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<mods:title id="CA82CCC29EAAFDDBBE601969B59274A2">Taxonomic Revision of the genus Mesomphalia Hope, 1839 (Insecta, Coleoptera, Chrysomelidae)</mods:title>
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<taxonomicName id="83B13FC04A43FFABFF2BF98CFE3D0B07" ID-CoL="62Z7Y" authority="Hope, 1839" authorityName="Hope" authorityYear="1839" box="[151,463,1553,1579]" class="Insecta" family="Chrysomelidae" genus="Mesomphalia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="genus">
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<emphasis id="76C598514A43FFABFF2BF98CFECE0B07" bold="true" box="[151,316,1553,1579]" italics="true" pageId="2" pageNumber="153">Mesomphalia</emphasis>
<bibRefCitation id="202039B24A43FFABFEFFF98CFE3D0B07" author="Hope" box="[323,463,1553,1579]" pageId="2" pageNumber="153" refString="Hope, F. W. (1839) Observations on the tortoise or shield beetles, commonly denominated Cassida by Linnaeus, with the characters of six new genera. Annals of Natural History, 3 (15), 92 - 100. http: // dx. doi. org / 10.1080 / 03745483909443203" type="journal article" year="1839">Hope, 1839</bibRefCitation>
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<taxonomicName id="83B13FC04A43FFABFF2BF9C5FE3E0B42" ID-CoL="62Z7Y" authority="Hope, 1839: 94" authorityName="Hope" authorityPageNumber="94" authorityYear="1839" box="[151,460,1624,1646]" class="Insecta" family="Chrysomelidae" genus="Mesomphalia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="genus">
<emphasis id="76C598514A43FFABFF2BF9C5FED30B42" box="[151,289,1624,1646]" italics="true" pageId="2" pageNumber="153">Mesomphalia</emphasis>
<treatmentCitation id="C51062524A43FFABFE95F9C4FE3E0B42" author="Hope" box="[297,460,1624,1646]" page="94" pageId="2" pageNumber="153" year="1839">
<bibRefCitation id="202039B24A43FFABFE95F9C4FE560B42" author="Hope" box="[297,420,1624,1646]" pageId="2" pageNumber="153" refString="Hope, F. W. (1839) Observations on the tortoise or shield beetles, commonly denominated Cassida by Linnaeus, with the characters of six new genera. Annals of Natural History, 3 (15), 92 - 100. http: // dx. doi. org / 10.1080 / 03745483909443203" type="journal article" year="1839">Hope, 1839</bibRefCitation>
: 94
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;
<treatmentCitation id="C51062524A43FFABFE65F9C5FD570B42" author="Chapuis" box="[473,677,1624,1646]" page="394" pageId="2" pageNumber="153" year="1875">
<bibRefCitation id="202039B24A43FFABFE65F9C5FD830B42" author="Chapuis" box="[473,625,1624,1646]" pageId="2" pageNumber="153" refString="Chapuis, F. (1875) Histoire naturelle des insects. Genera des coleopteres ou expos methodique et critique de tous les genres proposes jusqu'ici dans cet order d'insectes. Tome onzieme. Famille des phytophages. Librairie Encyclopedique de Roret, Paris, 420 pp." type="book" year="1875">Chapuis, 1875</bibRefCitation>
: 394
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<bibRefCitation id="202039B24A43FFABFD0DF9C5FCA50B42" author="Boheman" box="[689,855,1624,1646]" pageId="2" pageNumber="153" refString="Boheman, C. H. (1850) Monographia Cassididarum I. Officina Nordstedtiana, Holmiae, 452 pp." type="book" year="1850">Boheman, 1850</bibRefCitation>
: 112
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, 1856: 37;
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: 394
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;
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<bibRefCitation id="202039B24A43FFABFB6FF9C4FA9D0B42" author="Wagener" box="[1235,1391,1624,1646]" pageId="2" pageNumber="153" refString="Wagener, B. (1881) Cassididae. Mitteilungen Munchen Entomologischen Vereins, 1877, 49 - 79." type="journal article" year="1881">Wagener, 1881</bibRefCitation>
: 62
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;
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<bibRefCitation id="202039B24A43FFABFF7BF9EBFEBC0BA0" author="Spaeth" box="[199,334,1654,1676]" pageId="2" pageNumber="153" refString="Spaeth, F. (1914) Chrysomelidae: 16. Cassidinae. In: Junk, W. &amp; Schenkling S. (Eds.), Coleopterorum Catalogus. Pars 62. Berlin, pp. 1 - 182." type="journal article" year="1914">Spaeth, 1914</bibRefCitation>
: 32
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;
<treatmentCitation id="C51062524A43FFABFEC3F9EBFD100BA0" author="Jolivet" box="[383,738,1654,1676]" page="163" pageId="2" pageNumber="153" year="1995">
<bibRefCitation id="202039B24A43FFABFEC3F9EBFD5D0BA0" author="Jolivet" box="[383,687,1654,1676]" pageId="2" pageNumber="153" refString="Jolivet, P. H. &amp; Hawkeswood, T. J. (1995) Host-Plants of Chrysomelidae of the World. (an essay about the relationships between the leaf-beetles and their food plants). Backhuys Publishers, Leiden, 281 pp." type="book" year="1995">Jolivet &amp; Hawkeswood, 1995</bibRefCitation>
: 163
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;
<treatmentCitation id="C51062524A43FFABFD50F9EBFC310BA0" author="Borowiec" box="[748,963,1654,1676]" page="117" pageId="2" pageNumber="153" year="1999">
<bibRefCitation id="202039B24A43FFABFD50F9EBFC600BA0" author="Borowiec" box="[748,914,1654,1676]" pageId="2" pageNumber="153" refString="Borowiec, L. (1999) A world catalogue of the Cassidinae (Coleoptera: Chrysomelidae). Biologica Silesiae, Wroclaw, 476 pp." type="book" year="1999">Borowiec, 1999</bibRefCitation>
: 117
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;
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.
<taxonomicName id="83B13FC04A43FFABFF2BF908FF260B87" ID-CoL="7NTX" box="[151,212,1685,1707]" class="Insecta" family="Chrysomelidae" genus="Stolas" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="genus">
<emphasis id="76C598514A43FFABFF2BF908FF260B87" box="[151,212,1685,1707]" italics="true" pageId="2" pageNumber="153">Stolas</emphasis>
</taxonomicName>
subgenus
<taxonomicName id="83B13FC04A43FFABFE83F908FD610B87" ID-CoL="62Z7Y" authority="Hincks, 1952: 334" authorityName="Hincks" authorityPageNumber="334" authorityYear="1952" box="[319,659,1685,1707]" class="Insecta" family="Chrysomelidae" genus="Mesomphalia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="genus">
<emphasis id="76C598514A43FFABFE83F908FE3B0B87" box="[319,457,1685,1707]" italics="true" pageId="2" pageNumber="153">Mesomphalia</emphasis>
:
<treatmentCitation id="C51062524A43FFABFE6AF908FD610B87" author="Hincks" box="[470,659,1685,1707]" page="334" pageId="2" pageNumber="153" year="1952">
<bibRefCitation id="202039B24A43FFABFE6AF908FD920B87" author="Hincks" box="[470,608,1685,1707]" pageId="2" pageNumber="153" refString="Hincks, W. D. (1952) The genera of the Cassidinae (Coleoptera: Chrysomelidae). Transactions of the Royal Entomological Society of London, 103 (10), 327 - 358. http: // dx. doi. org / 10.1111 / j. 1365 - 2311.1952. tb 01061. x" type="journal article" year="1952">Hincks, 1952</bibRefCitation>
: 334
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</taxonomicName>
;
<treatmentCitation id="C51062524A43FFABFD21F908FC490B87" author="Seeno" box="[669,955,1685,1707]" page="174" pageId="2" pageNumber="153" year="1982">
<bibRefCitation id="202039B24A43FFABFD21F908FC7A0B87" author="Seeno" box="[669,904,1685,1707]" pageId="2" pageNumber="153" refString="Seeno, T. N. &amp; Wilcox, J. A. (1982) Leaf beetle genera (Coleoptera: Chrysomelidae). Entomography, 1, 1 - 221." type="journal article" year="1982">Seeno &amp; Wilcox, 1982</bibRefCitation>
: 174
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;
<treatmentCitation id="C51062524A43FFABFC79F908FB780B87" author="Chaboo" box="[965,1162,1685,1707]" page="237" pageId="2" pageNumber="153" year="2007">
<bibRefCitation id="202039B24A43FFABFC79F908FBA50B87" author="Chaboo" box="[965,1111,1685,1707]" pageId="2" pageNumber="153" refString="Chaboo, C. S. (2007) Biology and phylogeny of the Cassidinae Gyllenhal sensu lato (tortoise and leaf-mining beetles) (Coleoptera: Chrysomelidae). Bulletin of the American Museum of Natural History, 305, 1 - 250. http: // dx. doi. org / 10.1206 / 0003 - 0090 (2007) 305 [1: bapotc] 2.0. co; 2" type="journal article" year="2007">Chaboo, 2007</bibRefCitation>
: 237
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.
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<paragraph id="440E44434A43FFABFF2BF94AFC3E0BDC" blockId="2.[151,1436,1751,2029]" box="[151,972,1751,1776]" pageId="2" pageNumber="153">
<emphasis id="76C598514A43FFABFF2BF94AFEC00BDC" bold="true" box="[151,306,1751,1776]" pageId="2" pageNumber="153">
<typeStatus id="9B0AFAE14A43FFABFF2BF94AFF200BDC" box="[151,210,1751,1776]" pageId="2" pageNumber="153">Type</typeStatus>
species.
</emphasis>
<taxonomicName id="83B13FC04A43FFABFE85F94AFD5A0BDC" authority="Fabricius, 1781" authorityName="Fabricius" authorityYear="1781" box="[313,680,1751,1776]" class="Insecta" family="Chrysomelidae" genus="Cassida" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="species" species="gibbosa">
<emphasis id="76C598514A43FFABFE85F94AFE010BDC" box="[313,499,1751,1776]" italics="true" pageId="2" pageNumber="153">Cassida gibbosa</emphasis>
<bibRefCitation id="202039B24A43FFABFE46F94AFD5A0BDC" author="Fabricius" box="[506,680,1751,1776]" pageId="2" pageNumber="153" refString="Fabricius, J. C. (1781) Species Insectorum eorum differentias specificas, synonyma auctorum, loca natalia, metamorphosis adiectis observationibus, descriptionibus. Tomo I. Bohn, Hamburgi et Kilonii, 552 pp." type="book" year="1781">Fabricius, 1781</bibRefCitation>
</taxonomicName>
(by original designation).
</paragraph>
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<subSubSection id="0CAB17C84A43FFABFF7BF961FF280AC1" pageId="2" pageNumber="153" type="diagnosis">
<paragraph id="440E44434A43FFABFF7BF961FF280AC1" blockId="2.[151,1436,1751,2029]" pageId="2" pageNumber="153">
<emphasis id="76C598514A43FFABFF7BF961FEC80A39" bold="true" box="[199,314,1788,1813]" pageId="2" pageNumber="153">Diagnosis</emphasis>
.
<taxonomicName id="83B13FC04A43FFABFEF6F961FE100A39" box="[330,482,1788,1813]" class="Insecta" family="Chrysomelidae" genus="Mesomphalia" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="genus">
<emphasis id="76C598514A43FFABFEF6F961FE100A39" box="[330,482,1788,1813]" italics="true" pageId="2" pageNumber="153">Mesomphalia</emphasis>
</taxonomicName>
is distinguished from other
<taxonomicName id="83B13FC04A43FFABFC99F961FC3C0A39" box="[805,974,1788,1813]" class="Insecta" family="Chrysomelidae" kingdom="Animalia" order="Coleoptera" pageId="2" pageNumber="153" phylum="Arthropoda" rank="tribe" tribe="Mesomphaliini">Mesomphaliini</taxonomicName>
genera by the following combination of features: integument black and sculptured, 11 cylindrical antennomeres, with four basal subglabrous; short and curved mandibles; maxillary palpomere I long, twice as long as III; mentum short; labium membranous; prosternum Rounded with elongated and longitudinally grooved process; anterior third of elytra with shared acute conical gibbosity; prosternal collar with anterior margin curved, lateral extremities projected and divergent, followed medially by transverse sulcus; elytra, in lateral view, with abrupt anterior slope and gradual posterior slope.
</paragraph>
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<subSubSection id="0CAB17C84A42FFAAFF7BFF0AFF0408C6" pageId="3" pageNumber="154" type="description">
<paragraph id="440E44434A42FFAAFF7BFF0AFD1B0DD4" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="154">
<emphasis id="76C598514A42FFAAFF7BFF0AFEA30D9C" bold="true" box="[199,337,151,176]" pageId="3" pageNumber="154">Description</emphasis>
. Body almost circular (except
<taxonomicName id="83B13FC04A42FFAAFD17FF04FCCA0D9C" box="[683,824,151,176]" class="Insecta" family="Chrysomelidae" genus="Mesomphalia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="sublaevis">
<emphasis id="76C598514A42FFAAFD17FF04FCCA0D9C" box="[683,824,151,176]" italics="true" pageId="3" pageNumber="154">M. sublaevis</emphasis>
</taxonomicName>
, with length 1.3x width) in dorsal view; head partially concealed; pronotum and elytra slightly contiguous. In lateral view, body with gibbosity at the basal third of the elytra. Pronotal and elytral lateral margins explanate.
</paragraph>
<paragraph id="440E44434A42FFAAFF7BFE99FC040FA8" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="154">
Dorsal integument black and sculptured, abdominal ventrites smooth surfaced. Head (
<figureCitation id="DC8A58C64A42FFAAFB11FE99FAE80C30" box="[1197,1306,260,285]" captionStart="FIGURES 1 10" captionStartId="6.[151,264,1901,1924]" captionTargetBox="[224,1363,373,1880]" captionTargetId="figure@6.[224,1364,373,1881]" captionTargetPageId="6" captionText="FIGURES 1 10. Mesomphalia gibbosa (Fabricius, 1781) (male). 1 3, head: 1, dorsal view; 2, lateral view; 3, ventral view; 4 7, mandibles: 4, dorsal view; 5, ventral view; 6, superior lateral view; 7, inferior lateral view; 8, labrum, dorsal view; 9, labium, ventral view; 10, maxillae, dorsal view. Scale = 1 mm." httpUri="https://zenodo.org/record/249790/files/figure.png" pageId="3" pageNumber="154">Figs. 13</figureCitation>
) matt with erect, long and dense setae. Eyes (
<figureCitation id="DC8A58C64A42FFAAFD9AFEBAFD600C6C" box="[550,658,295,320]" captionStart="FIGURES 1 10" captionStartId="6.[151,264,1901,1924]" captionTargetBox="[224,1363,373,1880]" captionTargetId="figure@6.[224,1364,373,1881]" captionTargetPageId="6" captionText="FIGURES 1 10. Mesomphalia gibbosa (Fabricius, 1781) (male). 1 3, head: 1, dorsal view; 2, lateral view; 3, ventral view; 4 7, mandibles: 4, dorsal view; 5, ventral view; 6, superior lateral view; 7, inferior lateral view; 8, labrum, dorsal view; 9, labium, ventral view; 10, maxillae, dorsal view. Scale = 1 mm." httpUri="https://zenodo.org/record/249790/files/figure.png" pageId="3" pageNumber="154">Figs. 13</figureCitation>
) oval, 1.5x wider than long, finely faceted and laterally prominent. Vertex with erect, long and sparse setae, coarse and dense punctures. Frontal tubercles (
<figureCitation id="DC8A58C64A42FFAAFB17FED1FB000C48" box="[1195,1266,332,357]" captionStart="FIGURES 1 10" captionStartId="6.[151,264,1901,1924]" captionTargetBox="[224,1363,373,1880]" captionTargetId="figure@6.[224,1364,373,1881]" captionTargetPageId="6" captionText="FIGURES 1 10. Mesomphalia gibbosa (Fabricius, 1781) (male). 1 3, head: 1, dorsal view; 2, lateral view; 3, ventral view; 4 7, mandibles: 4, dorsal view; 5, ventral view; 6, superior lateral view; 7, inferior lateral view; 8, labrum, dorsal view; 9, labium, ventral view; 10, maxillae, dorsal view. Scale = 1 mm." httpUri="https://zenodo.org/record/249790/files/figure.png" pageId="3" pageNumber="154">Fig. 3</figureCitation>
) well-defined, slightly protuberant, and directed posteriorly. Coronal suture (
<figureCitation id="DC8A58C64A42FFAAFCC9FEF2FC4E0CA4" box="[885,956,367,392]" captionStart="FIGURES 1 10" captionStartId="6.[151,264,1901,1924]" captionTargetBox="[224,1363,373,1880]" captionTargetId="figure@6.[224,1364,373,1881]" captionTargetPageId="6" captionText="FIGURES 1 10. Mesomphalia gibbosa (Fabricius, 1781) (male). 1 3, head: 1, dorsal view; 2, lateral view; 3, ventral view; 4 7, mandibles: 4, dorsal view; 5, ventral view; 6, superior lateral view; 7, inferior lateral view; 8, labrum, dorsal view; 9, labium, ventral view; 10, maxillae, dorsal view. Scale = 1 mm." httpUri="https://zenodo.org/record/249790/files/figure.png" pageId="3" pageNumber="154">Fig. 1</figureCitation>
) deep, with smooth depression at apex. Frontoclypeus (
<figureCitation id="DC8A58C64A42FFAAFEFAFE09FE7B0C80" box="[326,393,404,429]" captionStart="FIGURES 1 10" captionStartId="6.[151,264,1901,1924]" captionTargetBox="[224,1363,373,1880]" captionTargetId="figure@6.[224,1364,373,1881]" captionTargetPageId="6" captionText="FIGURES 1 10. Mesomphalia gibbosa (Fabricius, 1781) (male). 1 3, head: 1, dorsal view; 2, lateral view; 3, ventral view; 4 7, mandibles: 4, dorsal view; 5, ventral view; 6, superior lateral view; 7, inferior lateral view; 8, labrum, dorsal view; 9, labium, ventral view; 10, maxillae, dorsal view. Scale = 1 mm." httpUri="https://zenodo.org/record/249790/files/figure.png" pageId="3" pageNumber="154">Fig. 3</figureCitation>
) protuberant anteriorly, with dense setae and coarse punctation; epistomal suture discernible. Mandibles short and curved; maxillary palpomere I long, twice as long as III; mentum short; labium membranous; prosternum rounded with elongated and longitudinally grooved process. Antennae (
<figureCitation id="DC8A58C64A42FFAAFBCCFE41FB230CD8" box="[1136,1233,476,501]" captionStart="FIGURES 11 15" captionStartId="7.[151,264,1913,1936]" captionTargetBox="[246,1340,193,1891]" captionTargetId="figure@7.[246,1341,193,1892]" captionTargetPageId="7" captionText="FIGURES 11 15. Mesomphalia gibbosa (Fabricius, 1781) (male). 11, antenna, dorsal view; 12 15, prothorax: 12, dorsal view; 13, ventral view; 14, lateral view; 15, frontal view (pr, proendosternite; ps, prosternal sulcus). Scale = 1 mm." httpUri="https://zenodo.org/record/249791/files/figure.png" pageId="3" pageNumber="154">Figs. 11</figureCitation>
,
<figureCitation id="DC8A58C64A42FFAAFB5FFE41FADB0CD8" box="[1251,1321,476,500]" captionStart="FIGURES 91 97" captionStartId="31.[151,264,1911,1934]" captionTargetBox="[220,1372,193,1893]" captionTargetId="figure@31.[220,1372,193,1894]" captionTargetPageId="31" captionText="FIGURES 91 97. Mesomphalia gibbosa (Fabricius, 1781). 91 93, antenna: 91, general aspect, arrows showing longitudinal grooves; 92, antennomere VI, with long and erect setae at the apex, and longitudinal suture next to the apex; 93, antennomere XI, with longitudinal suture at the median region; 94 97, pronotum: 94 95, median line smooth; 96 97, projection of posterior margin." httpUri="https://zenodo.org/record/249800/files/figure.png" pageId="3" pageNumber="154">9193</figureCitation>
) filiform, slightly flattened dorso-ventrally, wider at apex, with 11 cylindrical antennomeres, reaching anterior margin of elytra when reflexed. Scape, pedicel, and antennomeres IIIIV with long and sparse setae; VXI with short and dense setae on the sides and sparse setae at apex of antennomeres. Scape globose, 2.53.0x longer than pedicel; XI wider at post-median region, narrowing to truncate apex, 1.5
<date id="300F62834A42FFAAFCF0FDF1FCA80FA9" box="[844,858,620,645]" pageId="3" pageNumber="154">2</date>
x length of X.
</paragraph>
<paragraph id="440E44434A42FFAAFF7BFD12FD840E71" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="154">
Prothorax (
<figureCitation id="DC8A58C64A42FFAAFEF9FD12FE3F0F84" box="[325,461,655,680]" captionStart="FIGURES 11 15" captionStartId="7.[151,264,1913,1936]" captionTargetBox="[246,1340,193,1891]" captionTargetId="figure@7.[246,1341,193,1892]" captionTargetPageId="7" captionText="FIGURES 11 15. Mesomphalia gibbosa (Fabricius, 1781) (male). 11, antenna, dorsal view; 12 15, prothorax: 12, dorsal view; 13, ventral view; 14, lateral view; 15, frontal view (pr, proendosternite; ps, prosternal sulcus). Scale = 1 mm." httpUri="https://zenodo.org/record/249791/files/figure.png" pageId="3" pageNumber="154">Figs. 1215</figureCitation>
,
<figureCitation id="DC8A58C64A42FFAAFE66FD12FDD30F84" box="[474,545,655,680]" captionStart="FIGURES 91 97" captionStartId="31.[151,264,1911,1934]" captionTargetBox="[220,1372,193,1893]" captionTargetId="figure@31.[220,1372,193,1894]" captionTargetPageId="31" captionText="FIGURES 91 97. Mesomphalia gibbosa (Fabricius, 1781). 91 93, antenna: 91, general aspect, arrows showing longitudinal grooves; 92, antennomere VI, with long and erect setae at the apex, and longitudinal suture next to the apex; 93, antennomere XI, with longitudinal suture at the median region; 94 97, pronotum: 94 95, median line smooth; 96 97, projection of posterior margin." httpUri="https://zenodo.org/record/249800/files/figure.png" pageId="3" pageNumber="154">9496</figureCitation>
) matt and transverse. Pronotum (
<figureCitation id="DC8A58C64A42FFAAFC26FD12FC0A0F84" box="[922,1016,655,680]" captionStart="FIGURES 11 15" captionStartId="7.[151,264,1913,1936]" captionTargetBox="[246,1340,193,1891]" captionTargetId="figure@7.[246,1341,193,1892]" captionTargetPageId="7" captionText="FIGURES 11 15. Mesomphalia gibbosa (Fabricius, 1781) (male). 11, antenna, dorsal view; 12 15, prothorax: 12, dorsal view; 13, ventral view; 14, lateral view; 15, frontal view (pr, proendosternite; ps, prosternal sulcus). Scale = 1 mm." httpUri="https://zenodo.org/record/249791/files/figure.png" pageId="3" pageNumber="154">Figs. 12</figureCitation>
,
<figureCitation id="DC8A58C64A42FFAAFBB9FD12FBBE0F84" box="[1029,1100,655,680]" captionStart="FIGURES 91 97" captionStartId="31.[151,264,1911,1934]" captionTargetBox="[220,1372,193,1893]" captionTargetId="figure@31.[220,1372,193,1894]" captionTargetPageId="31" captionText="FIGURES 91 97. Mesomphalia gibbosa (Fabricius, 1781). 91 93, antenna: 91, general aspect, arrows showing longitudinal grooves; 92, antennomere VI, with long and erect setae at the apex, and longitudinal suture next to the apex; 93, antennomere XI, with longitudinal suture at the median region; 94 97, pronotum: 94 95, median line smooth; 96 97, projection of posterior margin." httpUri="https://zenodo.org/record/249800/files/figure.png" pageId="3" pageNumber="154">9496</figureCitation>
) about twice as long as wide; anterior margin slightly concave, exposing head; longitudinal median line (
<figureCitation id="DC8A58C64A42FFAAFC54FD29FB820FE1" box="[1000,1136,692,717]" captionStart="FIGURES 91 97" captionStartId="31.[151,264,1911,1934]" captionTargetBox="[220,1372,193,1893]" captionTargetId="figure@31.[220,1372,193,1894]" captionTargetPageId="31" captionText="FIGURES 91 97. Mesomphalia gibbosa (Fabricius, 1781). 91 93, antenna: 91, general aspect, arrows showing longitudinal grooves; 92, antennomere VI, with long and erect setae at the apex, and longitudinal suture next to the apex; 93, antennomere XI, with longitudinal suture at the median region; 94 97, pronotum: 94 95, median line smooth; 96 97, projection of posterior margin." httpUri="https://zenodo.org/record/249800/files/figure.png" pageId="3" pageNumber="154">Figs. 9495</figureCitation>
) smooth and glabrous, not reaching the margins; disc elevated; posterior angle (
<figureCitation id="DC8A58C64A42FFAAFCB9FD4AFC940FDC" box="[773,870,727,752]" captionStart="FIGURES 91 97" captionStartId="31.[151,264,1911,1934]" captionTargetBox="[220,1372,193,1893]" captionTargetId="figure@31.[220,1372,193,1894]" captionTargetPageId="31" captionText="FIGURES 91 97. Mesomphalia gibbosa (Fabricius, 1781). 91 93, antenna: 91, general aspect, arrows showing longitudinal grooves; 92, antennomere VI, with long and erect setae at the apex, and longitudinal suture next to the apex; 93, antennomere XI, with longitudinal suture at the median region; 94 97, pronotum: 94 95, median line smooth; 96 97, projection of posterior margin." httpUri="https://zenodo.org/record/249800/files/figure.png" pageId="3" pageNumber="154">Figs. 96</figureCitation>
) slightly projecting posteriorly. Prosternal collar (
<figureCitation id="DC8A58C64A42FFAAFF23FD61FEDB0E38" box="[159,297,764,789]" captionStart="FIGURES 11 15" captionStartId="7.[151,264,1913,1936]" captionTargetBox="[246,1340,193,1891]" captionTargetId="figure@7.[246,1341,193,1892]" captionTargetPageId="7" captionText="FIGURES 11 15. Mesomphalia gibbosa (Fabricius, 1781) (male). 11, antenna, dorsal view; 12 15, prothorax: 12, dorsal view; 13, ventral view; 14, lateral view; 15, frontal view (pr, proendosternite; ps, prosternal sulcus). Scale = 1 mm." httpUri="https://zenodo.org/record/249791/files/figure.png" pageId="3" pageNumber="154">Figs. 1314</figureCitation>
,
<figureCitation id="DC8A58C64A42FFAAFE85FD61FEA40E38" box="[313,342,764,788]" captionStart="FIGURES 98 100" captionStartId="32.[151,264,1439,1462]" captionTargetBox="[233,1391,176,1388]" captionTargetId="figure@32.[202,1397,143,1416]" captionTargetPageId="32" captionText="FIGURES 98 100. Mesomphalia variolaris Boheman, 1850. 98, ventral view; 99, dorsal view; 100, lateral view (pc, prosternal collar; ed, elytral disc; gb, gibbosity; lm, lateral margin; ts, transversal sulcus)." httpUri="https://zenodo.org/record/249801/files/figure.png" pageId="3" pageNumber="154">98</figureCitation>
) with anterior margin curved, lateral extremities projecting and divergent; median region with transverse sulcus. Prosternal process (
<figureCitation id="DC8A58C64A42FFAAFD82FC82FD7C0E14" box="[574,654,799,824]" captionStart="FIGURES 11 15" captionStartId="7.[151,264,1913,1936]" captionTargetBox="[246,1340,193,1891]" captionTargetId="figure@7.[246,1341,193,1892]" captionTargetPageId="7" captionText="FIGURES 11 15. Mesomphalia gibbosa (Fabricius, 1781) (male). 11, antenna, dorsal view; 12 15, prothorax: 12, dorsal view; 13, ventral view; 14, lateral view; 15, frontal view (pr, proendosternite; ps, prosternal sulcus). Scale = 1 mm." httpUri="https://zenodo.org/record/249791/files/figure.png" pageId="3" pageNumber="154">Fig. 13</figureCitation>
) narrowed medially with longitudinal sulcus; apical region expanded. Scutellum (Fig. 23) triangular and smooth.
</paragraph>
<paragraph id="440E44434A42FFAAFF7BFCFAFE650950" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="154">
Mesoventrite (
<figureCitation id="DC8A58C64A42FFAAFED5FCFAFE1C0EAC" box="[361,494,871,896]" captionStart="FIGURES 16 20" captionStartId="8.[151,264,1952,1975]" captionTargetBox="[234,1352,193,1834]" captionTargetId="figure@8.[234,1352,193,1932]" captionTargetPageId="8" captionText="FIGURES 16 20. Mesomphalia gibbosa (Fabricius, 1781) (male). 16, meso and metaventrites; 17, meso and metendosternite; 18, meso and metathorax; 19 20, metendosternite: 19, dorsal view; 20, ventral view (a, arm, la, lamina; me, mesendosternite; te, tendon). Scale = 1 mm." httpUri="https://zenodo.org/record/249792/files/figure.png" pageId="3" pageNumber="154">Figs. 16, 18</figureCitation>
) transverse. Mesoventral process depressed, with protuberant lateral margins. Elytra macroreticulate (Figs. 2122) with basal margin faintly crenulate; lateral margins slightly or markedly oblique on males and rounded on females; apical 2/3 of margin flattened and explanate; apical posterior angle slightly projected; anterior third with shared acute conical gibbosity; in lateral view, with abrupt anterior slope and gradual posterior slope. Humeral angle rounded, smooth and glabrous (except
<taxonomicName id="83B13FC04A42FFAAFC05FC64FBB6093C" box="[953,1092,1015,1040]" class="Insecta" family="Chrysomelidae" genus="Mesomphalia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="species" species="tumidula">
<emphasis id="76C598514A42FFAAFC05FC64FBB6093C" box="[953,1092,1015,1040]" italics="true" pageId="3" pageNumber="154">M. tumidula</emphasis>
</taxonomicName>
), slightly projected anteriorly. Disc with conical gibbosity at the basal third, height about twice the greatest elytral width; in lateral view, with abrupt anterior slope and gradual slope posteriorly. Metaventrite (
<figureCitation id="DC8A58C64A42FFAAFC33FBA2FBEE0974" box="[911,1052,1087,1112]" captionStart="FIGURES 16 20" captionStartId="8.[151,264,1952,1975]" captionTargetBox="[234,1352,193,1834]" captionTargetId="figure@8.[234,1352,193,1932]" captionTargetPageId="8" captionText="FIGURES 16 20. Mesomphalia gibbosa (Fabricius, 1781) (male). 16, meso and metaventrites; 17, meso and metendosternite; 18, meso and metathorax; 19 20, metendosternite: 19, dorsal view; 20, ventral view (a, arm, la, lamina; me, mesendosternite; te, tendon). Scale = 1 mm." httpUri="https://zenodo.org/record/249792/files/figure.png" pageId="3" pageNumber="154">Figs. 16, 18</figureCitation>
) with anterior margin excavated, adjacent to metacoxae.
</paragraph>
<paragraph id="440E44434A42FFAAFF7BFB14FCC50876" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="154">
Legs (
<figureCitation id="DC8A58C64A42FFAAFEB0FB1AFE61098C" box="[268,403,1159,1184]" captionStart="FIGURES 25 40" captionStartId="10.[151,264,1898,1921]" captionTargetBox="[241,1345,193,1841]" captionTargetId="figure@10.[241,1345,193,1932]" captionTargetPageId="10" captionText="FIGURES 25 40. Mesomphalia gibbosa (Fabricius, 1781) (male). 25, procoxa; 26, profemur; 27, protibia; 28 29, protarsomere: 28, dorsal view; 29, lateral view; 30, mesocoxa; 31, mesofemur; 32, mesotibia; 33 34, mesotarsomere: 33, dorsal view; 34, lateral view; 35, metacoxa; 36, metafemur; 37, metatibia; 38 39, metatarsomere: 38, dorsal view; 39, lateral view; 40, apendiculate claws. Scale = 1 mm." httpUri="https://zenodo.org/record/249793/files/figure.png" pageId="3" pageNumber="154">Figs. 2539</figureCitation>
) subequal in length and clothed in decumbent, long and sparse setae, except tibia, with setae becoming denser towards apex. Pro- and mesocoxae globose; metacoxae transverse; trochanters triangular; femora fusiform with basal third smooth, dilated, and apical 2/3 microreticulate and depressed; tibia slightly expanded to apex, with apicodorsal surface notched to receive tarsus. Tarsomeres (
<figureCitation id="DC8A58C64A42FFAAFC0AFB64FB18083E" box="[950,1258,1273,1298]" captionStart="FIGURES 25 40" captionStartId="10.[151,264,1898,1921]" captionTargetBox="[241,1345,193,1841]" captionTargetId="figure@10.[241,1345,193,1932]" captionTargetPageId="10" captionText="FIGURES 25 40. Mesomphalia gibbosa (Fabricius, 1781) (male). 25, procoxa; 26, profemur; 27, protibia; 28 29, protarsomere: 28, dorsal view; 29, lateral view; 30, mesocoxa; 31, mesofemur; 32, mesotibia; 33 34, mesotarsomere: 33, dorsal view; 34, lateral view; 35, metacoxa; 36, metafemur; 37, metatibia; 38 39, metatarsomere: 38, dorsal view; 39, lateral view; 40, apendiculate claws. Scale = 1 mm." httpUri="https://zenodo.org/record/249793/files/figure.png" pageId="3" pageNumber="154">Figs. 2829, 3334, 3839</figureCitation>
) IIIII bilobed; tarsomere I half the length of II; II as long III, but more robust; IV absent; V slightly longer than III. Tarsal claws (
<figureCitation id="DC8A58C64A42FFAAFF23FADCFF030876" box="[159,241,1345,1370]" captionStart="FIGURES 25 40" captionStartId="10.[151,264,1898,1921]" captionTargetBox="[241,1345,193,1841]" captionTargetId="figure@10.[241,1345,193,1932]" captionTargetPageId="10" captionText="FIGURES 25 40. Mesomphalia gibbosa (Fabricius, 1781) (male). 25, procoxa; 26, profemur; 27, protibia; 28 29, protarsomere: 28, dorsal view; 29, lateral view; 30, mesocoxa; 31, mesofemur; 32, mesotibia; 33 34, mesotarsomere: 33, dorsal view; 34, lateral view; 35, metacoxa; 36, metafemur; 37, metatibia; 38 39, metatarsomere: 38, dorsal view; 39, lateral view; 40, apendiculate claws. Scale = 1 mm." httpUri="https://zenodo.org/record/249793/files/figure.png" pageId="3" pageNumber="154">Fig. 40</figureCitation>
) reddish-brown, curved, tapered and appendiculate.
</paragraph>
<paragraph id="440E44434A42FFAAFF7BFAF8FF0408C6" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="154">
Abdomen completely covered by elytra. Ventrites shiny, except ventrite I with microreticulation on basal third to two-thirds; (
<figureCitation id="DC8A58C64A42FFAAFEFFFA14FE3F088E" box="[323,461,1417,1442]" captionStart="FIGURES 41 47" captionStartId="11.[151,264,1826,1849]" captionTargetBox="[209,1377,193,1663]" captionTargetId="figure@11.[209,1377,193,1805]" captionTargetPageId="11" captionText="FIGURES 41 47. Mesomphalia gibbosa (Fabricius, 1781) (male). 41, abdomen, ventral view; 42, ventrite V, general aspect; 43, tergite VIII, dorsal view; 44 45, median lobe; 44, lateral view; 45, dorsal view of the apex; 46 47, tegmen: 46, dorsal view; 47, lateral view. (ed, ejaculatory duct; ma, manubrium; o, ostium; te, tegmen). Scale = 1 mm" httpUri="https://zenodo.org/record/249794/files/figure.png" pageId="3" pageNumber="154">Figs. 4143</figureCitation>
,
<figureCitation id="DC8A58C64A42FFAAFE67FA17FDD1088E" box="[475,547,1418,1442]" captionStart="FIGURES 48 54" captionStartId="12.[151,264,1735,1758]" captionTargetBox="[239,1347,179,1714]" captionTargetId="figure@12.[239,1347,179,1714]" captionTargetPageId="12" captionText="FIGURES 48 54. Mesomphalia gibbosa (Fabricius, 1781) (female). 48, abdomen, ventral view; 49, sternite VII, general aspect; 50, sternite VIII; 51, sternite IX; 52, tergite X; 53, sternite IX and tergite X, lateral view; 54, spermatheca (am, ampulla; ed, spermatheca duct; Gl, ductus glandula auxiliaris; va, vasculum). Scale = 1 mm." httpUri="https://zenodo.org/record/249795/files/figure.png" pageId="3" pageNumber="154">4849</figureCitation>
) IIV subequal in length; V with erect, long setae, becoming denser towards median region and covering apical third, with flat posterior margin on males and preapical median depression on females.
</paragraph>
</subSubSection>
<subSubSection id="0CAB17C84A42FFADFF7BFA68FD760EE4" lastPageId="4" lastPageNumber="155" pageId="3" pageNumber="154" type="discussion">
<paragraph id="440E44434A42FFAAFF7BFA68FEC70BED" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="154">
<emphasis id="76C598514A42FFAAFF7BFA68FEC90B22" bold="true" box="[199,315,1525,1550]" pageId="3" pageNumber="154">Remarks.</emphasis>
<bibRefCitation id="202039B24A42FFAAFEFEFA6BFE220B22" author="Hope" box="[322,464,1526,1550]" pageId="3" pageNumber="154" refString="Hope, F. W. (1839) Observations on the tortoise or shield beetles, commonly denominated Cassida by Linnaeus, with the characters of six new genera. Annals of Natural History, 3 (15), 92 - 100. http: // dx. doi. org / 10.1080 / 03745483909443203" type="journal article" year="1839">Hope (1839)</bibRefCitation>
described
<taxonomicName id="83B13FC04A42FFAAFDF5FA68FD100B22" box="[585,738,1525,1550]" class="Insecta" family="Chrysomelidae" genus="Mesomphalia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="genus">
<emphasis id="76C598514A42FFAAFDF5FA68FD100B22" box="[585,738,1525,1550]" italics="true" pageId="3" pageNumber="154">Mesomphalia</emphasis>
</taxonomicName>
in detail.
<bibRefCitation id="202039B24A42FFAAFCECFA68FC1C0B22" author="Spaeth" box="[848,1006,1525,1550]" pageId="3" pageNumber="154" refString="Spaeth, F. (1901) Beschreibung neuer Cassididen nebst synonymischen Bemerkungen. IV. Verhandlungen der Zoologisch- Botanischen Gesellschaft, 51, 333 - 350." type="journal article" year="1901">Spaeth (1901)</bibRefCitation>
revised
<taxonomicName id="83B13FC04A42FFAAFBF1FA68FB140B22" box="[1101,1254,1525,1550]" class="Insecta" family="Chrysomelidae" genus="Mesomphalia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="genus">
<emphasis id="76C598514A42FFAAFBF1FA68FB140B22" box="[1101,1254,1525,1550]" italics="true" pageId="3" pageNumber="154">Mesomphalia</emphasis>
</taxonomicName>
and redefined it with the following attributes: prosternum with anterior margin projected anteriorly, with divergent anterolateral angles, followed by a transverse sulcus; elytra with gibbosity at basal third and an acuminate apex. In addition, all species of
<taxonomicName id="83B13FC04A42FFAAFE9CF9FCFE4A0B56" box="[288,440,1633,1658]" class="Insecta" family="Chrysomelidae" genus="Mesomphalia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="genus">
<emphasis id="76C598514A42FFAAFE9CF9FCFE4A0B56" box="[288,440,1633,1658]" italics="true" pageId="3" pageNumber="154">Mesomphalia</emphasis>
</taxonomicName>
have: black integument without metallic iridescence; deep coronal suture; basal antennomeres subglabrous, with long and sparse setae; pronotum and elytra with setae, always forming macules on the pronotum.
</paragraph>
<paragraph id="440E44434A42FFAAFF7BF950FE480AB6" blockId="3.[151,1437,151,2018]" pageId="3" pageNumber="154">
Within the tribe
<taxonomicName id="83B13FC04A42FFAAFE32F950FDC90BCA" box="[398,571,1741,1766]" class="Insecta" family="Chrysomelidae" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="tribe" tribe="Mesomphaliini">Mesomphaliini</taxonomicName>
,
<taxonomicName id="83B13FC04A42FFAAFDF7F950FD110BCA" box="[587,739,1741,1766]" class="Insecta" family="Chrysomelidae" genus="Mesomphalia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="genus">
<emphasis id="76C598514A42FFAAFDF7F950FD110BCA" box="[587,739,1741,1766]" italics="true" pageId="3" pageNumber="154">Mesomphalia</emphasis>
</taxonomicName>
resembles
<taxonomicName id="83B13FC04A42FFAAFCD0F953FB540BCA" authority="Chevrolat, 1836" authorityName="Chevrolat" authorityYear="1836" box="[876,1190,1741,1766]" class="Insecta" family="Chrysomelidae" genus="Cyrtonota" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="genus">
<emphasis id="76C598514A42FFAAFCD0F953FC2F0BC9" box="[876,989,1742,1765]" italics="true" pageId="3" pageNumber="154">Cyrtonota</emphasis>
Chevrolat, 1836
</taxonomicName>
,
<taxonomicName id="83B13FC04A42FFAAFB0AF950FF3D0A26" authority="Spaeth, 1902" authorityName="Spaeth" authorityYear="1902" class="Insecta" family="Chrysomelidae" genus="Trilaccodea" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="genus">
<emphasis id="76C598514A42FFAAFB0AF950FACE0BCA" box="[1206,1340,1741,1766]" italics="true" pageId="3" pageNumber="154">Trilaccodea</emphasis>
Spaeth, 1902
</taxonomicName>
and
<taxonomicName id="83B13FC04A42FFAAFEB1F96CFEA30A26" box="[269,337,1777,1802]" class="Insecta" family="Chrysomelidae" genus="Stolas" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="genus">
<emphasis id="76C598514A42FFAAFEB1F96CFEA30A26" box="[269,337,1777,1802]" italics="true" pageId="3" pageNumber="154">Stolas</emphasis>
</taxonomicName>
.
<taxonomicName id="83B13FC04A42FFAAFEDFF96FFE270A25" box="[355,469,1778,1801]" class="Insecta" family="Chrysomelidae" genus="Cyrtonota" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="genus">
<emphasis id="76C598514A42FFAAFEDFF96FFE270A25" box="[355,469,1778,1801]" italics="true" pageId="3" pageNumber="154">Cyrtonota</emphasis>
</taxonomicName>
and
<taxonomicName id="83B13FC04A42FFAAFDAFF96CFD590A26" box="[531,683,1777,1802]" class="Insecta" family="Chrysomelidae" genus="Mesomphalia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="genus">
<emphasis id="76C598514A42FFAAFDAFF96CFD590A26" box="[531,683,1777,1802]" italics="true" pageId="3" pageNumber="154">Mesomphalia</emphasis>
</taxonomicName>
share the following: subglabrous basal antennomeres, pronotum transverse with subparallel lateral margins, elytra with basal margin crenulate, disc convex, tarsal claws apendiculate and divaricate.
<taxonomicName id="83B13FC04A42FFAAFE6AF8A7FDBA0A7D" box="[470,584,1850,1873]" class="Insecta" family="Chrysomelidae" genus="Cyrtonota" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="genus">
<emphasis id="76C598514A42FFAAFE6AF8A7FDBA0A7D" box="[470,584,1850,1873]" italics="true" pageId="3" pageNumber="154">Cyrtonota</emphasis>
</taxonomicName>
is distinguished by: integument may be irridescent metallic; antennomere V subglabrous; pronotum varying from nearly impunctate to coarsely punctate; prosternal collar not projecting, and without transverse sulcus.
</paragraph>
<paragraph id="440E44434A42FFADFF7BF838FDF30C49" blockId="3.[151,1437,151,2018]" lastBlockId="4.[151,1437,151,968]" lastPageId="4" lastPageNumber="155" pageId="3" pageNumber="154">
<taxonomicName id="83B13FC04A42FFAAFF7BF838FE920A92" box="[199,352,1957,1982]" class="Insecta" family="Chrysomelidae" genus="Mesomphalia" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="genus">
<emphasis id="76C598514A42FFAAFF7BF838FE920A92" box="[199,352,1957,1982]" italics="true" pageId="3" pageNumber="154">Mesomphalia</emphasis>
</taxonomicName>
and
<taxonomicName id="83B13FC04A42FFAAFE26F838FDD20A92" box="[410,544,1957,1982]" class="Insecta" family="Chrysomelidae" genus="Trilaccodea" kingdom="Animalia" order="Coleoptera" pageId="3" pageNumber="154" phylum="Arthropoda" rank="genus">
<emphasis id="76C598514A42FFAAFE26F838FDD20A92" box="[410,544,1957,1982]" italics="true" pageId="3" pageNumber="154">Trilaccodea</emphasis>
</taxonomicName>
share: pronotum transversely pentagonal, with impunctate, dull, base on each side with distinct impression; surface of pronotum with setae on the surface; prosternal collar elongate, often angulate on sides. However, they differ greatly in other aspects.
<taxonomicName id="83B13FC04A45FFADFCDAFF0AFC1E0D9C" box="[870,1004,151,176]" class="Insecta" family="Chrysomelidae" genus="Trilaccodea" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="76C598514A45FFADFCDAFF0AFC1E0D9C" box="[870,1004,151,176]" italics="true" pageId="4" pageNumber="155">Trilaccodea</emphasis>
</taxonomicName>
has: integument with color patterns or metallic; elytra extremely broad, base around twice width base of pronotum; disc with large, blunt postscutellar tubercle; profile behind the tubercle sinuous or slightly concave with apical angle of elytra pointed downwards (Borowiec 2007).
<taxonomicName id="83B13FC04A45FFADFEDFFE99FE0E0C31" box="[355,508,260,285]" class="Insecta" family="Chrysomelidae" genus="Mesomphalia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="76C598514A45FFADFEDFFE99FE0E0C31" box="[355,508,260,285]" italics="true" pageId="4" pageNumber="155">Mesomphalia</emphasis>
</taxonomicName>
always has: integument strictly black, not metallic; elytra not wider than pronotum base; disc with sharply conical postscutellar gibbosity; profile behind the gibbosity slightly straight, with apical angle of elytra pointed upwards.
</paragraph>
<paragraph id="440E44434A45FFADFF7BFEF2FF140F4C" blockId="4.[151,1437,151,968]" pageId="4" pageNumber="155">
The similarity between
<taxonomicName id="83B13FC04A45FFADFE5DFEF2FDD70CA4" box="[481,549,367,392]" class="Insecta" family="Chrysomelidae" genus="Stolas" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="76C598514A45FFADFE5DFEF2FDD70CA4" box="[481,549,367,392]" italics="true" pageId="4" pageNumber="155">Stolas</emphasis>
</taxonomicName>
and
<taxonomicName id="83B13FC04A45FFADFDDBFEF2FCF20CA4" box="[615,768,367,392]" class="Insecta" family="Chrysomelidae" genus="Mesomphalia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="76C598514A45FFADFDDBFEF2FCF20CA4" box="[615,768,367,392]" italics="true" pageId="4" pageNumber="155">Mesomphalia</emphasis>
</taxonomicName>
is strong and led previous authors to synonymise them (
<bibRefCitation id="202039B24A45FFADFF22FE09FEC00C81" author="Hincks" box="[158,306,404,429]" pageId="4" pageNumber="155" refString="Hincks, W. D. (1952) The genera of the Cassidinae (Coleoptera: Chrysomelidae). Transactions of the Royal Entomological Society of London, 103 (10), 327 - 358. http: // dx. doi. org / 10.1111 / j. 1365 - 2311.1952. tb 01061. x" type="journal article" year="1952">Hincks 1952</bibRefCitation>
;
<bibRefCitation id="202039B24A45FFADFE81FE09FDCB0C80" author="Seeno" box="[317,569,404,429]" pageId="4" pageNumber="155" refString="Seeno, T. N. &amp; Wilcox, J. A. (1982) Leaf beetle genera (Coleoptera: Chrysomelidae). Entomography, 1, 1 - 221." type="journal article" year="1982">Seeno &amp; Wilcox 1982</bibRefCitation>
;
<bibRefCitation id="202039B24A45FFADFDF8FE09FD2C0C80" author="Chaboo" box="[580,734,404,429]" pageId="4" pageNumber="155" refString="Chaboo, C. S. (2007) Biology and phylogeny of the Cassidinae Gyllenhal sensu lato (tortoise and leaf-mining beetles) (Coleoptera: Chrysomelidae). Bulletin of the American Museum of Natural History, 305, 1 - 250. http: // dx. doi. org / 10.1206 / 0003 - 0090 (2007) 305 [1: bapotc] 2.0. co; 2" type="journal article" year="2007">Chaboo 2007</bibRefCitation>
), but without clear justification.
<taxonomicName id="83B13FC04A45FFADFBF4FE09FB7E0C81" box="[1096,1164,404,429]" class="Insecta" family="Chrysomelidae" genus="Stolas" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="76C598514A45FFADFBF4FE09FB7E0C81" box="[1096,1164,404,429]" italics="true" pageId="4" pageNumber="155">Stolas</emphasis>
</taxonomicName>
and
<taxonomicName id="83B13FC04A45FFADFB7EFE09FAA90C81" box="[1218,1371,404,429]" class="Insecta" family="Chrysomelidae" genus="Mesomphalia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="76C598514A45FFADFB7EFE09FAA90C81" box="[1218,1371,404,429]" italics="true" pageId="4" pageNumber="155">Mesomphalia</emphasis>
</taxonomicName>
share the following features: basal antennomeres subglabrous; pronotum in some species with subtriangular macules of yellowish-brown setae; elytra with basal margin faintly crenulate and disc with gibbosity at basal third. However, in
<taxonomicName id="83B13FC04A45FFADFF08FE62FF040F34" box="[180,246,511,536]" class="Insecta" family="Chrysomelidae" genus="Stolas" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="76C598514A45FFADFF08FE62FF040F34" box="[180,246,511,536]" italics="true" pageId="4" pageNumber="155">Stolas</emphasis>
</taxonomicName>
the integument may be metallic, the parascutellar disc may have macules of yellowish-brown setae (e.g.
<taxonomicName id="83B13FC04A45FFADFF2BFDB9FDEE0F10" authority="Spaeth, 1922" authorityName="Spaeth" authorityYear="1922" box="[151,540,548,573]" class="Insecta" family="Chrysomelidae" genus="Stolas" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="species" species="brevicuspis">
<emphasis id="76C598514A45FFADFF2BFDB9FE930F11" box="[151,353,548,573]" italics="true" pageId="4" pageNumber="155">Stolas brevicuspis</emphasis>
(Spaeth, 1922))
</taxonomicName>
, the prosternal collar is never projecting and the prosternal transverse sulcus is absent.
</paragraph>
<paragraph id="440E44434A45FFADFF7BFDF1FDD10E39" blockId="4.[151,1437,151,968]" pageId="4" pageNumber="155">
Outside
<taxonomicName id="83B13FC04A45FFADFE92FDF1FE280FA9" box="[302,474,620,645]" class="Insecta" family="Chrysomelidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="tribe" tribe="Mesomphaliini">Mesomphaliini</taxonomicName>
, the most similar genus is
<taxonomicName id="83B13FC04A45FFADFC80FDF1FB810FA9" authority="Weise, 1896" authorityName="Weise" authorityYear="1896" box="[828,1139,620,645]" class="Insecta" family="Chrysomelidae" genus="Goniochenia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="76C598514A45FFADFC80FDF1FC3E0FA9" box="[828,972,620,645]" italics="true" pageId="4" pageNumber="155">Goniochenia</emphasis>
Weise, 1896
</taxonomicName>
. It belongs to the tribe Goniocheniini Spaeth, 1942, which is characterized by the toothed claw and claw segment, while
<taxonomicName id="83B13FC04A45FFADFB4FFD12FA6F0F84" box="[1267,1437,655,680]" class="Insecta" family="Chrysomelidae" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="tribe" tribe="Mesomphaliini">Mesomphaliini</taxonomicName>
has untoothed claws (
<bibRefCitation id="202039B24A45FFADFE20FD29FDC40FE1" author="Hincks" box="[412,566,692,717]" pageId="4" pageNumber="155" refString="Hincks, W. D. (1952) The genera of the Cassidinae (Coleoptera: Chrysomelidae). Transactions of the Royal Entomological Society of London, 103 (10), 327 - 358. http: // dx. doi. org / 10.1111 / j. 1365 - 2311.1952. tb 01061. x" type="journal article" year="1952">Hincks 1952</bibRefCitation>
). Apart from variation in this tribal character,,
<taxonomicName id="83B13FC04A45FFADFBC8FD29FAF70FE1" box="[1140,1285,692,717]" class="Insecta" family="Chrysomelidae" genus="Goniochenia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="76C598514A45FFADFBC8FD29FAF70FE1" box="[1140,1285,692,717]" italics="true" pageId="4" pageNumber="155">Goniochenia</emphasis>
</taxonomicName>
(s. str.) and
<taxonomicName id="83B13FC04A45FFADFF2BFD4AFEC20FDC" box="[151,304,727,752]" class="Insecta" family="Chrysomelidae" genus="Mesomphalia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="76C598514A45FFADFF2BFD4AFEC20FDC" box="[151,304,727,752]" italics="true" pageId="4" pageNumber="155">Mesomphalia</emphasis>
</taxonomicName>
share integument without metallic iridescence, basal antennomeres subglabrous, and pronotum transverse, width at least
<date id="300F62834A45FFADFE0FFD61FE3C0E38" box="[435,462,764,788]" pageId="4" pageNumber="155">2x</date>
length.
</paragraph>
<paragraph id="440E44434A45FFADFF7BFC82FD760EE4" blockId="4.[151,1437,151,968]" pageId="4" pageNumber="155">
Little biological information is available for
<taxonomicName id="83B13FC04A45FFADFD6AFC82FC9C0E14" box="[726,878,799,824]" class="Insecta" family="Chrysomelidae" genus="Mesomphalia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="76C598514A45FFADFD6AFC82FC9C0E14" box="[726,878,799,824]" italics="true" pageId="4" pageNumber="155">Mesomphalia</emphasis>
</taxonomicName>
.
<bibRefCitation id="202039B24A45FFADFC3EFC82FBBB0E14" author="Haitlinger" box="[898,1097,799,824]" pageId="4" pageNumber="155" refString="Haitlinger, R. (1991) Canestriniid mites (Acari, Astigmata, Canestriniidae) occurring on Chrysomelidae (Insecta, Coleoptera) in Neotropical Region. I. Genera Grandiellina Fain, Arrunsithiana n. gen. Wiadomosci parazytologiczne, 37, 381 - 399." type="journal article" year="1991">Haitlinger (1991)</bibRefCitation>
reported an association of a canestriniid mite
<taxonomicName id="83B13FC04A45FFADFEE5FCD9FDB10E71" box="[345,579,836,861]" class="Arachnida" family="Canestriniidae" genus="Arrunsithiana" higherTaxonomySource="GBIF" kingdom="Animalia" order="Astigmata" pageId="4" pageNumber="155" phylum="Arthropoda" rank="species" species="battisi">
<emphasis id="76C598514A45FFADFEE5FCD9FDB10E71" box="[345,579,836,861]" italics="true" pageId="4" pageNumber="155">Arrunsithiana battisi</emphasis>
</taxonomicName>
with
<taxonomicName id="83B13FC04A45FFADFD3EFCD8FBFD0E70" authority="Fabricius, 1781" authorityName="Fabricius" authorityYear="1781" box="[642,1039,836,861]" class="Insecta" family="Chrysomelidae" genus="Cassida" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="species" species="gibbosa">
<emphasis id="76C598514A45FFADFD3EFCD8FCF20E71" box="[642,768,836,861]" italics="true" pageId="4" pageNumber="155">M. gibbosa</emphasis>
(
<bibRefCitation id="202039B24A45FFADFCB3FCD9FC330E70" author="Fabricius" box="[783,961,836,861]" pageId="4" pageNumber="155" refString="Fabricius, J. C. (1781) Species Insectorum eorum differentias specificas, synonyma auctorum, loca natalia, metamorphosis adiectis observationibus, descriptionibus. Tomo I. Bohn, Hamburgi et Kilonii, 552 pp." type="book" year="1781">Fabricius, 1781</bibRefCitation>
),
<bibRefCitation id="202039B24A45FFADFC69FCD9FBFD0E70" author="Fabricius" box="[981,1039,836,860]" pageId="4" pageNumber="155" refString="Fabricius, J. C. (1781) Species Insectorum eorum differentias specificas, synonyma auctorum, loca natalia, metamorphosis adiectis observationibus, descriptionibus. Tomo I. Bohn, Hamburgi et Kilonii, 552 pp." type="book" year="1781">1781</bibRefCitation>
</taxonomicName>
, and 14 species of
<taxonomicName id="83B13FC04A45FFADFB5FFCD9FAD50E71" box="[1251,1319,836,861]" class="Insecta" family="Chrysomelidae" genus="Stolas" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="76C598514A45FFADFB5FFCD9FAD50E71" box="[1251,1319,836,861]" italics="true" pageId="4" pageNumber="155">Stolas</emphasis>
</taxonomicName>
.
<bibRefCitation id="202039B24A45FFADFA88FCD9FE730EAC" author="Jolivet" pageId="4" pageNumber="155" refString="Jolivet, P. H. &amp; Hawkeswood, T. J. (1995) Host-Plants of Chrysomelidae of the World. (an essay about the relationships between the leaf-beetles and their food plants). Backhuys Publishers, Leiden, 281 pp." type="book" year="1995">Jolivet &amp; Hawkeswood (1995)</bibRefCitation>
recorded
<taxonomicName id="83B13FC04A45FFADFE4EFCF4FDB40EAC" box="[498,582,873,896]" class="Magnoliopsida" family="Convolvulaceae" genus="Ipomea" higherTaxonomySource="GBIF" kingdom="Plantae" order="Solanales" pageId="4" pageNumber="155" phylum="Magnoliophyta" rank="genus">
<emphasis id="76C598514A45FFADFE4EFCF4FDB40EAC" box="[498,582,873,896]" italics="true" pageId="4" pageNumber="155">Ipomea</emphasis>
</taxonomicName>
L. (
<taxonomicName id="83B13FC04A45FFADFDC9FCFAFCDF0EAC" box="[629,813,871,896]" class="Magnoliopsida" family="Convolvulaceae" kingdom="Plantae" order="Solanales" pageId="4" pageNumber="155" phylum="Tracheophyta" rank="family">Convolvulaceae</taxonomicName>
) as the host plant of
<taxonomicName id="83B13FC04A45FFADFBA5FCFAFB430EAC" box="[1049,1201,871,896]" class="Insecta" family="Chrysomelidae" genus="Mesomphalia" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="genus">
<emphasis id="76C598514A45FFADFBA5FCFAFB430EAC" box="[1049,1201,871,896]" italics="true" pageId="4" pageNumber="155">Mesomphalia</emphasis>
</taxonomicName>
, but without citing a reference for this. The only confirmed host plant species is
<taxonomicName id="83B13FC04A45FFADFCE8FC11FADE0E88" authority="Spreng." authorityName="Spreng." box="[852,1324,908,933]" class="Magnoliopsida" family="Asteraceae" genus="Mikania" kingdom="Plantae" order="Asterales" pageId="4" pageNumber="155" phylum="Tracheophyta" rank="species" species="glomerata">
<emphasis id="76C598514A45FFADFCE8FC11FBDD0E89" box="[852,1071,908,933]" italics="true" pageId="4" pageNumber="155">Mikania glomerata</emphasis>
Spreng. (Asteraceae)
</taxonomicName>
, which is attributed to
<taxonomicName id="83B13FC04A45FFADFE9AFC2CFD720EE4" authority="Flinte et al. 2008" authorityName="Flinte et al." authorityYear="2008" box="[294,640,943,968]" class="Insecta" family="Chrysomelidae" genus="Cassida" kingdom="Animalia" order="Coleoptera" pageId="4" pageNumber="155" phylum="Arthropoda" rank="species" species="gibbosa">
<emphasis id="76C598514A45FFADFE9AFC2CFE560EE4" box="[294,420,943,968]" italics="true" pageId="4" pageNumber="155">M. gibbosa</emphasis>
(
<bibRefCitation id="202039B24A45FFADFE0EFC32FD8B0EE4" author="Flinte" box="[434,633,943,968]" pageId="4" pageNumber="155" refString="Flinte, V., de Macedo, M. V. &amp; Monteiro, R. F. (2008) Tortoise Beetles (Chrysomelidae: Cassidinae) of a tropical rain forest in Rio de Janeiro, Brazil. In: Jolivet, P., Santiago-Blay, J. &amp; Schmidt, M. (Eds.), Research on Chrysomelidae. Brill Academic Publishers, Leiden, pp. 195 - 209." type="book chapter" year="2008">
Flinte
<emphasis id="76C598514A45FFADFE40FC2CFDDC0EE4" box="[508,558,943,968]" italics="true" pageId="4" pageNumber="155">et al</emphasis>
. 2008
</bibRefCitation>
)
</taxonomicName>
.
</paragraph>
</subSubSection>
</treatment>
</document>