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<mods:title id="8DF1FE2FE0077B2A37C1EFC9C265089D">Taxonomic review of Cratocerus Dejean, 1829 (Coleoptera, Carabidae) with the description of six new species</mods:title>
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<mods:namePart id="29F630F5E00AD60F058FE42CC283F1AE">Grzymala, Traci L.</mods:namePart>
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<mods:namePart id="42C3DF34452EE25C217DE67AD0996C81">Will, Kipling W.</mods:namePart>
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<treatment id="3401E6D113FB84FC432E04716D2F68C6" ID-GBIF-Taxon="152053639" LSID="urn:lsid:plazi:treatment:3401E6D113FB84FC432E04716D2F68C6" httpUri="http://treatment.plazi.org/id/3401E6D113FB84FC432E04716D2F68C6" lastPageId="5" lastPageNumber="82" pageId="4" pageNumber="81">
<subSubSection id="7C33072B4E4E45C2C501CE852CE59A99" pageId="4" pageNumber="81" type="multiple">
<paragraph id="CA45B46E939EA63762C1C8BFF47FE931" pageId="4" pageNumber="81">Taxon classification Animalia Coleoptera Carabidae</paragraph>
</subSubSection>
<subSubSection id="2C5774B930D3CBF6FD89E762141CBE40" pageId="4" pageNumber="81" type="nomenclature">
<paragraph id="956F2CD782D379796B8D3BEEC8BFCD1F" pageId="4" pageNumber="81">
<taxonomicName id="F85FC4184AAEBC6DC32CFC2C0B36D487" ID-CoL="Z9KR" authority="Dejean, 1829" authorityName="Dejean" authorityYear="1829" class="Insecta" family="Carabidae" genus="Cratocerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cratocerus monilicornis" order="Coleoptera" pageId="4" pageNumber="81" phylum="Arthropoda" rank="species" species="monilicornis">Cratocerus monilicornis Dejean, 1829</taxonomicName>
Figs 1, 9a, 11c
</paragraph>
</subSubSection>
<subSubSection id="99564B2266404F9FFB21BAA2929D175A" pageId="4" pageNumber="81" type="reference_group">
<paragraph id="84C36AA55C4980678A040D3BE52BB625" pageId="4" pageNumber="81">
<taxonomicName id="06F18E7D19E1BDFA50179E6C7A41E798" class="Insecta" family="Carabidae" genus="Cratocerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cratocerus monilicornis" order="Coleoptera" pageId="4" pageNumber="81" phylum="Arthropoda" rank="species" species="monilicornis">Cratocerus monilicornis</taxonomicName>
<bibRefCitation id="3BE2316BDA3EC5DD5A6216A27B172B44" author="Dejean, P" journalOrPublisher="Mequignon-Marvis, Paris" pageId="23" pageNumber="100" title="Species general des Coleopteres de la collection de M. le Comte Dejean. Tome Quatrieme." year="1829">Dejean 1829</bibRefCitation>
: 14-15;
<bibRefCitation id="700794EE2FF44A28AF1AB834BC5F264D" author="Dejean, P" journalOrPublisher="Mequignon-Marvis, Paris" pageId="23" pageNumber="100" title="Iconographie et histoire naturelle Coleopteres d'Europe. Tome Quatrieme." year="1834">Dejean and Boisduval 1834</bibRefCitation>
: 17;
<bibRefCitation id="1B069A2F3E41FA6D17A57B19405911C0" author="Hope, FW" journalOrPublisher="Bohn, H. G., London" pageId="24" pageNumber="101" title="The Coleopterist'S, manual, the second part, containing the predaceous land and water beetles of Linnaeus and Fabricius" year="1838">Hope 1838</bibRefCitation>
: 84;
<bibRefCitation id="663EEFC0053BDAB9B8D4C3EEB666A579" author="Lacordaire, JT" journalOrPublisher="Encyclopedique de Roret, Paris" pageId="24" pageNumber="101" title="Histoire naturelle des insects. Genera des Coleopteres ou expose methodique et critique de tous les genres proposes, jusqu'ici dans et ordre d'insectes. Tome premier contenant les familles des cicindeletes, carabiques, dytiscides, gyrinides et palpicornes." year="1854">Lacordaire 1854</bibRefCitation>
: 263-264;
<bibRefCitation id="93FA713065FA95A36E53335D3D0CCCA0" author="Gemminger, M" journalOrPublisher="EH Gummi, Munich" pageId="24" pageNumber="101" title="Catalogus Coleopterorum hucusque descriptorum synonymicus et systematicus. Tomus I. Cicindelidae-Carabidae." year="1868">Gemminger and Harold 1868</bibRefCitation>
: 250;
<bibRefCitation id="DC5CD579C3BBB6741E494AC9EB6CB0B8" author="Chaudoir, M" journalOrPublisher="Annales de la Societe entomologique de Belgique" pageId="23" pageNumber="100" pagination="5 - 24" title="Essai monographique sur les Drimostomides et les Cratocerides et description d'un genre nouveau de Morionides." volume="15" year="1872">Chaudoir 1872</bibRefCitation>
: 18;
<bibRefCitation id="94F3093766E8F85F7A31E359E72CE845" author="Csiki," editor="Junk, W" journalOrPublisher="Schenkling, S., Berlin" pageId="23" pageNumber="100" title="Coleopterorum Catalogus." year="1929">Csiki 1929</bibRefCitation>
: 493;
<bibRefCitation id="B814D1BD072817CFE1E3B4500BD8D5FF" author="Blackwelder, RE" journalOrPublisher="National Museum, Washington, D. C." pageId="23" pageNumber="100" title="Checklist of the Coleopterous insects of Mexico, Central America, The West Indies, and South America. Part 1. Bulletin of the U. S." year="1944">Blackwelder 1944</bibRefCitation>
: 34; Straneo 1977: 115;
<bibRefCitation id="20D595F7214FD2CB5DD265BD68B685A4" author="Lorenz, W" journalOrPublisher="Lorenz, W., Tutzing, Germany" pageId="24" pageNumber="101" title="A systematic list of extant ground beetles of the world (Coleoptera &quot; Geadephaga &quot;: Trachypachidae and Carabidae incl. Paussinae, Cicindelinae, Rhysodinae)" year="2005">Lorenz 2005</bibRefCitation>
: 248.
</paragraph>
</subSubSection>
<subSubSection id="10CD6A24FE51056AE57DEC7F1A0C9352" pageId="4" pageNumber="81" type="material">
<paragraph id="D47DBB6390210F5D671B2C9942DAEA7E" pageId="4" pageNumber="81">Material.</paragraph>
<paragraph id="9B6F38148986EF8B456D1EE02C0E697D" pageId="4" pageNumber="81">
1 male, labeled: &quot;Museum Paris Montagnes des Orgues Prov de Rio-Janeiro Env. De la Tijuca Massif du Bico Do Papagaio E.R. Wagner 1902 700 m alt. Dicembre&quot; (EMEC654896) borrowed from MNHN; 1 male, labeled: &quot;Brazil, [Santa Catarina] Nova Teutonia,
<geoCoordinate id="678850CA404BD530DF0DBF7D1AC08D20" direction="south" orientation="latitude" precision="925" value="-27.183332">27°11'S</geoCoordinate>
,
<geoCoordinate id="3162313750F84DACBAFB76243531FD70" direction="west" orientation="longitude" precision="925" value="-52.383335">52°23'W</geoCoordinate>
, [-27.183 -52.383], Fritz Plaumann, 4.xi.1939&quot; (no EMEC# assigned for this specimen, examined at MNSM); 1 female, labeled: &quot;Corcovado Brazil GB 27-IV-1962 J. &amp; B. Bechyne leg.&quot; (EMEC654898) borrowed from CUIC; 1 female, labeled: &quot;Argt. Misiones Bemberg 29-1-45 Hayward-Wilink 60 bach&quot; (EMEC654897) borrowed from MNHN.
</paragraph>
</subSubSection>
<subSubSection id="C32AF79CD6029292008906A44A37D3D5" pageId="4" pageNumber="81" type="diagnosis">
<paragraph id="C338607406A9A0F7B1255B76179C3E0B" pageId="4" pageNumber="81">Diagnosis.</paragraph>
<paragraph id="EF420B4DF65A9E796D3A53BEEAD4A4EA" pageId="4" pageNumber="81">This species is easily distinguished from all other congeners by its relatively large overall body size (9.5-12.0 mm), two pairs of supraorbital setae, and rounded lateral lobes of the mentum.</paragraph>
</subSubSection>
<subSubSection id="DF874971AEE177126536A7E19B04884E" lastPageId="5" lastPageNumber="82" pageId="4" pageNumber="81" type="description">
<paragraph id="75D9F515F7E2A34CFB7CA4FA8ADA4844" pageId="4" pageNumber="81">Description.</paragraph>
<paragraph id="6BA62F82175B2A6A49A395CE59ED74A0" lastPageId="5" lastPageNumber="82" pageId="4" pageNumber="81">
Size large for the genus, ABL 9.5-12.0 mm, TW 4.0-5.0 mm.
<pageBreakToken id="BBB84418F518C2A3C3F2971BE3886920" pageId="5" pageNumber="82" start="start">Color</pageBreakToken>
. Head, clypeus, mandibles, pronotum, elytra, and venter rufo-piceous; labrum, antennae, remaining mouthparts, and legs fusco-ferrugineous. Head. HL 1.6-1.9 mm, HW 1.4-1.7 mm. Two supraorbital setae present over each eye. Apices of mandibles slightly curved, apices at less than 45 dedgree angle to mandibular midline. Apical maxillary palpomere length 5
<normalizedToken id="BE115AF8953F55901400FFCAD2285DCC" originalValue="×">x</normalizedToken>
width at base. Mentum with rounded lateral lobes. Thorax. Pronotum (Fig. 9a) with distinct median impression, one distinct basal fovea laterad; one lateral seta located on apicolateral pronotal bead; one posterior seta present on posterior bead; pronotal anterior margin convex, greatest width at center; pronotal width 1.3-1.4
<normalizedToken id="CCDE964B3AF6476A6F8083B4942BB828" originalValue="×">x</normalizedToken>
pronotal length. Elytral length 1.4-1.5
<normalizedToken id="AD815C78D3EEFF01C291DA668E5F7B5A" originalValue="×">x</normalizedToken>
combined elytral width. Legs. Protibia with five distinct, stout spines confined to distal half on lateral margin. Genitalia, male. Median lobe curved sharply, between forty-five and ninety degrees, with respect to basal 1/3, remaining 2/3 with continuing curvature; not expanded apically; length average for genus. Left paramere complexly curved (Figure 11c). Genitalia, female. Spermatheca not examined for this species.
</paragraph>
<caption id="3CAC8D264621BC79F0105433CFEF832D" pageId="5" pageNumber="82">
<paragraph id="0119778DA511A61B63E668F23D157E25" pageId="5" pageNumber="82">
Figure 1. Dorsal habitus image of
<taxonomicName id="526A023FBC6AD3DF81B4660343CBC347" class="Insecta" family="Carabidae" genus="Cratocerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cratocerus monilicornis" order="Coleoptera" pageId="5" pageNumber="82" phylum="Arthropoda" rank="species" species="monilicornis">Cratocerus monilicornis</taxonomicName>
Dejean. Scale bar = 1.0 mm.
</paragraph>
</caption>
</subSubSection>
<subSubSection id="5B725692E7DD3AD0A65CB47A5E5BBA22" pageId="5" pageNumber="82" type="distribution">
<paragraph id="D7AE5662F1804A54BEE0D5C5F59C66A6" pageId="5" pageNumber="82">Distribution.</paragraph>
<paragraph id="5C3E3D0CCFF9B1DB7A15BCEF997CE1C4" pageId="5" pageNumber="82">
This species is geographically distributed in Brazil and northeastern Argentina (Fig. 13). According to
<bibRefCitation id="0F92B0388AA83FAC8B8A3E34A9F9799F" author="Csiki," editor="Junk, W" journalOrPublisher="Schenkling, S., Berlin" pageId="23" pageNumber="100" title="Coleopterorum Catalogus." year="1929">Csiki 1929</bibRefCitation>
and
<bibRefCitation id="E4C8DD7EA8B345C5B49ED1EE75928655" author="Blackwelder, RE" journalOrPublisher="National Museum, Washington, D. C." pageId="23" pageNumber="100" title="Checklist of the Coleopterous insects of Mexico, Central America, The West Indies, and South America. Part 1. Bulletin of the U. S." year="1944">Blackwelder 1944</bibRefCitation>
,
<taxonomicName id="AFC481D5D01331C988D627D21227405C" class="Insecta" family="Carabidae" genus="Cratocerus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Cratocerus monilicornis" order="Coleoptera" pageId="5" pageNumber="82" phylum="Arthropoda" rank="species" species="monilicornis">Cratocerus monilicornis</taxonomicName>
has also been recorded from Venezuela, but specimens from this locality were not examined during this study.
</paragraph>
</subSubSection>
<subSubSection id="A6F5F7E2799345072258478675542571" pageId="5" pageNumber="82" type="additional literature">
<paragraph id="27D2AA12DEC099973BA33B86913CB91B" pageId="5" pageNumber="82">Additional literature.</paragraph>
<paragraph id="D30C858886CCE182637F835C7AC5ADC5" pageId="5" pageNumber="82">
<bibRefCitation id="55704AE8176B8D094012CA34DE4FC365" author="Dejean, P" journalOrPublisher="Mequignon-Marvis, Paris" pageId="23" pageNumber="100" title="Iconographie et histoire naturelle Coleopteres d'Europe. Tome Troisieme." year="1832">Dejean and Boisduval 1832</bibRefCitation>
: fig. 3;
<bibRefCitation id="7282D9EA4E9F44F8163A9EBC47D8E8C3" author="Ragusa," journalOrPublisher="Il Naturalist a Siciliano" pageId="24" pageNumber="101" pagination="201 - 210" title="Catalogo Ragionato dei Coleotteri di Sicilia." volume="6" year="1887">Ragusa 1887</bibRefCitation>
: 210.
</paragraph>
</subSubSection>
</treatment>
</document>