. &amp; Soldati, F. (2014) Updating the CD-ROM on Coleoptera Tenebrionidae of Italy and the check-list of the same family. Biodiversity Journal, 5, 429 - 442." type="journal article" year="2014">Aliquò &amp; Soldati 2014</bibRefCitation>
)
</taxonomicName>
. The investigation of the newly designated 
<typeStatus id="382CC0AB406AFFD65604BB737370FF16" box="[277,380,1591,1617]" pageId="1" pageNumber="576" type="lectotype">lectotype</typeStatus>
of 
<taxonomicName id="2097058A406AFFD6568FBB7C705AFF15" authority="(Rossi)" baseAuthorityName="Rossi" baseAuthorityYear="1790" box="[414,598,1591,1618]" class="Insecta" family="Tenebrionidae" genus="Helops" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="576" phylum="Arthropoda" rank="species" species="tristis">
<emphasis id="D5E3A21B406AFFD6568FBB7C73F3FF16" box="[414,511,1592,1617]" italics="true" pageId="1" pageNumber="576">D. tristis</emphasis>
(Rossi)
</taxonomicName>
clearly indicates that it is not synonymous with 
<taxonomicName id="2097058A406AFFD65378BB7C7336FF31" authority="Mulsant, 1854" authorityName="Mulsant. The" authorityYear="1854" class="Hexanauplia" family="Pandaridae" genus="Pandarus" kingdom="Animalia" order="Siphonostomatoida" pageId="1" pageNumber="576" phylum="Arthropoda" rank="species" species="coarcticollis" subGenus="Dendarus">
<emphasis id="D5E3A21B406AFFD65378BB7C7689FF16" box="[1129,1157,1592,1617]" italics="true" pageId="1" pageNumber="576">D.</emphasis>
(
<emphasis id="D5E3A21B406AFFD65385BB73770EFF16" box="[1172,1282,1591,1617]" italics="true" pageId="1" pageNumber="576">Dendarus</emphasis>
) 
<emphasis id="D5E3A21B406AFFD65200BB737790FF16" box="[1297,1436,1591,1617]" italics="true" pageId="1" pageNumber="576">coarcticollis</emphasis>
Mulsant, 1854
</taxonomicName>
(
<figureCitation id="7FAC628C406AFFD6565BBB1F7387FF32" box="[330,395,1627,1653]" captionStart="FIGURE 1" captionStartId="2.[151,250,1073,1097]" captionTargetBox="[151,1436,181,1049]" captionTargetId="figure-20@2.[151,1436,181,1049]" captionTargetPageId="2" captionText="FIGURE 1. Habitus photographs of selected Dendarus species. Lectotype (A) and paralectotype (B) of Helops tristis, lectotype (C) of Pandarus coarcticollis (photos by MNHN/Christophe Rivier)." figureDoi="http://doi.org/10.5281/zenodo.5556809" httpUri="https://zenodo.org/record/5556809/files/figure.png" pageId="1" pageNumber="576">Fig. 1</figureCitation>
). In fact, it is fully convergent with the taxonomic concept of 
<taxonomicName id="2097058A406AFFD6535FBB1876B6FF32" baseAuthorityName="Mulsant &amp; Rey" baseAuthorityYear="1854" box="[1102,1210,1627,1653]" class="Insecta" family="Tenebrionidae" genus="Dendarus" higherTaxonomySource="GBIF" kingdom="Animalia" order="Coleoptera" pageId="1" pageNumber="576" phylum="Arthropoda" rank="species" species="lugens">
<emphasis id="D5E3A21B406AFFD6535FBB1876B6FF32" box="[1102,1210,1627,1653]" italics="true" pageId="1" pageNumber="576">D. lugens</emphasis>
</taxonomicName>
, and as a result it is hereby considered as a senior synonym of this species.
</paragraph>
</subSubSection>
</treatment>
</document>