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<document id="42722445AE69FCFA239B02F2E645B91A" ID-DOI="10.1111/zoj.12077" ID-ISSN="0024-4082" ID-Zenodo-Dep="5290979" IM.materialsCitations_approvedBy="felipe" IM.metadata_approvedBy="felipe" IM.tables_requiresApprovalFor="existingObjects,plazi" IM.taxonomicNames_approvedBy="felipe" checkinTime="1630051897396" checkinUser="marcus" docAuthor="Sluys, Ronald, Solà, Eduard, Gritzalis, Konstantinos, Vila-Farré, Miquel, Mateos, Eduardo &amp; Riutort, Marta" docDate="2013" docId="03DB7724E212837AFC88FC1AFBC7F8BB" docLanguage="en" docName="zoj12077.pdf" docOrigin="Zoological Journal of the Linnean Society 169 (3)" docSource="http://dx.doi.org/10.1111/zoj.12077" docStyle="DocumentStyle:FBF0BF18BB2B256F1F8008F27D80BF37.3:ZoolJLinnSoc.2010-2016.journal_article" docStyleId="FBF0BF18BB2B256F1F8008F27D80BF37" docStyleName="ZoolJLinnSoc.2010-2016.journal_article" docStyleVersion="3" docTitle="Recurva postrema Sluys &amp; Sola 2013, SP. NOV." docType="treatment" docVersion="4" lastPageNumber="541" masterDocId="FFE20F5CE2028368FFD9FFDEFFEFFFD4" masterDocTitle="Integrative delineation of species of Mediterranean freshwater planarians (Platyhelminthes: Tricladida: Dugesiidae)" masterLastPageNumber="547" masterPageNumber="523" pageNumber="539" updateTime="1699140483653" updateUser="plazi">
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<mods:title id="F02B0C0576AE3A6C3179F08BE7CD0EF7">Integrative delineation of species of Mediterranean freshwater planarians (Platyhelminthes: Tricladida: Dugesiidae)</mods:title>
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<mods:namePart id="2F1D803D836C7E29E5DF202819F461F0">Sluys, Ronald</mods:namePart>
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<mods:namePart id="6C8207B6C981908797D04BCC35337525">Solà, Eduard</mods:namePart>
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<mods:namePart id="A99C57CABE13E761BDFB700BF99553C6">Gritzalis, Konstantinos</mods:namePart>
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<mods:namePart id="686C1CBB8A83935199AD219E0A155E73">Vila-Farré, Miquel</mods:namePart>
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<mods:namePart id="3B77AEAAEA4B918AB93F50878E19A82B">Mateos, Eduardo</mods:namePart>
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<mods:namePart id="02966FA2FDD16EA4F77E442C17EFB443">Riutort, Marta</mods:namePart>
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<mods:title id="54BC5295924BBA6E7817508350441553">Zoological Journal of the Linnean Society</mods:title>
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<mods:number id="DC3DA4EDA2748316072D40DE40577D0C">169</mods:number>
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<subSubSection id="C36895B9E2128378FC88FC1AFA8DFC0D" box="[849,1378,964,985]" pageId="16" pageNumber="539" type="nomenclature">
<paragraph id="8BCDC632E2128378FC88FC1AFA8DFC0D" blockId="16.[849,1378,964,1018]" box="[849,1378,964,985]" pageId="16" pageNumber="539">
<heading id="D085715EE2128378FC88FC1AFA8DFC0D" box="[849,1378,964,985]" centered="true" fontSize="9" level="2" pageId="16" pageNumber="539" reason="4">
<taxonomicName id="4C72BDB1E2128378FC88FC1AFB16FC0D" authority="Sluys &amp; Sola" authorityName="Sluys &amp; Sola" authorityYear="2013" box="[849,1273,964,985]" class="Turbellaria" family="Dugesiidae" genus="Recurva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="16" pageNumber="541" phylum="Platyhelminthes" rank="species" species="postrema" status="sp. nov.">
<emphasis id="B9061A20E2128378FC88FC1AFBACFC0C" bold="true" box="[849,1091,964,985]" italics="true" pageId="16" pageNumber="539">RECURVA POSTREMA</emphasis>
SLUYS &amp; SOLÀ
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<taxonomicNameLabel id="A235A75BE2128378FADAFC19FA8DFC0D" box="[1283,1378,964,985]" pageId="16" pageNumber="539" rank="species">SP. NOV.</taxonomicNameLabel>
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<subSubSection id="C36895B9E212837AFBD7FC3AFBC7F8BB" lastPageId="18" lastPageNumber="541" pageId="16" pageNumber="539" type="description">
<paragraph id="8BCDC632E2128378FBD7FC3AFB4AFC2E" blockId="16.[849,1378,964,1018]" box="[1038,1189,996,1018]" pageId="16" pageNumber="539">
(
<figureCitation id="1349DAB7E2128378FBCEFC3AFB73FC2E" box="[1047,1180,996,1018]" captionStart-0="Figure 14" captionStart-1="Figures 15, 16" captionStartId-0="17.[164,243,1103,1122]" captionStartId-1="18.[144,234,1555,1574]" captionTargetBox-0="[217,726,196,1073]" captionTargetBox-1="[302,1259,221,1524]" captionTargetId-0="figure-637@17.[217,726,196,1073]" captionTargetPageId-0="17" captionTargetPageId-1="18" captionText-0="Figure 14. Recurva postrema Sluys &amp; Solà sp. nov. Photograph of external features (scale bar not available)." captionText-1="Figures 15, 16. Recurva postrema Sluys &amp; Solà sp. nov. 15. ZMA V.Pl. 7122.4. Sagittal reconstruction of the copulatory apparatus. 16. ZMA V.Pl. 7122.1. Sagittal reconstruction of the copulatory apparatus." figureDoi-0="http://doi.org/10.5281/zenodo.5297139" figureDoi-1="http://doi.org/10.5281/zenodo.5297141" httpUri-0="https://zenodo.org/record/5297139/files/figure.png" httpUri-1="https://zenodo.org/record/5297141/files/figure.png" pageId="16" pageNumber="539">FIGS 1416</figureCitation>
)
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<paragraph id="8BCDC632E2128378FCFFFBD4FC81FB4F" blockId="16.[806,1421,1034,1454]" pageId="16" pageNumber="539">
<emphasis id="B9061A20E2128378FCFFFBD4FBFFFBCB" box="[806,1040,1034,1055]" italics="true" pageId="16" pageNumber="539">Material examined:</emphasis>
<materialsCitation id="3B1ACC6FE2128378FBFCFBD4FC81FB4F" collectingDate="2009-04-05" collectionCode="ZMA, V" collectorName="NE Laerma &amp; Eduardo Mateos &amp; Eduard Sola" country="Greece" elevation="135" latitude="36.168545" location="Rhodes" longLatPrecision="1" longitude="27.959599" pageId="16" pageNumber="539" specimenCount="1" typeStatus="holotype">
<typeStatus id="54C97890E2128378FBFCFBD4FB60FBF4" box="[1061,1167,1034,1056]" pageId="16" pageNumber="539" type="holotype">Holotype</typeStatus>
:
<collectionCode id="ED635EF7E2128378FB7EFBD4FB30FBCB" box="[1191,1247,1034,1055]" country="Netherlands" httpUri="http://grbio.org/cool/a65z-wv6r" name="Universiteit van Amsterdam, Zoologisch Museum" pageId="16" pageNumber="539">ZMA</collectionCode>
<collectionCode id="ED635EF7E2128378FB2BFBD4FAE8FBF4" box="[1266,1287,1034,1056]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="16" pageNumber="539" type="Museum">V</collectionCode>
.Pl. 7122.1,
<collectorName id="2687A3E4E2128378FCFFFBF7FC21FBEA" box="[806,974,1065,1086]" pageId="16" pageNumber="539">NE Laerma</collectorName>
,
<location id="8EAD90E9E2128378FC2EFBF7FBA2FBEB" LSID="urn:lsid:plazi:treatment:03DB7724E212837AFC88FC1AFBC7F8BB:8EAD90E9E2128378FC2EFBF7FBA2FBEB" box="[1015,1101,1065,1087]" country="Greece" latitude="36.168545" longLatPrecision="1" longitude="27.959599" name="Rhodes" pageId="16" pageNumber="539">Rhodes</location>
,
<collectingCountry id="F36586A2E2128378FBAEFBF7FB27FBEA" box="[1143,1224,1065,1086]" name="Greece" pageId="16" pageNumber="539">Greece</collectingCountry>
,
<geoCoordinate id="EE46A0F5E2128378FB28FBF7FA67FBEA" box="[1265,1416,1064,1087]" degrees="36" direction="north" minutes="10" orientation="latitude" pageId="16" pageNumber="539" precision="1" seconds="6.76" value="36.168545">36°106.76″N</geoCoordinate>
,
<geoCoordinate id="EE46A0F5E2128378FCFFFB99FC26FB89" box="[806,969,1095,1117]" degrees="27" direction="east" minutes="57" orientation="longitude" pageId="16" pageNumber="539" precision="1" seconds="34.55" value="27.959599">27°5734.55″E</geoCoordinate>
, alt.
<quantity id="4C8A6BD7E2128378FBDAFB99FBA4FB89" box="[1027,1099,1095,1117]" metricMagnitude="2" metricUnit="m" metricValue="1.35" pageId="16" pageNumber="539" unit="m" value="135.0">
<elevation id="005F2101E2128378FBDAFB99FBA4FB89" box="[1027,1099,1095,1117]" metricMagnitude="2" metricUnit="m" metricValue="1.35" pageId="16" pageNumber="539" unit="m" value="135.0">135 m</elevation>
</quantity>
,
<date id="FFCCE0F2E2128378FB81FB99FB07FB88" box="[1112,1256,1095,1117]" pageId="16" pageNumber="539" value="2009-04-05">
<collectingDate id="EF88191AE2128378FB81FB99FB07FB88" box="[1112,1256,1095,1117]" pageId="16" pageNumber="539" value="2009-04-05">5 April 2009</collectingDate>
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, coll.
<collectorName id="2687A3E4E2128378FAF2FB99FC96FBAF" pageId="16" pageNumber="539">Eduardo Mateos</collectorName>
and
<collectorName id="2687A3E4E2128378FC1AFBB8FBB2FBA8" box="[963,1117,1126,1148]" pageId="16" pageNumber="539">Eduard Solà</collectorName>
, sagittal sections on six slides.
</materialsCitation>
</paragraph>
<paragraph id="8BCDC632E2128378FCE6FB7AFB93FA7A" blockId="16.[806,1421,1034,1454]" pageId="16" pageNumber="539">
<materialsCitation id="3B1ACC6FE2128378FCE6FB7AFB16FB0C" collectionCode="ZMA, V" pageId="16" pageNumber="539" specimenCount="1" typeStatus="paratype">
<typeStatus id="54C97890E2128378FCE6FB7AFC59FB6D" box="[831,950,1188,1209]" pageId="16" pageNumber="539" type="paratype">Paratypes</typeStatus>
:
<collectionCode id="ED635EF7E2128378FC19FB7DFC17FB6C" box="[960,1016,1187,1208]" country="Netherlands" httpUri="http://grbio.org/cool/a65z-wv6r" name="Universiteit van Amsterdam, Zoologisch Museum" pageId="16" pageNumber="539">ZMA</collectionCode>
<collectionCode id="ED635EF7E2128378FC26FB7DFBFBFB6D" box="[1023,1044,1187,1209]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="16" pageNumber="539" type="Museum">V</collectionCode>
.Pl. 7122.2, ibid., sagittal sections on four slides (not fully mature specimen)
</materialsCitation>
;
<materialsCitation id="3B1ACC6FE2128378FAD8FB1CFB00FB22" collectionCode="V" pageId="16" pageNumber="539" specimenCount="1" typeStatus="paratype">
<collectionCode id="ED635EF7E2128378FAD8FB1CFAF9FB0C" box="[1281,1302,1218,1240]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="16" pageNumber="539" type="Museum">V</collectionCode>
.Pl. 7122.3, ibid., sagittal sections on six slides
</materialsCitation>
;
<materialsCitation id="3B1ACC6FE2128378FADFFB3EFB56FAC1" collectionCode="V" pageId="16" pageNumber="539" specimenCount="1" typeStatus="paratype">
<collectionCode id="ED635EF7E2128378FADFFB3EFAF5FB22" box="[1286,1306,1248,1270]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="16" pageNumber="539" type="Museum">V</collectionCode>
.Pl.7122.4, ibid., sagittal sections on four slides
</materialsCitation>
;
<materialsCitation id="3B1ACC6FE2128378FB1AFB21FB3CFAE7" collectionCode="V" pageId="16" pageNumber="539" specimenCount="1" typeStatus="paratype">
<collectionCode id="ED635EF7E2128378FB1AFB21FB37FAC1" box="[1219,1240,1279,1301]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="16" pageNumber="539" type="Museum">V</collectionCode>
.Pl. 7122.5, ibid., sagittal sections on seven slides
</materialsCitation>
;
<materialsCitation id="3B1ACC6FE2128378FB36FAC3FB01FA86" collectionCode="V" pageId="16" pageNumber="539" specimenCount="1" typeStatus="paratype">
<collectionCode id="ED635EF7E2128378FB36FAC3FAEBFAE7" box="[1263,1284,1309,1331]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="16" pageNumber="539" type="Museum">V</collectionCode>
.Pl. 7122.6, ibid., horizontal sections on four slides
</materialsCitation>
;
<materialsCitation id="3B1ACC6FE2128378FB25FAE2FB5CFAA5" collectionCode="V" pageId="16" pageNumber="539" specimenCount="1" typeStatus="paratype">
<collectionCode id="ED635EF7E2128378FB25FAE2FAFEFA86" box="[1276,1297,1340,1362]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="16" pageNumber="539" type="Museum">V</collectionCode>
.Pl. 7122.7, ibid., sagittal sections on six slides
</materialsCitation>
;
<materialsCitation id="3B1ACC6FE2128378FB66FA85FB4CFA5B" collectionCode="V" pageId="16" pageNumber="539" specimenCount="1" typeStatus="paratype">
<collectionCode id="ED635EF7E2128378FB66FA85FB3BFAA5" box="[1215,1236,1371,1393]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="16" pageNumber="539" type="Museum">V</collectionCode>
.Pl. 7122.8, ibid., sagittal sections on eight slides
</materialsCitation>
;
<materialsCitation id="3B1ACC6FE2128378FB6DFAA7FB97FA7A" collectionCode="V" pageId="16" pageNumber="539" specimenCount="1" typeStatus="paratype">
<collectionCode id="ED635EF7E2128378FB6DFAA7FB26FA5B" box="[1204,1225,1401,1423]" country="Canada" lsid="urn:lsid:biocol.org:col:13946" name="Royal British Columbia Museum - Herbarium" pageId="16" pageNumber="539" type="Museum">V</collectionCode>
.Pl. 7122.9, ibid., sagittal sections on six slides
</materialsCitation>
.
</paragraph>
<paragraph id="8BCDC632E2128378FCFFFA01FBCCF985" blockId="16.[806,1421,1503,1617]" pageId="16" pageNumber="539">
<emphasis id="B9061A20E2128378FCFFFA01FC4BFA20" box="[806,932,1503,1524]" italics="true" pageId="16" pageNumber="539">Etymology:</emphasis>
The specific epithet is derived from the Latin adjective
<emphasis id="B9061A20E2128378FC30FA21FBB2F9C0" box="[1001,1117,1535,1556]" italics="true" pageId="16" pageNumber="539">postremus</emphasis>
, located posteriorly, and alludes to the far posteriorly located position of the copulatory apparatus.
</paragraph>
<paragraph id="8BCDC632E2128379FCFFF95DFF14FA08" blockId="16.[806,1422,1666,1903]" lastBlockId="17.[164,779,1203,1500]" lastPageId="17" lastPageNumber="540" pageId="16" pageNumber="539">
<emphasis id="B9061A20E2128378FCFFF95DFC72F94C" box="[806,925,1667,1688]" italics="true" pageId="16" pageNumber="539">Diagnosis:</emphasis>
Animals slender, with rotund head. Pharynx and copulatory apparatus situated in the far posterior end of the body. Dorsal testes, distributed throughout the body length but anteriormost testes located at a considerable distance behind the brain. Vasa deferentia open asymmetrically into intrabulbar seminal vesicle. Penis papilla asymmetrical, with more or less vertical orientation in the male atrium. Ejaculatory duct opening at the anterior or ventroanterior side of the penis papilla. Ventral or ventroanterior, muscular penial fold present at the point of insertion of the penis papilla. Ovaries located at about 1/4th the distance between the brain and the root of the pharynx. Distal, posterior parts of the oviducts increase in diameter before communicating with an equally wide common oviduct. Bursal canal is surrounded by a well-developed coat of circular muscle.
</paragraph>
<caption id="DF0D96BAE2138379FF7DFB91FCE4FBAB" ID-DOI="http://doi.org/10.5281/zenodo.5297139" ID-Zenodo-Dep="5297139" httpUri="https://zenodo.org/record/5297139/files/figure.png" pageId="17" pageNumber="540" startId="17.[164,243,1103,1122]" targetBox="[217,726,196,1073]" targetPageId="17">
<paragraph id="8BCDC632E2138379FF7DFB91FCE4FBAB" blockId="17.[164,779,1103,1152]" pageId="17" pageNumber="540">
<emphasis id="B9061A20E2138379FF7DFB91FEF6FBB6" bold="true" box="[164,281,1103,1122]" pageId="17" pageNumber="540">Figure 14.</emphasis>
<taxonomicName id="4C72BDB1E2138379FEFDFB91FD4FFBB6" authority="Sluys &amp; Sola" authorityName="Sluys &amp; Sola" authorityYear="2013" box="[292,672,1103,1122]" class="Turbellaria" family="Dugesiidae" genus="Recurva" higherTaxonomySource="GBIF" kingdom="Animalia" order="Tricladida" pageId="17" pageNumber="540" phylum="Platyhelminthes" rank="species" species="postrema" status="sp. nov.">
<emphasis id="B9061A20E2138379FEFDFB91FE10FBB6" bold="true" box="[292,511,1103,1122]" italics="true" pageId="17" pageNumber="540">Recurva postrema</emphasis>
Sluys &amp; Solà
</taxonomicName>
<emphasis id="B9061A20E2138379FD77FB91FCE4FBB6" bold="true" box="[686,779,1103,1122]" pageId="17" pageNumber="540">
<taxonomicNameLabel id="A235A75BE2138379FD77FB91FCE4FBB6" box="[686,779,1103,1122]" pageId="17" pageNumber="540" rank="species">sp. nov.</taxonomicNameLabel>
</emphasis>
Photograph of external features (scale bar not available).
</paragraph>
</caption>
<paragraph id="8BCDC632E2138379FF7DF9D8FE9FF961" blockId="17.[164,779,1542,1717]" pageId="17" pageNumber="540">
<emphasis id="B9061A20E2138379FF7DF9D8FE25F9CF" box="[164,458,1542,1563]" italics="true" pageId="17" pageNumber="540">Ecology and distribution:</emphasis>
The species is known only from the
<typeStatus id="54C97890E2138379FEC1F9FBFEA6F9EE" box="[280,329,1573,1594]" pageId="17" pageNumber="540">type</typeStatus>
locality, where it was collected from stagnant water in a rather dry creek. Specimens were found in high numbers, gliding on the substrate, together with other small, white flatworms of an unknown species.
</paragraph>
<paragraph id="8BCDC632E2138379FF7DF901FAFDFE47" blockId="17.[164,779,1758,1903]" lastBlockId="17.[826,1442,198,1902]" pageId="17" pageNumber="540">
<emphasis id="B9061A20E2138379FF7DF901FEC3F920" box="[164,300,1759,1780]" italics="true" pageId="17" pageNumber="540">Description:</emphasis>
Preserved specimens measure up to
<quantity id="4C8A6BD7E2138379FF7DF923FF16F8C6" box="[164,249,1789,1811]" metricMagnitude="-3" metricUnit="m" metricValue="9.5" pageId="17" pageNumber="540" unit="mm" value="9.5">9.5 mm</quantity>
in length and
<quantity id="4C8A6BD7E2138379FE7DF923FDE8F8C6" box="[420,519,1789,1811]" metricMagnitude="-3" metricUnit="m" metricValue="2.25" pageId="17" pageNumber="540" unit="mm" value="2.25">2.25 mm</quantity>
in width. Notably live specimens are very slender (
<figureCitation id="1349DAB7E2138379FE22F8C5FDA4F8E6" box="[507,587,1819,1842]" captionStart="Figure 14" captionStartId="17.[164,243,1103,1122]" captionTargetBox="[217,726,196,1073]" captionTargetId="figure-637@17.[217,726,196,1073]" captionTargetPageId="17" captionText="Figure 14. Recurva postrema Sluys &amp; Solà sp. nov. Photograph of external features (scale bar not available)." figureDoi="http://doi.org/10.5281/zenodo.5297139" httpUri="https://zenodo.org/record/5297139/files/figure.png" pageId="17" pageNumber="540">Fig. 14</figureCitation>
), with a rotund head that is provided with a pair of close-set eyes, situated in pigment-free patches. Each eye cup houses numerous retinal cells. Behind the eyes, along the lateral margins of the body, there is an auricular streak on either side, at the level of which the head narrows so that there is a more slender neck region. The dorsal surface is finely pigmented pale brown, with notable accumulations of pigment around the pharyngeal pocket. Ventral surface pale.
</paragraph>
<paragraph id="8BCDC632E2138379FC8AFE42FAE9FD5C" blockId="17.[826,1442,198,1902]" pageId="17" pageNumber="540">
The pharynx measures between 1/6th and 1/8th of the body length and is positioned far into the posterior part of the body. The musculature of the pharynx conforms to the planariid
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. This highly posterior location of the pharynx means that the copulatory apparatus is pushed far into the tail end of the animal. The mouth opening is located at the posterior end of the pharyngeal pocket.
</paragraph>
<paragraph id="8BCDC632E2138379FC8AFD4FFC52F94C" blockId="17.[826,1442,198,1902]" pageId="17" pageNumber="540">
The testes are located dorsally, extending from directly behind the ovaries to almost the posterior margin of the body. After having penetrated the penis bulb, the vasa deferentia open separately into the intrabulbar seminal vesicle. The openings of the seminal ducts are asymmetrical in that one vas deferens opens into the ventral section and the other in a more dorsal section of the seminal vesicle (
<figureCitation id="1349DAB7E2138379FC9BFC5BFC2AFC4F" box="[834,965,901,923]" captionStart="Figures 15, 16" captionStartId="18.[144,234,1555,1574]" captionTargetBox="[302,1259,221,1524]" captionTargetPageId="18" captionText="Figures 15, 16. Recurva postrema Sluys &amp; Solà sp. nov. 15. ZMA V.Pl. 7122.4. Sagittal reconstruction of the copulatory apparatus. 16. ZMA V.Pl. 7122.1. Sagittal reconstruction of the copulatory apparatus." figureDoi="http://doi.org/10.5281/zenodo.5297141" httpUri="https://zenodo.org/record/5297141/files/figure.png" pageId="17" pageNumber="540">Figs 15, 16</figureCitation>
). The latter, lined with a nucleated epithelium and surrounded by a coat of intermingled muscle, communicates with the ejaculatory duct, which in most of the specimens examined exhibits an S-shaped loop before curving downwards to follow its central course through the penis papilla. The papilla is more or less cylindrical in shape and has a more or less vertical orientation in the male atrium. The penis papilla is highly asymmetrical in the sense that in its distal, ventral section the ejaculatory duct shows a sharp, anteriorly directed, knee-shaped bend, after which it opens at the anterior or ventroanterior side of the penis papilla (
<figureCitation id="1349DAB7E2138379FB0DFB2AFAB8FADE" box="[1236,1367,1268,1290]" captionStart="Figures 15, 16" captionStartId="18.[144,234,1555,1574]" captionTargetBox="[302,1259,221,1524]" captionTargetPageId="18" captionText="Figures 15, 16. Recurva postrema Sluys &amp; Solà sp. nov. 15. ZMA V.Pl. 7122.4. Sagittal reconstruction of the copulatory apparatus. 16. ZMA V.Pl. 7122.1. Sagittal reconstruction of the copulatory apparatus." figureDoi="http://doi.org/10.5281/zenodo.5297141" httpUri="https://zenodo.org/record/5297141/files/figure.png" pageId="17" pageNumber="540">Figs 15, 16</figureCitation>
). This course of the ejaculatory duct results in the situation that the anterior portion or lip of the penis papilla is shorter and smaller, in some specimens much shorter and smaller, than the posterior section. At the base of this anterior or ventro-anterior lip of the penis papilla, at its point of insertion, a penial fold is present. This fold is characterized by a more or less developed outbulging and is provided with its own musculature. It is a penial fold, in contrast to an atrial fold, because it is located entally to the point of attachment of the musculature of the penis bulb. The penis papilla is covered with a thin, nucleated epithelium.
</paragraph>
<paragraph id="8BCDC632E2138379FC8AF97FFAD8F8E5" blockId="17.[826,1442,198,1902]" pageId="17" pageNumber="540">
The ovaries are located at about 1/4th the distance between the brain and the root of the pharynx. This implies that also the row of testes starts at a considerable distance posterior to the brain, as may be observed even in living specimens (
<figureCitation id="1349DAB7E2138379FB00F8C5FAC5F8E5" box="[1241,1322,1819,1841]" captionStart="Figure 14" captionStartId="17.[164,243,1103,1122]" captionTargetBox="[217,726,196,1073]" captionTargetId="figure-637@17.[217,726,196,1073]" captionTargetPageId="17" captionText="Figure 14. Recurva postrema Sluys &amp; Solà sp. nov. Photograph of external features (scale bar not available)." figureDoi="http://doi.org/10.5281/zenodo.5297139" httpUri="https://zenodo.org/record/5297139/files/figure.png" pageId="17" pageNumber="540">Fig. 14</figureCitation>
).
</paragraph>
<paragraph id="8BCDC632E2138379FC8AF8E4FA4EF8BA" blockId="17.[826,1442,198,1902]" pageId="17" pageNumber="540">Directly posterior to the gonopore the oviducts turn dorso-medially, while their diameter increases</paragraph>
<paragraph id="8BCDC632E210837AFC61FF15FBC7F8BB" blockId="18.[952,1119,199,222]" lastBlockId="18.[806,1421,1697,1903]" pageId="18" pageNumber="541">sv cb considerably. Subsequently, the oviducts fuse to form a common oviduct, with an equally wide diameter, that opens into the ventral section of the bursal canal. The latter starts at the copulatory bursa as a rather narrow duct that gradually widens and posterior to the gonopore makes a sharp anteriorly directed bend before opening into the rather dorsal section of the atrium. The more or less horizontally running and widened part of the bursal canal receives the openings of the shell glands anteriorly to the opening of the common oviduct. The nucleated bursal canal is surrounded by a well-developed coat of circular muscle. The copulatory bursa sits immediately anterior to the penis bulb.</paragraph>
</subSubSection>
</treatment>
</document>