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<document id="561F74C3BBBC4484420A35263A9DC46A" ID-CLB-Dataset="9177" ID-DOI="10.11646/zootaxa.4820.3.6" ID-GBIF-Dataset="566f9922-d257-41d4-acdc-2afbdcf1b495" ID-ISSN="1175-5326" ID-Zenodo-Dep="4398178" ID-ZooBank="21DA765A-133B-4419-B108-2E5E6AE0667D" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1609212809856" checkinUser="plazi" docAuthor="Anker, Arthur" docDate="2020" docId="0C397C6DFFF9FF9228D2FA92F918CAA2" docLanguage="en" docName="zootaxa.4820.3.6.pdf" docOrigin="Zootaxa 4820 (3)" docStyle="DocumentStyle:647186512141C8FC8976D5BCC54AEB7D.9:Zootaxa.2013-.journal_article" docStyleId="647186512141C8FC8976D5BCC54AEB7D" docStyleName="Zootaxa.2013-.journal_article" docStyleVersion="9" docTitle="Heteroaxianassa heardi" docType="treatment" docVersion="8" lastPageNumber="536" masterDocId="F0000415FFF2FF9F2845FFD8FA4ACE44" masterDocTitle="Strianassa lerayi gen. et sp. nov., a new laomediid mud-shrimp from the eastern Pacific, with new records of Axianassa ngochoae Anker, 2010 and Heteroaxianassa heardi (Anker, 2011) in the western Pacific (Malacostraca: Decapoda: Gebiidea)" masterLastPageNumber="539" masterPageNumber="523" pageNumber="534" updateTime="1698848381255" updateUser="ExternalLinkService" zenodo-license-document="CLOSED">
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<mods:title id="5B7C30DA10B66C68202F154CBFE9482B">Strianassa lerayi gen. et sp. nov., a new laomediid mud-shrimp from the eastern Pacific, with new records of Axianassa ngochoae Anker, 2010 and Heteroaxianassa heardi (Anker, 2011) in the western Pacific (Malacostraca: Decapoda: Gebiidea)</mods:title>
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<mods:namePart id="2D906207DD07C30FAB05A8C7B746D8A9">Anker, Arthur</mods:namePart>
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<mods:title id="68EEA030E5FB68DA3B8D58150090AB10">Zootaxa</mods:title>
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<mods:date id="BDFE0E1F104BF0F93655DD56BA0D5CCE">2020</mods:date>
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<mods:number id="F4078831A342F738FF537F423A4FB10D">2020-07-29</mods:number>
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<mods:classification id="1A490A5549E5B2434BADA5D8A5BF384B">journal article</mods:classification>
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<mods:identifier id="6FE71218A7C2C3BB8BA6F5B7AD9B025E" type="DOI">10.11646/zootaxa.4820.3.6</mods:identifier>
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<treatment id="0C397C6DFFF9FF9228D2FA92F918CAA2" ID-DOI="http://doi.org/10.5281/zenodo.4398152" ID-GBIF-Taxon="176342419" ID-Zenodo-Dep="4398152" LSID="urn:lsid:plazi:treatment:0C397C6DFFF9FF9228D2FA92F918CAA2" httpUri="http://treatment.plazi.org/id/0C397C6DFFF9FF9228D2FA92F918CAA2" lastPageId="13" lastPageNumber="536" pageId="11" pageNumber="534">
<subSubSection id="CC8A9EF0FFF9FF9428D2FA92F810CB20" box="[151,602,1354,1381]" pageId="11" pageNumber="534" type="nomenclature">
<paragraph id="842FCD7BFFF9FF9428D2FA92F810CB20" blockId="11.[151,602,1354,1416]" box="[151,602,1354,1381]" pageId="11" pageNumber="534">
<heading id="DF677A17FFF9FF9428D2FA92F810CB20" bold="true" box="[151,602,1354,1381]" fontSize="11" level="1" pageId="11" pageNumber="534" reason="1">
<emphasis id="B6E41169FFF9FF9428D2FA92F810CB20" bold="true" box="[151,602,1354,1381]" pageId="11" pageNumber="534">
<emphasis id="B6E41169FFF9FF9428D2FA92FB1ECB20" bold="true" box="[151,340,1354,1380]" italics="true" pageId="11" pageNumber="534">Heteroxianassa</emphasis>
<taxonomicName id="4390B6F8FFF9FF942919FA92F810CB20" authority="(Anker, 2011)" baseAuthorityName="Anker" baseAuthorityYear="2011" box="[348,602,1354,1381]" class="Malacostraca" family="Axianassidae" genus="Heteroaxianassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="534" phylum="Arthropoda" rank="species" species="heardi">
<emphasis id="B6E41169FFF9FF942919FA92FBE0CB21" bold="true" box="[348,426,1354,1381]" italics="true" pageId="11" pageNumber="534">heardi</emphasis>
(
<bibRefCitation id="E001B08AFFF9FF9429FCFA92F818CB20" author="Anker, A." box="[441,594,1354,1380]" pageId="11" pageNumber="534" pagination="1 - 22" refId="ref7779" refString="Anker, A. (2011) Four new infaunal decapod crustaceans (Caridea: Alpheidae and Gebiidea: Axianassidae) from Lizard Island, Australia, one of them also occurring in Moorea, French Polynesia. Zootaxa, 2734 (1), 1 - 22. https: // doi. org / 10.11646 / zootaxa. 2734.1.1" type="journal article" year="2011">Anker, 2011</bibRefCitation>
)
</taxonomicName>
</emphasis>
</heading>
</paragraph>
</subSubSection>
<subSubSection id="CC8A9EF0FFF9FF9428D2FAB6FAA1CBCC" box="[151,235,1390,1416]" pageId="11" pageNumber="534" type="description">
<paragraph id="842FCD7BFFF9FF9428D2FAB6FAA1CBCC" blockId="11.[151,602,1354,1416]" box="[151,235,1390,1416]" pageId="11" pageNumber="534">
(
<figureCitation id="1CABD1FEFFF9FF9428E5FAB6FAA9CBCC" box="[160,227,1390,1416]" captionStart="FIGURE 9" captionStartId="12.[151,250,1932,1956]" captionTargetBox="[151,1436,489,1909]" captionTargetId="figure-182@12.[151,1436,489,1909]" captionTargetPageId="12" captionText="FIGURE 9. Heteroaxianassa heardi (Anker, 2011), male (cl 4.3 mm), MNHN-IU-2013-1466, from Madang, Papua New Guinea: A—living individual, dorsal view; B—same, lateral view. Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.4398200" httpUri="https://zenodo.org/record/4398200/files/figure.png" pageId="11" pageNumber="534">Fig. 9</figureCitation>
)
</paragraph>
</subSubSection>
<subSubSection id="CC8A9EF0FFF9FF9428D2FA6BF805CBAD" pageId="11" pageNumber="534" type="reference_group">
<paragraph id="842FCD7BFFF9FF9428D2FA6BF829CB8F" blockId="11.[151,611,1459,1513]" box="[151,611,1459,1483]" pageId="11" pageNumber="534">
<treatmentCitationGroup id="A480EA55FFF9FF9428D2FA6BF829CB8F" box="[151,611,1459,1483]" pageId="11" pageNumber="534">
<treatmentCitation id="0531EB6AFFF9FF9428D2FA6BFBA6CB8F" ID-DOI="http://doi.org/10.5281/zenodo.6186746" ID-Zenodo-Dep="6186746" author="Anker, A." box="[151,492,1459,1483]" httpUri="http://treatment.plazi.org/id/667AC75A174CFF9A66E0B17447AFC88A" page="14" pageId="11" pageNumber="534" year="2011">
<taxonomicName id="4390B6F8FFF9FF9428D2FA6BFBA6CB8F" ID-CoL="a5b9e104-656f-4a9e-87f7-a9c650984dd3" authority="Anker 2011: 14" authorityName="Anker" authorityPageNumber="14" authorityYear="2011" box="[151,492,1459,1483]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="534" phylum="Arthropoda" rank="species" species="heardi">
<emphasis id="B6E41169FFF9FF9428D2FA6BFB0DCB8F" box="[151,327,1459,1483]" italics="true" pageId="11" pageNumber="534">Axianassa heardi</emphasis>
<bibRefCitation id="E001B08AFFF9FF942909FA6BFBA6CB8F" author="Anker, A." box="[332,492,1459,1483]" pageId="11" pageNumber="534" pagination="1 - 22" refId="ref7779" refString="Anker, A. (2011) Four new infaunal decapod crustaceans (Caridea: Alpheidae and Gebiidea: Axianassidae) from Lizard Island, Australia, one of them also occurring in Moorea, French Polynesia. Zootaxa, 2734 (1), 1 - 22. https: // doi. org / 10.11646 / zootaxa. 2734.1.1" type="journal article" year="2011">Anker 2011: 14</bibRefCitation>
</taxonomicName>
</treatmentCitation>
, figs. 9–14.
</treatmentCitationGroup>
</paragraph>
<paragraph id="842FCD7BFFF9FF9428D2FA0AF805CBAD" blockId="11.[151,611,1459,1513]" box="[151,591,1489,1513]" pageId="11" pageNumber="534">
<treatmentCitationGroup id="A480EA55FFF9FF9428D2FA0AF805CBAD" box="[151,591,1489,1513]" pageId="11" pageNumber="534">
<taxonomicName id="4390B6F8FFF9FF9428D2FA0AFBC3CBAD" baseAuthorityName="Anker" baseAuthorityYear="2011" box="[151,393,1489,1513]" class="Malacostraca" family="Axianassidae" genus="Heteroaxianassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="534" phylum="Arthropoda" rank="species" species="heardi">
<emphasis id="B6E41169FFF9FF9428D2FA0AFBC3CBAD" box="[151,393,1489,1513]" italics="true" pageId="11" pageNumber="534">Heteroaxianassa heardi</emphasis>
</taxonomicName>
—
<treatmentCitation id="0531EB6AFFF9FF9429E6FA09F800CBAD" author="Sakai, K." box="[419,586,1489,1513]" page="597" pageId="11" pageNumber="534" year="2016">
<bibRefCitation id="E001B08AFFF9FF9429E6FA09F800CBAD" author="Sakai, K." box="[419,586,1489,1513]" pageId="11" pageNumber="534" pagination="595 - 602" refId="ref9550" refString="Sakai, K. (2016) An aberrant specimen of the genus Laomedia De Haan, 1841, with an additional description of Laomedia astacina De Haan, 1841 and a new genus, Heteroaxianassa (Decapoda, Thalassinidea 1, Laomediidae). Crustaceana, 89, 595 - 602. https: // doi. org / 10.1163 / 15685403 - 00003535" type="journal article" year="2016">Sakai 2016: 597</bibRefCitation>
</treatmentCitation>
.
</treatmentCitationGroup>
</paragraph>
</subSubSection>
<subSubSection id="CC8A9EF0FFF9FF9428D2F9CFF832C83E" pageId="11" pageNumber="534" type="materials_examined">
<paragraph id="842FCD7BFFF9FF9428D2F9CFF832C83E" blockId="11.[151,1437,1559,2018]" pageId="11" pageNumber="534">
<emphasis id="B6E41169FFF9FF9428D2F9CFFB31C875" bold="true" box="[151,379,1559,1585]" pageId="11" pageNumber="534">Material examined</emphasis>
.
<materialsCitation id="34F8C726FFF9FF9429CEF9C0F83EC83E" ID-GBIF-Occurrence="3005592303" collectingDate="2012-12-07" collectorName="A. Anker" country="Papua New Guinea" elevation="3" latitude="-5.1716666" location="Panab Island" longLatPrecision="130" longitude="145.80833" pageId="11" pageNumber="534" specimenCount="1" specimenCount-male="1" stateProvince="Madang">
<specimenCount id="929606F2FFF9FF9429CEF9C0FB92C875" box="[395,472,1559,1586]" pageId="11" pageNumber="534" type="male">1 male</specimenCount>
(cl
<quantity id="4368609EFFF9FF942A42F9C0F829C876" box="[519,611,1560,1586]" metricMagnitude="-3" metricUnit="m" metricValue="4.3" pageId="11" pageNumber="534" unit="mm" value="4.3">4.3 mm</quantity>
), MNHN-IU-2013-1466,
<collectingCountry id="FC878DEBFFF9FF942BC3F9C0FE2FC875" box="[902,1125,1559,1586]" name="Papua New Guinea" pageId="11" pageNumber="534">Papua New Guinea</collectingCountry>
,
<collectingRegion id="46540399FFF9FF942C34F9CFFE87C875" box="[1137,1229,1559,1585]" country="Papua New Guinea" name="Madang" pageId="11" pageNumber="534">Madang</collectingRegion>
lagoon,
<location id="814F9BA0FFF9FF942D77F9CFFA80C811" LSID="urn:lsid:plazi:treatment:0C397C6DFFF9FF9228D2FA92F918CAA2:814F9BA0FFF9FF942D77F9CFFA80C811" country="Papua New Guinea" latitude="-5.1716666" longLatPrecision="130" longitude="145.80833" name="Panab Island" pageId="11" pageNumber="534" stateProvince="Madang">Panab Island</location>
, sta. PR203,
<geoCoordinate id="E1A4ABBCFFF9FF942918F9E5FB93C81C" box="[349,473,1597,1624]" degrees="05" direction="south" minutes="10.3" orientation="latitude" pageId="11" pageNumber="534" precision="92" value="-5.1716666">05°10.3′S–</geoCoordinate>
<geoCoordinate id="E1A4ABBCFFF9FF94299CF9E5F812C81C" box="[473,600,1597,1624]" degrees="145" direction="east" minutes="48.5" orientation="longitude" pageId="11" pageNumber="534" precision="92" value="145.80833">145°48.5′E</geoCoordinate>
, sandy slope, depth:
<quantity id="4368609EFFF9FF942B07F9E3F9C1C812" box="[834,907,1595,1622]" metricMagnitude="0" metricUnit="m" metricValue="3.0" metricValueMax="5.0" metricValueMin="1.0" pageId="11" pageNumber="534" unit="m" value="3.0" valueMax="5.0" valueMin="1.0">
<elevation id="0FBD2A48FFF9FF942B07F9E3F9C1C812" box="[834,907,1595,1622]" metricMagnitude="0" metricUnit="m" metricValue="3.0" metricValueMax="5.0" metricValueMin="1.0" pageId="11" pageNumber="534" unit="m" value="3.0" valueMax="5.0" valueMin="1.0">1–5 m</elevation>
</quantity>
, scuba diving, suction pump, in burrow, leg.
<collectorName id="2965A8ADFFF9FF942DC4F9E4FA95C83D" pageId="11" pageNumber="534">A. Anker</collectorName>
,
<date id="F02EEBBBFFF9FF9428AFF9B8FBFBC83E" box="[234,433,1631,1658]" pageId="11" pageNumber="534" value="2012-12-07">
<collectingDate id="E06A1253FFF9FF9428AFF9B8FBFBC83E" box="[234,433,1631,1658]" pageId="11" pageNumber="534" value="2012-12-07">7 December 2012</collectingDate>
</date>
[fcn PZD-600C]
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="CC8A9EF0FFF9FF922882F95CF918CAA2" lastPageId="13" lastPageNumber="536" pageId="11" pageNumber="534" type="discussion">
<paragraph id="842FCD7BFFF9FF942882F95CFF28C916" blockId="11.[151,1437,1559,2018]" pageId="11" pageNumber="534">
<emphasis id="B6E41169FFF9FF942882F95CFB7EC8DA" bold="true" box="[199,308,1668,1694]" pageId="11" pageNumber="534">Remarks</emphasis>
. The relatively young but complete male specimen from
<collectingRegion id="46540399FFF9FF942BEEF95BFE4DC8D9" box="[939,1031,1667,1693]" country="Papua New Guinea" name="Madang" pageId="11" pageNumber="534">Madang</collectingRegion>
(
<figureCitation id="1CABD1FEFFF9FF942C50F95BFE1DC8DA" box="[1045,1111,1667,1694]" captionStart="FIGURE 9" captionStartId="12.[151,250,1932,1956]" captionTargetBox="[151,1436,489,1909]" captionTargetId="figure-182@12.[151,1436,489,1909]" captionTargetPageId="12" captionText="FIGURE 9. Heteroaxianassa heardi (Anker, 2011), male (cl 4.3 mm), MNHN-IU-2013-1466, from Madang, Papua New Guinea: A—living individual, dorsal view; B—same, lateral view. Photographs by the author." figureDoi="http://doi.org/10.5281/zenodo.4398200" httpUri="https://zenodo.org/record/4398200/files/figure.png" pageId="11" pageNumber="534">Fig. 9</figureCitation>
) represents the first record of
<taxonomicName id="4390B6F8FFF9FF9428D2F970FB48C885" baseAuthorityName="Anker" baseAuthorityYear="2011" box="[151,258,1703,1729]" class="Malacostraca" family="Axianassidae" genus="Heteroaxianassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="534" phylum="Arthropoda" rank="species" species="heardi">
<emphasis id="B6E41169FFF9FF9428D2F970FB48C885" box="[151,258,1703,1729]" italics="true" pageId="11" pageNumber="534">H. heardi</emphasis>
</taxonomicName>
from
<collectingCountry id="FC878DEBFFF9FF942903F970F869C885" box="[326,547,1703,1730]" name="Papua New Guinea" pageId="11" pageNumber="534">Papua New Guinea</collectingCountry>
. The species was previously known only from the
<typeStatus id="5B2B73D9FFF9FF942C1FF97FFEF6C885" box="[1114,1212,1703,1729]" pageId="11" pageNumber="534" type="holotype">holotype</typeStatus>
from Lizard Island, Great Barrier Reef,
<collectingCountry id="FC878DEBFFF9FF94293DF913FBA8C8A1" box="[376,482,1739,1765]" name="Australia" pageId="11" pageNumber="534">Australia</collectingCountry>
, and Moorea Island, Society Islands,
<collectingCountry id="FC878DEBFFF9FF942BC8F913FE1AC8A1" box="[909,1104,1739,1765]" name="French Polynesia" pageId="11" pageNumber="534">French Polynesia</collectingCountry>
(
<bibRefCitation id="E001B08AFFF9FF942C1AF913FEA6C8A2" author="Anker, A." box="[1119,1260,1739,1766]" pageId="11" pageNumber="534" pagination="1 - 22" refId="ref7779" refString="Anker, A. (2011) Four new infaunal decapod crustaceans (Caridea: Alpheidae and Gebiidea: Axianassidae) from Lizard Island, Australia, one of them also occurring in Moorea, French Polynesia. Zootaxa, 2734 (1), 1 - 22. https: // doi. org / 10.11646 / zootaxa. 2734.1.1" type="journal article" year="2011">Anker 2011</bibRefCitation>
). The relatively broad and not anteriorly produced rostrum, the transverse suture of the uropodal exopod with a row of numerous small teeth and the unarmed distolateral margin of the uropodal exopod are the most obvious features that separate
<taxonomicName id="4390B6F8FFF9FF9428D2F8E0FB4BC915" baseAuthorityName="Anker" baseAuthorityYear="2011" box="[151,257,1847,1873]" class="Malacostraca" family="Axianassidae" genus="Heteroaxianassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="534" phylum="Arthropoda" rank="species" species="heardi">
<emphasis id="B6E41169FFF9FF9428D2F8E0FB4BC915" box="[151,257,1847,1873]" italics="true" pageId="11" pageNumber="534">H. heardi</emphasis>
</taxonomicName>
from its only congener,
<taxonomicName id="4390B6F8FFF9FF942A51F8E0F97EC916" authority="Komai, 2014" authorityName="Komai" authorityYear="2014" box="[532,820,1847,1874]" class="Malacostraca" family="Axianassidae" genus="Heteroaxianassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="534" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B6E41169FFF9FF942A51F8E0F8D2C915" box="[532,664,1848,1873]" italics="true" pageId="11" pageNumber="534">H. japonica</emphasis>
<bibRefCitation id="E001B08AFFF9FF942ADAF8EFF97EC916" author="Komai, T." box="[671,820,1847,1874]" pageId="11" pageNumber="534" pagination="141 - 149" refId="ref8898" refString="Komai, T. (2014) A new species of the mud shrimp genus Axianassa (Crustacea: Decapoda: Gebiidea: Axianassidae) from Japan. Species Diversity, 19, 141 - 149. https: // doi. org / 10.12782 / sd. 19.2.141 s" type="journal article" year="2014">Komai, 2014</bibRefCitation>
</taxonomicName>
, presently known only from
<collectingCountry id="FC878DEBFFF9FF942C33F8E0FEFCC916" box="[1142,1206,1848,1874]" name="Japan" pageId="11" pageNumber="534">Japan</collectingCountry>
(
<bibRefCitation id="E001B08AFFF9FF942C81F8EFFF1CC916" author="Komai, T." box="[1220,1366,1847,1874]" pageId="11" pageNumber="534" pagination="141 - 149" refId="ref8898" refString="Komai, T. (2014) A new species of the mud shrimp genus Axianassa (Crustacea: Decapoda: Gebiidea: Axianassidae) from Japan. Species Diversity, 19, 141 - 149. https: // doi. org / 10.12782 / sd. 19.2.141 s" type="journal article" year="2014">Komai 2014</bibRefCitation>
).
</paragraph>
<paragraph id="842FCD7BFFF9FF932882F883FEC5CF95" blockId="11.[151,1437,1559,2018]" lastBlockId="12.[151,1437,151,465]" lastPageId="12" lastPageNumber="535" pageId="11" pageNumber="534">
The present author tentatively follows
<bibRefCitation id="E001B08AFFF9FF942A36F883F93CC932" author="Komai, T. &amp; Fujita" box="[627,886,1883,1910]" pageId="11" pageNumber="534" pagination="452 - 460" refId="ref9008" refString="Komai, T. &amp; Fujita, (2019) A new species of the mud shrimp genus Axianassa Schmitt, 1924 (Decapoda: Gebiidea: Laomediidae) from the Ryukyu Islands, southern Japan. Zootaxa, 4658 (3), 452 - 460. https: // doi. org / 10.11646 / zootaxa. 4658.3.2" type="journal article" year="2019">Komai &amp; Fujita (2019)</bibRefCitation>
, who recognised the validity of the genus
<taxonomicName id="4390B6F8FFF9FF942D03F884FB4DC9DD" authorityName="Sakai" authorityYear="2016" class="Malacostraca" family="Axianassidae" genus="Heteroaxianassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="534" phylum="Arthropoda" rank="genus">
<emphasis id="B6E41169FFF9FF942D03F884FB4DC9DD" italics="true" pageId="11" pageNumber="534">Heteroaxianassa</emphasis>
</taxonomicName>
.
<bibRefCitation id="E001B08AFFF9FF942953F8A7FBECC9DE" author="Sakai, K." box="[278,422,1919,1946]" pageId="11" pageNumber="534" pagination="595 - 602" refId="ref9550" refString="Sakai, K. (2016) An aberrant specimen of the genus Laomedia De Haan, 1841, with an additional description of Laomedia astacina De Haan, 1841 and a new genus, Heteroaxianassa (Decapoda, Thalassinidea 1, Laomediidae). Crustaceana, 89, 595 - 602. https: // doi. org / 10.1163 / 15685403 - 00003535" type="journal article" year="2016">Sakai (2016)</bibRefCitation>
separated
<taxonomicName id="4390B6F8FFF9FF942A5BF858F891C9DD" authorityName="Sakai" authorityYear="2016" box="[542,731,1920,1945]" class="Malacostraca" family="Axianassidae" genus="Heteroaxianassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="534" phylum="Arthropoda" rank="genus">
<emphasis id="B6E41169FFF9FF942A5BF858F891C9DD" box="[542,731,1920,1945]" italics="true" pageId="11" pageNumber="534">Heteroaxianassa</emphasis>
</taxonomicName>
from
<taxonomicName id="4390B6F8FFF9FF942B65F858F9D9C9DE" authorityName="Schmitt" authorityYear="1924" box="[800,915,1920,1946]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="534" phylum="Arthropoda" rank="genus">
<emphasis id="B6E41169FFF9FF942B65F858F9D9C9DE" box="[800,915,1920,1946]" italics="true" pageId="11" pageNumber="534">Axianassa</emphasis>
</taxonomicName>
based on two differences, the marginally dentate rostrum and the presence of a dentate transverse suture on the uropodal exopod. Both
<bibRefCitation id="E001B08AFFF9FF942CC1F87BFF50C9FA" author="Anker, A." box="[1156,1306,1955,1982]" pageId="11" pageNumber="534" pagination="1 - 22" refId="ref7779" refString="Anker, A. (2011) Four new infaunal decapod crustaceans (Caridea: Alpheidae and Gebiidea: Axianassidae) from Lizard Island, Australia, one of them also occurring in Moorea, French Polynesia. Zootaxa, 2734 (1), 1 - 22. https: // doi. org / 10.11646 / zootaxa. 2734.1.1" type="journal article" year="2011">Anker (2011)</bibRefCitation>
and
<bibRefCitation id="E001B08AFFF9FF942D15F87BFAAEC9A6" author="Komai, T." pageId="11" pageNumber="534" pagination="141 - 149" refId="ref8898" refString="Komai, T. (2014) A new species of the mud shrimp genus Axianassa (Crustacea: Decapoda: Gebiidea: Axianassidae) from Japan. Species Diversity, 19, 141 - 149. https: // doi. org / 10.12782 / sd. 19.2.141 s" type="journal article" year="2014">Komai (2014)</bibRefCitation>
, describing
<taxonomicName id="4390B6F8FFF9FF94292FF810FB9AC9A5" baseAuthorityName="Anker" baseAuthorityYear="2011" box="[362,464,1991,2018]" class="Malacostraca" family="Axianassidae" genus="Heteroaxianassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="534" phylum="Arthropoda" rank="species" species="heardi">
<emphasis id="B6E41169FFF9FF94292FF810FB9AC9A5" box="[362,464,1991,2018]" italics="true" pageId="11" pageNumber="534">A. heardi</emphasis>
</taxonomicName>
and
<taxonomicName id="4390B6F8FFF9FF942A40F810F8CFC9A5" authorityName="Komai" authorityYear="2014" box="[517,645,1992,2018]" class="Malacostraca" family="Axianassidae" genus="Heteroaxianassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="11" pageNumber="534" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B6E41169FFF9FF942A40F810F8CFC9A5" box="[517,645,1992,2018]" italics="true" pageId="11" pageNumber="534">A. japonica</emphasis>
</taxonomicName>
, respectively, were well aware of these differences, but preferred not to establish a new genus awaiting a more comprehensive revision of the axianassid (now laomediid) genera.
<bibRefCitation id="E001B08AFFFEFF932D7CFF4FFB0CCE92" author="Anker, A. &amp; Pachelle, P. P. G." pageId="12" pageNumber="535" pagination="101 - 125" refId="ref8131" refString="Anker, A. &amp; Pachelle, P. P. G. (2016) Mud-shrimps of the genus Axianassa Schmitt, 1924 from Panama, with description of two new species (Decapoda: Gebiidea: Laomediidae). Zootaxa, 4111, 101 - 125. https: // doi. org / 10.11646 / zootaxa. 4111.2.1" type="journal article" year="2016">Anker &amp; Pachelle (2016)</bibRefCitation>
stated: “The exact position of
<taxonomicName id="4390B6F8FFFEFF932AD1FF63F8B0CE91" baseAuthorityName="Anker" baseAuthorityYear="2011" box="[660,762,187,213]" class="Malacostraca" family="Axianassidae" genus="Heteroaxianassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="535" phylum="Arthropoda" rank="species" species="heardi">
<emphasis id="B6E41169FFFEFF932AD1FF63F8B0CE91" box="[660,762,187,213]" italics="true" pageId="12" pageNumber="535">A. heardi</emphasis>
</taxonomicName>
and
<taxonomicName id="4390B6F8FFFEFF932B68FF63F9E6CE90" authorityName="Komai" authorityYear="2014" box="[813,940,187,213]" class="Malacostraca" family="Axianassidae" genus="Heteroaxianassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="535" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B6E41169FFFEFF932B68FF63F9E6CE90" box="[813,940,187,213]" italics="true" pageId="12" pageNumber="535">A. japonica</emphasis>
</taxonomicName>
, as well as
<emphasis id="B6E41169FFFEFF932C6DFF63FE77CE91" box="[1064,1085,187,213]" italics="true" pageId="12" pageNumber="535">S.</emphasis>
[
<taxonomicName id="4390B6F8FFFEFF932C0EFF63FF4CCE91" authorityName="Paiva, Tavares &amp; Silva-Nero" authorityYear="2010" box="[1099,1286,187,213]" class="Malacostraca" family="Laomediidae" genus="Saintlaurentiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="535" phylum="Arthropoda" rank="genus">
<emphasis id="B6E41169FFFEFF932C0EFF63FF4CCE91" box="[1099,1286,187,213]" italics="true" pageId="12" pageNumber="535">Saintlaurentiella</emphasis>
</taxonomicName>
]
<emphasis id="B6E41169FFFEFF932D50FF63FFDFCE91" box="[1301,1429,187,213]" italics="true" pageId="12" pageNumber="535">heterocheir</emphasis>
, can only be clarified through a more comprehensive molecular analysis and complementarily, a rigorous cladistic analysis of morphological characters of all or most taxa currently assigned to the
<taxonomicName id="4390B6F8FFFEFF932C79FEDBFE84CF59" authorityName="Borradaile" authorityYear="1903" box="[1084,1230,259,285]" class="Malacostraca" family="Laomediidae" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="535" phylum="Arthropoda" rank="family">Laomediidae</taxonomicName>
and
<taxonomicName id="4390B6F8FFFEFF932D42FEDBFFD6CF59" authorityName="Schmitt" authorityYear="1924" box="[1287,1436,259,285]" class="Malacostraca" family="Axianassidae" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="535" phylum="Arthropoda" rank="family">Axianassidae</taxonomicName>
… However, we feel that splitting
<taxonomicName id="4390B6F8FFFEFF932A59FEFFF8C5CF05" authorityName="Schmitt" authorityYear="1924" box="[540,655,295,321]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="535" phylum="Arthropoda" rank="genus">
<emphasis id="B6E41169FFFEFF932A59FEFFF8C5CF05" box="[540,655,295,321]" italics="true" pageId="12" pageNumber="535">Axianassa</emphasis>
</taxonomicName>
and describing a new genus for
<taxonomicName id="4390B6F8FFFEFF932BB9FEFFFE2ECF05" baseAuthorityName="Anker" baseAuthorityYear="2011" box="[1020,1124,295,321]" class="Malacostraca" family="Axianassidae" genus="Heteroaxianassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="535" phylum="Arthropoda" rank="species" species="heardi">
<emphasis id="B6E41169FFFEFF932BB9FEFFFE2ECF05" box="[1020,1124,295,321]" italics="true" pageId="12" pageNumber="535">A. heardi</emphasis>
</taxonomicName>
and
<taxonomicName id="4390B6F8FFFEFF932CDEFEFFFF56CF04" authorityName="Komai" authorityYear="2014" box="[1179,1308,295,321]" class="Malacostraca" family="Axianassidae" genus="Heteroaxianassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="535" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B6E41169FFFEFF932CDEFEFFFF56CF04" box="[1179,1308,295,321]" italics="true" pageId="12" pageNumber="535">A. japonica</emphasis>
</taxonomicName>
based only on two characters, one of them being rather insignificant (dentate rostral margin) on a broader phylogenetic scale is not really necessary.” However, the same year,
<bibRefCitation id="E001B08AFFFEFF932AE1FEB7F97FCFCE" author="Sakai, K." box="[676,821,367,394]" pageId="12" pageNumber="535" pagination="595 - 602" refId="ref9550" refString="Sakai, K. (2016) An aberrant specimen of the genus Laomedia De Haan, 1841, with an additional description of Laomedia astacina De Haan, 1841 and a new genus, Heteroaxianassa (Decapoda, Thalassinidea 1, Laomediidae). Crustaceana, 89, 595 - 602. https: // doi. org / 10.1163 / 15685403 - 00003535" type="journal article" year="2016">Sakai (2016)</bibRefCitation>
, clearly in yet another act of his typical “nomenclatural mihilism” (
<bibRefCitation id="E001B08AFFFEFF932953FE4BFBE1CFE9" author="Dubois, A." box="[278,427,403,429]" pageId="12" pageNumber="535" pagination="857 - 863" refId="ref8539" refString="Dubois, A. (2008) A partial but radical solution to the problem of nomenclatural taxonomic inflation and synonymy load. Biological Journal of the Linnean Society, 93, 857 - 863. https: // doi. org / 10.1111 / j. 1095 - 8312.2007.00900. x" type="journal article" year="2008">Dubois 2008</bibRefCitation>
;
<bibRefCitation id="E001B08AFFFEFF9329F2FE4BF828CFE9" author="Evenhuis, N." box="[439,610,403,429]" pageId="12" pageNumber="535" pagination="59 - 68" refId="ref8665" refString="Evenhuis, N. (2008) The &quot; Mihi Itch &quot; - a brief history. Zootaxa, 1890 (1), 59 - 68. https: // doi. org / 10.11646 / zootaxa. 1890.1.3" type="journal article" year="2008">Evenhuis 2008</bibRefCitation>
), quickly claimed the genus
<taxonomicName id="4390B6F8FFFEFF932BE6FE4BFE2ACFE8" authorityName="Sakai" authorityYear="2016" box="[931,1120,403,428]" class="Malacostraca" family="Axianassidae" genus="Heteroaxianassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="535" phylum="Arthropoda" rank="genus">
<emphasis id="B6E41169FFFEFF932BE6FE4BFE2ACFE8" box="[931,1120,403,428]" italics="true" pageId="12" pageNumber="535">Heteroaxianassa</emphasis>
</taxonomicName>
, using the previous authors’ hard descriptive work and without even examining material of the two component species.
</paragraph>
<caption id="D0EF9DF3FFFEFF9328D2F854FE50C98D" ID-DOI="http://doi.org/10.5281/zenodo.4398200" ID-Zenodo-Dep="4398200" httpUri="https://zenodo.org/record/4398200/files/figure.png" pageId="12" pageNumber="535" startId="12.[151,250,1932,1956]" targetBox="[151,1436,489,1909]" targetPageId="12">
<paragraph id="842FCD7BFFFEFF9328D2F854FE50C98D" blockId="12.[151,1436,1932,1993]" pageId="12" pageNumber="535">
<emphasis id="B6E41169FFFEFF9328D2F854FB46C9E1" bold="true" box="[151,268,1932,1957]" pageId="12" pageNumber="535">FIGURE 9</emphasis>
.
<taxonomicName id="4390B6F8FFFEFF93295DF855F8D4C9E1" authority="(Anker, 2011)" baseAuthorityName="Anker" baseAuthorityYear="2011" box="[280,670,1933,1957]" class="Malacostraca" family="Axianassidae" genus="Heteroaxianassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="12" pageNumber="535" phylum="Arthropoda" rank="species" species="heardi">
<emphasis id="B6E41169FFFEFF93295DF855F843C9E1" box="[280,521,1933,1957]" italics="true" pageId="12" pageNumber="535">Heteroaxianassa heardi</emphasis>
(
<bibRefCitation id="E001B08AFFFEFF932A53F855F8DDC9E1" author="Anker, A." box="[534,663,1933,1957]" pageId="12" pageNumber="535" pagination="1 - 22" refId="ref7779" refString="Anker, A. (2011) Four new infaunal decapod crustaceans (Caridea: Alpheidae and Gebiidea: Axianassidae) from Lizard Island, Australia, one of them also occurring in Moorea, French Polynesia. Zootaxa, 2734 (1), 1 - 22. https: // doi. org / 10.11646 / zootaxa. 2734.1.1" type="journal article" year="2011">Anker, 2011</bibRefCitation>
)
</taxonomicName>
, male (cl 4.3 mm), MNHN-IU-2013-1466, from Madang, Papua New Guinea: A—living individual, dorsal view; B—same, lateral view. Photographs by the author.
</paragraph>
</caption>
<paragraph id="842FCD7BFFFFFF922882FF4FFE04CD83" blockId="13.[151,1437,151,1254]" pageId="13" pageNumber="536">
The two characters used by
<bibRefCitation id="E001B08AFFFFFF922A47FF4FF8D9CEF6" author="Sakai, K." box="[514,659,151,178]" pageId="13" pageNumber="536" pagination="595 - 602" refId="ref9550" refString="Sakai, K. (2016) An aberrant specimen of the genus Laomedia De Haan, 1841, with an additional description of Laomedia astacina De Haan, 1841 and a new genus, Heteroaxianassa (Decapoda, Thalassinidea 1, Laomediidae). Crustaceana, 89, 595 - 602. https: // doi. org / 10.1163 / 15685403 - 00003535" type="journal article" year="2016">Sakai (2016)</bibRefCitation>
to separate
<taxonomicName id="4390B6F8FFFFFF922B5FFF4FF99DCEF4" authorityName="Sakai" authorityYear="2016" box="[794,983,151,176]" class="Malacostraca" family="Axianassidae" genus="Heteroaxianassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="536" phylum="Arthropoda" rank="genus">
<emphasis id="B6E41169FFFFFF922B5FFF4FF99DCEF4" box="[794,983,151,176]" italics="true" pageId="13" pageNumber="536">Heteroaxianassa</emphasis>
</taxonomicName>
from
<taxonomicName id="4390B6F8FFFFFF922C59FF4FFEC5CEF5" authorityName="Schmitt" authorityYear="1924" box="[1052,1167,151,177]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="536" phylum="Arthropoda" rank="genus">
<emphasis id="B6E41169FFFFFF922C59FF4FFEC5CEF5" box="[1052,1167,151,177]" italics="true" pageId="13" pageNumber="536">Axianassa</emphasis>
</taxonomicName>
are the same characters discussed by
<bibRefCitation id="E001B08AFFFFFF92296AFF63FB80CE91" author="Anker, A." box="[303,458,187,213]" pageId="13" pageNumber="536" pagination="1 - 22" refId="ref7779" refString="Anker, A. (2011) Four new infaunal decapod crustaceans (Caridea: Alpheidae and Gebiidea: Axianassidae) from Lizard Island, Australia, one of them also occurring in Moorea, French Polynesia. Zootaxa, 2734 (1), 1 - 22. https: // doi. org / 10.11646 / zootaxa. 2734.1.1" type="journal article" year="2011">Anker (2011)</bibRefCitation>
, Komai (2011) and
<bibRefCitation id="E001B08AFFFFFF922AF5FF63F999CE92" author="Anker, A. &amp; Pachelle, P. P. G." box="[688,979,187,214]" pageId="13" pageNumber="536" pagination="101 - 125" refId="ref8131" refString="Anker, A. &amp; Pachelle, P. P. G. (2016) Mud-shrimps of the genus Axianassa Schmitt, 1924 from Panama, with description of two new species (Decapoda: Gebiidea: Laomediidae). Zootaxa, 4111, 101 - 125. https: // doi. org / 10.11646 / zootaxa. 4111.2.1" type="journal article" year="2016">Anker &amp; Pachelle (2016)</bibRefCitation>
, i.e. the marginally dentate rostrum and the presence of a transverse suture (diaeresis) on the uropodal exopod. These two characters need to be reassessed taking into the account all genera currently placed in the
<taxonomicName id="4390B6F8FFFFFF922B56FEDBF9E3CF59" authorityName="Borradaile" authorityYear="1903" box="[787,937,259,285]" class="Malacostraca" family="Laomediidae" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="536" phylum="Arthropoda" rank="family">Laomediidae</taxonomicName>
. In most species of
<taxonomicName id="4390B6F8FFFFFF922CC0FEDBFEBFCF59" authorityName="De Haan" authorityYear="1944" box="[1157,1269,259,285]" class="Malacostraca" family="Laomediidae" genus="Laomedia" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="536" phylum="Arthropoda" rank="genus">
<emphasis id="B6E41169FFFFFF922CC0FEDBFEBFCF59" box="[1157,1269,259,285]" italics="true" pageId="13" pageNumber="536">Laomedia</emphasis>
</taxonomicName>
, the rostrum is dentate, i.e. the lateral margins of the rostrum are furnished with small, widely spaced teeth (e.g.
<bibRefCitation id="E001B08AFFFFFF922C90FEFFFF1DCF05" author="Sakai, K." box="[1237,1367,295,321]" pageId="13" pageNumber="536" pagination="27 - 34" refId="ref9501" refString="Sakai, K. (1962) Systematic studies on Thalassinidea. I. Laomedia astacina De Haan. Publications of the Seto Marine Biological Laboratory, 10, 27 - 34, pls. 5 - 7. https: // doi. org / 10.5134 / 175300" type="journal article" year="1962">Sakai 1962</bibRefCitation>
: pl. V, fig. 4;
<bibRefCitation id="E001B08AFFFFFF92289AFE93FBC3CF21" author="Ngoc-Ho, N." box="[223,393,331,357]" pageId="13" pageNumber="536" pagination="729 - 747" refId="ref9300" refString="Ngoc-Ho, N. (1997) The genus Laomedia De Haan, 1841 with description of a new species from Vietnam (Crustacea, Thalassinidea, Laomediidae), Zoosystema, 19, 729 - 747." type="journal article" year="1997">Ngoc-Ho 1997</bibRefCitation>
: fig. 1A). However, in one species, more precisely
<taxonomicName id="4390B6F8FFFFFF922BF9FE93FE27CF20" authorityName="Ngoc-Ho" authorityYear="1997" box="[956,1133,331,356]" class="Malacostraca" family="Laomediidae" genus="Laomedia" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="536" phylum="Arthropoda" rank="species" species="paucispinosa">
<emphasis id="B6E41169FFFFFF922BF9FE93FE27CF20" box="[956,1133,331,356]" italics="true" pageId="13" pageNumber="536">L. paucispinosa</emphasis>
</taxonomicName>
<bibRefCitation id="E001B08AFFFFFF922C37FE93FF69CF21" author="Ngoc-Ho, N." box="[1138,1315,331,357]" pageId="13" pageNumber="536" pagination="729 - 747" refId="ref9300" refString="Ngoc-Ho, N. (1997) The genus Laomedia De Haan, 1841 with description of a new species from Vietnam (Crustacea, Thalassinidea, Laomediidae), Zoosystema, 19, 729 - 747." type="journal article" year="1997">Ngoc-Ho, 1997</bibRefCitation>
, the rostral margins are largely smooth, except for one minute apical tooth (
<bibRefCitation id="E001B08AFFFFFF922B28FEB7FE51CFCD" author="Ngoc-Ho, N." box="[877,1051,367,393]" pageId="13" pageNumber="536" pagination="729 - 747" refId="ref9300" refString="Ngoc-Ho, N. (1997) The genus Laomedia De Haan, 1841 with description of a new species from Vietnam (Crustacea, Thalassinidea, Laomediidae), Zoosystema, 19, 729 - 747." type="journal article" year="1997">Ngoc-Ho 1997</bibRefCitation>
: figs. 2C, 4C). Thus, the marginal dentition of the rostrum may be intragenerically variable within the family, as in the case of
<taxonomicName id="4390B6F8FFFFFF922CEEFE4BFF51CFE9" authorityName="De Haan" authorityYear="1944" box="[1195,1307,403,429]" class="Malacostraca" family="Laomediidae" genus="Laomedia" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="536" phylum="Arthropoda" rank="genus">
<emphasis id="B6E41169FFFFFF922CEEFE4BFF51CFE9" box="[1195,1307,403,429]" italics="true" pageId="13" pageNumber="536">Laomedia</emphasis>
</taxonomicName>
(with most of the marginal teeth on the rostrum apparently secondarily lost in
<taxonomicName id="4390B6F8FFFFFF922BDAFE6FFE19CF94" authorityName="Ngoc-Ho" authorityYear="1997" box="[927,1107,439,464]" class="Malacostraca" family="Laomediidae" genus="Laomedia" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="536" phylum="Arthropoda" rank="species" species="paucispinosa">
<emphasis id="B6E41169FFFFFF922BDAFE6FFE19CF94" box="[927,1107,439,464]" italics="true" pageId="13" pageNumber="536">L. paucispinosa</emphasis>
</taxonomicName>
). Furthermore, in
<taxonomicName id="4390B6F8FFFFFF922D6CFE6FF85ACFB6" authority="Kensley &amp; Heard, 1990" authorityName="Kensley" authorityYear="1990" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="536" phylum="Arthropoda" rank="species" species="arenaria">
<emphasis id="B6E41169FFFFFF922D6CFE6FFAB2CFB6" italics="true" pageId="13" pageNumber="536">Axianassa arenaria</emphasis>
<bibRefCitation id="E001B08AFFFFFF922945FE00F85ACFB6" author="Kensley, B. &amp; Heard, R." box="[256,528,472,498]" pageId="13" pageNumber="536" pagination="558 - 572" refId="ref8858" refString="Kensley, B. &amp; Heard, R. (1990) The genus Axianassa (Crustacea: Decapoda: Thalassinidea) in the Americas. Proceedings of the Biological Society of Washington, 103, 558 - 572." type="journal article" year="1990">Kensley &amp; Heard, 1990</bibRefCitation>
</taxonomicName>
, the rostrum also bears a minute apical tooth, described as “papilla” by
<bibRefCitation id="E001B08AFFFFFF922D05FE00FB0CCC52" author="Kensley, B. &amp; Heard, R." pageId="13" pageNumber="536" pagination="558 - 572" refId="ref8858" refString="Kensley, B. &amp; Heard, R. (1990) The genus Axianassa (Crustacea: Decapoda: Thalassinidea) in the Americas. Proceedings of the Biological Society of Washington, 103, 558 - 572." type="journal article" year="1990">
Kensley &amp;
<collectingCountry id="FC878DEBFFFFFF9228F1FE24FAB2CC52" box="[180,248,508,534]" name="Heard Island and McDonald Islands" pageId="13" pageNumber="536">Heard</collectingCountry>
(1990
</bibRefCitation>
: fig. 5B). Consequently, the use of this character in the separation between the laomediid genera is at least questionable, as already pointed out by
<bibRefCitation id="E001B08AFFFFFF922AE8FDF8F987CC7F" author="Anker, A. &amp; Pachelle, P. P. G." box="[685,973,544,571]" pageId="13" pageNumber="536" pagination="101 - 125" refId="ref8131" refString="Anker, A. &amp; Pachelle, P. P. G. (2016) Mud-shrimps of the genus Axianassa Schmitt, 1924 from Panama, with description of two new species (Decapoda: Gebiidea: Laomediidae). Zootaxa, 4111, 101 - 125. https: // doi. org / 10.11646 / zootaxa. 4111.2.1" type="journal article" year="2016">Anker &amp; Pachelle (2016)</bibRefCitation>
. Importantly,
<taxonomicName id="4390B6F8FFFFFF922C2FFDF9FEAACC7F" authority="Anker, 2020" authorityName="Anker" authorityYear="2020" box="[1130,1248,545,571]" class="Malacostraca" family="Laomediidae" genus="Strianassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="536" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B6E41169FFFFFF922C2FFDF9FEAACC7F" box="[1130,1248,545,571]" italics="true" pageId="13" pageNumber="536">Strianassa</emphasis>
</taxonomicName>
<emphasis id="B6E41169FFFFFF922CADFDF8FF18CC7E" bold="true" box="[1256,1362,544,570]" pageId="13" pageNumber="536">
<taxonomicNameLabel id="ADD7AC12FFFFFF922CADFDF8FF18CC7E" box="[1256,1362,544,570]" pageId="13" pageNumber="536" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
is different from
<taxonomicName id="4390B6F8FFFFFF92295AFD9CFBC5CC1A" authorityName="De Haan" authorityYear="1944" box="[287,399,580,606]" class="Malacostraca" family="Laomediidae" genus="Laomedia" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="536" phylum="Arthropoda" rank="genus">
<emphasis id="B6E41169FFFFFF92295AFD9CFBC5CC1A" box="[287,399,580,606]" italics="true" pageId="13" pageNumber="536">Laomedia</emphasis>
</taxonomicName>
and
<taxonomicName id="4390B6F8FFFFFF922982FD9DF8CECC1A" authorityName="Sakai" authorityYear="2016" box="[455,644,581,606]" class="Malacostraca" family="Axianassidae" genus="Heteroaxianassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="536" phylum="Arthropoda" rank="genus">
<emphasis id="B6E41169FFFFFF922982FD9DF8CECC1A" box="[455,644,581,606]" italics="true" pageId="13" pageNumber="536">Heteroaxianassa</emphasis>
</taxonomicName>
by the presence of both marginal and dorsal dentition on the rostrum, this configuration being unique within the family. On the other hand, the presence or absence of a transverse suture (diaeresis) on the uropodal exopod may represent a more reliable character for the generic diagnoses in the
<taxonomicName id="4390B6F8FFFFFF922D00FD55FA95CC8E" authorityName="Borradaile" authorityYear="1903" class="Malacostraca" family="Laomediidae" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="536" phylum="Arthropoda" rank="family">Laomediidae</taxonomicName>
. In
<emphasis id="B6E41169FFFFFF922940FD68FB89CC8E" box="[261,451,688,714]" italics="true" pageId="13" pageNumber="536">
<taxonomicName id="4390B6F8FFFFFF922940FD68FB32CC8E" authorityName="De Haan" authorityYear="1944" box="[261,376,688,714]" class="Malacostraca" family="Laomediidae" genus="Laomedia" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="536" phylum="Arthropoda" rank="genus">Laomedia</taxonomicName>
,
<taxonomicName id="4390B6F8FFFFFF9229C7FD69FB89CC8E" authorityName="Nardo" authorityYear="1847" box="[386,451,689,714]" class="Malacostraca" family="Laomediidae" genus="Jaxea" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="536" phylum="Arthropoda" rank="genus">Jaxea</taxonomicName>
</emphasis>
and
<taxonomicName id="4390B6F8FFFFFF9229B2FD68F83ACC8E" authorityName="Kingsley" authorityYear="1897" box="[503,624,688,714]" class="Malacostraca" family="Laomediidae" genus="Naushonia" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="536" phylum="Arthropoda" rank="genus">
<emphasis id="B6E41169FFFFFF9229B2FD68F83ACC8E" box="[503,624,688,714]" italics="true" pageId="13" pageNumber="536">Naushonia</emphasis>
</taxonomicName>
both uropodal rami have a well-developed, complete, i.e. extending from the mesial to the lateral margin of the ramus, transverse suture, typically armed with small teeth (e.g.
<bibRefCitation id="E001B08AFFFFFF922D4AFD0CFFDDCCAA" author="Selbie, C. M." box="[1295,1431,724,750]" pageId="13" pageNumber="536" refId="ref9668" refString="Selbie, C. M. (1914) The Decapoda Reptantia of the coasts of Ireland. Part I. Palinura, Astacura and Anomura (except Pagu- ridea). Dublin: Browne &amp; Nolan, 116 pp., 15 pls. https: // doi. org / 10.5962 / bhl. title. 53734" type="book" year="1914">Selbie 1914</bibRefCitation>
; Hgoc-Ho 1997;
<bibRefCitation id="E001B08AFFFFFF92290FFD20FB98CD56" author="Anker, A." box="[330,466,760,786]" pageId="13" pageNumber="536" pagination="755 - 770" refId="ref7840" refString="Anker, A. (2014) Naushonia draconis sp. nov., a heavily armoured mud shrimp from the Caribbean Sea, and taxonomic status of Espeleonaushonia Juarrero &amp; Martinez-Iglesias, 1997 (Decapoda: Gebiidea: Laomediidae). Marine Biology Research, 10, 755 - 770. https: // doi. org / 10.1080 / 17451000.2013.852684" type="journal article" year="2014">Anker 2014</bibRefCitation>
;
<bibRefCitation id="E001B08AFFFFFF922999FD20F89BCD57" author="Komai, T. &amp; Anker, A." box="[476,721,760,787]" pageId="13" pageNumber="536" pagination="341 - 360" refId="ref8948" refString="Komai, T. &amp; Anker, A. (2015) Additional records of the laomediid mud-shrimp genus Naushonia Kingsley, 1897 (Crustacea: Decapoda: Gebiidea), with a revised identification key. Zootaxa, 3974 (3), 341 - 360. https: // doi. org / 10.11646 / zootaxa. 3974.3.3" type="journal article" year="2015">Komai &amp; Anker 2015</bibRefCitation>
). In
<taxonomicName id="4390B6F8FFFFFF922B45FD20F9F1CD56" authorityName="Paiva, Tavares &amp; Silva-Nero" authorityYear="2010" box="[768,955,760,786]" class="Malacostraca" family="Laomediidae" genus="Saintlaurentiella" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="536" phylum="Arthropoda" rank="genus">
<emphasis id="B6E41169FFFFFF922B45FD20F9F1CD56" box="[768,955,760,786]" italics="true" pageId="13" pageNumber="536">Saintlaurentiella</emphasis>
</taxonomicName>
and
<emphasis id="B6E41169FFFFFF922BB5FD21FED5CD56" box="[1008,1183,761,786]" italics="true" pageId="13" pageNumber="536">Heteroaxianssa</emphasis>
, only the exopod has a complete transverse suture armed with more or less widely spaced teeth; however, in
<taxonomicName id="4390B6F8FFFFFF922C0BFCC5FE98CD72" authorityName="Komai" authorityYear="2014" box="[1102,1234,797,822]" class="Malacostraca" family="Axianassidae" genus="Heteroaxianassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="536" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B6E41169FFFFFF922C0BFCC5FE98CD72" box="[1102,1234,797,822]" italics="true" pageId="13" pageNumber="536">H. japonica</emphasis>
</taxonomicName>
, the endopod also has an incomplete and unarmed suture running from the lateral tooth to about mid-width of the ramus (LeLoeuff &amp; Intès 1974;
<bibRefCitation id="E001B08AFFFFFF92295DFCBCFBD4CD3A" author="Anker, A." box="[280,414,868,894]" pageId="13" pageNumber="536" pagination="1 - 22" refId="ref7779" refString="Anker, A. (2011) Four new infaunal decapod crustaceans (Caridea: Alpheidae and Gebiidea: Axianassidae) from Lizard Island, Australia, one of them also occurring in Moorea, French Polynesia. Zootaxa, 2734 (1), 1 - 22. https: // doi. org / 10.11646 / zootaxa. 2734.1.1" type="journal article" year="2011">Anker 2011</bibRefCitation>
;
<bibRefCitation id="E001B08AFFFFFF9229EDFCBCF87DCD3A" author="Komai, T." box="[424,567,868,894]" pageId="13" pageNumber="536" pagination="141 - 149" refId="ref8898" refString="Komai, T. (2014) A new species of the mud shrimp genus Axianassa (Crustacea: Decapoda: Gebiidea: Axianassidae) from Japan. Species Diversity, 19, 141 - 149. https: // doi. org / 10.12782 / sd. 19.2.141 s" type="journal article" year="2014">Komai 2014</bibRefCitation>
). Finally, in
<taxonomicName id="4390B6F8FFFFFF922AFAFCBDF978CD3B" authorityName="Schmitt" authorityYear="1924" box="[703,818,869,895]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="536" phylum="Arthropoda" rank="genus">
<emphasis id="B6E41169FFFFFF922AFAFCBDF978CD3B" box="[703,818,869,895]" italics="true" pageId="13" pageNumber="536">Axianassa</emphasis>
</taxonomicName>
and
<taxonomicName id="4390B6F8FFFFFF922B23FCBDF996CD3B" authority="Anker, 2020" authorityName="Anker" authorityYear="2020" box="[870,988,869,895]" class="Malacostraca" family="Laomediidae" genus="Strianassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="536" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B6E41169FFFFFF922B23FCBDF996CD3B" box="[870,988,869,895]" italics="true" pageId="13" pageNumber="536">Strianassa</emphasis>
</taxonomicName>
<emphasis id="B6E41169FFFFFF922BA7FCBCFE00CD3A" bold="true" box="[994,1098,868,894]" pageId="13" pageNumber="536">
<taxonomicNameLabel id="ADD7AC12FFFFFF922BA7FCBCFE00CD3A" box="[994,1098,868,894]" pageId="13" pageNumber="536" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
, the uropodal rami are lacking a transverse suture; however, in
<taxonomicName id="4390B6F8FFFFFF922A45FC51F817CDE6" authority="Anker, 2020" authorityName="Anker" authorityYear="2020" box="[512,605,904,931]" class="Malacostraca" family="Laomediidae" genus="Strianassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="536" phylum="Arthropoda" rank="species" species="lerayi" status="sp. nov.">
<emphasis id="B6E41169FFFFFF922A45FC51F817CDE6" box="[512,605,904,931]" italics="true" pageId="13" pageNumber="536">S. lerayi</emphasis>
</taxonomicName>
<emphasis id="B6E41169FFFFFF922A20FC50F8F4CDE6" bold="true" box="[613,702,904,930]" pageId="13" pageNumber="536">
<taxonomicNameLabel id="ADD7AC12FFFFFF922A20FC50F8F4CDE6" box="[613,702,904,930]" pageId="13" pageNumber="536" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
, the exopod bears a row of transversally aligned teeth adjacent to its posterior margin (
<bibRefCitation id="E001B08AFFFFFF9229C4FC74F8C2CD82" author="Kensley, B. &amp; Heard, R." box="[385,648,940,966]" pageId="13" pageNumber="536" pagination="558 - 572" refId="ref8858" refString="Kensley, B. &amp; Heard, R. (1990) The genus Axianassa (Crustacea: Decapoda: Thalassinidea) in the Americas. Proceedings of the Biological Society of Washington, 103, 558 - 572." type="journal article" year="1990">
Kensley &amp;
<collectingCountry id="FC878DEBFFFFFF922A47FC74F80CCD82" box="[514,582,940,966]" name="Heard Island and McDonald Islands" pageId="13" pageNumber="536">Heard</collectingCountry>
1990
</bibRefCitation>
;
<bibRefCitation id="E001B08AFFFFFF922AD7FC74F9D4CD82" author="Anker, A. &amp; Pachelle, P. P. G." box="[658,926,940,966]" pageId="13" pageNumber="536" pagination="101 - 125" refId="ref8131" refString="Anker, A. &amp; Pachelle, P. P. G. (2016) Mud-shrimps of the genus Axianassa Schmitt, 1924 from Panama, with description of two new species (Decapoda: Gebiidea: Laomediidae). Zootaxa, 4111, 101 - 125. https: // doi. org / 10.11646 / zootaxa. 4111.2.1" type="journal article" year="2016">Anker &amp; Pachelle 2016</bibRefCitation>
; present study).
</paragraph>
<paragraph id="842FCD7BFFFFFF922882FC08F918CAA2" blockId="13.[151,1437,151,1254]" pageId="13" pageNumber="536">
Based on the present morphological evidence, the two western Pacific species currently placed in
<taxonomicName id="4390B6F8FFFFFF922D61FC09FAAFCA4A" authorityName="Sakai" authorityYear="2016" class="Malacostraca" family="Axianassidae" genus="Heteroaxianassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="536" phylum="Arthropoda" rank="genus">
<emphasis id="B6E41169FFFFFF922D61FC09FAAFCA4A" italics="true" pageId="13" pageNumber="536">Heteroaxianassa</emphasis>
</taxonomicName>
, viz.
<taxonomicName id="4390B6F8FFFFFF922962FC2DFBDECA4A" baseAuthorityName="Anker" baseAuthorityYear="2011" box="[295,404,1012,1038]" class="Malacostraca" family="Axianassidae" genus="Heteroaxianassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="536" phylum="Arthropoda" rank="species" species="heardi">
<emphasis id="B6E41169FFFFFF922962FC2DFBDECA4A" box="[295,404,1012,1038]" italics="true" pageId="13" pageNumber="536">H. heardi</emphasis>
</taxonomicName>
and
<taxonomicName id="4390B6F8FFFFFF92298BFC2DF81ECA4A" authorityName="Komai" authorityYear="2014" box="[462,596,1013,1038]" class="Malacostraca" family="Axianassidae" genus="Heteroaxianassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="536" phylum="Arthropoda" rank="species" species="japonica">
<emphasis id="B6E41169FFFFFF92298BFC2DF81ECA4A" box="[462,596,1013,1038]" italics="true" pageId="13" pageNumber="536">H. japonica</emphasis>
</taxonomicName>
, undoubtedly represent a small clade within the
<taxonomicName id="4390B6F8FFFFFF922CC1FC2CFF50CA4A" authorityName="Borradaile" authorityYear="1903" box="[1156,1306,1012,1038]" class="Malacostraca" family="Laomediidae" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="536" phylum="Arthropoda" rank="family">Laomediidae</taxonomicName>
. In the two most likely phylogenetic scenarios, this clade either forms a sister clade to the remaining species of
<taxonomicName id="4390B6F8FFFFFF922CB7FBC1FF2FCA77" authorityName="Schmitt" authorityYear="1924" box="[1266,1381,1049,1075]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="536" phylum="Arthropoda" rank="genus">
<emphasis id="B6E41169FFFFFF922CB7FBC1FF2FCA77" box="[1266,1381,1049,1075]" italics="true" pageId="13" pageNumber="536">Axianassa</emphasis>
</taxonomicName>
or is embedded within
<taxonomicName id="4390B6F8FFFFFF922924FBE5FB9ECA13" authorityName="Schmitt" authorityYear="1924" box="[353,468,1085,1111]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="536" phylum="Arthropoda" rank="genus">
<emphasis id="B6E41169FFFFFF922924FBE5FB9ECA13" box="[353,468,1085,1111]" italics="true" pageId="13" pageNumber="536">Axianassa</emphasis>
</taxonomicName>
. Only in the first case, a generic status for this clade, i.e.
<taxonomicName id="4390B6F8FFFFFF922C1AFBE5FF56CA12" authorityName="Sakai" authorityYear="2016" box="[1119,1308,1085,1110]" class="Malacostraca" family="Axianassidae" genus="Heteroaxianassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="536" phylum="Arthropoda" rank="genus">
<emphasis id="B6E41169FFFFFF922C1AFBE5FF56CA12" box="[1119,1308,1085,1110]" italics="true" pageId="13" pageNumber="536">Heteroaxianassa</emphasis>
</taxonomicName>
, is perhaps justifiable, defined mainly by the presence of a complete transverse suture on the uropodal exopod as the only intergenerically valid differentiating character. The description of
<taxonomicName id="4390B6F8FFFFFF922B2AFB5DF9AFCADB" authority="Anker, 2020" authorityName="Anker" authorityYear="2020" box="[879,997,1157,1183]" class="Malacostraca" family="Laomediidae" genus="Strianassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="536" phylum="Arthropoda" rank="genus" status="gen. nov.">
<emphasis id="B6E41169FFFFFF922B2AFB5DF9AFCADB" box="[879,997,1157,1183]" italics="true" pageId="13" pageNumber="536">Strianassa</emphasis>
</taxonomicName>
<emphasis id="B6E41169FFFFFF922BABFB5CFE12CADA" bold="true" box="[1006,1112,1156,1182]" pageId="13" pageNumber="536">
<taxonomicNameLabel id="ADD7AC12FFFFFF922BABFB5CFE12CADA" box="[1006,1112,1156,1182]" pageId="13" pageNumber="536" rank="genus">gen. nov.</taxonomicNameLabel>
</emphasis>
makes the phylogeny of the
<taxonomicName id="4390B6F8FFFFFF9228D2FB70FB63CA86" authorityName="Borradaile" authorityYear="1903" box="[151,297,1192,1218]" class="Malacostraca" family="Laomediidae" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="536" phylum="Arthropoda" rank="family">Laomediidae</taxonomicName>
even more intriguing as the new genus shows clear affinities to both
<taxonomicName id="4390B6F8FFFFFF922C6AFB71FEE8CA87" authorityName="Schmitt" authorityYear="1924" box="[1071,1186,1193,1219]" class="Malacostraca" family="Laomediidae" genus="Axianassa" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="536" phylum="Arthropoda" rank="genus">
<emphasis id="B6E41169FFFFFF922C6AFB71FEE8CA87" box="[1071,1186,1193,1219]" italics="true" pageId="13" pageNumber="536">Axianassa</emphasis>
</taxonomicName>
and
<taxonomicName id="4390B6F8FFFFFF922C9CFB71FFDCCA86" authorityName="Sakai" authorityYear="2016" box="[1241,1430,1193,1218]" class="Malacostraca" family="Axianassidae" genus="Heteroaxianassa" higherTaxonomySource="GBIF" kingdom="Animalia" order="Decapoda" pageId="13" pageNumber="536" phylum="Arthropoda" rank="genus">
<emphasis id="B6E41169FFFFFF922C9CFB71FFDCCA86" box="[1241,1430,1193,1218]" italics="true" pageId="13" pageNumber="536">Heteroaxianassa</emphasis>
</taxonomicName>
, while also exhibiting several, possibly plesiomorphic features.
</paragraph>
</subSubSection>
</treatment>
</document>
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