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<document ID-DOI="http://dx.doi.org/10.3897/zookeys.955.51983" ID-GBIF-Dataset="45a98afc-af5a-4734-93d1-205e1ac3694b" ID-PMC="PMC7423785" ID-Pensoft-Pub="1313-2970-955-1" ID-Pensoft-UUID="F7018CF19C6158309D57C83A2F5E9478" ID-PubMed="32855592" ID-ZooBank="C8EF4A4B6F054621B9B5AE47FEE7C217" ModsDocID="1313-2970-955-1" checkinTime="1596691600214" checkinUser="pensoft" docAuthor="Grego, Jozef, Mumladze, Levan, Falniowski, Andrzej, Osikowski, Artur, Rysiewska, Aleksandra, Palatov, Dimitry M. &amp; Hofman, Sebastian" docDate="2020" docId="28ACD3799F4557609D11D2E37785C07B" docLanguage="en" docName="ZooKeys 955: 1-77" docOrigin="ZooKeys 955" docSource="http://dx.doi.org/10.3897/zookeys.955.51983" docTitle="Imeretiopsis iazoni Grego &amp; Mumladze 2020, sp. nov." docType="treatment" docUuid="39F50FD0-D0AC-4DD5-9298-90CFE998515F" docUuidSource="ZooBank" docVersion="5" id="F7018CF19C6158309D57C83A2F5E9478" lastPageNumber="1" masterDocId="F7018CF19C6158309D57C83A2F5E9478" masterDocTitle="Revealing the stygobiotic and crenobiotic molluscan biodiversity hotspot in Caucasus: Part I. The phylogeny of stygobiotic Sadlerianinae Szarowska, 2006 (Mollusca, Gastropoda, Hydrobiidae) from Georgia with descriptions of five new genera and twenty-one new species" masterLastPageNumber="77" masterPageNumber="1" pageNumber="1" updateTime="1668169198162" updateUser="ExternalLinkService">
<mods:mods xmlns:mods="http://www.loc.gov/mods/v3">
<mods:titleInfo>
<mods:title>Revealing the stygobiotic and crenobiotic molluscan biodiversity hotspot in Caucasus: Part I. The phylogeny of stygobiotic Sadlerianinae Szarowska, 2006 (Mollusca, Gastropoda, Hydrobiidae) from Georgia with descriptions of five new genera and twenty-one new species</mods:title>
</mods:titleInfo>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Grego, Jozef</mods:namePart>
<mods:affiliation>Horna Micina 219, 97401, Banska Bystrica, Slovakia</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Mumladze, Levan</mods:namePart>
<mods:affiliation>Institute of Zoology, Ilia State University, Kakutsa Cholokashvili Ave 3 / 5, Tbilisi 0162, Georgia</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0002-2172-6973</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Falniowski, Andrzej</mods:namePart>
<mods:affiliation>Department of Malacology, Institute of Zoology and Biomedical Research, Jagiellonian University, ul. Gronostajowa 9, 30 - 387, Krakow, Poland</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Osikowski, Artur</mods:namePart>
<mods:affiliation>Department of Animal Reproduction, Anatomy and Genomics, University of Agriculture in Krakow, al. Mickiewicza 24 / 28, 30 - 059, Krakow, Poland</mods:affiliation>
<mods:nameIdentifier type="orcid">https://orcid.org/0000-0001-6646-2687</mods:nameIdentifier>
<mods:nameIdentifier type="email">a.osikowski@urk.edu.pl</mods:nameIdentifier>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Rysiewska, Aleksandra</mods:namePart>
<mods:affiliation>Department of Malacology, Institute of Zoology and Biomedical Research, Jagiellonian University, ul. Gronostajowa 9, 30 - 387, Krakow, Poland</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Palatov, Dimitry M.</mods:namePart>
<mods:affiliation>Department of Hydrobiology, Biological Faculty, Moscow State University, 1 - 12 Leninskie Gory, 119991, Moscow, Russian Federation</mods:affiliation>
</mods:name>
<mods:name type="personal">
<mods:role>
<mods:roleTerm>Author</mods:roleTerm>
</mods:role>
<mods:namePart>Hofman, Sebastian</mods:namePart>
<mods:affiliation>Department of Comparative Anatomy, Institute of Zoology and Biomedical Research, Jagiellonian University, ul. Gronostajowa 9, 30 - 387, Krakow, Poland</mods:affiliation>
</mods:name>
<mods:typeOfResource>text</mods:typeOfResource>
<mods:relatedItem type="host">
<mods:titleInfo>
<mods:title>ZooKeys</mods:title>
</mods:titleInfo>
<mods:part>
<mods:date>2020</mods:date>
<mods:detail type="volume">
<mods:number>955</mods:number>
</mods:detail>
<mods:extent unit="page">
<mods:start>1</mods:start>
<mods:end>77</mods:end>
</mods:extent>
</mods:part>
</mods:relatedItem>
<mods:location>
<mods:url>http://dx.doi.org/10.3897/zookeys.955.51983</mods:url>
</mods:location>
<mods:classification>journal article</mods:classification>
<mods:identifier type="DOI">http://dx.doi.org/10.3897/zookeys.955.51983</mods:identifier>
<mods:identifier type="Pensoft-Pub">1313-2970-955-1</mods:identifier>
<mods:identifier type="ZooBank">C8EF4A4B6F054621B9B5AE47FEE7C217</mods:identifier>
<mods:identifier type="Pensoft-UUID">F7018CF19C6158309D57C83A2F5E9478</mods:identifier>
</mods:mods>
<treatment ID-GBIF-Taxon="166096063" LSID="urn:lsid:zoobank.org:act:39F50FD0-D0AC-4DD5-9298-90CFE998515F" httpUri="http://treatment.plazi.org/id/28ACD3799F4557609D11D2E37785C07B" lastPageNumber="1" pageId="0" pageNumber="1">
<subSubSection pageId="0" pageNumber="1" type="nomenclature">
<paragraph pageId="0" pageNumber="1">
<taxonomicName LSID="28ACD379-9F45-5760-9D11-D2E37785C07B" authority="Grego &amp; Mumladze" authorityName="Grego &amp; Mumladze" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Imeretiopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Imeretiopsis iazoni" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="iazoni" status="sp. nov.">Imeretiopsis iazoni Grego &amp; Mumladze</taxonomicName>
<taxonomicNameLabel pageId="0" pageNumber="1">sp. nov.</taxonomicNameLabel>
<figureCitation captionStart="Plate 8" captionStartId="F20" captionText="Plate 8. 1 - 4 Imeretiopsis gorgoleti sp. nov., Racha, Gorgoleti, small spring above the road to Tsakhi: 1 holotype 2 paratype NHMW 113279 3, 4 paratypes 5, 6 I. iazoni sp. nov., Imereti, Kutaisi, Iazoni Cave: 5 holotype 6 paratype 7, 8 I. cameroni sp. nov., Imereti, Kutaisi, Iazoni Cave, paratypes 9 - 13 I. gorgoleti sp. nov., specimens used for molecular and anatomical studies 14 I. gorgoleti sp. nov., operculum. The numbers correspond to individuals, and the letters represent the different views of the same individual. Photograph Sara Schnedl, NHMW, and A. Rysiewska." figureDoi="10.3897/zookeys.955.51983.plate8" httpUri="https://binary.pensoft.net/fig/439170" pageId="0" pageNumber="1">Plate 8 (5-6)</figureCitation>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="type locality">
<paragraph pageId="0" pageNumber="1">Type locality.</paragraph>
<paragraph pageId="0" pageNumber="1">
Georgia • Imereti, Kuatisi, Iazoni (Tskhal-Tsiteli) Cave Spring (იაზონის
<normalizedToken originalValue="იგივე">იგივე</normalizedToken>
<normalizedToken originalValue="წყალ-წითელას">წყალ-წითელას</normalizedToken>
<normalizedToken originalValue="მღვიმე">მღვიმე</normalizedToken>
), right bank of Tskalsitela River Canyon;
<geoCoordinate degrees="42" direction="north" minutes="16" orientation="latitude" precision="15" seconds="18" value="42.271667">42°16'18&quot;N</geoCoordinate>
,
<geoCoordinate degrees="42" direction="east" minutes="44" orientation="longitude" precision="15" seconds="02" value="42.73389">42°44'02&quot;E</geoCoordinate>
; 145 m a.s.l.; sandy sediment inside the cave.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="materials_examined">
<paragraph pageId="0" pageNumber="1">Material.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype</emphasis>
: Georgia • 1 adult, dry; type locality; J. Grego, L. Mumladze and G. Bananashvili leg.; ISU FM-T014-H.
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Paratypes</emphasis>
: same as for holotype, ISU FM-T014-P1/1 dry, coll. JG F1409/1dry.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="diagnosis">
<paragraph pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph pageId="0" pageNumber="1">
The species differs from the most closely related
<taxonomicName authorityName="Grego &amp; Mumladze" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Imeretiopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Imeretiopsis gorgoleti" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="gorgoleti">
<emphasis italics="true" pageId="0" pageNumber="1">Imeretiopsis gorgoleti</emphasis>
</taxonomicName>
sp. nov. by its much smaller, less inflated shells with proportionally smaller and less expanded aperture and by the smaller umbilicus. From the other stygobiotic gastropods of the region with similar shell shape it differs by its smaller shell with the sinuated lateral labral profile. From the sympatric
<taxonomicName lsidName="I. cameroni" pageId="0" pageNumber="1" rank="species" species="cameroni">
<emphasis italics="true" pageId="0" pageNumber="1">I. cameroni</emphasis>
</taxonomicName>
sp. nov. it differs by the much smaller shell, more inflated whorls, flatter apex and more open umbilicus. Measurement comparison
<taxonomicName authorityName="Grego &amp; Mumladze" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Imeretiopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Imeretiopsis" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Imeretiopsis</emphasis>
</taxonomicName>
species is given in Table
<tableCitation captionStart="Table 5" captionStartId="T5" captionText="Table 5. Measurement comparison of species from genus Imeretiopsis gen. nov." httpUri="http://table.plazi.org/id/AF6D0360C47946F5854BAA8037A2EDC3" pageId="0" pageNumber="1" tableUuid="AF6D0360C47946F5854BAA8037A2EDC3">5</tableCitation>
.
</paragraph>
<caption ID-Table-UUID="AF6D0360C47946F5854BAA8037A2EDC3" httpUri="http://table.plazi.org/id/AF6D0360C47946F5854BAA8037A2EDC3" pageId="0" pageNumber="1" start="Table 5" startId="T5">
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" pageId="0" pageNumber="1">Table 5.</emphasis>
Measurement comparison of species from genus
<taxonomicName authorityName="Grego &amp; Mumladze" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Imeretiopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Imeretiopsis" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Imeretiopsis</emphasis>
</taxonomicName>
gen. nov.
</paragraph>
</caption>
<paragraph pageId="0" pageNumber="1">
<table pageId="0" pageNumber="1">
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="2">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Genus species</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="1">H</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="1">W</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="1">BH</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="1">BW</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="1">AH</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="1">AW</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="1">CA</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="2">
<emphasis bold="true" pageId="0" pageNumber="1">H/W</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="2">
<emphasis bold="true" pageId="0" pageNumber="1">AH / AW</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="2">
<emphasis bold="true" pageId="0" pageNumber="1">W / BW</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="2">
<emphasis bold="true" pageId="0" pageNumber="1">H/BH</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="2">
<emphasis bold="true" pageId="0" pageNumber="1">H/AH</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="2">
<emphasis bold="true" pageId="0" pageNumber="1">W / AW</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="2">
<emphasis bold="true" pageId="0" pageNumber="1">H/(W- BW)</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="1">mm</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="1">mm</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="1">mm</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="1">mm</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="1">mm</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="1">mm</emphasis>
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<emphasis bold="true" pageId="0" pageNumber="1">deg.</emphasis>
</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName authorityName="Grego &amp; Mumladze" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Imeretiopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Imeretiopsis gorgoleti" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="gorgoleti">
<emphasis italics="true" pageId="0" pageNumber="1">Imeretiopsis gorgoleti</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="1">Holotype</emphasis>
LT
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2.18</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.44</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.35</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.15</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.9 4</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.82</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">38</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.51</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.15</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.25</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.61</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2.32</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.76</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">7.52</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="2">
<taxonomicName authorityName="Grego &amp; Mumladze" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Imeretiopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Imeretiopsis gorgoleti" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="gorgoleti">
<emphasis italics="true" pageId="0" pageNumber="1">Imeretiopsis gorgoleti</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="1">Paratype</emphasis>
LT
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.64</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.03</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.09</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.85</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.71</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.62</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">35</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.59</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.15</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.21</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.50</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2.31</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.66</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">9.11</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.85</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.02</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.18</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.94</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.76</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.62</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">30</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.82</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.23</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.08</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.57</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2.43</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.64</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">23.72</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName authorityName="Grego &amp; Mumladze" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Imeretiopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Imeretiopsis iazoni" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="iazoni">
<emphasis italics="true" pageId="0" pageNumber="1">Imeretiopsis iazoni</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="1">Holotype</emphasis>
LT
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.47</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.74</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.85</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.68</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.50</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.47</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">35</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.99</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.06</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.09</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.73</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2.94</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.57</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">24.50</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName authorityName="Grego &amp; Mumladze" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Imeretiopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Imeretiopsis iazoni" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="iazoni">
<emphasis italics="true" pageId="0" pageNumber="1">Imeretiopsis iazoni</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="1">Paratype</emphasis>
LT
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.35</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.88</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.82</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.68</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.50</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.41</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">32</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.53</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.22</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.29</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.65</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2.70</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2.15</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">6.75</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="2">
<taxonomicName authorityName="Grego &amp; Mumladze" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Imeretiopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Imeretiopsis cameroni" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="cameroni">
<emphasis italics="true" pageId="0" pageNumber="1">Imeretiopsis cameroni</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="1">Paratype</emphasis>
LT
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2.29</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.94</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.15</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.79</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.71</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.53</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">34</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2.44</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.34</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.19</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.99</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3.23</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.77</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">15.27</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2.12</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.85</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.06</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.71</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.62</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.50</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">32</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2.49</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.24</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.20</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2.00</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3.42</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.70</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">15.14</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName authorityName="Grego &amp; Mumladze" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Imeretiopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Imeretiopsis cameroni" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="cameroni">
<emphasis italics="true" pageId="0" pageNumber="1">Imeretiopsis cameroni</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="1">Holotype</emphasis>
LT
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2.00</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.81</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.96</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.72</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.62</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.49</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">35</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2.47</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.26</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.12</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2.09</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3.24</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.65</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">23.50</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName authorityName="Grego &amp; Palatov" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Imeretiopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Imeretiopsis prometheus" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="prometheus">
<emphasis italics="true" pageId="0" pageNumber="1">Imeretiopsis prometheus</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="1">Holotype</emphasis>
LT
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.66</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.53</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.87</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.72</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.60</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.47</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">33</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3.12</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.27</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.74</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.90</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2.79</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.14</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">-8.67</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="5">
<taxonomicName authorityName="Grego &amp; Palatov" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Imeretiopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Imeretiopsis prometheus" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="prometheus">
<emphasis italics="true" pageId="0" pageNumber="1">Imeretiopsis prometheus</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="1">Topotype</emphasis>
LT
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.63</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.85</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.88</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.75</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.55</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.48</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">32</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.91</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.16</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.13</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.86</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2.95</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.79</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">16.25</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.55</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.63</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.90</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.70</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.53</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.45</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">31</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.95</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.17</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2.32</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.72</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2.95</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3.61</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.68</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.43</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.75</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.75</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.63</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.48</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.38</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">32</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.90</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.27</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.20</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.90</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3.00</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2.00</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">11.40</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.48</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.88</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.85</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.70</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.58</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.45</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">30</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.69</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.28</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.25</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.74</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2.57</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.94</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">8.43</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.70</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.88</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.88</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.73</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.53</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.46</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">34</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.94</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.14</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.21</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.94</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3.24</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.89</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">11.33</td>
</tr>
<tr pageId="0" pageNumber="1">
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">
<taxonomicName authorityName="Grego &amp; Mumladze" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Imeretiopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Imeretiopsis nakeralaensis" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="nakeralaensis">
<emphasis italics="true" pageId="0" pageNumber="1">Imeretiopsis nakeralaensis</emphasis>
</taxonomicName>
sp. nov.
<emphasis bold="true" pageId="0" pageNumber="1">Holotype</emphasis>
LT
</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2.00</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.09</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.02</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.85</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.64</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">0.53</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">29</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.84</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.20</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.28</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">1.96</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">3.13</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">2.04</td>
<td colspan="1" pageId="0" pageNumber="1" rowspan="1">8.55</td>
</tr>
</table>
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="description">
<paragraph pageId="0" pageNumber="1">Description.</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Shell</emphasis>
: rather small, 1.38-1.47 mm high, elongate-conical with four whorls, blunt and flat apex, inflated whorls and deep suture. Umbilicus narrow, almost closed. Shell surface glossy, milky white with irregular growth lines, randomly forming faint, rib-like structures. Aperture irregularly oval, slightly depressed from columellar side and slightly expanded. Lateral labral profile very weakly sinuated, columellar profile rather straight.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Operculum</emphasis>
: not known.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Animal body</emphasis>
: not known.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Holotype measurements</emphasis>
: H-1.47 mm; W-0.74 mm; BH-0.85 mm; BW-0.68 mm; AH-0.50 mm; AW-0.47 mm; CA: 35°.
</paragraph>
<paragraph pageId="0" pageNumber="1">
<emphasis bold="true" italics="true" pageId="0" pageNumber="1">Anatomy</emphasis>
: not known.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="etymology">
<paragraph pageId="0" pageNumber="1">Etymology.</paragraph>
<paragraph pageId="0" pageNumber="1">
Named after the type locality, Iazoni Cave (იაზონის
<normalizedToken originalValue="მღვიმე">მღვიმე</normalizedToken>
) (= Tskal-Tsiteli = Rioni Cave (=
<normalizedToken originalValue="წყალწითელას">წყალწითელას</normalizedToken>
=
<normalizedToken originalValue="რიონის">რიონის</normalizedToken>
<normalizedToken originalValue="მღვიმე">მღვიმე</normalizedToken>
)) in Kutaisi.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="habitat">
<paragraph pageId="0" pageNumber="1">Habitat.</paragraph>
<paragraph pageId="0" pageNumber="1">
Stygobiotic species. See habitat of
<taxonomicName authorityName="Grego &amp; Mumladze" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Imeretiopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Imeretiopsis cameroni" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="cameroni">
<emphasis italics="true" pageId="0" pageNumber="1">Imeretiopsis cameroni</emphasis>
</taxonomicName>
sp. nov.
</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="distribution">
<paragraph pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph pageId="0" pageNumber="1">Only known from the type locality.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="conservation status">
<paragraph pageId="0" pageNumber="1">Conservation status.</paragraph>
<paragraph pageId="0" pageNumber="1">The number of known locations (1) is no more than 5 and EOO is smaller than 20 km2. There is no reason to suppose that AOO, EOO, number of locations, number of subpopulations or the number or mature individuals are declining however due to its extremely small EOO we assessed as Vulnerable (VU) D2.</paragraph>
</subSubSection>
<subSubSection pageId="0" pageNumber="1" type="remarks">
<paragraph pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph pageId="0" pageNumber="1">
The assignment to the genus
<taxonomicName authorityName="Grego &amp; Mumladze" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Imeretiopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Imeretiopsis" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="genus">
<emphasis italics="true" pageId="0" pageNumber="1">Imeretiopsis</emphasis>
</taxonomicName>
gen. nov. is only provisional due to sinuated aperture margins and resemblance to
<taxonomicName lsidName="I. gorgoleti" pageId="0" pageNumber="1" rank="species" species="gorgoleti">
<emphasis italics="true" pageId="0" pageNumber="1">I. gorgoleti</emphasis>
</taxonomicName>
. sp. nov.; molecular data will be needed to determine its true taxonomic status. The type locality has indications of occasional pollution, and most of the stygobiotic
<taxonomicName lsidName="" pageId="0" pageNumber="1" phylum="Mollusca" rank="phylum">Mollusca</taxonomicName>
endemic to the cave (
<taxonomicName lsidName="M. borutzkii" pageId="0" pageNumber="1" rank="species" species="borutzkii">
<emphasis italics="true" pageId="0" pageNumber="1">M. borutzkii</emphasis>
</taxonomicName>
(Shadin, 1932),
<taxonomicName class="Bivalvia" family="Sphaeriidae" genus="Euglesa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Euglesa subterranea" order="Sphaeriida" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="subterranea">
<emphasis italics="true" pageId="0" pageNumber="1">Euglesa subterranea</emphasis>
</taxonomicName>
(Shadin, 1932) and
<taxonomicName authorityName="Grego &amp; Mumladze" authorityYear="2020" class="Gastropoda" family="Hydrobiidae" genus="Imeretiopsis" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Imeretiopsis cameroni" order="Littorinimorpha" pageId="0" pageNumber="1" phylum="Mollusca" rank="species" species="cameroni">
<emphasis italics="true" pageId="0" pageNumber="1">Imeretiopsis cameroni</emphasis>
</taxonomicName>
sp. nov.) have shown declining populations. The new species is scarcer than all of the sympatric species.
</paragraph>
</subSubSection>
</treatment>
</document>
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