treatments-xml/data/65/09/87/650987BF272CFFFFCBEAFA29D5E5FD37.xml

<document id="A06459ED2308C473FA6016D80824F5DC" ID-CLB-Dataset="27571" ID-DOI="10.11646/zootaxa.4554.2.3" ID-GBIF-Dataset="e5164349-3dfb-491f-ae2a-af56edc829f0" ID-ISSN="1175-5326" ID-Zenodo-Dep="2623746" ID-ZooBank="81247C5A-18DD-4B13-B40E-EE0E0AB0D15C" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1554271384300" checkinUser="plazi" docAuthor="Jiménez-Arcos, Víctor H., Calzada-Arciniega, Rafael Alejandro, Alfaro-Juantorena, Liz A., Vázquez-Reyes, Leopoldo D., Blair, Christopher &amp; Parra-Olea, Gabriela" docDate="2019" docId="650987BF272CFFFFCBEAFA29D5E5FD37" docLanguage="en" docName="zootaxa.4554.2.3.pdf" docOrigin="Zootaxa 4554 (2)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Charadrahyla sakbah Jiménez-Arcos &amp; Calzada-Arciniega &amp; Alfaro-Juantorena &amp; Vázquez-Reyes &amp; Blair &amp; Parra-Olea 2019, sp. nov." docType="treatment" docVersion="8" lastPageNumber="380" masterDocId="9930FFC7272EFFF6CB7DFFB1D17FFF9F" masterDocTitle="A new species of Charadrahyla (Anura: Hylidae) from the cloud forest of western Oaxaca, Mexico" masterLastPageNumber="385" masterPageNumber="371" pageNumber="373" updateTime="1698716275540" updateUser="ExternalLinkService">
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<mods:title id="B6C3D191155C7073EC85B41BC89DF304">A new species of Charadrahyla (Anura: Hylidae) from the cloud forest of western Oaxaca, Mexico</mods:title>
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<mods:namePart id="C1D57E0E4B99067853E9C09AD05253ED">Blair, Christopher</mods:namePart>
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<heading id="B65781C5272CFFF4CBEAFA29D37DFA2C" bold="true" box="[151,514,1432,1459]" fontSize="11" level="1" pageId="2" pageNumber="373" reason="1">
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<taxonomicName id="2AA04D2A272CFFF4CBEAFA29D0E2FA2D" ID-CoL="TMW2" authority="Jiménez-Arcos &amp; Calzada-Arciniega &amp; Alfaro-Juantorena &amp; Vázquez-Reyes &amp; Blair &amp; Parra-Olea, 2019" authorityName="Jiménez-Arcos &amp; Calzada-Arciniega &amp; Alfaro-Juantorena &amp; Vázquez-Reyes &amp; Blair &amp; Parra-Olea" authorityYear="2019" box="[151,413,1432,1458]" class="Amphibia" family="Hylidae" genus="Charadrahyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="2" pageNumber="373" phylum="Chordata" rank="species" species="sakbah" status="sp. nov.">
<emphasis id="DFD4EABB272CFFF4CBEAFA29D0E2FA2D" bold="true" box="[151,413,1432,1458]" italics="true" pageId="2" pageNumber="373">Charadrahyla sakbah</emphasis>
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<taxonomicNameLabel id="C4E757C0272CFFF4CAD8FA28D37DFA2C" box="[421,514,1433,1459]" pageId="2" pageNumber="373" rank="species">sp. nov.</taxonomicNameLabel>
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<paragraph id="ED1F36A9272CFFF4CBEAFA0AD2A4FA4B" blockId="2.[151,987,1432,1528]" box="[151,987,1467,1492]" pageId="2" pageNumber="373">Jiménez-Arcos, Calzada-Arciniega, Alfaro-Juantorena, Blair &amp; Parra-Olea</paragraph>
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<subSubSection id="A5BA6522272CFFF4CBEAFA6ED230FA67" box="[151,847,1503,1528]" pageId="2" pageNumber="373" type="description">
<paragraph id="ED1F36A9272CFFF4CBEAFA6ED230FA67" blockId="2.[151,987,1432,1528]" box="[151,847,1503,1528]" pageId="2" pageNumber="373">Mixteca Cloud-forest Treefrog, Rana arborícola de la Mixteca</paragraph>
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<paragraph id="ED1F36A9272CFFF4CBEAF996D470F918" blockId="2.[151,1437,1575,1996]" pageId="2" pageNumber="373">
<materialsCitation id="5DC83CF4272CFFF4CBEAF996D470F918" ID-GBIF-Occurrence="2234214348" collectingDate="2018-01-26" collectionCode="IBH" collectorName="Victor H. Jimenez Arcos &amp; Liz A. Alfaro Juantorena." country="Mexico" county="Santa Maria Yucuhiti" latitude="17.074902" location="Adult" longLatPrecision="1" longitude="-97.834755" municipality="Progreso" pageId="2" pageNumber="373" specimenCount="1" stateProvince="Oaxaca" typeStatus="holotype">
<emphasis id="DFD4EABB272CFFF4CBEAF996D074F9DF" bold="true" box="[151,267,1575,1600]" pageId="2" pageNumber="373">
<typeStatus id="321B880B272CFFF4CBEAF996D079F9DF" box="[151,262,1575,1600]" pageId="2" pageNumber="373" type="holotype">Holotype</typeStatus>
.
</emphasis>
(
<figureCitation id="759B2A2C272CFFF4CA60F996D009F9DF" box="[285,374,1575,1600]" captionStart="FIGURE 2" captionStartId="3.[151,250,1510,1532]" captionTargetBox="[181,1401,221,1464]" captionTargetId="figure@3.[150,1436,193,1489]" captionTargetPageId="3" captionText="FIGURE 2. Specimens in life of C haradrahyla sakbah. (A) male holotype (IBH 30989). (B) male paratype (IBH 30990). (C) Female paratype (IBH 30992). (D) Female paratype (IBH 30993). (E) Female paratype (IBH 30994). (F) Enlarged vent tubercles in ventral view of C. sakbah of male holotype (IBH 30989) in life." httpUri="https://zenodo.org/record/2623750/files/figure.png" pageId="2" pageNumber="373">Fig. 2A</figureCitation>
in life; 
<figureCitation id="759B2A2C272CFFF4CAA8F996D32EF9A0" box="[469,593,1575,1600]" captionStart="FIGURE 4" captionStartId="6.[151,250,1923,1945]" captionTargetBox="[152,1435,373,1901]" captionTargetId="figure@6.[152,1435,373,1901]" captionTargetPageId="6" captionText="FIGURE 4. Dorsal (A) and ventral view (B) of the preserved holotype of Charadrahyla sakbah. (C) Ventral aspect of hand and (D) foot of male holotype (IBH 30989)." httpUri="https://zenodo.org/record/2623754/files/figure.png" pageId="2" pageNumber="373">Fig. 4A, B</figureCitation>
in preservative). 
<collectionCode id="8BB1AE6C272CFFF4C861F999D233F9A0" box="[796,844,1576,1599]" country="Mexico" httpUri="http://grbio.org/cool/p644-mj54" name="Universidad Nacional Autonoma de Mexico, Instituto de Biologia" pageId="2" pageNumber="373">IBH</collectionCode>
30989. 
<location id="E87F6072272CFFF4C8D1F996D293F9DF" LSID="urn:lsid:plazi:treatment:650987BF272CFFFFCBEAFA29D5E5FD37:E87F6072272CFFF4C8D1F996D293F9DF" box="[940,1004,1575,1600]" country="Mexico" county="Santa Maria Yucuhiti" latitude="17.074902" longLatPrecision="1" longitude="-97.834755" municipality="Progreso" name="Adult" pageId="2" pageNumber="373" stateProvince="Oaxaca">Adult</location>
male. 
<location id="E87F6072272CFFF4CF40F999D5D4F9DF" LSID="urn:lsid:plazi:treatment:650987BF272CFFFFCBEAFA29D5E5FD37:E87F6072272CFFF4CF40F999D5D4F9DF" box="[1085,1195,1575,1600]" country="Mexico" county="Santa Maria Yucuhiti" latitude="17.074902" longLatPrecision="1" longitude="-97.834755" municipality="Progreso" name="Rio Chite" pageId="2" pageNumber="373" stateProvince="Oaxaca">Río Chite</location>
ku'e (Río de las Mil Cascadas), 
<location id="E87F6072272CFFF4CA62F9FDD0ADF9FC" LSID="urn:lsid:plazi:treatment:650987BF272CFFFFCBEAFA29D5E5FD37:E87F6072272CFFF4CA62F9FDD0ADF9FC" box="[287,466,1611,1636]" country="Mexico" county="Santa Maria Yucuhiti" latitude="17.074902" longLatPrecision="1" longitude="-97.834755" municipality="Progreso" name="San Isidro Paz" pageId="2" pageNumber="373" stateProvince="Oaxaca">San Isidro Paz</location>
y 
<collectingMunicipality id="0D7BACD3272CFFF4C97DF9FDD319F9FB" box="[512,614,1612,1636]" pageId="2" pageNumber="373">Progreso</collectingMunicipality>
, 
<collectingCounty id="047E4E25272CFFF4C907F9FDD202F9FB" box="[634,893,1611,1636]" pageId="2" pageNumber="373">Santa Maria Yucuhiti</collectingCounty>
, 
<collectingRegion id="2F64F84B272CFFF4C8ECF9FDD297F9FB" box="[913,1000,1612,1636]" country="Mexico" name="Oaxaca" pageId="2" pageNumber="373">Oaxaca</collectingRegion>
, 
<collectingCountry id="95B77639272CFFF4C886F9FAD52AF9FB" box="[1019,1109,1611,1636]" name="Mexico" pageId="2" pageNumber="373">Mexico</collectingCountry>
. 1390 mts. 
<geoCoordinate id="8894506E272CFFF4CF92F9FDD4E7F9FC" box="[1263,1432,1612,1636]" direction="north" orientation="latitude" pageId="2" pageNumber="373" precision="1" value="17.074902">17.074902° N</geoCoordinate>
, 
<geoCoordinate id="8894506E272CFFF4CBEAF9DED044F918" box="[151,315,1647,1672]" direction="west" orientation="longitude" pageId="2" pageNumber="373" precision="1" value="-97.834755">97.834758° W</geoCoordinate>
, 
<date id="991E1069272CFFF4CA3BF9DED37CF917" box="[326,515,1647,1672]" pageId="2" pageNumber="373" value="2018-01-26">
<collectingDate id="895AE981272CFFF4CA3BF9DED37CF917" box="[326,515,1647,1672]" pageId="2" pageNumber="373" value="2018-01-26">26 January 2018</collectingDate>
</date>
. Collected by 
<collectorName id="4055537F272CFFF4C9DFF9C1D2C6F917" box="[674,953,1647,1672]" pageId="2" pageNumber="373">Víctor H. Jiménez Arcos</collectorName>
and 
<collectorName id="4055537F272CFFF4C892F9DED470F918" box="[1007,1295,1647,1672]" pageId="2" pageNumber="373">Liz A. Alfaro Juantorena.</collectorName>
</materialsCitation>
</paragraph>
<paragraph id="ED1F36A9272CFFF4CBBAF922D26CF96B" blockId="2.[151,1437,1575,1996]" pageId="2" pageNumber="373">
<materialsCitation id="5DC83CF4272CFFF4CBBAF922D348F933" ID-GBIF-Occurrence="2234214346" box="[199,567,1683,1708]" collectionCode="IBH" pageId="2" pageNumber="373" specimenCount="1" typeStatus="paratype">
<emphasis id="DFD4EABB272CFFF4CBBAF922D039F933" bold="true" box="[199,326,1683,1708]" pageId="2" pageNumber="373">
<typeStatus id="321B880B272CFFF4CBBAF922D03EF933" box="[199,321,1683,1708]" pageId="2" pageNumber="373" type="paratype">Paratypes</typeStatus>
.
</emphasis>
<collectionCode id="8BB1AE6C272CFFF4CA31F925D003F934" box="[332,380,1684,1707]" country="Mexico" httpUri="http://grbio.org/cool/p644-mj54" name="Universidad Nacional Autonoma de Mexico, Instituto de Biologia" pageId="2" pageNumber="373">IBH</collectionCode>
30990 (
<figureCitation id="759B2A2C272CFFF4CAAAF922D351F933" box="[471,558,1683,1708]" captionStart="FIGURE 2" captionStartId="3.[151,250,1510,1532]" captionTargetBox="[181,1401,221,1464]" captionTargetId="figure@3.[150,1436,193,1489]" captionTargetPageId="3" captionText="FIGURE 2. Specimens in life of C haradrahyla sakbah. (A) male holotype (IBH 30989). (B) male paratype (IBH 30990). (C) Female paratype (IBH 30992). (D) Female paratype (IBH 30993). (E) Female paratype (IBH 30994). (F) Enlarged vent tubercles in ventral view of C. sakbah of male holotype (IBH 30989) in life." httpUri="https://zenodo.org/record/2623750/files/figure.png" pageId="2" pageNumber="373">Fig. 2B</figureCitation>
)
</materialsCitation>
, 
<materialsCitation id="5DC83CF4272CFFF4C93FF925D26FF96B" ID-GBIF-Occurrence="2234214350" collectingDate="2018-01-26" collectionCode="IBH" collectorName="Victor H. Jimenez-Arcos &amp; Rafael Alejandro Calzada Arciniega" pageId="2" pageNumber="373" specimenCount="1" typeStatus="holotype">
<collectionCode id="8BB1AE6C272CFFF4C93FF925D30DF934" box="[578,626,1684,1707]" country="Mexico" httpUri="http://grbio.org/cool/p644-mj54" name="Universidad Nacional Autonoma de Mexico, Instituto de Biologia" pageId="2" pageNumber="373">IBH</collectionCode>
30991. Adult males. 
<collectionCode id="8BB1AE6C272CFFF4C81CF925D2EEF934" box="[865,913,1684,1707]" country="Mexico" httpUri="http://grbio.org/cool/p644-mj54" name="Universidad Nacional Autonoma de Mexico, Instituto de Biologia" pageId="2" pageNumber="373">IBH</collectionCode>
30992, 
<collectionCode id="8BB1AE6C272CFFF4C896F925D564F934" box="[1003,1051,1684,1707]" country="Mexico" httpUri="http://grbio.org/cool/p644-mj54" name="Universidad Nacional Autonoma de Mexico, Instituto de Biologia" pageId="2" pageNumber="373">IBH</collectionCode>
30993, 
<collectionCode id="8BB1AE6C272CFFF4CF08F925D5DAF934" box="[1141,1189,1684,1707]" country="Mexico" httpUri="http://grbio.org/cool/p644-mj54" name="Universidad Nacional Autonoma de Mexico, Instituto de Biologia" pageId="2" pageNumber="373">IBH</collectionCode>
30994. Adult females (
<figureCitation id="759B2A2C272CFFF4CBE2F906D035F94F" box="[159,330,1719,1744]" captionStart="FIGURE 2" captionStartId="3.[151,250,1510,1532]" captionTargetBox="[181,1401,221,1464]" captionTargetId="figure@3.[150,1436,193,1489]" captionTargetPageId="3" captionText="FIGURE 2. Specimens in life of C haradrahyla sakbah. (A) male holotype (IBH 30989). (B) male paratype (IBH 30990). (C) Female paratype (IBH 30992). (D) Female paratype (IBH 30993). (E) Female paratype (IBH 30994). (F) Enlarged vent tubercles in ventral view of C. sakbah of male holotype (IBH 30989) in life." httpUri="https://zenodo.org/record/2623750/files/figure.png" pageId="2" pageNumber="373">Fig 2 C, D, E</figureCitation>
). All bearing identical locality data as the 
<typeStatus id="321B880B272CFFF4C83AF906D2D2F94F" box="[839,941,1719,1744]" pageId="2" pageNumber="373" type="holotype">holotype</typeStatus>
. 
<date id="991E1069272CFFF4C8C6F906D5FEF94F" box="[955,1153,1719,1744]" pageId="2" pageNumber="373" value="2018-01-26">
<collectingDate id="895AE981272CFFF4C8C6F906D5FEF94F" box="[955,1153,1719,1744]" pageId="2" pageNumber="373" value="2018-01-26">26 January 2018</collectingDate>
</date>
. Collected by 
<collectorName id="4055537F272CFFF4CE53F909D040F96B" pageId="2" pageNumber="373">Víctor H. Jiménez-Arcos</collectorName>
and 
<collectorName id="4055537F272CFFF4CA08F96AD26FF96B" box="[373,784,1755,1780]" pageId="2" pageNumber="373">Rafael Alejandro Calzada Arciniega</collectorName>
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A5BA6522272CFFF3CBBAF94ED0EAFE32" lastPageId="5" lastPageNumber="376" pageId="2" pageNumber="373" type="diagnosis">
<paragraph id="ED1F36A9272CFFF4CBBAF94ED414F853" blockId="2.[151,1437,1575,1996]" pageId="2" pageNumber="373">
<emphasis id="DFD4EABB272CFFF4CBBAF94ED03EF887" bold="true" box="[199,321,1791,1816]" pageId="2" pageNumber="373">Diagnosis.</emphasis>
A large member of the genus 
<taxonomicName id="2AA04D2A272CFFF4C9DCF94ED23FF887" authorityName="Faivovich, Haddad, Garcia, Frost, Campbell &amp; Wheeler" authorityYear="2005" box="[673,832,1791,1816]" class="Amphibia" family="Hylidae" genus="Charadrahyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="2" pageNumber="373" phylum="Chordata" rank="genus">
<emphasis id="DFD4EABB272CFFF4C9DCF94ED23FF887" box="[673,832,1791,1816]" italics="true" pageId="2" pageNumber="373">Charadrahyla</emphasis>
</taxonomicName>
with females showing larger SVL (mean 
<quantity id="2A589B4C272CFFF4CE5CF8B1D4E7F888" box="[1313,1432,1792,1816]" metricMagnitude="-2" metricUnit="m" metricValue="8.404" pageId="2" pageNumber="373" unit="mm" value="84.04">84.04 mm</quantity>
; range 
<quantity id="2A589B4C272CFFF4CB9CF892D0D9F8A3" box="[225,422,1827,1852]" metricMagnitude="-2" metricUnit="m" metricValue="8.345" metricValueMax="8.575" metricValueMin="8.115" pageId="2" pageNumber="373" unit="mm" value="83.45" valueMax="85.75" valueMin="81.15">81.15–85.75 mm</quantity>
) than males (
<quantity id="2A589B4C272CFFF4C937F895D3BAF8A4" box="[586,709,1828,1852]" metricMagnitude="-2" metricUnit="m" metricValue="7.007" pageId="2" pageNumber="373" unit="mm" value="70.07">70.07 mm</quantity>
; 
<quantity id="2A589B4C272CFFF4C9A8F892D2E2F8A3" box="[725,925,1827,1852]" metricMagnitude="-2" metricUnit="m" metricValue="7.056" metricValueMax="7.321" metricValueMin="6.7909999999999995" pageId="2" pageNumber="373" unit="mm" value="70.56" valueMax="73.21" valueMin="67.91">67.91–73.21 mm</quantity>
). 
<taxonomicName id="2AA04D2A272CFFF4C8C9F892D5D1F8A3" authorityName="Jiménez-Arcos &amp; Calzada-Arciniega &amp; Alfaro-Juantorena &amp; Vázquez-Reyes &amp; Blair &amp; Parra-Olea" authorityYear="2019" box="[948,1198,1827,1852]" class="Amphibia" family="Hylidae" genus="Charadrahyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="2" pageNumber="373" phylum="Chordata" rank="species" species="sakbah">
<emphasis id="DFD4EABB272CFFF4C8C9F892D5D1F8A3" box="[948,1198,1827,1852]" italics="true" pageId="2" pageNumber="373">Charadrahyla sakbah</emphasis>
</taxonomicName>
differs for all other species of the genus by the presence of the following character combination: (1) axillary membrane present in male and female adults (
<figureCitation id="759B2A2C272CFFF4CA12F8DAD094F81B" box="[367,491,1899,1924]" captionStart="FIGURE 3" captionStartId="4.[151,250,1571,1593]" captionTargetBox="[169,1418,241,1512]" captionTargetId="figure@4.[151,1435,223,1530]" captionTargetPageId="4" captionText="FIGURE 3. Morphological characters of Charadrahyla sakbah. (A) Axillary membrane of male holotype (IBH 30989). (B) Axillary membrane of female paratype (IBH 30992). (C) Retracted and (D) extended hypertrophied webbing in male holotype (IBH 30989). (E) Nuptial excrescences in dorsal view and (F) view of the palm of the hand without nuptial excrescences in Finger I in male holotype (IBH 30989)." httpUri="https://zenodo.org/record/2623752/files/figure.png" pageId="2" pageNumber="373">Fig. 3A, B</figureCitation>
), (2) hypertrophied webbings between toes I and II in adult males (
<figureCitation id="759B2A2C272CFFF4CF97F8DAD419F81B" box="[1258,1382,1899,1924]" captionStart="FIGURE 3" captionStartId="4.[151,250,1571,1593]" captionTargetBox="[169,1418,241,1512]" captionTargetId="figure@4.[151,1435,223,1530]" captionTargetPageId="4" captionText="FIGURE 3. Morphological characters of Charadrahyla sakbah. (A) Axillary membrane of male holotype (IBH 30989). (B) Axillary membrane of female paratype (IBH 30992). (C) Retracted and (D) extended hypertrophied webbing in male holotype (IBH 30989). (E) Nuptial excrescences in dorsal view and (F) view of the palm of the hand without nuptial excrescences in Finger I in male holotype (IBH 30989)." httpUri="https://zenodo.org/record/2623752/files/figure.png" pageId="2" pageNumber="373">Fig. 3C, D</figureCitation>
), (3) one conical tubercle on either side of the vent in females and males (
<figureCitation id="759B2A2C272CFFF4C8C4F83ED56EF837" box="[953,1041,1935,1960]" captionStart="FIGURE 2" captionStartId="3.[151,250,1510,1532]" captionTargetBox="[181,1401,221,1464]" captionTargetId="figure@3.[150,1436,193,1489]" captionTargetPageId="3" captionText="FIGURE 2. Specimens in life of C haradrahyla sakbah. (A) male holotype (IBH 30989). (B) male paratype (IBH 30990). (C) Female paratype (IBH 30992). (D) Female paratype (IBH 30993). (E) Female paratype (IBH 30994). (F) Enlarged vent tubercles in ventral view of C. sakbah of male holotype (IBH 30989) in life." httpUri="https://zenodo.org/record/2623750/files/figure.png" pageId="2" pageNumber="373">Fig. 2F</figureCitation>
), (4) nuptial excrescences in adult males (
<figureCitation id="759B2A2C272CFFF4CB9AF802D042F853" box="[231,317,1971,1996]" captionStart="FIGURE 3" captionStartId="4.[151,250,1571,1593]" captionTargetBox="[169,1418,241,1512]" captionTargetId="figure@4.[151,1435,223,1530]" captionTargetPageId="4" captionText="FIGURE 3. Morphological characters of Charadrahyla sakbah. (A) Axillary membrane of male holotype (IBH 30989). (B) Axillary membrane of female paratype (IBH 30992). (C) Retracted and (D) extended hypertrophied webbing in male holotype (IBH 30989). (E) Nuptial excrescences in dorsal view and (F) view of the palm of the hand without nuptial excrescences in Finger I in male holotype (IBH 30989)." httpUri="https://zenodo.org/record/2623752/files/figure.png" pageId="2" pageNumber="373">Fig. 3E</figureCitation>
), (5) vocal slits absent, and (6) females with snout dorsal profile rounded and males acuminate.
</paragraph>
<caption id="B9DF6621272DFFF5CBEAFA57D2E4F9A5" httpUri="https://zenodo.org/record/2623750/files/figure.png" pageId="3" pageNumber="374" startId="3.[151,250,1510,1532]" targetBox="[181,1401,221,1464]" targetPageId="3">
<paragraph id="ED1F36A9272DFFF5CBEAFA57D2E4F9A5" blockId="3.[151,1436,1510,1594]" pageId="3" pageNumber="374">
<emphasis id="DFD4EABB272DFFF5CBEAFA57D072FA63" bold="true" box="[151,269,1510,1532]" pageId="3" pageNumber="374">FIGURE 2</emphasis>
. Specimens in life of 
<taxonomicName id="2AA04D2A272DFFF5CA91FA56D3AFFA63" box="[492,720,1510,1532]" class="Amphibia" family="Hylidae" genus="Charadrahyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="374" phylum="Chordata" rank="subSpecies" species="haradrahyla" subSpecies="sakbah">
<emphasis id="DFD4EABB272DFFF5CA91FA56D3AFFA63" box="[492,720,1510,1532]" italics="true" pageId="3" pageNumber="374">C haradrahyla sakbah</emphasis>
</taxonomicName>
. (A) male holotype (IBH 30989). (B) male paratype (IBH 30990). (C) Female paratype (IBH 30992). (D) Female paratype (IBH 30993). (E) Female paratype (IBH 30994). (F) Enlarged vent tubercles in ventral view of 
<taxonomicName id="2AA04D2A272DFFF5CACFF994D367F9A5" authorityName="Jiménez-Arcos &amp; Calzada-Arciniega &amp; Alfaro-Juantorena &amp; Vázquez-Reyes &amp; Blair &amp; Parra-Olea" authorityYear="2019" box="[434,536,1572,1594]" class="Amphibia" family="Hylidae" genus="Charadrahyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="374" phylum="Chordata" rank="species" species="sakbah">
<emphasis id="DFD4EABB272DFFF5CACFF994D367F9A5" box="[434,536,1572,1594]" italics="true" pageId="3" pageNumber="374">C. sakbah</emphasis>
</taxonomicName>
of male holotype (IBH 30989) in life.
</paragraph>
</caption>
<paragraph id="ED1F36A9272DFFF5CBBAF9D0D027F821" blockId="3.[151,1436,1633,1982]" pageId="3" pageNumber="374">
<emphasis id="DFD4EABB272DFFF5CBBAF9D0D33BF9E5" bold="true" box="[199,580,1633,1658]" pageId="3" pageNumber="374">Comparison with other species.</emphasis>
Character states of 
<taxonomicName id="2AA04D2A272DFFF5C851F9D0D55CF9E5" authorityName="Jiménez-Arcos &amp; Calzada-Arciniega &amp; Alfaro-Juantorena &amp; Vázquez-Reyes &amp; Blair &amp; Parra-Olea" authorityYear="2019" box="[812,1059,1633,1658]" class="Amphibia" family="Hylidae" genus="Charadrahyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="374" phylum="Chordata" rank="species" species="sakbah">
<emphasis id="DFD4EABB272DFFF5C851F9D0D55CF9E5" box="[812,1059,1633,1658]" italics="true" pageId="3" pageNumber="374">Charadrahyla sakbah</emphasis>
</taxonomicName>
in parentheses. Specimens of 
<taxonomicName id="2AA04D2A272DFFF5CEFFF9D3D0BAF901" authority="(Gunther 1901)" authorityName="Gunther" authorityYear="1901" baseAuthorityName="Gunther" baseAuthorityYear="1901" class="Amphibia" family="Hylidae" genus="Charadrahyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="374" phylum="Chordata" rank="species" species="taeniopus">
<emphasis id="DFD4EABB272DFFF5CEFFF9D3D07BF902" italics="true" pageId="3" pageNumber="374">C. taeniopus</emphasis>
(
<bibRefCitation id="89314B58272DFFF5CA6AF934D0C3F901" author="Gunther, C. L. G. A." box="[279,444,1669,1694]" pageId="3" pageNumber="374" pagination="1 - 326" refId="ref7741" refString="Gunther, C. L. G. A. (1901) Reptilia and Batrachia. In: Godman, F. &amp; Salvin, O. (Ed.), Biologia Centrali-Americana, London, Zoology, 1901, pp. 1 - 326." type="journal article" year="1901">Günther 1901</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="2AA04D2A272DFFF5CAABF937D243F901" authority="(Duellman 1961)" authorityName="Duellman" authorityYear="1961" baseAuthorityName="Duellman" baseAuthorityYear="1961" box="[470,828,1669,1694]" class="Amphibia" family="Hylidae" genus="Charadrahyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="374" phylum="Chordata" rank="species" species="chaneque">
<emphasis id="DFD4EABB272DFFF5CAABF937D317F901" box="[470,616,1669,1694]" italics="true" pageId="3" pageNumber="374">C. chaneque</emphasis>
(
<bibRefCitation id="89314B58272DFFF5C906F934D24DF901" author="Duellman, W. E." box="[635,818,1669,1694]" pageId="3" pageNumber="374" pagination="1 - 5" refId="ref7236" refString="Duellman, W. E. (1961) Description of a New Species of Tree Frog from Mexico Studies of American Hylid Frogs VI. Herpetologica, 17, 1 - 5." type="journal article" year="1961">Duellman 1961</bibRefCitation>
)
</taxonomicName>
, 
<taxonomicName id="2AA04D2A272DFFF5C831F937D43EF901" authority="(Mendelson &amp; Campbell 1999)" authorityName="Mendelson &amp; Campbell" authorityYear="1999" baseAuthorityName="Mendelson &amp; Campbell" baseAuthorityYear="1999" box="[844,1345,1669,1694]" class="Amphibia" family="Hylidae" genus="Charadrahyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="374" phylum="Chordata" rank="species" species="nephila">
<emphasis id="DFD4EABB272DFFF5C831F937D2B8F901" box="[844,967,1669,1694]" italics="true" pageId="3" pageNumber="374">C. nephila</emphasis>
(
<bibRefCitation id="89314B58272DFFF5C8A7F934D445F901" author="Mendelson, J. R. III &amp; Campbell, J. A." box="[986,1338,1669,1694]" pageId="3" pageNumber="374" pagination="80" refId="ref8096" refString="Mendelson, J. R. III &amp; Campbell, J. A. (1999) The taxonomic status of populations referred to Hyla chaneque in southern Mexico, with the description of a new treefrog from Oaxaca. Journal of Herpetology, 33, 80 ‾ 86. https: // doi. org / 10.2307 / 1565545" type="journal article" year="1999">Mendelson &amp; Campbell 1999</bibRefCitation>
)
</taxonomicName>
and 
<taxonomicName id="2AA04D2A272DFFF5CEFFF937D564F95D" authority="(Canseco-Marquez, Ramirez-Gonzalez &amp; Gonzalez-Bernal 2017)" authorityName="Canseco-Marquez, Ramirez-Gonzalez &amp; Gonzalez-Bernal" authorityYear="2017" baseAuthorityName="Canseco-Marquez, Ramirez-Gonzalez &amp; Gonzalez-Bernal" baseAuthorityYear="2017" class="Amphibia" family="Hylidae" genus="Charadrahyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="374" phylum="Chordata" rank="species" species="esperancensis">
<emphasis id="DFD4EABB272DFFF5CEFFF937D04BF95E" italics="true" pageId="3" pageNumber="374">C. esperancensis</emphasis>
(
<bibRefCitation id="89314B58272DFFF5CA3FF91BD56CF95D" author="Canseco-Marquez, L. &amp; Ramirez-Gonzalez, C. G. &amp; Gonzalez-Bernal, E." box="[322,1043,1705,1730]" pageId="3" pageNumber="374" pagination="64 - 72" refId="ref6989" refString="Canseco-Marquez, L., Ramirez-Gonzalez, C. G. &amp; Gonzalez-Bernal, E. (2017) Discovery of another new species of Charadrahyla (Anura, Hylidae) from the cloud forest of northern Oaxaca, Mexico. Zootaxa, 4329 (1), 64 - 72. https: // doi. org / 10.11646 / zootaxa. 4329.1.2" type="journal article" year="2017">Canseco-Márquez, Ramírez-González &amp; González-Bernal 2017</bibRefCitation>
)
</taxonomicName>
lack axillary membrane (present), lack conical tubercles in both sides of the vent (present) and have normal webbings between toes I and II in adult males (hypertrophied). Moreover, these species are distributed in the Atlantic versant. 
<taxonomicName id="2AA04D2A272DFFF5CF20F940D40FF896" baseAuthorityName="Gunther" baseAuthorityYear="1901" box="[1117,1392,1777,1802]" class="Amphibia" family="Hylidae" genus="Charadrahyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="374" phylum="Chordata" rank="species" species="taeniopus">
<emphasis id="DFD4EABB272DFFF5CF20F940D40FF896" box="[1117,1392,1777,1802]" italics="true" pageId="3" pageNumber="374">Charadrahyla taeniopus</emphasis>
</taxonomicName>
has been recorded in localities in the states of 
<collectingRegion id="2F64F84B272DFFF5C9D5F8A4D27AF8B1" box="[680,773,1813,1838]" country="Mexico" name="Hidalgo" pageId="3" pageNumber="374">Hidalgo</collectingRegion>
, 
<collectingRegion id="2F64F84B272DFFF5C86AF8A7D201F8B2" box="[791,894,1814,1837]" country="Mexico" name="Veracruz de Ignacio de la Llave" pageId="3" pageNumber="374">Veracruz</collectingRegion>
, and 
<collectingRegion id="2F64F84B272DFFF5C8BAF8A4D56CF8B1" box="[967,1043,1813,1838]" country="Mexico" name="Puebla" pageId="3" pageNumber="374">Puebla</collectingRegion>
(
<bibRefCitation id="89314B58272DFFF5CF55F8A4D59CF8B1" author="Duellman, W. E." box="[1064,1251,1813,1838]" pageId="3" pageNumber="374" refId="ref7389" refString="Duellman, W. E. (2001) The Hylid Frogs of Middle America, Society for the Study of Amphibians and Reptiles, Ithaca, New York, 1170 pp." type="book" year="2001">Duellman 2001</bibRefCitation>
). 
<taxonomicName id="2AA04D2A272DFFF5CF80F8A4D07CF8CD" baseAuthorityName="Duellman" baseAuthorityYear="1961" class="Amphibia" family="Hylidae" genus="Charadrahyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="374" phylum="Chordata" rank="species" species="chaneque">
<emphasis id="DFD4EABB272DFFF5CF80F8A4D07CF8CD" italics="true" pageId="3" pageNumber="374">Charadrahyla chaneque</emphasis>
</taxonomicName>
is restricted to the east of the Isthmus of Tehuantepec in 
<collectingRegion id="2F64F84B272DFFF5C8B2F88BD55CF8CD" box="[975,1059,1850,1874]" country="Mexico" name="Oaxaca" pageId="3" pageNumber="374">Oaxaca</collectingRegion>
and 
<collectingRegion id="2F64F84B272DFFF5CF1AF888D5BEF8CD" box="[1127,1217,1849,1874]" country="Mexico" name="Chiapas" pageId="3" pageNumber="374">Chiapas</collectingRegion>
(
<bibRefCitation id="89314B58272DFFF5CFABF888D4EFF8CD" author="Duellman, W. E." box="[1238,1424,1849,1874]" pageId="3" pageNumber="374" refId="ref7389" refString="Duellman, W. E. (2001) The Hylid Frogs of Middle America, Society for the Study of Amphibians and Reptiles, Ithaca, New York, 1170 pp." type="book" year="2001">Duellman 2001</bibRefCitation>
). 
<taxonomicName id="2AA04D2A272DFFF5CBEAF8ECD0EEF8E9" baseAuthorityName="Mendelson &amp; Campbell" baseAuthorityYear="1999" box="[151,401,1885,1910]" class="Amphibia" family="Hylidae" genus="Charadrahyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="374" phylum="Chordata" rank="species" species="nephila">
<emphasis id="DFD4EABB272DFFF5CBEAF8ECD0EEF8E9" box="[151,401,1885,1910]" italics="true" pageId="3" pageNumber="374">Charadrahyla nephila</emphasis>
</taxonomicName>
is recorded in the Sierra de Juarez and Sierra Mixe (
<bibRefCitation id="89314B58272DFFF5C8A6F8ECD452F8E9" author="Mendelson, J. R. III &amp; Campbell, J. A." box="[987,1325,1885,1910]" pageId="3" pageNumber="374" pagination="80" refId="ref8096" refString="Mendelson, J. R. III &amp; Campbell, J. A. (1999) The taxonomic status of populations referred to Hyla chaneque in southern Mexico, with the description of a new treefrog from Oaxaca. Journal of Herpetology, 33, 80 ‾ 86. https: // doi. org / 10.2307 / 1565545" type="journal article" year="1999">Mendelson &amp; Campbell 1999</bibRefCitation>
), whereas 
<taxonomicName id="2AA04D2A272DFFF5CBEAF833D02AF806" baseAuthorityName="Canseco-Marquez, Ramirez-Gonzalez &amp; Gonzalez-Bernal" baseAuthorityYear="2017" box="[151,341,1922,1945]" class="Amphibia" family="Hylidae" genus="Charadrahyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="3" pageNumber="374" phylum="Chordata" rank="species" species="esperancensis">
<emphasis id="DFD4EABB272DFFF5CBEAF833D02AF806" box="[151,341,1922,1945]" italics="true" pageId="3" pageNumber="374">C. esperancensis</emphasis>
</taxonomicName>
is restricted to Sierra de Juarez (
<bibRefCitation id="89314B58272DFFF5C9BEF833D56CF805" author="Canseco-Marquez, L. &amp; Ramirez-Gonzalez, C. G. &amp; Gonzalez-Bernal, E." box="[707,1043,1921,1946]" pageId="3" pageNumber="374" pagination="64 - 72" refId="ref6989" refString="Canseco-Marquez, L., Ramirez-Gonzalez, C. G. &amp; Gonzalez-Bernal, E. (2017) Discovery of another new species of Charadrahyla (Anura, Hylidae) from the cloud forest of northern Oaxaca, Mexico. Zootaxa, 4329 (1), 64 - 72. https: // doi. org / 10.11646 / zootaxa. 4329.1.2" type="journal article" year="2017">
Canseco-Márquez 
<emphasis id="DFD4EABB272DFFF5C8E5F833D2AFF805" box="[920,976,1921,1946]" italics="true" pageId="3" pageNumber="374">et al.</emphasis>
2017
</bibRefCitation>
), both mountain systems located at northern 
<collectingRegion id="2F64F84B272DFFF5CB81F817D02BF821" box="[252,340,1958,1982]" country="Mexico" name="Oaxaca" pageId="3" pageNumber="374">Oaxaca</collectingRegion>
.
</paragraph>
<caption id="B9DF6621272AFFF2CBEAF992D357F90A" httpUri="https://zenodo.org/record/2623752/files/figure.png" pageId="4" pageNumber="375" startId="4.[151,250,1571,1593]" targetBox="[169,1418,241,1512]" targetPageId="4">
<paragraph id="ED1F36A9272AFFF2CBEAF992D357F90A" blockId="4.[151,1436,1571,1685]" pageId="4" pageNumber="375">
<emphasis id="DFD4EABB272AFFF2CBEAF992D071F9A7" bold="true" box="[151,270,1571,1593]" pageId="4" pageNumber="375">FIGURE 3</emphasis>
. Morphological characters of 
<taxonomicName id="2AA04D2A272AFFF2C93AF992D258F9A6" authorityName="Jiménez-Arcos &amp; Calzada-Arciniega &amp; Alfaro-Juantorena &amp; Vázquez-Reyes &amp; Blair &amp; Parra-Olea" authorityYear="2019" box="[583,807,1571,1593]" class="Amphibia" family="Hylidae" genus="Charadrahyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="4" pageNumber="375" phylum="Chordata" rank="species" species="sakbah">
<emphasis id="DFD4EABB272AFFF2C93AF992D258F9A6" box="[583,807,1571,1593]" italics="true" pageId="4" pageNumber="375">Charadrahyla sakbah</emphasis>
</taxonomicName>
. (A) Axillary membrane of male holotype (IBH 30989). (B) Axillary membrane of female paratype (IBH 30992). (C) Retracted and (D) extended hypertrophied webbing in male holotype (IBH 30989). (E) Nuptial excrescences in dorsal view and (F) view of the palm of the hand without nuptial excrescences in Finger I in male holotype (IBH 30989).
</paragraph>
</caption>
<paragraph id="ED1F36A9272AFFF3CBBAF970D0EAFE32" blockId="4.[151,1436,1729,2006]" lastBlockId="5.[151,1437,151,2013]" lastPageId="5" lastPageNumber="376" pageId="4" pageNumber="375">
The only adult female recorded in the literature of 
<taxonomicName id="2AA04D2A272AFFF2C876F973D2F3F945" box="[779,908,1730,1754]" class="Amphibia" family="Hylidae" genus="Charadrahyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="4" pageNumber="375" phylum="Chordata" rank="species" species="pinorum">
<emphasis id="DFD4EABB272AFFF2C876F973D2F3F945" box="[779,908,1730,1754]" italics="true" pageId="4" pageNumber="375">C. pinorum</emphasis>
</taxonomicName>
has a SVL of 
<quantity id="2A589B4C272AFFF2CF48F970D5E5F946" box="[1077,1178,1729,1754]" metricMagnitude="-2" metricUnit="m" metricValue="3.46" pageId="4" pageNumber="375" unit="mm" value="34.6">34.6 mm</quantity>
and adult males range from 
<quantity id="2A589B4C272AFFF2CBA9F954D0F2F962" box="[212,397,1765,1790]" metricMagnitude="-2" metricUnit="m" metricValue="3.08" metricValueMax="3.31" metricValueMin="2.85" pageId="4" pageNumber="375" unit="mm" value="30.8" valueMax="33.1" valueMin="28.5">28.5 to 33.1 mm</quantity>
(81.15–85.75 and 
<quantity id="2A589B4C272AFFF2C922F954D261F962" box="[607,798,1765,1790]" metricMagnitude="-2" metricUnit="m" metricValue="7.056" metricValueMax="7.321" metricValueMin="6.7909999999999995" pageId="4" pageNumber="375" unit="mm" value="70.56" valueMax="73.21" valueMin="67.91">67.91–73.21 mm</quantity>
females and males respectively), webbings between toes I and II normal (hypertrophied), vocal slits present (absent), male snout shape in dorsal profile rounded (acuminate), and tympanum not evident (visible). The adult females of 
<taxonomicName id="2AA04D2A272AFFF2C8C7F89FD548F8DA" box="[954,1079,1838,1861]" class="Amphibia" family="Hylidae" genus="Charadrahyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="4" pageNumber="375" phylum="Chordata" rank="species" species="juanitae">
<emphasis id="DFD4EABB272AFFF2C8C7F89FD548F8DA" box="[954,1079,1838,1861]" italics="true" pageId="4" pageNumber="375">C. juanitae</emphasis>
</taxonomicName>
have a SVL range from 37.6 to 39.8 and adult males from 
<quantity id="2A589B4C272AFFF2CAB6F8E3D3F6F8F6" box="[459,649,1873,1898]" metricMagnitude="-2" metricUnit="m" metricValue="3.175" metricValueMax="3.5799999999999996" metricValueMin="2.77" pageId="4" pageNumber="375" unit="mm" value="31.75" valueMax="35.8" valueMin="27.7">27.7 to 35.8 mm</quantity>
(81.15–85.75 and 
<quantity id="2A589B4C272AFFF2C81CF8E0D55EF8F6" box="[865,1057,1873,1898]" metricMagnitude="-2" metricUnit="m" metricValue="7.056" metricValueMax="7.321" metricValueMin="6.7909999999999995" pageId="4" pageNumber="375" unit="mm" value="70.56" valueMax="73.21" valueMin="67.91">67.91–73.21 mm</quantity>
females and males respectively), webbings between toes I and II normal (hypertrophied), nuptial excrescences underdeveloped (present in finger I and isolated patches in finger II), vocal slits present (absent), male snout shape in dorsal profile rounded (acuminate), and tympanum not evident (visible). Specimens of 
<taxonomicName id="2AA04D2A272AFFF2C806F80FD573F849" baseAuthorityName="Duellman" baseAuthorityYear="1968" box="[891,1036,1981,2006]" class="Amphibia" family="Hylidae" genus="Charadrahyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="4" pageNumber="375" phylum="Chordata" rank="species" species="altipotens">
<emphasis id="DFD4EABB272AFFF2C806F80FD573F849" box="[891,1036,1981,2006]" italics="true" pageId="4" pageNumber="375">C. altipotens</emphasis>
</taxonomicName>
present normal webbings between toes I and II (hypertrophied), absent nuptial excrescences (present), yellow belly coloration in males (white to pale brown), and adult male dorsal coloration without blotches (darker blotches). Specimens of 
<taxonomicName id="2AA04D2A272BFFF3CFC9FF0CD47BFF4B" authorityName=": Adler &amp; Dennis" authorityYear="1972" box="[1204,1284,189,212]" class="Amphibia" family="Hylidae" genus="Charadrahyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="376" phylum="Chordata" rank="species" species="trux">
<emphasis id="DFD4EABB272BFFF3CFC9FF0CD47BFF4B" box="[1204,1284,189,212]" italics="true" pageId="5" pageNumber="376">C. trux</emphasis>
</taxonomicName>
lack axillary membrane (present), conical tubercles on either side of vent absent (two conical tubercles present), vocal slits present (absent), and quadratojugals not articulating with the maxillaries (articulated) in five of nine specimens (see 
<bibRefCitation id="89314B58272BFFF3CBEAFE96D0EBFEDF" author="Adler, K. &amp; Dennis, D. M." box="[151,404,295,320]" pageId="5" pageNumber="376" pagination="1 - 19" refId="ref6564" refString="Adler, K. &amp; Dennis, D. M. (1972) New tree frogs of the genus Hyla from the cloud forests of western Guerrero, Mexico. Occasional Papers of The Museum of Natural History, University of Kansas, 7, 1 - 19." type="journal article" year="1972">Adler &amp; Dennis 1972</bibRefCitation>
). 
<taxonomicName id="2AA04D2A272BFFF3CAD7FE96D3DBFEDF" box="[426,676,295,320]" class="Amphibia" family="Hylidae" genus="Charadrahyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="376" phylum="Chordata" rank="species" species="tecuani">
<emphasis id="DFD4EABB272BFFF3CAD7FE96D3DBFEDF" box="[426,676,295,320]" italics="true" pageId="5" pageNumber="376">Charadrahyla tecuani</emphasis>
</taxonomicName>
adult males have SVL range from 
<quantity id="2A589B4C272BFFF3CF43FE96D581FEA0" box="[1086,1278,295,320]" metricMagnitude="-2" metricUnit="m" metricValue="5.515" metricValueMax="5.779999999999999" metricValueMin="5.25" pageId="5" pageNumber="376" unit="mm" value="55.15" valueMax="57.8" valueMin="52.5">52.5 to 57.8 mm</quantity>
(
<quantity id="2A589B4C272BFFF3CE72FE96D1BCFEFB" metricMagnitude="-2" metricUnit="m" metricValue="7.056" metricValueMax="7.321" metricValueMin="6.7909999999999995" pageId="5" pageNumber="376" unit="mm" value="70.56" valueMax="73.21" valueMin="67.91">67.91–73.21 mm</quantity>
adult male), axillary membrane absent (present), vocal slits present (absent), and yellow belly coloration in males (white to pale brown). 
<tableCitation id="A0220312272BFFF3CA83FEDED328FE17" box="[510,599,367,392]" captionStart="TABLE 1" captionStartId="7.[147,228,678,700]" captionText="TABLE 1. Comparison of ten morphological characters used in the diagnosis of species in the genus Charadrahyla. Data were taken from Adler and Dennis (1972), Duellman (1970, 2001), Mendelson &amp; Campbell (1999), Campbell et al. (2009), Canseco-Márquez et al. (2017) and specimens examined." pageId="5" pageNumber="376">Table 1</tableCitation>
summarizes the comparison of the main diagnostic characters among 
<taxonomicName id="2AA04D2A272BFFF3CBEAFE25D049FE32" authorityName="Faivovich, Haddad, Garcia, Frost, Campbell &amp; Wheeler" authorityYear="2005" box="[151,310,404,429]" class="Amphibia" family="Hylidae" genus="Charadrahyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="5" pageNumber="376" phylum="Chordata" rank="genus">
<emphasis id="DFD4EABB272BFFF3CBEAFE25D049FE32" box="[151,310,404,429]" italics="true" pageId="5" pageNumber="376">Charadrahyla</emphasis>
</taxonomicName>
species.
</paragraph>
</subSubSection>
<subSubSection id="A5BA6522272BFFFECBBAFE06D0ECFA60" lastPageId="8" lastPageNumber="379" pageId="5" pageNumber="376" type="description">
<paragraph id="ED1F36A9272BFFF3CBBAFE06D23CFC8A" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="376">
<emphasis id="DFD4EABB272BFFF3CBBAFE06D09AFE4F" bold="true" box="[199,485,439,464]" pageId="5" pageNumber="376">
Description of 
<typeStatus id="321B880B272BFFF3CA0BFE06D0A0FE4F" box="[374,479,439,464]" pageId="5" pageNumber="376" type="holotype">holotype</typeStatus>
.
</emphasis>
Head wider than the body; body shape slender towards the cloaca (
<figureCitation id="759B2A2C272BFFF3CFA4FE06D42DFE4F" box="[1241,1362,439,464]" captionStart="FIGURE 4" captionStartId="6.[151,250,1923,1945]" captionTargetBox="[152,1435,373,1901]" captionTargetId="figure@6.[152,1435,373,1901]" captionTargetPageId="6" captionText="FIGURE 4. Dorsal (A) and ventral view (B) of the preserved holotype of Charadrahyla sakbah. (C) Ventral aspect of hand and (D) foot of male holotype (IBH 30989)." httpUri="https://zenodo.org/record/2623754/files/figure.png" pageId="5" pageNumber="376">Fig. 4A, B</figureCitation>
), SVL 
<quantity id="2A589B4C272BFFF3CBEAFE6DD074FE6B" box="[151,267,476,500]" metricMagnitude="-2" metricUnit="m" metricValue="7.321" pageId="5" pageNumber="376" unit="mm" value="73.21">73.21 mm</quantity>
, tibia length 39.98, foot length 33.61, head length 22.35, head width 21.66, diameter of tympanum 2.85, diameter of eye 6.85, interorbital distance 7.22, eye–tympanum distance 3.31. head slightly longer than wide; lateral profile rounded with snout sloping downward posterior to the nostril; snout shape in dorsal profile acuminate; canthus rostralis distinct and angular; loreal region concave; slightly protuberant nostril oval, which are directed posterolaterally; internarial region concave; vocal slits absent. Top of the head flat; interorbital region 33.33% of head width; eye diameter 31.63% of head width. Supratympanic fold distinct, thick, extending posteroventrally from posterior margin of orbit until anterior insertion of forearm; tympanum distinct, round; tympanic annulus distinguishable, obscured by supratympanic fold dorsally; width of tympanum 44.82% diameter of eye; width of tympanum 92.75% eye–tympanum distance.
</paragraph>
<paragraph id="ED1F36A9272BFFF3CBBAFCAED45EFC72" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="376">
Axillary membranes present extending slightly beyond the first third of the forearm (more visible in life); thoracic fold absent; dermal fold on wrist present. Fingers long and slender, with slight lateral fringes, bearing large rounded terminal discs, less developed on the Finger I; relative finger lengths: I &lt;II &lt;IV &lt;III; discs on Fingers II, III, IV of similar size (
<figureCitation id="759B2A2C272BFFF3CAEFFC3DD095FC3B" box="[402,490,908,933]" captionStart="FIGURE 4" captionStartId="6.[151,250,1923,1945]" captionTargetBox="[152,1435,373,1901]" captionTargetId="figure@6.[152,1435,373,1901]" captionTargetPageId="6" captionText="FIGURE 4. Dorsal (A) and ventral view (B) of the preserved holotype of Charadrahyla sakbah. (C) Ventral aspect of hand and (D) foot of male holotype (IBH 30989)." httpUri="https://zenodo.org/record/2623754/files/figure.png" pageId="5" pageNumber="376">Fig. 4B</figureCitation>
), slightly wider than the tympanum; disc on Finger I smaller, width 62.54% width of tympanum. Subarticular tubercles large, diameter about two-third width of terminal disc on same finger, rounded, none bifid; supernumerary tubercules smaller than subarticular tubercules, rounded and distinguishable.
</paragraph>
<paragraph id="ED1F36A9272BFFF3CBBAFC46D3BEFA02" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="376">
Hand webbing formula: 
<emphasis id="DFD4EABB272BFFF3CAA5FC46D385FB8F" bold="true" box="[472,762,1015,1040]" pageId="5" pageNumber="376">
<date id="991E1069272BFFF3CAA5FC46D372FB8F" box="[472,525,1015,1040]" pageId="5" pageNumber="376">I 2-2</date>
½ 
<date id="991E1069272BFFF3C957FC46D32CFB8F" box="[554,595,1015,1040]" pageId="5" pageNumber="376">II 1</date>
¼-2 
<date id="991E1069272BFFF3C9F5FC46D3ABFB8F" box="[648,724,1015,1040]" pageId="5" pageNumber="376">III 2-1</date>
IV
</emphasis>
(
<figureCitation id="759B2A2C272BFFF3C877FC46D21DFB8F" box="[778,866,1015,1040]" captionStart="FIGURE 4" captionStartId="6.[151,250,1923,1945]" captionTargetBox="[152,1435,373,1901]" captionTargetId="figure@6.[152,1435,373,1901]" captionTargetPageId="6" captionText="FIGURE 4. Dorsal (A) and ventral view (B) of the preserved holotype of Charadrahyla sakbah. (C) Ventral aspect of hand and (D) foot of male holotype (IBH 30989)." httpUri="https://zenodo.org/record/2623754/files/figure.png" pageId="5" pageNumber="376">Fig. 4C</figureCitation>
). Tarsal fold absent; tibia length 54.61% SVL; foot length 45.91% SVL. Inner metatarsal tubercle distinct, large, ovoid, 2.1 times larger than subarticular tubercules; outer metatarsal small but distinct; subarticular tubercules distinct, elongated, elevated, diameter about one-half width of terminal disc on the same toe; supernumerary tubercles small, circular, arranged in row along axis of proximal portions of phalanges. Nuptial excrescences present, dark-colored with cone-shaped papillae (see 
<bibRefCitation id="89314B58272BFFF3CE35FB38D18AFB5B" author="Luna, M. C. &amp; McDiarmid, R. W. &amp; Faivovich, J." pageId="5" pageNumber="376" pagination="403 - 446" refId="ref7975" refString="Luna, M. C., McDiarmid, R. W. &amp; Faivovich, J. (2018) From erotic excrescences to pheromone shots: structure and diversity of nuptial pads in anurans. Biological Journal of the Linnean Society, 124, 403 - 446. https: // doi. org / 10.1093 / biolinnean / bly 048" type="journal article" year="2018">
Luna 
<emphasis id="DFD4EABB272BFFF3CEF5FB38D1CBFB5A" italics="true" pageId="5" pageNumber="376">et al.</emphasis>
2018
</bibRefCitation>
) (
<figureCitation id="759B2A2C272BFFF3CA71FB1DD01EFB5B" box="[268,353,1196,1221]" captionStart="FIGURE 3" captionStartId="4.[151,250,1571,1593]" captionTargetBox="[169,1418,241,1512]" captionTargetId="figure@4.[151,1435,223,1530]" captionTargetPageId="4" captionText="FIGURE 3. Morphological characters of Charadrahyla sakbah. (A) Axillary membrane of male holotype (IBH 30989). (B) Axillary membrane of female paratype (IBH 30992). (C) Retracted and (D) extended hypertrophied webbing in male holotype (IBH 30989). (E) Nuptial excrescences in dorsal view and (F) view of the palm of the hand without nuptial excrescences in Finger I in male holotype (IBH 30989)." httpUri="https://zenodo.org/record/2623752/files/figure.png" pageId="5" pageNumber="376">Fig. 3E</figureCitation>
) Toes long and slender, bearing rounded: I &lt;II &lt;V &lt;III &lt;IV; terminal discs smaller than discs on fingers. Foot webbing formula: 
<emphasis id="DFD4EABB272BFFF3C951FB7ED2D8FB77" bold="true" box="[556,935,1231,1256]" pageId="5" pageNumber="376">
I ¼-½ 
<date id="991E1069272BFFF3C903FB7ED265FB77" box="[638,794,1231,1256]" pageId="5" pageNumber="376">II 1-2 III 1-2</date>
½ 
<date id="991E1069272BFFF3C841FB7ED2F5FB77" box="[828,906,1231,1256]" pageId="5" pageNumber="376">IV 2-1</date>
V
</emphasis>
(
<figureCitation id="759B2A2C272BFFF3C8C4FB7ED568FB77" box="[953,1047,1231,1256]" captionStart="FIGURE 4" captionStartId="6.[151,250,1923,1945]" captionTargetBox="[152,1435,373,1901]" captionTargetId="figure@6.[152,1435,373,1901]" captionTargetPageId="6" captionText="FIGURE 4. Dorsal (A) and ventral view (B) of the preserved holotype of Charadrahyla sakbah. (C) Ventral aspect of hand and (D) foot of male holotype (IBH 30989)." httpUri="https://zenodo.org/record/2623754/files/figure.png" pageId="5" pageNumber="376">Fig. 4D</figureCitation>
). Webbings between toes I and II hypertrophied; in life the edge of the web rolled up dorsally forming a flap (
<figureCitation id="759B2A2C272BFFF3C88CFB45D537FA93" box="[1009,1096,1268,1293]" captionStart="FIGURE 3" captionStartId="4.[151,250,1571,1593]" captionTargetBox="[169,1418,241,1512]" captionTargetId="figure@4.[151,1435,223,1530]" captionTargetPageId="4" captionText="FIGURE 3. Morphological characters of Charadrahyla sakbah. (A) Axillary membrane of male holotype (IBH 30989). (B) Axillary membrane of female paratype (IBH 30992). (C) Retracted and (D) extended hypertrophied webbing in male holotype (IBH 30989). (E) Nuptial excrescences in dorsal view and (F) view of the palm of the hand without nuptial excrescences in Finger I in male holotype (IBH 30989)." httpUri="https://zenodo.org/record/2623752/files/figure.png" pageId="5" pageNumber="376">Fig. 3C</figureCitation>
), which can be extended (
<figureCitation id="759B2A2C272BFFF3CE12FB45D1C3FAAF" captionStart="FIGURE 3" captionStartId="4.[151,250,1571,1593]" captionTargetBox="[169,1418,241,1512]" captionTargetId="figure@4.[151,1435,223,1530]" captionTargetPageId="4" captionText="FIGURE 3. Morphological characters of Charadrahyla sakbah. (A) Axillary membrane of male holotype (IBH 30989). (B) Axillary membrane of female paratype (IBH 30992). (C) Retracted and (D) extended hypertrophied webbing in male holotype (IBH 30989). (E) Nuptial excrescences in dorsal view and (F) view of the palm of the hand without nuptial excrescences in Finger I in male holotype (IBH 30989)." httpUri="https://zenodo.org/record/2623752/files/figure.png" pageId="5" pageNumber="376">Fig. 3D</figureCitation>
); was extended for preservation; larger and with irregular edges than the other webbings (
<figureCitation id="759B2A2C272BFFF3CFAEFAA6D44EFAAF" box="[1235,1329,1303,1328]" captionStart="FIGURE 4" captionStartId="6.[151,250,1923,1945]" captionTargetBox="[152,1435,373,1901]" captionTargetId="figure@6.[152,1435,373,1901]" captionTargetPageId="6" captionText="FIGURE 4. Dorsal (A) and ventral view (B) of the preserved holotype of Charadrahyla sakbah. (C) Ventral aspect of hand and (D) foot of male holotype (IBH 30989)." httpUri="https://zenodo.org/record/2623754/files/figure.png" pageId="5" pageNumber="376">Fig. 4D</figureCitation>
). Cloacal opening directed posteroventrally at mid-level of thighs; conical tubercle on either side of vent, partially covered by distinctive vent sheath. Skin on dorsal surfaces smooth; skin on ventral surfaces distinctly granular; skin on flanks between forelimbs and hind limbs smooth.
</paragraph>
<paragraph id="ED1F36A9272BFFF3CBBAFA18D4E3F9B3" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="376">
Tongue cordiform, and slightly free in the back and less anteriorly. Vocal slits absent. Vomerine odontoids 6–5 on each side, left-right respectively, situated on transverse dentigerous process at anterior level of choanae, separated medially by a length slightly smaller than odontoid process; choanae subtriangular, widely separated. The quadratojugals ossified and in contact with the maxillaries (determined by radiographs; 
<bibRefCitation id="89314B58272BFFF3CFDCF9A5D4EBF9B3" author="Adler, K. &amp; Dennis, D. M." box="[1185,1428,1556,1581]" pageId="5" pageNumber="376" pagination="1 - 19" refId="ref6564" refString="Adler, K. &amp; Dennis, D. M. (1972) New tree frogs of the genus Hyla from the cloud forests of western Guerrero, Mexico. Occasional Papers of The Museum of Natural History, University of Kansas, 7, 1 - 19." type="journal article" year="1972">Adler &amp; Dennis 1972</bibRefCitation>
)
</paragraph>
<paragraph id="ED1F36A9272BFFF3CBBAF986D44CF8EF" blockId="5.[151,1437,151,2013]" pageId="5" pageNumber="376">
<emphasis id="DFD4EABB272BFFF3CBBAF986D06FF9CF" bold="true" box="[199,272,1591,1616]" pageId="5" pageNumber="376">Color.</emphasis>
In life, dorsum of body and head, including lateral surfaces of head, are dark yellowish with golden tones (
<figureCitation id="759B2A2C272BFFF3CB9CF9EDD043F9EB" box="[225,316,1628,1653]" captionStart="FIGURE 2" captionStartId="3.[151,250,1510,1532]" captionTargetBox="[181,1401,221,1464]" captionTargetId="figure@3.[150,1436,193,1489]" captionTargetPageId="3" captionText="FIGURE 2. Specimens in life of C haradrahyla sakbah. (A) male holotype (IBH 30989). (B) male paratype (IBH 30990). (C) Female paratype (IBH 30992). (D) Female paratype (IBH 30993). (E) Female paratype (IBH 30994). (F) Enlarged vent tubercles in ventral view of C. sakbah of male holotype (IBH 30989) in life." httpUri="https://zenodo.org/record/2623750/files/figure.png" pageId="5" pageNumber="376">Fig. 2A</figureCitation>
). A brown stripe begins almost near the nostrils, continues to the middle of the eye and surrounding it below, continuing through supratympanic fold. Dorsum with irregular dark brown blotch, extending up to twothirds of the back; the last third with brown spots. Dorsal surfaces of forearm dark yellowish, with distinct brown transverse bars, four in forearms; thighs yellow through the internal area towards the body up to the middle zone; the middle zone of the thighs, towards the tibia, greenish with transverse bars dark brown; discs on fingers III and IV darker that fingers. Flanks of the body yellow with three brown spots on each side. Venter of throat pale cream. Chest pale cream with smooth skin. Belly white in the middle; granular skin towards the area of the hindlimbs which are pale pink to yellowish; palpebral membrane clear on three-quarters and gray in the last quarter.
</paragraph>
<paragraph id="ED1F36A9272BFFF0CBBAF8CCD5CDFEA0" blockId="5.[151,1437,151,2013]" lastBlockId="6.[151,1436,151,320]" lastPageId="6" lastPageNumber="377" pageId="5" pageNumber="376">
In preservative, dorsum of body and head, including lateral surfaces of head, are pale gray. A dark gray mask in the lateral surfaces of head that begins almost at the tip of the snout and continues until anterior insertion of forearm. Dorsum with irregular dark gray blotch, extending up to two-thirds of the back; the last third of the back with smaller spots. Dorsal surfaces of limbs pale gray with distinct dark gray transverse bars, four on forearm, three and five (left and right) on thigh, and six on tibia (both sides); discs of fingers III and IV darker than fingers, fingers and disc I and II whitish; flanks whitish with three irregular dark gray spots in each side (
<figureCitation id="759B2A2C2728FFF0CFE2FF6ED463FF67" box="[1183,1308,223,248]" captionStart="FIGURE 4" captionStartId="6.[151,250,1923,1945]" captionTargetBox="[152,1435,373,1901]" captionTargetId="figure@6.[152,1435,373,1901]" captionTargetPageId="6" captionText="FIGURE 4. Dorsal (A) and ventral view (B) of the preserved holotype of Charadrahyla sakbah. (C) Ventral aspect of hand and (D) foot of male holotype (IBH 30989)." httpUri="https://zenodo.org/record/2623754/files/figure.png" pageId="6" pageNumber="377">Fig. 4A, B</figureCitation>
). Venter of throat whitish. Chest white with smooth skin. Belly light brown, granular. Undersurface of thighs white, granular area light brown; palpebral membrane clear on first three-quarters and gray in the last quarter.
</paragraph>
<caption id="B9DF66212728FFF0CBEAF832D355F828" httpUri="https://zenodo.org/record/2623754/files/figure.png" pageId="6" pageNumber="377" startId="6.[151,250,1923,1945]" targetBox="[152,1435,373,1901]" targetPageId="6">
<paragraph id="ED1F36A92728FFF0CBEAF832D355F828" blockId="6.[151,1435,1923,1975]" pageId="6" pageNumber="377">
<emphasis id="DFD4EABB2728FFF0CBEAF832D073F807" bold="true" box="[151,268,1923,1945]" pageId="6" pageNumber="377">FIGURE 4</emphasis>
. Dorsal (A) and ventral view (B) of the preserved holotype of 
<taxonomicName id="2AA04D2A2728FFF0C8FCF832D521F806" authorityName="Jiménez-Arcos &amp; Calzada-Arciniega &amp; Alfaro-Juantorena &amp; Vázquez-Reyes &amp; Blair &amp; Parra-Olea" authorityYear="2019" box="[897,1118,1923,1945]" class="Amphibia" family="Hylidae" genus="Charadrahyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="6" pageNumber="377" phylum="Chordata" rank="species" species="sakbah">
<emphasis id="DFD4EABB2728FFF0C8FCF832D521F806" box="[897,1118,1923,1945]" italics="true" pageId="6" pageNumber="377">Charadrahyla sakbah</emphasis>
</taxonomicName>
. (C) Ventral aspect of hand and (D) foot of male holotype (IBH 30989).
</paragraph>
</caption>
<caption id="B9DF66212729FFF1CBEEFD17D424FD46" pageId="7" pageNumber="378">
<paragraph id="ED1F36A92729FFF1CBEEFD17D424FD46" blockId="7.[147,1824,677,729]" pageId="7" pageNumber="378">
<emphasis id="DFD4EABB2729FFF1CBEEFD17D183FD23" bold="true" box="[147,252,678,700]" pageId="7" pageNumber="378">TABLE 1.</emphasis>
Comparison of ten morphological characters used in the diagnosis of species in the genus 
<taxonomicName id="2AA04D2A2729FFF1CF23FD14D599FD23" authorityName="Faivovich, Haddad, Garcia, Frost, Campbell &amp; Wheeler" authorityYear="2005" box="[1118,1254,677,700]" class="Amphibia" family="Hylidae" genus="Charadrahyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="7" pageNumber="378" phylum="Chordata" rank="genus">
<emphasis id="DFD4EABB2729FFF1CF23FD14D599FD23" box="[1118,1254,677,700]" italics="true" pageId="7" pageNumber="378">Charadrahyla</emphasis>
</taxonomicName>
. Data were taken from Adler and Dennis (1972), Duellman (1970, 2001), Mendelson &amp; Campbell (1999), Campbell 
<emphasis id="DFD4EABB2729FFF1C9C9FD73D39AFD46" box="[692,741,706,729]" italics="true" pageId="7" pageNumber="378">et al.</emphasis>
(2009), Canseco-Márquez 
<emphasis id="DFD4EABB2729FFF1C897FD73D56BFD46" box="[1002,1044,706,729]" italics="true" pageId="7" pageNumber="378">et al</emphasis>
. (2017) and specimens examined.
</paragraph>
</caption>
<paragraph id="ED1F36A92729FFF1CAD5FD4FD65DFA12" pageId="7" pageNumber="378">
<table id="9FA0C40927290009CBEEFD4FD635FA03" box="[147,1866,766,1436]" gridcols="11" gridrows="11" pageId="7" pageNumber="378">
<tr id="539034EB27290009CBEEFD4FD635FCAD" box="[147,1866,766,818]" gridrow="0" pageId="7" pageNumber="378" rowspan-0="1">
<th id="10415D9727290009CA13FD4FD086FCAD" box="[366,505,766,818]" gridcol="1" gridrow="0" pageId="7" pageNumber="378">
<taxonomicName id="2AA04D2A2729FFF1CAD5FD4FD0A8FCAD" authorityName="Jiménez-Arcos &amp; Calzada-Arciniega &amp; Alfaro-Juantorena &amp; Vázquez-Reyes &amp; Blair &amp; Parra-Olea" authorityYear="2019" class="Amphibia" family="Hylidae" genus="Charadrahyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="7" pageNumber="378" phylum="Chordata" rank="species" species="sakbah">
<emphasis id="DFD4EABB2729FFF1CAD5FD4FD0A8FCAD" bold="true" italics="true" pageId="7" pageNumber="378">C. sakbah</emphasis>
</taxonomicName>
</th>
<th id="10415D9727290009C966FD4FD3FBFCAD" box="[539,644,766,818]" gridcol="2" gridrow="0" pageId="7" pageNumber="378">
<taxonomicName id="2AA04D2A2729FFF1C939FD4FD300FCAD" baseAuthorityName="Gunther" baseAuthorityYear="1901" class="Amphibia" family="Hylidae" genus="Charadrahyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="7" pageNumber="378" phylum="Chordata" rank="species" species="taeniopus">
<emphasis id="DFD4EABB2729FFF1C939FD4FD300FCAD" bold="true" italics="true" pageId="7" pageNumber="378">C. taeniopus</emphasis>
</taxonomicName>
</th>
<th id="10415D9727290009C9D6FD4FD27EFCAD" box="[683,769,766,818]" gridcol="3" gridrow="0" pageId="7" pageNumber="378">
<taxonomicName id="2AA04D2A2729FFF1C9B6FD4FD27FFCAD" class="Amphibia" family="Hylidae" genus="Charadrahyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="7" pageNumber="378" phylum="Chordata" rank="species" species="pinorum">
<emphasis id="DFD4EABB2729FFF1C9B6FD4FD27FFCAD" bold="true" italics="true" pageId="7" pageNumber="378">C. pinorum</emphasis>
</taxonomicName>
</th>
<th id="10415D9727290009C857FD4FD2F5FCAD" box="[810,906,766,818]" gridcol="4" gridrow="0" pageId="7" pageNumber="378">
<taxonomicName id="2AA04D2A2729FFF1C832FD4FD2F5FCAD" baseAuthorityName="Duellman" baseAuthorityYear="1961" class="Amphibia" family="Hylidae" genus="Charadrahyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="7" pageNumber="378" phylum="Chordata" rank="species" species="chaneque">
<emphasis id="DFD4EABB2729FFF1C832FD4FD2F5FCAD" bold="true" italics="true" pageId="7" pageNumber="378">C. chaneque</emphasis>
</taxonomicName>
</th>
<th id="10415D9727290009C8CCFD4FD565FCAD" box="[945,1050,766,818]" gridcol="5" gridrow="0" pageId="7" pageNumber="378">
<taxonomicName id="2AA04D2A2729FFF1C8A6FD4FD569FCAD" baseAuthorityName="Duellman" baseAuthorityYear="1968" class="Amphibia" family="Hylidae" genus="Charadrahyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="7" pageNumber="378" phylum="Chordata" rank="species" species="altipotens">
<emphasis id="DFD4EABB2729FFF1C8A6FD4FD569FCAD" bold="true" italics="true" pageId="7" pageNumber="378">C. altipotens</emphasis>
</taxonomicName>
</th>
<th id="10415D9727290009CF38FD4FD5ACFCAD" box="[1093,1235,766,818]" gridcol="6" gridrow="0" pageId="7" pageNumber="378">
<taxonomicName id="2AA04D2A2729FFF1CFFCFD4FD5DFFCAD" authorityName=": Adler &amp; Dennis" authorityYear="1972" class="Amphibia" family="Hylidae" genus="Charadrahyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="7" pageNumber="378" phylum="Chordata" rank="species" species="trux">
<emphasis id="DFD4EABB2729FFF1CFFCFD4FD5DFFCAD" bold="true" italics="true" pageId="7" pageNumber="378">C. trux</emphasis>
</taxonomicName>
</th>
<th id="10415D9727290009CE77FD4FD41FFCAD" box="[1290,1376,766,818]" gridcol="7" gridrow="0" pageId="7" pageNumber="378">
<taxonomicName id="2AA04D2A2729FFF1CE57FD4FD422FCAD" class="Amphibia" family="Hylidae" genus="Charadrahyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="7" pageNumber="378" phylum="Chordata" rank="species" species="juanitae">
<emphasis id="DFD4EABB2729FFF1CE57FD4FD422FCAD" bold="true" italics="true" pageId="7" pageNumber="378">C. juanitae</emphasis>
</taxonomicName>
</th>
<th id="10415D9727290009CEEBFD4FD493FCAD" box="[1430,1516,766,818]" gridcol="8" gridrow="0" pageId="7" pageNumber="378">
<taxonomicName id="2AA04D2A2729FFF1CECBFD4FD499FCAD" baseAuthorityName="Mendelson &amp; Campbell" baseAuthorityYear="1999" class="Amphibia" family="Hylidae" genus="Charadrahyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="7" pageNumber="378" phylum="Chordata" rank="species" species="nephila">
<emphasis id="DFD4EABB2729FFF1CECBFD4FD499FCAD" bold="true" italics="true" pageId="7" pageNumber="378">C. nephila</emphasis>
</taxonomicName>
</th>
<th id="10415D9727290009CD6FFD4FD7E2FCAD" box="[1554,1693,766,818]" gridcol="9" gridrow="0" pageId="7" pageNumber="378">
<taxonomicName id="2AA04D2A2729FFF1CD31FD4FD703FCAD" class="Amphibia" family="Hylidae" genus="Charadrahyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="7" pageNumber="378" phylum="Chordata" rank="species" species="tecuani">
<emphasis id="DFD4EABB2729FFF1CD31FD4FD703FCAD" bold="true" italics="true" pageId="7" pageNumber="378">C. tecuani</emphasis>
</taxonomicName>
</th>
<th id="10415D9727290009CDBCFD4FD635FCAD" box="[1729,1866,766,818]" gridcol="10" gridrow="0" pageId="7" pageNumber="378">
<taxonomicName id="2AA04D2A2729FFF1CD87FD4FD635FCAD" baseAuthorityName="Canseco-Marquez, Ramirez-Gonzalez &amp; Gonzalez-Bernal" baseAuthorityYear="2017" class="Amphibia" family="Hylidae" genus="Charadrahyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="7" pageNumber="378" phylum="Chordata" rank="species" species="esperancensis">
<emphasis id="DFD4EABB2729FFF1CD87FD4FD635FCAD" bold="true" italics="true" pageId="7" pageNumber="378">C. esperancensis</emphasis>
</taxonomicName>
</th>
</tr>
<tr id="539034EB27290009CBEEFC88D635FCF2" box="[147,1866,825,877]" gridrow="1" pageId="7" pageNumber="378">
<th id="10415D9727290009CBEEFC88D02FFCF2" box="[147,336,825,877]" gridcol="0" gridrow="1" pageId="7" pageNumber="378">
<emphasis id="DFD4EABB2729FFF1CBEEFC8BD180FCF2" bold="true" pageId="7" pageNumber="378">1. Maximum adult SVL (mm)</emphasis>
</th>
<td id="10415D9727290009CA13FC88D086FCF2" box="[366,505,825,877]" gridcol="1" gridrow="1" pageId="7" pageNumber="378">85.75 ♀ 73.21 ³</td>
<td id="10415D9727290009C966FC88D3FBFCF2" box="[539,644,825,877]" gridcol="2" gridrow="1" pageId="7" pageNumber="378">70.0 ♀ 65.9 ³</td>
<td id="10415D9727290009C9D6FC88D27EFCF2" box="[683,769,825,877]" gridcol="3" gridrow="1" pageId="7" pageNumber="378">34.6 ♀ 33.1 ³</td>
<td id="10415D9727290009C857FC88D2F5FCF2" box="[810,906,825,877]" gridcol="4" gridrow="1" pageId="7" pageNumber="378">79.3 ♀ 60.7 ³</td>
<td id="10415D9727290009C8CCFC88D565FCF2" box="[945,1050,825,877]" gridcol="5" gridrow="1" pageId="7" pageNumber="378">78.8 ♀ 80.6 ³</td>
<td id="10415D9727290009CF38FC88D5ACFCF2" box="[1093,1235,825,877]" gridcol="6" gridrow="1" pageId="7" pageNumber="378">– ♀ 81.0 ³</td>
<td id="10415D9727290009CE77FC88D41FFCF2" box="[1290,1376,825,877]" gridcol="7" gridrow="1" pageId="7" pageNumber="378">39.8 ♀ 35.8 ³</td>
<td id="10415D9727290009CEEBFC88D493FCF2" box="[1430,1516,825,877]" gridcol="8" gridrow="1" pageId="7" pageNumber="378">80.7 ♀ 70.9 ³</td>
<td id="10415D9727290009CD6FFC88D7E2FCF2" box="[1554,1693,825,877]" gridcol="9" gridrow="1" pageId="7" pageNumber="378">– ♀ 57.8 ³</td>
<td id="10415D9727290009CDBCFC88D635FCF2" box="[1729,1866,825,877]" gridcol="10" gridrow="1" pageId="7" pageNumber="378">– ♀ 48.9 ³</td>
</tr>
<tr id="539034EB27290009CBEEFCC5D635FC38" box="[147,1866,884,935]" gridrow="2" pageId="7" pageNumber="378">
<th id="10415D9727290009CBEEFCC5D02FFC38" box="[147,336,884,935]" gridcol="0" gridrow="2" pageId="7" pageNumber="378">
<emphasis id="DFD4EABB2729FFF1CBEEFCC5D075FC38" bold="true" pageId="7" pageNumber="378">2. Axillary membranes</emphasis>
</th>
<td id="10415D9727290009CA13FCC5D086FC38" box="[366,505,884,935]" gridcol="1" gridrow="2" pageId="7" pageNumber="378">Present</td>
<td id="10415D9727290009C966FCC5D3FBFC38" box="[539,644,884,935]" gridcol="2" gridrow="2" pageId="7" pageNumber="378">Absent</td>
<td id="10415D9727290009C9D6FCC5D27EFC38" box="[683,769,884,935]" gridcol="3" gridrow="2" pageId="7" pageNumber="378">Present</td>
<td id="10415D9727290009C857FCC5D2F5FC38" box="[810,906,884,935]" gridcol="4" gridrow="2" pageId="7" pageNumber="378">Absent</td>
<td id="10415D9727290009C8CCFCC5D565FC38" box="[945,1050,884,935]" gridcol="5" gridrow="2" pageId="7" pageNumber="378">Present</td>
<td id="10415D9727290009CF38FCC5D5ACFC38" box="[1093,1235,884,935]" gridcol="6" gridrow="2" pageId="7" pageNumber="378">Absent</td>
<td id="10415D9727290009CE77FCC5D41FFC38" box="[1290,1376,884,935]" gridcol="7" gridrow="2" pageId="7" pageNumber="378">Present</td>
<td id="10415D9727290009CEEBFCC5D493FC38" box="[1430,1516,884,935]" gridcol="8" gridrow="2" pageId="7" pageNumber="378">Absent</td>
<td id="10415D9727290009CD6FFCC5D7E2FC38" box="[1554,1693,884,935]" gridcol="9" gridrow="2" pageId="7" pageNumber="378">Absent</td>
<td id="10415D9727290009CDBCFCC5D635FC38" box="[1729,1866,884,935]" gridcol="10" gridrow="2" pageId="7" pageNumber="378">Absent</td>
</tr>
<tr id="539034EB27290009CBEEFC1FD635FC7E" box="[147,1866,942,993]" gridrow="3" pageId="7" pageNumber="378">
<th id="10415D9727290009CBEEFC1FD02FFC7E" box="[147,336,942,993]" gridcol="0" gridrow="3" pageId="7" pageNumber="378">
<emphasis id="DFD4EABB2729FFF1CBEEFC1FD1B2FC7E" bold="true" pageId="7" pageNumber="378">3. Webbing foot males</emphasis>
</th>
<td id="10415D9727290009CA13FC1FD086FC7E" box="[366,505,942,993]" gridcol="1" gridrow="3" pageId="7" pageNumber="378">Hypertrophied</td>
<td id="10415D9727290009C966FC1FD3FBFC7E" box="[539,644,942,993]" gridcol="2" gridrow="3" pageId="7" pageNumber="378">Normal</td>
<td id="10415D9727290009C9D6FC1FD27EFC7E" box="[683,769,942,993]" gridcol="3" gridrow="3" pageId="7" pageNumber="378">Normal</td>
<td id="10415D9727290009C857FC1FD2F5FC7E" box="[810,906,942,993]" gridcol="4" gridrow="3" pageId="7" pageNumber="378">Normal</td>
<td id="10415D9727290009C8CCFC1FD565FC7E" box="[945,1050,942,993]" gridcol="5" gridrow="3" pageId="7" pageNumber="378">Normal</td>
<td id="10415D9727290009CF38FC1FD5ACFC7E" box="[1093,1235,942,993]" gridcol="6" gridrow="3" pageId="7" pageNumber="378">Hypertrophied</td>
<td id="10415D9727290009CE77FC1FD41FFC7E" box="[1290,1376,942,993]" gridcol="7" gridrow="3" pageId="7" pageNumber="378">Normal</td>
<td id="10415D9727290009CEEBFC1FD493FC7E" box="[1430,1516,942,993]" gridcol="8" gridrow="3" pageId="7" pageNumber="378">Normal</td>
<td id="10415D9727290009CD6FFC1FD7E2FC7E" box="[1554,1693,942,993]" gridcol="9" gridrow="3" pageId="7" pageNumber="378">Hypertrophied</td>
<td id="10415D9727290009CDBCFC1FD635FC7E" box="[1729,1866,942,993]" gridcol="10" gridrow="3" pageId="7" pageNumber="378">Normal</td>
</tr>
<tr id="539034EB27290009CBEEFC59D635FB84" box="[147,1866,1000,1051]" gridrow="4" pageId="7" pageNumber="378">
<th id="10415D9727290009CBEEFC59D02FFB84" box="[147,336,1000,1051]" gridcol="0" gridrow="4" pageId="7" pageNumber="378">
<emphasis id="DFD4EABB2729FFF1CBEEFC59D044FB84" bold="true" pageId="7" pageNumber="378">4. Conical tubercles in vent</emphasis>
</th>
<td id="10415D9727290009CA13FC59D086FB84" box="[366,505,1000,1051]" gridcol="1" gridrow="4" pageId="7" pageNumber="378">Present</td>
<td id="10415D9727290009C966FC59D3FBFB84" box="[539,644,1000,1051]" gridcol="2" gridrow="4" pageId="7" pageNumber="378">Absent</td>
<td id="10415D9727290009C9D6FC59D27EFB84" box="[683,769,1000,1051]" gridcol="3" gridrow="4" pageId="7" pageNumber="378">Present</td>
<td id="10415D9727290009C857FC59D2F5FB84" box="[810,906,1000,1051]" gridcol="4" gridrow="4" pageId="7" pageNumber="378">Absent</td>
<td id="10415D9727290009C8CCFC59D565FB84" box="[945,1050,1000,1051]" gridcol="5" gridrow="4" pageId="7" pageNumber="378">Absent</td>
<td id="10415D9727290009CF38FC59D5ACFB84" box="[1093,1235,1000,1051]" gridcol="6" gridrow="4" pageId="7" pageNumber="378">Absent</td>
<td id="10415D9727290009CE77FC59D41FFB84" box="[1290,1376,1000,1051]" gridcol="7" gridrow="4" pageId="7" pageNumber="378">
Present 
<emphasis id="DFD4EABB2729FFF1CE29FC42D423FB9D" bold="true" box="[1364,1372,1011,1026]" pageId="7" pageNumber="378">1</emphasis>
</td>
<td id="10415D9727290009CEEBFC59D493FB84" box="[1430,1516,1000,1051]" gridcol="8" gridrow="4" pageId="7" pageNumber="378">Absent</td>
<td id="10415D9727290009CD6FFC59D7E2FB84" box="[1554,1693,1000,1051]" gridcol="9" gridrow="4" pageId="7" pageNumber="378">Present</td>
<td id="10415D9727290009CDBCFC59D635FB84" box="[1729,1866,1000,1051]" gridcol="10" gridrow="4" pageId="7" pageNumber="378">Absent</td>
</tr>
<tr id="539034EB27290009CBEEFB93D635FBCA" box="[147,1866,1058,1109]" gridrow="5" pageId="7" pageNumber="378">
<th id="10415D9727290009CBEEFB93D02FFBCA" box="[147,336,1058,1109]" gridcol="0" gridrow="5" pageId="7" pageNumber="378">
<emphasis id="DFD4EABB2729FFF1CBEEFB93D06BFBCA" bold="true" pageId="7" pageNumber="378">5. Nuptial excrescences</emphasis>
</th>
<td id="10415D9727290009CA13FB93D086FBCA" box="[366,505,1058,1109]" gridcol="1" gridrow="5" pageId="7" pageNumber="378">Present</td>
<td id="10415D9727290009C966FB93D3FBFBCA" box="[539,644,1058,1109]" gridcol="2" gridrow="5" pageId="7" pageNumber="378">Present</td>
<td id="10415D9727290009C9D6FB93D27EFBCA" box="[683,769,1058,1109]" gridcol="3" gridrow="5" pageId="7" pageNumber="378">Absent</td>
<td id="10415D9727290009C857FB93D2F5FBCA" box="[810,906,1058,1109]" gridcol="4" gridrow="5" pageId="7" pageNumber="378">Present</td>
<td id="10415D9727290009C8CCFB93D565FBCA" box="[945,1050,1058,1109]" gridcol="5" gridrow="5" pageId="7" pageNumber="378">Absent</td>
<td id="10415D9727290009CF38FB93D5ACFBCA" box="[1093,1235,1058,1109]" gridcol="6" gridrow="5" pageId="7" pageNumber="378">Present</td>
<td id="10415D9727290009CE77FB93D41FFBCA" box="[1290,1376,1058,1109]" gridcol="7" gridrow="5" pageId="7" pageNumber="378">Present</td>
<td id="10415D9727290009CEEBFB93D493FBCA" box="[1430,1516,1058,1109]" gridcol="8" gridrow="5" pageId="7" pageNumber="378">Present</td>
<td id="10415D9727290009CD6FFB93D7E2FBCA" box="[1554,1693,1058,1109]" gridcol="9" gridrow="5" pageId="7" pageNumber="378">Present</td>
<td id="10415D9727290009CDBCFB93D635FBCA" box="[1729,1866,1058,1109]" gridcol="10" gridrow="5" pageId="7" pageNumber="378">Present</td>
</tr>
<tr id="539034EB27290009CBEEFBD5D635FBE4" box="[147,1866,1124,1147]" gridrow="6" pageId="7" pageNumber="378">
<th id="10415D9727290009CBEEFBD5D02FFBE4" box="[147,336,1124,1147]" gridcol="0" gridrow="6" pageId="7" pageNumber="378">
<emphasis id="DFD4EABB2729FFF1CBEEFBD4D06CFBE4" bold="true" box="[147,275,1125,1147]" pageId="7" pageNumber="378">6. Vocal slits</emphasis>
</th>
<td id="10415D9727290009CA13FBD5D086FBE4" box="[366,505,1124,1147]" gridcol="1" gridrow="6" pageId="7" pageNumber="378">Absent</td>
<td id="10415D9727290009C966FBD5D3FBFBE4" box="[539,644,1124,1147]" gridcol="2" gridrow="6" pageId="7" pageNumber="378">Present</td>
<td id="10415D9727290009C9D6FBD5D27EFBE4" box="[683,769,1124,1147]" gridcol="3" gridrow="6" pageId="7" pageNumber="378">Present</td>
<td id="10415D9727290009C857FBD5D2F5FBE4" box="[810,906,1124,1147]" gridcol="4" gridrow="6" pageId="7" pageNumber="378">Absent</td>
<td id="10415D9727290009C8CCFBD5D565FBE4" box="[945,1050,1124,1147]" gridcol="5" gridrow="6" pageId="7" pageNumber="378">Absent</td>
<td id="10415D9727290009CF38FBD5D5ACFBE4" box="[1093,1235,1124,1147]" gridcol="6" gridrow="6" pageId="7" pageNumber="378">Present</td>
<td id="10415D9727290009CE77FBD5D41FFBE4" box="[1290,1376,1124,1147]" gridcol="7" gridrow="6" pageId="7" pageNumber="378">Present</td>
<td id="10415D9727290009CEEBFBD5D493FBE4" box="[1430,1516,1124,1147]" gridcol="8" gridrow="6" pageId="7" pageNumber="378">Present</td>
<td id="10415D9727290009CD6FFBD5D7E2FBE4" box="[1554,1693,1124,1147]" gridcol="9" gridrow="6" pageId="7" pageNumber="378">Present</td>
<td id="10415D9727290009CDBCFBD5D635FBE4" box="[1729,1866,1124,1147]" gridcol="10" gridrow="6" pageId="7" pageNumber="378">Absent</td>
</tr>
<tr id="539034EB27290009CBEEFB3DD635FB66" box="[147,1866,1164,1273]" gridrow="7" pageId="7" pageNumber="378">
<th id="10415D9727290009CBEEFB3DD02FFB66" box="[147,336,1164,1273]" gridcol="0" gridrow="7" pageId="7" pageNumber="378">
<emphasis id="DFD4EABB2729FFF1CBEEFB3DD063FB66" bold="true" pageId="7" pageNumber="378">7. Quadratojugal in contact with maxillary 8. Male snout</emphasis>
</th>
<td id="10415D9727290009CA13FB3DD086FB66" box="[366,505,1164,1273]" gridcol="1" gridrow="7" pageId="7" pageNumber="378">Yes</td>
<td id="10415D9727290009C966FB3DD3FBFB66" box="[539,644,1164,1273]" gridcol="2" gridrow="7" pageId="7" pageNumber="378">Yes</td>
<td id="10415D9727290009C9D6FB3DD27EFB66" box="[683,769,1164,1273]" gridcol="3" gridrow="7" pageId="7" pageNumber="378">–</td>
<td id="10415D9727290009C857FB3DD2F5FB66" box="[810,906,1164,1273]" gridcol="4" gridrow="7" pageId="7" pageNumber="378">Yes</td>
<td id="10415D9727290009C8CCFB3DD565FB66" box="[945,1050,1164,1273]" gridcol="5" gridrow="7" pageId="7" pageNumber="378">Yes</td>
<td id="10415D9727290009CF38FB3DD5ACFB66" box="[1093,1235,1164,1273]" gridcol="6" gridrow="7" pageId="7" pageNumber="378">
Variable 
<emphasis id="DFD4EABB2729FFF1CFCCFB14D5C6FB2B" bold="true" box="[1201,1209,1189,1204]" pageId="7" pageNumber="378">2</emphasis>
</td>
<td id="10415D9727290009CE77FB3DD41FFB66" box="[1290,1376,1164,1273]" gridcol="7" gridrow="7" pageId="7" pageNumber="378">–</td>
<td id="10415D9727290009CEEBFB3DD493FB66" box="[1430,1516,1164,1273]" gridcol="8" gridrow="7" pageId="7" pageNumber="378">Yes</td>
<td id="10415D9727290009CD6FFB3DD7E2FB66" box="[1554,1693,1164,1273]" gridcol="9" gridrow="7" pageId="7" pageNumber="378">Yes</td>
<td id="10415D9727290009CDBCFB3DD635FB66" box="[1729,1866,1164,1273]" gridcol="10" gridrow="7" pageId="7" pageNumber="378">–</td>
</tr>
<tr id="539034EB27290009CBEEFB4FD635FAAC" box="[147,1866,1278,1331]" gridrow="8" pageId="7" pageNumber="378">
<th id="10415D9727290009CBEEFB4FD02FFAAC" box="[147,336,1278,1331]" gridcol="0" gridrow="8" pageId="7" pageNumber="378">
<emphasis id="DFD4EABB2729FFF1CBEEFAB1D1BEFAAC" bold="true" pageId="7" pageNumber="378">shape in dorsal view</emphasis>
</th>
<td id="10415D9727290009CA13FB4FD086FAAC" box="[366,505,1278,1331]" gridcol="1" gridrow="8" pageId="7" pageNumber="378">Acuminate</td>
<td id="10415D9727290009C966FB4FD3FBFAAC" box="[539,644,1278,1331]" gridcol="2" gridrow="8" pageId="7" pageNumber="378">Acuminate</td>
<td id="10415D9727290009C9D6FB4FD27EFAAC" box="[683,769,1278,1331]" gridcol="3" gridrow="8" pageId="7" pageNumber="378">Rounded</td>
<td id="10415D9727290009C857FB4FD2F5FAAC" box="[810,906,1278,1331]" gridcol="4" gridrow="8" pageId="7" pageNumber="378">Rounded</td>
<td id="10415D9727290009C8CCFB4FD565FAAC" box="[945,1050,1278,1331]" gridcol="5" gridrow="8" pageId="7" pageNumber="378">Acuminate</td>
<td id="10415D9727290009CF38FB4FD5ACFAAC" box="[1093,1235,1278,1331]" gridcol="6" gridrow="8" pageId="7" pageNumber="378">Rounded</td>
<td id="10415D9727290009CE77FB4FD41FFAAC" box="[1290,1376,1278,1331]" gridcol="7" gridrow="8" pageId="7" pageNumber="378">Rounded</td>
<td id="10415D9727290009CEEBFB4FD493FAAC" box="[1430,1516,1278,1331]" gridcol="8" gridrow="8" pageId="7" pageNumber="378">Rounded</td>
<td id="10415D9727290009CD6FFB4FD7E2FAAC" box="[1554,1693,1278,1331]" gridcol="9" gridrow="8" pageId="7" pageNumber="378">Acuminate</td>
<td id="10415D9727290009CDBCFB4FD635FAAC" box="[1729,1866,1278,1331]" gridcol="10" gridrow="8" pageId="7" pageNumber="378">Pointed</td>
</tr>
<tr id="539034EB27290009CBEEFAF0D635FAC6" box="[147,1866,1345,1369]" gridrow="9" pageId="7" pageNumber="378">
<th id="10415D9727290009CBEEFAF0D02FFAC6" box="[147,336,1345,1369]" gridcol="0" gridrow="9" pageId="7" pageNumber="378">
<emphasis id="DFD4EABB2729FFF1CBEEFAF2D06AFAC6" bold="true" box="[147,277,1347,1369]" pageId="7" pageNumber="378">9. Testes size</emphasis>
</th>
<td id="10415D9727290009CA13FAF0D086FAC6" box="[366,505,1345,1369]" gridcol="1" gridrow="9" pageId="7" pageNumber="378">Enlarged</td>
<td id="10415D9727290009C966FAF0D3FBFAC6" box="[539,644,1345,1369]" gridcol="2" gridrow="9" pageId="7" pageNumber="378">Enlarged</td>
<td id="10415D9727290009C9D6FAF0D27EFAC6" box="[683,769,1345,1369]" gridcol="3" gridrow="9" pageId="7" pageNumber="378">–</td>
<td id="10415D9727290009C857FAF0D2F5FAC6" box="[810,906,1345,1369]" gridcol="4" gridrow="9" pageId="7" pageNumber="378">Normal</td>
<td id="10415D9727290009C8CCFAF0D565FAC6" box="[945,1050,1345,1369]" gridcol="5" gridrow="9" pageId="7" pageNumber="378">Enlarged</td>
<td id="10415D9727290009CF38FAF0D5ACFAC6" box="[1093,1235,1345,1369]" gridcol="6" gridrow="9" pageId="7" pageNumber="378">Enlarged</td>
<td id="10415D9727290009CE77FAF0D41FFAC6" box="[1290,1376,1345,1369]" gridcol="7" gridrow="9" pageId="7" pageNumber="378">–</td>
<td id="10415D9727290009CEEBFAF0D493FAC6" box="[1430,1516,1345,1369]" gridcol="8" gridrow="9" pageId="7" pageNumber="378">Normal</td>
<td id="10415D9727290009CD6FFAF0D7E2FAC6" box="[1554,1693,1345,1369]" gridcol="9" gridrow="9" pageId="7" pageNumber="378">Enlarged</td>
<td id="10415D9727290009CDBCFAF0D635FAC6" box="[1729,1866,1345,1369]" gridcol="10" gridrow="9" pageId="7" pageNumber="378">–</td>
</tr>
<tr id="539034EB27290009CBEEFAD9D635FA03" box="[147,1866,1384,1436]" gridrow="10" pageId="7" pageNumber="378">
<th id="10415D9727290009CBEEFAD9D02FFA03" box="[147,336,1384,1436]" gridcol="0" gridrow="10" pageId="7" pageNumber="378">
<emphasis id="DFD4EABB2729FFF1CBEEFAD8D184FA03" bold="true" pageId="7" pageNumber="378">10. Belly coloration</emphasis>
</th>
<td id="10415D9727290009CA13FAD9D086FA03" box="[366,505,1384,1436]" gridcol="1" gridrow="10" pageId="7" pageNumber="378">White-pale brown</td>
<td id="10415D9727290009C966FAD9D3FBFA03" box="[539,644,1384,1436]" gridcol="2" gridrow="10" pageId="7" pageNumber="378">Brownish- black</td>
<td id="10415D9727290009C9D6FAD9D27EFA03" box="[683,769,1384,1436]" gridcol="3" gridrow="10" pageId="7" pageNumber="378">Creamy white</td>
<td id="10415D9727290009C857FAD9D2F5FA03" box="[810,906,1384,1436]" gridcol="4" gridrow="10" pageId="7" pageNumber="378">Creamy brown</td>
<td id="10415D9727290009C8CCFAD9D565FA03" box="[945,1050,1384,1436]" gridcol="5" gridrow="10" pageId="7" pageNumber="378">Yellow</td>
<td id="10415D9727290009CF38FAD9D5ACFA03" box="[1093,1235,1384,1436]" gridcol="6" gridrow="10" pageId="7" pageNumber="378">Creamy brown</td>
<td id="10415D9727290009CE77FAD9D41FFA03" box="[1290,1376,1384,1436]" gridcol="7" gridrow="10" pageId="7" pageNumber="378">Creamy white</td>
<td id="10415D9727290009CEEBFAD9D493FA03" box="[1430,1516,1384,1436]" gridcol="8" gridrow="10" pageId="7" pageNumber="378">Purplish tan</td>
<td id="10415D9727290009CD6FFAD9D7E2FA03" box="[1554,1693,1384,1436]" gridcol="9" gridrow="10" pageId="7" pageNumber="378">Yellow</td>
<td id="10415D9727290009CDBCFAD9D635FA03" box="[1729,1866,1384,1436]" gridcol="10" gridrow="10" pageId="7" pageNumber="378">White</td>
</tr>
</table>
</paragraph>
<paragraph id="ED1F36A92729FFF1CBEEFA0CD5B7FA48" blockId="7.[147,1270,1469,1524]" box="[147,1224,1469,1495]" pageId="7" pageNumber="378">
<superScript id="1AD59BE12729FFF1CBEEFA0CD1E4FA53" attach="right" box="[147,155,1469,1484]" fontSize="6" pageId="7" pageNumber="378">1</superScript>
<bibRefCitation id="89314B582729FFF1CBE6FA71D040FA48" author="Duellman, W. E." box="[155,319,1472,1495]" pageId="7" pageNumber="378" refId="ref7389" refString="Duellman, W. E. (2001) The Hylid Frogs of Middle America, Society for the Study of Amphibians and Reptiles, Ithaca, New York, 1170 pp." type="book" year="2001">Duellman (2001)</bibRefCitation>
describes slightly larger pigmented tubercles present immediately around the cloacal opening.
</paragraph>
<paragraph id="ED1F36A92729FFF1CBEEFA6BD589FA6B" blockId="7.[147,1270,1469,1524]" box="[147,1270,1498,1524]" pageId="7" pageNumber="378">
<superScript id="1AD59BE12729FFF1CBEEFA6BD1E4FA76" attach="right" box="[147,155,1498,1513]" fontSize="6" pageId="7" pageNumber="378">2</superScript>
According to 
<bibRefCitation id="89314B582729FFF1CA63FA6CD080FA6B" author="Adler, K. &amp; Dennis, D. M." box="[286,511,1501,1524]" pageId="7" pageNumber="378" pagination="1 - 19" refId="ref6564" refString="Adler, K. &amp; Dennis, D. M. (1972) New tree frogs of the genus Hyla from the cloud forests of western Guerrero, Mexico. Occasional Papers of The Museum of Natural History, University of Kansas, 7, 1 - 19." type="journal article" year="1972">Adler &amp; Dennis (1972)</bibRefCitation>
, four of nine individuals is articulated in one or both sides with the maxillaries.
</paragraph>
<paragraph id="ED1F36A92726FFFECBBAFF26D32DFD87" blockId="8.[151,1437,151,536]" pageId="8" pageNumber="379">
<emphasis id="DFD4EABB2726FFFECBBAFF26D043FF2F" bold="true" box="[199,316,151,176]" pageId="8" pageNumber="379">Variation.</emphasis>
All the males show a lateral profile rounded with snout sloping downward posterior to the nostril and snout dorsal profile acuminate; in contrast females are bluntly rounded, in both lateral and dorsal profiles. The axillary membrane is more noticeable in life (
<figureCitation id="759B2A2C2726FFFEC9EBFF6ED271FF67" box="[662,782,223,248]" captionStart="FIGURE 3" captionStartId="4.[151,250,1571,1593]" captionTargetBox="[169,1418,241,1512]" captionTargetId="figure@4.[151,1435,223,1530]" captionTargetPageId="4" captionText="FIGURE 3. Morphological characters of Charadrahyla sakbah. (A) Axillary membrane of male holotype (IBH 30989). (B) Axillary membrane of female paratype (IBH 30992). (C) Retracted and (D) extended hypertrophied webbing in male holotype (IBH 30989). (E) Nuptial excrescences in dorsal view and (F) view of the palm of the hand without nuptial excrescences in Finger I in male holotype (IBH 30989)." httpUri="https://zenodo.org/record/2623752/files/figure.png" pageId="8" pageNumber="379">Fig. 3A, B</figureCitation>
) and in the females, possibly due to their larger body size. The three males (
<typeStatus id="321B880B2726FFFECA21FEB5D0BFFE82" box="[348,448,260,285]" pageId="8" pageNumber="379" type="holotype">holotype</typeStatus>
and two 
<typeStatus id="321B880B2726FFFEC951FEB5D3E6FE83" box="[556,665,260,284]" pageId="8" pageNumber="379" type="paratype">paratypes</typeStatus>
) show hypertrophied webbing. The nuptial excrescences are almost indistinguishable in the males, perhaps by the preservation process, but brown to black in life (
<figureCitation id="759B2A2C2726FFFECFC9FE96D475FEDF" box="[1204,1290,295,320]" captionStart="FIGURE 3" captionStartId="4.[151,250,1571,1593]" captionTargetBox="[169,1418,241,1512]" captionTargetId="figure@4.[151,1435,223,1530]" captionTargetPageId="4" captionText="FIGURE 3. Morphological characters of Charadrahyla sakbah. (A) Axillary membrane of male holotype (IBH 30989). (B) Axillary membrane of female paratype (IBH 30992). (C) Retracted and (D) extended hypertrophied webbing in male holotype (IBH 30989). (E) Nuptial excrescences in dorsal view and (F) view of the palm of the hand without nuptial excrescences in Finger I in male holotype (IBH 30989)." httpUri="https://zenodo.org/record/2623752/files/figure.png" pageId="8" pageNumber="379">Fig. 3E</figureCitation>
). In the three males of the type series the nuptial excrescences correspond to dark-colored nuptial pads with cone-shaped papillae. The testes of two males (
<typeStatus id="321B880B2726FFFEC96FFEC1D3FCFE17" box="[530,643,368,392]" pageId="8" pageNumber="379" type="paratype">paratypes</typeStatus>
: IBH 30990-91) were ovoid, white and not visibly granular (mean 
<quantity id="2A589B4C2726FFFECE16FEC1D1B9FE33" metricMagnitude="-3" metricUnit="m" metricValue="8.48" pageId="8" pageNumber="379" unit="mm" value="8.48">8.48 mm</quantity>
; range 
<quantity id="2A589B4C2726FFFECA68FE25D0C4FE33" box="[277,443,404,429]" metricMagnitude="-3" metricUnit="m" metricValue="8.37" metricValueMax="8.62" metricValueMin="8.12" pageId="8" pageNumber="379" unit="mm" value="8.37" valueMax="8.62" valueMin="8.12">8.12–8.62 mm</quantity>
; 12.49; 12.32–12.66% of SVL). The females and males show a conical tubercle on either side of vent like the 
<typeStatus id="321B880B2726FFFECA06FE06D0A2FE4F" box="[379,477,439,464]" pageId="8" pageNumber="379" type="holotype">holotype</typeStatus>
(
<figureCitation id="759B2A2C2726FFFECA91FE06D33DFE4F" box="[492,578,439,464]" captionStart="FIGURE 2" captionStartId="3.[151,250,1510,1532]" captionTargetBox="[181,1401,221,1464]" captionTargetId="figure@3.[150,1436,193,1489]" captionTargetPageId="3" captionText="FIGURE 2. Specimens in life of C haradrahyla sakbah. (A) male holotype (IBH 30989). (B) male paratype (IBH 30990). (C) Female paratype (IBH 30992). (D) Female paratype (IBH 30993). (E) Female paratype (IBH 30994). (F) Enlarged vent tubercles in ventral view of C. sakbah of male holotype (IBH 30989) in life." httpUri="https://zenodo.org/record/2623750/files/figure.png" pageId="8" pageNumber="379">Fig. 2F</figureCitation>
). In four specimens examined (IBH 30989, IBH 30991, IBH 30992 and IBH 30993) the quadratojugals are present, ossified and in contact with the maxilla. The variation in the proportions for females and males are given in 
<tableCitation id="A02203122726FFFECA85FE4ED331FD87" box="[504,590,511,536]" captionStart="TABLE 2" captionStartId="8.[151,239,580,603]" captionText="TABLE 2. Mean and range in parenthesis of morphological proportions recorded in Charadrahyla sakbah type series. HW: head width and SVL: snout-vent length." pageId="8" pageNumber="379">Table 2</tableCitation>
.
</paragraph>
<caption id="B9DF66212726FFFECBEAFDF5D304FDE4" pageId="8" pageNumber="379">
<paragraph id="ED1F36A92726FFFECBEAFDF5D304FDE4" blockId="8.[151,1437,579,635]" pageId="8" pageNumber="379">
<emphasis id="DFD4EABB2726FFFECBEAFDF5D074FDC4" bold="true" box="[151,267,580,603]" pageId="8" pageNumber="379">TABLE 2.</emphasis>
Mean and range in parenthesis of morphological proportions recorded in 
<taxonomicName id="2AA04D2A2726FFFECF52FDF2D465FDC4" authorityName="Jiménez-Arcos &amp; Calzada-Arciniega &amp; Alfaro-Juantorena &amp; Vázquez-Reyes &amp; Blair &amp; Parra-Olea" authorityYear="2019" box="[1071,1306,579,603]" class="Amphibia" family="Hylidae" genus="Charadrahyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="8" pageNumber="379" phylum="Chordata" rank="species" species="sakbah">
<emphasis id="DFD4EABB2726FFFECF52FDF2D465FDC4" box="[1071,1306,579,603]" italics="true" pageId="8" pageNumber="379">Charadrahyla sakbah</emphasis>
</taxonomicName>
type series. HW: head width and SVL: snout-vent length.
</paragraph>
</caption>
<paragraph id="ED1F36A92726FFFECBE1FD27D440FC51" pageId="8" pageNumber="379">
<table id="9FA0C40927260009CBE1FD27D440FC51" box="[156,1343,662,974]" gridcols="3" gridrows="8" pageId="8" pageNumber="379">
<tr id="539034EB27260009CBE1FD27D440FD33" box="[156,1343,662,684]" gridrow="0" pageId="8" pageNumber="379">
<th id="10415D9727260009CBE1FD27D3F9FD33" box="[156,646,662,684]" gridcol="0" gridrow="0" pageId="8" pageNumber="379">Morphological proportions</th>
<th id="10415D9727260009C844FD27D577FD33" box="[825,1032,662,684]" gridcol="1" gridrow="0" pageId="8" pageNumber="379">Females</th>
<th id="10415D9727260009CF0DFD27D440FD33" box="[1136,1343,662,684]" gridcol="2" gridrow="0" pageId="8" pageNumber="379">Males</th>
</tr>
<tr id="539034EB27260009CBE1FD71D440FD49" box="[156,1343,704,726]" gridrow="1" pageId="8" pageNumber="379">
<th id="10415D9727260009CBE1FD71D3F9FD49" box="[156,646,704,726]" gridcol="0" gridrow="1" pageId="8" pageNumber="379">Interorbital / HW</th>
<td id="10415D9727260009C844FD71D577FD49" box="[825,1032,704,726]" gridcol="1" gridrow="1" pageId="8" pageNumber="379">30.15 (29.04–32.35)</td>
<td id="10415D9727260009CF0DFD71D440FD49" box="[1136,1343,704,726]" gridcol="2" gridrow="1" pageId="8" pageNumber="379">33.68 (33.22–34.48)</td>
</tr>
<tr id="539034EB27260009CBE1FD58D440FD60" box="[156,1343,745,767]" gridrow="2" pageId="8" pageNumber="379">
<th id="10415D9727260009CBE1FD58D3F9FD60" box="[156,646,745,767]" gridcol="0" gridrow="2" pageId="8" pageNumber="379">Diameter of eye / HW</th>
<td id="10415D9727260009C844FD58D577FD60" box="[825,1032,745,767]" gridcol="1" gridrow="2" pageId="8" pageNumber="379">33.19 (30.19–37.26)</td>
<td id="10415D9727260009CF0DFD58D440FD60" box="[1136,1343,745,767]" gridcol="2" gridrow="2" pageId="8" pageNumber="379">32.18 (31.63–32.91)</td>
</tr>
<tr id="539034EB27260009CBE1FCA3D440FCB7" box="[156,1343,786,808]" gridrow="3" pageId="8" pageNumber="379">
<th id="10415D9727260009CBE1FCA3D3F9FCB7" box="[156,646,786,808]" gridcol="0" gridrow="3" pageId="8" pageNumber="379">Diameter of tympanum/Diameter of eye</th>
<td id="10415D9727260009C844FCA3D577FCB7" box="[825,1032,786,808]" gridcol="1" gridrow="3" pageId="8" pageNumber="379">34.16 (31.47–36.33)</td>
<td id="10415D9727260009CF0DFCA3D440FCB7" box="[1136,1343,786,808]" gridcol="2" gridrow="3" pageId="8" pageNumber="379">40.21 (35.54–44.82)</td>
</tr>
<tr id="539034EB27260009CBE1FC8DD440FCCD" box="[156,1343,828,850]" gridrow="4" pageId="8" pageNumber="379">
<th id="10415D9727260009CBE1FC8DD3F9FCCD" box="[156,646,828,850]" gridcol="0" gridrow="4" pageId="8" pageNumber="379">Diameter of tympanum/Distance eye-tympanum</th>
<td id="10415D9727260009C844FC8DD577FCCD" box="[825,1032,828,850]" gridcol="1" gridrow="4" pageId="8" pageNumber="379">60.92 (60.34–61.71)</td>
<td id="10415D9727260009CF0DFC8DD440FCCD" box="[1136,1343,828,850]" gridcol="2" gridrow="4" pageId="8" pageNumber="379">75.62 (67.54–86.11)</td>
</tr>
<tr id="539034EB27260009CBE1FCD4D440FCE4" box="[156,1343,869,891]" gridrow="5" pageId="8" pageNumber="379">
<th id="10415D9727260009CBE1FCD4D3F9FCE4" box="[156,646,869,891]" gridcol="0" gridrow="5" pageId="8" pageNumber="379">Width of Disc Finger I/Diameter of tympanum</th>
<td id="10415D9727260009C844FCD4D577FCE4" box="[825,1032,869,891]" gridcol="1" gridrow="5" pageId="8" pageNumber="379">88.51 (81.12–98.94)</td>
<td id="10415D9727260009CF0DFCD4D440FCE4" box="[1136,1343,869,891]" gridcol="2" gridrow="5" pageId="8" pageNumber="379">66.10 (64.50–67.37)</td>
</tr>
<tr id="539034EB27260009CBE1FC3FD440FC3B" box="[156,1343,910,932]" gridrow="6" pageId="8" pageNumber="379">
<th id="10415D9727260009CBE1FC3FD3F9FC3B" box="[156,646,910,932]" gridcol="0" gridrow="6" pageId="8" pageNumber="379">Tibia length/SVL</th>
<td id="10415D9727260009C844FC3FD577FC3B" box="[825,1032,910,932]" gridcol="1" gridrow="6" pageId="8" pageNumber="379">52.19 (51.56–53.23)</td>
<td id="10415D9727260009CF0DFC3FD440FC3B" box="[1136,1343,910,932]" gridcol="2" gridrow="6" pageId="8" pageNumber="379">53.76 (52.74–54.61)</td>
</tr>
<tr id="539034EB27260009CBE1FC09D440FC51" box="[156,1343,952,974]" gridrow="7" pageId="8" pageNumber="379">
<th id="10415D9727260009CBE1FC09D3F9FC51" box="[156,646,952,974]" gridcol="0" gridrow="7" pageId="8" pageNumber="379">Foot length/SVL</th>
<td id="10415D9727260009C844FC09D577FC51" box="[825,1032,952,974]" gridcol="1" gridrow="7" pageId="8" pageNumber="379">43.42 (41.92–44.95)</td>
<td id="10415D9727260009CF0DFC09D440FC51" box="[1136,1343,952,974]" gridcol="2" gridrow="7" pageId="8" pageNumber="379">46.17 (45.82–46.77)</td>
</tr>
</table>
</paragraph>
<paragraph id="ED1F36A92726FFFECBBAFBA3D0ECFA60" blockId="8.[151,1437,1042,2003]" pageId="8" pageNumber="379">
Considerable color variations are present mainly in females. One female 
<typeStatus id="321B880B2726FFFEC884FBA2D526FBB4" box="[1017,1113,1043,1067]" pageId="8" pageNumber="379" type="paratype">paratype</typeStatus>
(IBH 30992) shows a dorsal ground color yellow to greenish, with large irregular dark green blotches on the dorsum (
<figureCitation id="759B2A2C2726FFFECF08FB87D5B2FBD0" box="[1141,1229,1078,1103]" captionStart="FIGURE 2" captionStartId="3.[151,250,1510,1532]" captionTargetBox="[181,1401,221,1464]" captionTargetId="figure@3.[150,1436,193,1489]" captionTargetPageId="3" captionText="FIGURE 2. Specimens in life of C haradrahyla sakbah. (A) male holotype (IBH 30989). (B) male paratype (IBH 30990). (C) Female paratype (IBH 30992). (D) Female paratype (IBH 30993). (E) Female paratype (IBH 30994). (F) Enlarged vent tubercles in ventral view of C. sakbah of male holotype (IBH 30989) in life." httpUri="https://zenodo.org/record/2623750/files/figure.png" pageId="8" pageNumber="379">Fig. 2C</figureCitation>
). A second female 
<typeStatus id="321B880B2726FFFECBEAFBEAD188FBEC" box="[151,247,1115,1139]" pageId="8" pageNumber="379" type="paratype">paratype</typeStatus>
(IBH 30993) shows a dark cream to brownish dorsal ground color. Irregular blotches are also present, with greenish-brown tonalities (
<figureCitation id="759B2A2C2726FFFEC97DFBCFD323FB08" box="[512,604,1150,1175]" captionStart="FIGURE 2" captionStartId="3.[151,250,1510,1532]" captionTargetBox="[181,1401,221,1464]" captionTargetId="figure@3.[150,1436,193,1489]" captionTargetPageId="3" captionText="FIGURE 2. Specimens in life of C haradrahyla sakbah. (A) male holotype (IBH 30989). (B) male paratype (IBH 30990). (C) Female paratype (IBH 30992). (D) Female paratype (IBH 30993). (E) Female paratype (IBH 30994). (F) Enlarged vent tubercles in ventral view of C. sakbah of male holotype (IBH 30989) in life." httpUri="https://zenodo.org/record/2623750/files/figure.png" pageId="8" pageNumber="379">Fig. 2D</figureCitation>
). The last female in the type series (IBH 30994) exhibits a dorsal ground color brownish-orange, and blotches with a chocolate color (
<figureCitation id="759B2A2C2726FFFEC844FB13D2F0FB24" box="[825,911,1186,1211]" captionStart="FIGURE 2" captionStartId="3.[151,250,1510,1532]" captionTargetBox="[181,1401,221,1464]" captionTargetId="figure@3.[150,1436,193,1489]" captionTargetPageId="3" captionText="FIGURE 2. Specimens in life of C haradrahyla sakbah. (A) male holotype (IBH 30989). (B) male paratype (IBH 30990). (C) Female paratype (IBH 30992). (D) Female paratype (IBH 30993). (E) Female paratype (IBH 30994). (F) Enlarged vent tubercles in ventral view of C. sakbah of male holotype (IBH 30989) in life." httpUri="https://zenodo.org/record/2623750/files/figure.png" pageId="8" pageNumber="379">Fig. 2E</figureCitation>
). The three females show transverse bars in the limbs of a similar color to the dorsal blotches of each (
<figureCitation id="759B2A2C2726FFFEC87AFB77D2A9FB40" box="[775,982,1222,1247]" captionStart="FIGURE 2" captionStartId="3.[151,250,1510,1532]" captionTargetBox="[181,1401,221,1464]" captionTargetId="figure@3.[150,1436,193,1489]" captionTargetPageId="3" captionText="FIGURE 2. Specimens in life of C haradrahyla sakbah. (A) male holotype (IBH 30989). (B) male paratype (IBH 30990). (C) Female paratype (IBH 30992). (D) Female paratype (IBH 30993). (E) Female paratype (IBH 30994). (F) Enlarged vent tubercles in ventral view of C. sakbah of male holotype (IBH 30989) in life." httpUri="https://zenodo.org/record/2623750/files/figure.png" pageId="8" pageNumber="379">Fig. 2 C, D and E</figureCitation>
). A similar pattern was observed in one female not collected—the dorsal ground color was green with blotches and transverse bars a darker green (
<figureCitation id="759B2A2C2726FFFECE12FB5BD1C2FAB8" captionStart="FIGURE 5" captionStartId="10.[151,250,1976,1998]" captionTargetBox="[161,1425,193,1959]" captionTargetId="figure@10.[161,1425,193,1959]" captionTargetPageId="10" captionText="FIGURE 5. (A) A female (not collected) of C. sakbah in the riverside. (B) Panoramic view of the Chite ku'e river. (C) Typical pool and waterfall where C. sakbah individuals are usually found." httpUri="https://zenodo.org/record/2623756/files/figure.png" pageId="8" pageNumber="379">Fig. 5A</figureCitation>
). All the females observed show lateral pattern of enlarged spots dark brown to black, like the female shown in 
<figureCitation id="759B2A2C2726FFFECBEAFA83D18FFAD4" box="[151,240,1330,1355]" captionStart="FIGURE 5" captionStartId="10.[151,250,1976,1998]" captionTargetBox="[161,1425,193,1959]" captionTargetId="figure@10.[161,1425,193,1959]" captionTargetPageId="10" captionText="FIGURE 5. (A) A female (not collected) of C. sakbah in the riverside. (B) Panoramic view of the Chite ku'e river. (C) Typical pool and waterfall where C. sakbah individuals are usually found." httpUri="https://zenodo.org/record/2623756/files/figure.png" pageId="8" pageNumber="379">Fig. 5A</figureCitation>
. The throats in all female 
<typeStatus id="321B880B2726FFFEC961FA82D3F8FAD4" box="[540,647,1331,1355]" pageId="8" pageNumber="379" type="paratype">paratypes</typeStatus>
show irregular blotches dark brown to black. For males, the blotches are less conspicuous. One male (IBH 30990) shows a similar dorsal ground color in life as the 
<typeStatus id="321B880B2726FFFECFC4FAE7D461FAF0" box="[1209,1310,1366,1391]" pageId="8" pageNumber="379" type="holotype">holotype</typeStatus>
, but differs in that coloration is gray, blotches and transverse bars in limbs are darker gray, tibias and lower part of the back green with more intense yellow in the flanks (
<figureCitation id="759B2A2C2726FFFEC9BBFA2FD25CFA28" box="[710,803,1438,1463]" captionStart="FIGURE 2" captionStartId="3.[151,250,1510,1532]" captionTargetBox="[181,1401,221,1464]" captionTargetId="figure@3.[150,1436,193,1489]" captionTargetPageId="3" captionText="FIGURE 2. Specimens in life of C haradrahyla sakbah. (A) male holotype (IBH 30989). (B) male paratype (IBH 30990). (C) Female paratype (IBH 30992). (D) Female paratype (IBH 30993). (E) Female paratype (IBH 30994). (F) Enlarged vent tubercles in ventral view of C. sakbah of male holotype (IBH 30989) in life." httpUri="https://zenodo.org/record/2623750/files/figure.png" pageId="8" pageNumber="379">Fig. 2B</figureCitation>
). The other 
<typeStatus id="321B880B2726FFFEC8C1FA2ED563FA28" box="[956,1052,1439,1463]" pageId="8" pageNumber="379" type="paratype">paratype</typeStatus>
male (IBH 30991; not showed) presented a coloration almost identical to 
<typeStatus id="321B880B2726FFFEC90EFA73D3A6FA44" box="[627,729,1474,1499]" pageId="8" pageNumber="379" type="holotype">holotype</typeStatus>
. The three males show throats white in life and cream to light brown in preservative.
</paragraph>
</subSubSection>
<subSubSection id="A5BA65222726FFFECBBAF9BBD01AF9F5" pageId="8" pageNumber="379" type="etymology">
<paragraph id="ED1F36A92726FFFECBBAF9BBD01AF9F5" blockId="8.[151,1437,1042,2003]" pageId="8" pageNumber="379">
<emphasis id="DFD4EABB2726FFFECBBAF9BBD031F9BC" bold="true" box="[199,334,1546,1571]" pageId="8" pageNumber="379">Etymology.</emphasis>
The specific epithet is taken from the Mixteco language word “sa’bah” which mean “frog”. This is recognition to the San Isidro Paz y Progreso community for their conservation efforts towards their natural and cultural resources.
</paragraph>
</subSubSection>
<subSubSection id="A5BA65222726FFFFCBBAF9C7D5E5FD37" lastPageId="9" lastPageNumber="380" pageId="8" pageNumber="379" type="distribution">
<paragraph id="ED1F36A92726FFFFCBBAF9C7D358FF4A" blockId="8.[151,1437,1042,2003]" lastBlockId="9.[151,1437,151,680]" lastPageId="9" lastPageNumber="380" pageId="8" pageNumber="379">
<emphasis id="DFD4EABB2726FFFECBBAF9C7D32FF910" bold="true" box="[199,592,1654,1679]" pageId="8" pageNumber="379">Distribution and natural history.</emphasis>
<taxonomicName id="2AA04D2A2726FFFEC92AF9C7D233F910" authorityName="Jiménez-Arcos &amp; Calzada-Arciniega &amp; Alfaro-Juantorena &amp; Vázquez-Reyes &amp; Blair &amp; Parra-Olea" authorityYear="2019" box="[599,844,1654,1679]" class="Amphibia" family="Hylidae" genus="Charadrahyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="8" pageNumber="379" phylum="Chordata" rank="species" species="sakbah">
<emphasis id="DFD4EABB2726FFFEC92AF9C7D233F910" box="[599,844,1654,1679]" italics="true" pageId="8" pageNumber="379">Charadrahyla sakbah</emphasis>
</taxonomicName>
is only known from one locality in San Isidro Paz y Progreso, on the river Chite ku'e at 
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in elevation. This river is surrounded by cloud forest, presents several rocky outcrops, with steep slopes that generate numerous waterfalls. In some points the river flow extends to more than 
<quantity id="2A589B4C2726FFFECBB0F952D079F965" box="[205,262,1763,1787]" metricMagnitude="1" metricUnit="m" metricValue="1.0" pageId="8" pageNumber="379" unit="m" value="10.0">10 m</quantity>
wide (
<figureCitation id="759B2A2C2726FFFECA2DF953D0C3F964" box="[336,444,1762,1787]" captionStart="FIGURE 5" captionStartId="10.[151,250,1976,1998]" captionTargetBox="[161,1425,193,1959]" captionTargetId="figure@10.[161,1425,193,1959]" captionTargetPageId="10" captionText="FIGURE 5. (A) A female (not collected) of C. sakbah in the riverside. (B) Panoramic view of the Chite ku'e river. (C) Typical pool and waterfall where C. sakbah individuals are usually found." httpUri="https://zenodo.org/record/2623756/files/figure.png" pageId="8" pageNumber="379">Fig 5B,C</figureCitation>
). The mean temperature in the area is 20°C and the rainfall annually is 
<quantity id="2A589B4C2726FFFECFACF952D42FF965" box="[1233,1360,1763,1787]" metricMagnitude="0" metricUnit="m" metricValue="1.8933" pageId="8" pageNumber="379" unit="mm" value="1893.3">1893.3 mm</quantity>
(
<bibRefCitation id="89314B582726FFFECE23F952D1ABF880" author="SMN" pageId="8" pageNumber="379" refId="ref8972" refString="SMN (2018) Informacion climatologica, database. Servicio Meteorologico Nacional, Mexico. Available from: http: // smn. cna. gob. mx / es / component / content / article? id = 42 (accessed 1 June 2018)" type="url" year="2018">SMN 2018</bibRefCitation>
). The first specimen was found at night (00.10 hrs CST) on a rock wall about 
<quantity id="2A589B4C2726FFFECF3EF8B6D5FBF881" box="[1091,1156,1799,1823]" metricMagnitude="0" metricUnit="m" metricValue="1.8" pageId="8" pageNumber="379" unit="m" value="1.8">1.8 m</quantity>
high and less than a 
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from a waterfall. The air temperature at the time of discovery was 16.9°C with a relative humidity of 70%. Of the 12 observed individuals, 10 were found on a rocky-wall similar to the first specimen, one inside of a pool by the riverside, and one more in vegetation 
<quantity id="2A589B4C2726FFFEC92AF8C2D3F8F815" box="[599,647,1907,1931]" metricMagnitude="0" metricUnit="m" metricValue="2.0" pageId="8" pageNumber="379" unit="m" value="2.0">2 m</quantity>
high from the substrate. The mean air temperature, considering all encounters, was 16.4°C (15.3-17.6) with 75% (70-80%) relative humidity. We recorded two other sympatric frogs, 
<taxonomicName id="2AA04D2A2726FFFECBEAF80BD09AF84C" baseAuthorityName="Ahl" baseAuthorityYear="1934" box="[151,485,1978,2003]" class="Amphibia" family="Hylidae" genus="Ptychohyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="8" pageNumber="379" phylum="Chordata" rank="species" species="leonhardschultzei">
<emphasis id="DFD4EABB2726FFFECBEAF80BD09AF84C" box="[151,485,1978,2003]" italics="true" pageId="8" pageNumber="379">Ptychohyla leonhardschultzei</emphasis>
</taxonomicName>
(categorized as endangered by IUCN; Santos-Barrera 
<emphasis id="DFD4EABB2726FFFECF1CF80DD5E3F84C" box="[1121,1180,1978,2003]" italics="true" pageId="8" pageNumber="379">et al.</emphasis>
2006) and 
<taxonomicName id="2AA04D2A2726FFFFCE58F80BD033FF2F" baseAuthorityName="Taylor" baseAuthorityYear="1939" class="Amphibia" family="Ranidae" genus="Lithobates" higherTaxonomySource="GBIF" kingdom="Animalia" lastPageId="9" lastPageNumber="380" order="Anura" pageId="8" pageNumber="379" phylum="Chordata" rank="species" species="sierramadrensis">
<emphasis id="DFD4EABB2726FFFFCE58F80BD033FF2F" italics="true" lastPageId="9" lastPageNumber="380" pageId="8" pageNumber="379">Lithobates sierramadrensis</emphasis>
</taxonomicName>
, as well as the salamander 
<taxonomicName id="2AA04D2A2727FFFFC9F3FF26D229FF2F" box="[654,854,151,176]" class="Amphibia" family="Plethodontidae" genus="Isthmura" higherTaxonomySource="GBIF" kingdom="Animalia" order="Caudata" pageId="9" pageNumber="380" phylum="Chordata" rank="species" species="maxima">
<emphasis id="DFD4EABB2727FFFFC9F3FF26D229FF2F" box="[654,854,151,176]" italics="true" pageId="9" pageNumber="380">Isthmura maxima</emphasis>
</taxonomicName>
(categorized as endangered by 
<bibRefCitation id="89314B582727FFFFCFBBFF29D364FF4A" author="IUCN SSC Amphibian Specialist Group" pageId="9" pageNumber="380" pagination="53989368" refId="ref7789" refString="IUCN SSC Amphibian Specialist Group. 2016. Isthmura maxima. The IUCN Red List of Threatened Species, 2016, e. T 61907 A 53989368." type="journal article" year="2016">IUCN; IUCN SSC Amphibian Specialist Group 2016</bibRefCitation>
).
</paragraph>
<paragraph id="ED1F36A92727FFFFCBBAFF6ED476FE32" blockId="9.[151,1437,151,680]" pageId="9" pageNumber="380">
During collecting, females and males made low intensity “beeps”, similar to the warning sounds of other treefrogs (e.g. 
<taxonomicName id="2AA04D2A2727FFFFCA44FEB5D34AFE82" box="[313,565,260,285]" class="Amphibia" family="Hylidae" genus="Agalychnis" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="380" phylum="Chordata" rank="species" species="dacnicolor">
<emphasis id="DFD4EABB2727FFFFCA44FEB5D34AFE82" box="[313,565,260,285]" italics="true" pageId="9" pageNumber="380">Agalychnis dacnicolor</emphasis>
</taxonomicName>
; 
<bibRefCitation id="89314B582727FFFFC939FEB5D38BFE83" author="Duellman, W. E." box="[580,756,260,285]" pageId="9" pageNumber="380" refId="ref7389" refString="Duellman, W. E. (2001) The Hylid Frogs of Middle America, Society for the Study of Amphibians and Reptiles, Ithaca, New York, 1170 pp." type="book" year="2001">Duellman 2001</bibRefCitation>
). We observed that the color and tonality varied substantially within the same individual, perhaps due to the temperature or stress of the collection. Moreover, 
<taxonomicName id="2AA04D2A2727FFFFCFB1FE99D443FEDF" authorityName="Jiménez-Arcos &amp; Calzada-Arciniega &amp; Alfaro-Juantorena &amp; Vázquez-Reyes &amp; Blair &amp; Parra-Olea" authorityYear="2019" box="[1228,1340,295,320]" class="Amphibia" family="Hylidae" genus="Charadrahyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="380" phylum="Chordata" rank="species" species="sakbah">
<emphasis id="DFD4EABB2727FFFFCFB1FE99D443FEDF" box="[1228,1340,295,320]" italics="true" pageId="9" pageNumber="380">C. sakbah</emphasis>
</taxonomicName>
shows a marked sexual dimorphism in body size, snout shape, coloration, and hypertrophied webbings. For the other two species with hypertrophied webbings (
<taxonomicName id="2AA04D2A2727FFFFC915FEC1D3C1FE17" authorityName=": Adler &amp; Dennis" authorityYear="1972" box="[616,702,368,392]" class="Amphibia" family="Hylidae" genus="Charadrahyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="380" phylum="Chordata" rank="species" species="trux">
<emphasis id="DFD4EABB2727FFFFC915FEC1D3C1FE17" box="[616,702,368,392]" italics="true" pageId="9" pageNumber="380">C. trux</emphasis>
</taxonomicName>
and 
<taxonomicName id="2AA04D2A2727FFFFC879FEC1D200FE17" box="[772,895,368,392]" class="Amphibia" family="Hylidae" genus="Charadrahyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="380" phylum="Chordata" rank="species" species="tecuani">
<emphasis id="DFD4EABB2727FFFFC879FEC1D200FE17" box="[772,895,368,392]" italics="true" pageId="9" pageNumber="380">C. tecuani</emphasis>
</taxonomicName>
) the adult females are unknown. Therefore, 
<taxonomicName id="2AA04D2A2727FFFFCBEAFE25D0F3FE32" authorityName="Jiménez-Arcos &amp; Calzada-Arciniega &amp; Alfaro-Juantorena &amp; Vázquez-Reyes &amp; Blair &amp; Parra-Olea" authorityYear="2019" box="[151,396,404,429]" class="Amphibia" family="Hylidae" genus="Charadrahyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="380" phylum="Chordata" rank="species" species="sakbah">
<emphasis id="DFD4EABB2727FFFFCBEAFE25D0F3FE32" box="[151,396,404,429]" italics="true" pageId="9" pageNumber="380">Charadrahyla sakbah</emphasis>
</taxonomicName>
is the first species of the genus where the dimorphism in this trait is confirmed.
</paragraph>
<paragraph id="ED1F36A92727FFFFCBBAFE06D5E5FD37" blockId="9.[151,1437,151,680]" pageId="9" pageNumber="380">
Considering the limited knowledge about reproductive biology in frogs of the genus 
<taxonomicName id="2AA04D2A2727FFFFCF05FE06D468FE4F" authorityName="Faivovich, Haddad, Garcia, Frost, Campbell &amp; Wheeler" authorityYear="2005" box="[1144,1303,439,464]" class="Amphibia" family="Hylidae" genus="Charadrahyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="380" phylum="Chordata" rank="genus">
<emphasis id="DFD4EABB2727FFFFCF05FE06D468FE4F" box="[1144,1303,439,464]" italics="true" pageId="9" pageNumber="380">Charadrahyla</emphasis>
</taxonomicName>
, we made a dissection of one of the female 
<typeStatus id="321B880B2727FFFFCA84FE6CD31BFE6A" box="[505,612,477,501]" pageId="9" pageNumber="380" type="paratype">paratypes</typeStatus>
(IBH 30992) in order to determine the oocyte number and pigmentation. The oocytes number estimated was 
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both parts of the oviduct. The pigmentation was yellow and we observed different developmental stages, but no mature oocyte. We found the size of the testes in 
<taxonomicName id="2AA04D2A2727FFFFCF0FFD94D59CFDA2" authorityName="Jiménez-Arcos &amp; Calzada-Arciniega &amp; Alfaro-Juantorena &amp; Vázquez-Reyes &amp; Blair &amp; Parra-Olea" authorityYear="2019" box="[1138,1251,548,573]" class="Amphibia" family="Hylidae" genus="Charadrahyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="380" phylum="Chordata" rank="species" species="sakbah">
<emphasis id="DFD4EABB2727FFFFCF0FFD94D59CFDA2" box="[1138,1251,548,573]" italics="true" pageId="9" pageNumber="380">C. sakbah</emphasis>
</taxonomicName>
(ranging 12.32– 12.66% of SVL) to be smaller than reported in 
<taxonomicName id="2AA04D2A2727FFFFC9C1FDF9D233FDFF" baseAuthorityName="Gunther" baseAuthorityYear="1901" box="[700,844,584,608]" class="Amphibia" family="Hylidae" genus="Charadrahyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="380" phylum="Chordata" rank="species" species="taeniopus">
<emphasis id="DFD4EABB2727FFFFC9C1FDF9D233FDFF" box="[700,844,584,608]" italics="true" pageId="9" pageNumber="380">C. taeniopus</emphasis>
</taxonomicName>
(without measurements but see 
<figureCitation id="759B2A2C2727FFFFCFBCFDF6D47AFDFF" box="[1217,1285,583,608]" captionStart="FIGURE 5" captionStartId="10.[151,250,1976,1998]" captionTargetBox="[161,1425,193,1959]" captionTargetId="figure@10.[161,1425,193,1959]" captionTargetPageId="10" captionText="FIGURE 5. (A) A female (not collected) of C. sakbah in the riverside. (B) Panoramic view of the Chite ku'e river. (C) Typical pool and waterfall where C. sakbah individuals are usually found." httpUri="https://zenodo.org/record/2623756/files/figure.png" pageId="9" pageNumber="380">
Fig. 
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</figureCitation>
in 
<bibRefCitation id="89314B582727FFFFCE53FDF6D1ADFD1A" author="Duellman, W. E." pageId="9" pageNumber="380" pagination="159 - 168" refId="ref7270" refString="Duellman, W. E. (1965) Frogs of the Hyla taeniopus Group. Copeia, 2, 159 - 168. https: // doi. org / 10.2307 / 1440718" type="journal article" year="1965">Duellman 1965</bibRefCitation>
), 
<taxonomicName id="2AA04D2A2727FFFFCB9BFDDCD012FD1A" baseAuthorityName="Duellman" baseAuthorityYear="1968" box="[230,365,620,645]" class="Amphibia" family="Hylidae" genus="Charadrahyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="380" phylum="Chordata" rank="species" species="altipotens">
<emphasis id="DFD4EABB2727FFFFCB9BFDDCD012FD1A" box="[230,365,620,645]" italics="true" pageId="9" pageNumber="380">C altipotens</emphasis>
</taxonomicName>
(average of five males 20.65% of SVL; 
<bibRefCitation id="89314B582727FFFFC84DFDDDD29FFD1A" author="Duellman, W. E." box="[816,992,620,645]" pageId="9" pageNumber="380" pagination="559 - 578" refId="ref7304" refString="Duellman, W. E. (1968) Descriptions of New Hylids Frogs from Mexico and Central America. University of Kansas Publications, Museum of Natural History, 17 (13), 559 - 578. https: // doi. org / 10.5962 / bhl. part. 7138" type="journal article" year="1968">Duellman 1968</bibRefCitation>
), 
<taxonomicName id="2AA04D2A2727FFFFC889FDDCD53DFD1B" authorityName=": Adler &amp; Dennis" authorityYear="1972" box="[1012,1090,621,644]" class="Amphibia" family="Hylidae" genus="Charadrahyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="380" phylum="Chordata" rank="species" species="trux">
<emphasis id="DFD4EABB2727FFFFC889FDDCD53DFD1B" box="[1012,1090,621,644]" italics="true" pageId="9" pageNumber="380">C. trux</emphasis>
</taxonomicName>
(average of two males 20.49% of SVL; 
<bibRefCitation id="89314B582727FFFFCB84FD3ED090FD37" author="Adler, K. &amp; Dennis, D. M." box="[249,495,655,680]" pageId="9" pageNumber="380" pagination="1 - 19" refId="ref6564" refString="Adler, K. &amp; Dennis, D. M. (1972) New tree frogs of the genus Hyla from the cloud forests of western Guerrero, Mexico. Occasional Papers of The Museum of Natural History, University of Kansas, 7, 1 - 19." type="journal article" year="1972">Adler &amp; Dennis 1972</bibRefCitation>
;) and 
<taxonomicName id="2AA04D2A2727FFFFC94FFD21D3E1FD37" box="[562,670,656,680]" class="Amphibia" family="Hylidae" genus="Charadrahyla" higherTaxonomySource="GBIF" kingdom="Animalia" order="Anura" pageId="9" pageNumber="380" phylum="Chordata" rank="species" species="tecuani">
<emphasis id="DFD4EABB2727FFFFC94FFD21D3E1FD37" box="[562,670,656,680]" italics="true" pageId="9" pageNumber="380">C tecuani</emphasis>
</taxonomicName>
(23.2–24.1% of SVL; 
<bibRefCitation id="89314B582727FFFFC8E2FD3ED5F1FD37" author="Campbell, J. A. &amp; Blancas-Hernandez, J. C. &amp; Smith, E. N." box="[927,1166,655,680]" pageId="9" pageNumber="380" pagination="287 - 295" refId="ref6800" refString="Campbell, J. A., Blancas-Hernandez, J. C. &amp; Smith, E. N. (2009) A New Species of Stream-breeding Treefrog of the Genus Charadrahyla (Hylidae) from the Sierra Madre del Sur of Guerrero, Mexico. Copeia, 2009 (2), 287 - 295. https: // doi. org / 10.1643 / CH- 08 - 143" type="journal article" year="2009">
Campbell 
<emphasis id="DFD4EABB2727FFFFCF6EFD20D53CFD37" box="[1043,1091,655,680]" italics="true" pageId="9" pageNumber="380">et al</emphasis>
. 2009
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).
</paragraph>
</subSubSection>
</treatment>
</document>