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<mods:title id="8E6FDAD7CA0C67BB68EB2BC1CE613711">Revision of the freshwater crabs of the genus Tehuana Rodriguez &amp; Smalley in Smalley 1970 (Decapoda, Pseudothelphusidae), with the descriptions of two new species</mods:title>
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<taxonomicName id="7A9334E6A0F1BE76ECA83941F6E04E90" ID-CoL="554MG" LSID="27C50A5D-F13E-5283-A541-2709123F12BE" authority="(Rathbun, 1905)" baseAuthorityName="Rathbun" baseAuthorityYear="1905" class="Malacostraca" family="Pseudothelphusidae" genus="Tehuana" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Tehuana complanata" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="complanata">Tehuana complanata (Rathbun, 1905)</taxonomicName>
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<figureCitation id="DE1E091E262EC6CF9E3A6BE3086C6B15" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Mesial view of the apical portion of the G 1 of the species of Tehuana A T. ayotzintepecensis sp. nov., CNCR 34628 B T. chontalpaensis, CNCR 17093 C T. col sp. nov., CNCR 33928 D T. complanata, CNCR 11957 E T. diabolis, CNCR 12056 F T. lamellifrons, CNCR 33939 G T. lamothei, CNCR 8812 H T. jacatepecensis, CNCR 11920 I T. poglayenorum, CNCR 33931 J T. veracruzana, CNCR 33934. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1117.85362.figure8" httpUri="https://binary.pensoft.net/fig/725904" pageId="0" pageNumber="1">Figs 8D</figureCitation>
<figureCitation id="CCEDA2EA3C4A44FDFA54EAD97279A77B" captionStart="Figure 9" captionStartId="F9" captionText="Figure 9. Cephalic view of the apical portion of the G 1 of the species of Tehuana A T. ayotzintepecensis sp. nov., CNCR 34628 B T. chontalpaensis, CNCR 17093 C T. col sp. nov., CNCR 33928 D T. complanata, CNCR 11957 E T. diabolis, CNCR 12056 F T. lamellifrons, CNCR 33939 G T. lamothei, CNCR 8812 H T. jacatepecensis, CNCR 11920 I T. poglayenorum, CNCR 33931 J T. veracruzana, CNCR 33934. Scale bars: 1 mm (A, C, H, J); 0.5 mm (B, D, E, F, G, I)." figureDoi="10.3897/zookeys.1117.85362.figure9" httpUri="https://binary.pensoft.net/fig/725905" pageId="0" pageNumber="1">, 9D</figureCitation>
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<paragraph id="19AF22C927D73E9532CA9F0F5CB2E390" pageId="0" pageNumber="1">
<taxonomicName id="4BB37FB42A3A43C025A102CF166A067F" class="Magnoliopsida" family="Asteraceae" genus="Pseudothelphusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudothelphusa bocourti" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="bocourti">? Pseudothelphusa bocourti</taxonomicName>
.-Rathbun, 1898: 512 (in key), 513, 533 (list) [not 
<taxonomicName id="0181510E63BBC2ED65CF63A0A8EBF9EE" authorityName="A. Milne-Edwards" authorityYear="1866" class="Magnoliopsida" family="Asteraceae" genus="Boscia" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Boscia bocourti" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="bocourti">Boscia bocourti</taxonomicName>
A. Milne-Edwards, 1866].
</paragraph>
<paragraph id="762261163890F2524B31A8A3F1B0E6F0" pageId="0" pageNumber="1">
<taxonomicName id="CEB9758C890A1FEC7760DC59B43397E0" authorityName="Rathbun" authorityYear="1905" class="Magnoliopsida" family="Asteraceae" genus="Pseudothelphusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudothelphusa complanata" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="complanata">Pseudothelphusa complanata</taxonomicName>
Rathbun, 1905: 303, fig. 3.- 
<bibRefCitation id="3820C4F3EE3473F11C90E92656505C93" author="Coifmann, I" journalOrPublisher="Archivo Zoologico Italiano" pageId="0" pageNumber="1" pagination="93 - 116" refId="B20" refString="Coifmann, I, 1939. Potamonidi dell R. Museo Zoologico di Torino. Archivo Zoologico Italiano 27: 93 - 116" title="Potamonidi dell R. Museo Zoologico di Torino." volume="27" year="1939">Coifmann 1939</bibRefCitation>
: 107 (list).
</paragraph>
<paragraph id="937F1AA13F0F0BBB93DBDFCE4FFF1F0A" pageId="0" pageNumber="1">
<taxonomicName id="4DDCBDEE93148A0FA4FA732855E5E10E" class="Magnoliopsida" family="Asteraceae" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="“Pseudothelphusa” complanata" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="complanata">
<normalizedToken id="2B0DBC156E9002925A8DC2D339F7D2AD" originalValue="“Pseudothelphusa”">&quot;Pseudothelphusa&quot;</normalizedToken>
complanata
</taxonomicName>
.-Pretzmann, 1965: 10 (list).
</paragraph>
<paragraph id="545422ED33C6EEB8B5BB25EABFA7896D" pageId="0" pageNumber="1">
<taxonomicName id="76FD2A8D64C6E7DBA2B539C3F9FFAC7C" authorityName="de Saussure" authorityYear="1857" class="Magnoliopsida" family="Asteraceae" genus="Pseudothelphusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudothelphusa" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Pseudothelphusa</taxonomicName>
<taxonomicName id="9A485BF696B7A8F82F3F6ABDE38143F3" authorityName="de Saussure" authorityYear="1857" class="Magnoliopsida" family="Asteraceae" genus="Pseudothelphusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudothelphusa" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Pseudothelphusa</taxonomicName>
(
<taxonomicName id="CAB433BF847D2B64667342DE7B6E6910" authorityName="de Saussure" authorityYear="1857" class="Magnoliopsida" family="Asteraceae" genus="Pseudothelphusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudothelphusa" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Pseudothelphusa</taxonomicName>
) 
<taxonomicName id="CBDA88DDAC2C5DED08D038074A33C652" authorityName="Pretzmann" authorityYear="1968" class="Magnoliopsida" family="Asteraceae" genus="Pseudothelphusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudothelphusa lamellifrons subsp. gruneri" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="subSpecies" species="lamellifrons" subSpecies="gruneri">Pseudothelphusa lamellifrons gruneri</taxonomicName>
.-Pretzmann, 1968: 7.- 
<bibRefCitation id="60B9AAB549BF542D73FF1E76DBBBC7AF" author="Pretzmann, G" journalOrPublisher="Sitzungsberichte der Mathematisch-naturwissenschaftlichen Klasse der Oesterreichischen Akademie der Wissenschaften, Wien" pageId="0" pageNumber="1" pagination="15 - 24" publicationUrl="https://decapoda.nhm.org/pdfs/27495/27495.pdf" refId="B47" refString="Pretzmann, G, 1971. Fortschritte in der Klassifizierung der Pseudothelphusidae. Sitzungsberichte der Mathematisch-naturwissenschaftlichen Klasse der Oesterreichischen Akademie der Wissenschaften, Wien 179: 15 - 24, https://decapoda.nhm.org/pdfs/27495/27495.pdf" title="Fortschritte in der Klassifizierung der Pseudothelphusidae." url="https://decapoda.nhm.org/pdfs/27495/27495.pdf" volume="179" year="1971">Pretzmann 1971</bibRefCitation>
: 22 (list).
</paragraph>
<paragraph id="1151C6149596597CDF9965262B1531E3" pageId="0" pageNumber="1">
<taxonomicName id="4A6D3F028C8C1ECBDDF9A8280936E369" class="Magnoliopsida" family="Asteraceae" genus="Pseudothelphusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudothelphusa (Pseudothelphusa) americana subsp. lamellifrons" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="subSpecies" species="americana" subGenus="Pseudothelphusa" subSpecies="lamellifrons">Pseudothelphusa (Pseudothelphusa) americana lamellifrons</taxonomicName>
.-Bott, 1970: 334, pl. 2, figs 11-13.
</paragraph>
<paragraph id="90FF388EAA79955F669BA57021F3186D" pageId="0" pageNumber="1">
<taxonomicName id="EEF3B05B376EE1ED27E2017AE8A0DB47" class="Magnoliopsida" family="Asteraceae" genus="Potamocarcinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Potamocarcinus (Raddaus) bocourti subsp. complanata" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="subSpecies" species="bocourti" subGenus="Raddaus" subSpecies="complanata">Potamocarcinus (Raddaus) bocourti complanata</taxonomicName>
.-Pretzmann, 1971: 20.(list).
</paragraph>
<paragraph id="1AA139102DDC126134E5FE2ECD454EA2" pageId="0" pageNumber="1">
<taxonomicName id="F95682D91B40E678655C4A3EB8CC4EB0" class="Magnoliopsida" family="Asteraceae" genus="Potamocarcinus" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Potamocarcinus (Raddaus) bocourti subsp. complanatus" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="subSpecies" species="bocourti" subGenus="Raddaus" subSpecies="complanatus">Potamocarcinus (Raddaus) bocourti complanatus</taxonomicName>
.-Pretzmann, 1972: 78, text fig. 18, fig. 542.
</paragraph>
<paragraph id="998696F567963A3F44721FA2E0FAA05A" pageId="0" pageNumber="1">
<taxonomicName id="7EA677264752C5825870E57A3C0E3E53" authorityName="de Saussure" authorityYear="1857" class="Magnoliopsida" family="Asteraceae" genus="Pseudothelphusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudothelphusa" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="genus">Pseudothelphusa</taxonomicName>
<taxonomicName id="B298FE1A8B26AA96BF13294E63784A68" authorityName="Pretzmann" authorityYear="1968" class="Magnoliopsida" family="Asteraceae" genus="Pseudothelphusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudothelphusa (Tehuana) lamellifrons subsp. gruneri" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="subSpecies" species="lamellifrons" subGenus="Tehuana" subSpecies="gruneri">Pseudothelphusa (Tehuana) lamellifrons gruneri</taxonomicName>
.-Pretzmann, 1972: 108, figs 621-623, 674-677.
</paragraph>
<paragraph id="804CB36B8CAF8530A17B6BC8DEE26120" pageId="0" pageNumber="1">
<taxonomicName id="4638E5A53CCB66EF0C67349E3F9A2CC4" authorityName="Rodriguez &amp; Smalley" authorityYear="1972" class="Magnoliopsida" family="Asteraceae" genus="Pseudothelphusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudothelphusa (Tehuana) cordobensis" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="cordobensis" subGenus="Tehuana">Pseudothelphusa (Tehuana) cordobensis</taxonomicName>
<normalizedToken id="36937708857B3E647E8B4679D9B9D9CB" originalValue="Rodríguez">Rodriguez</normalizedToken>
&amp; Smalley, 1972: 77, fig. 8, pl. 5.
</paragraph>
<paragraph id="552D2A60DCE7905D613AF821C410C616" pageId="0" pageNumber="1">
<taxonomicName id="B3DFD540A14CAC8EDA2AE6A8C6F52863" authorityName="Rathbun" authorityYear="1905" class="Magnoliopsida" family="Asteraceae" genus="Pseudothelphusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudothelphusa (Tehuana) complanata" order="Asterales" pageId="0" pageNumber="1" phylum="Tracheophyta" rank="species" species="complanata" subGenus="Tehuana">Pseudothelphusa (Tehuana) complanata</taxonomicName>
<normalizedToken id="9FB35EB3D9933582123A7A647C548655" originalValue="Türkay">Tuerkay</normalizedToken>
, 1978: 145, figs 2a, b, 3.
</paragraph>
<paragraph id="AC97F7E1E44676870E3B22F9267CE038" pageId="0" pageNumber="1">
<taxonomicName id="1C1B0DECDF9773D21C0559E413E3AA2E" baseAuthorityName="Rathbun" baseAuthorityYear="1905" class="Malacostraca" family="Pseudothelphusidae" genus="Tehuana" higherTaxonomySource="treatment-meta" kingdom="Animalia" lsidName="Tehuana complanata" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="complanata">Tehuana complanata</taxonomicName>
Rodriguez, 1982: 131, fig. 85.- 
<bibRefCitation id="F64060A80E53E2DD7465D95EF626262D" author="Villalobos, A" journalOrPublisher="Lectures on Mathematics in the Life Sciences" pageId="0" pageNumber="1" refId="B66" refString="Villalobos, A, 1982. Decapoda. In: Hulbert SH, Villalobos A (Eds) Aquatic Biota of Mexico, Central America and the West Indies, San Diego University, California, 215-239." title="Decapoda. In: Hulbert SH, Villalobos A (Eds) Aquatic Biota of Mexico, Central America and the West Indies, San Diego University, California, 215 - 239." year="1982">Villalobos 1982</bibRefCitation>
: 221 (list).- 
<bibRefCitation id="7C83DDA00D2828E3C4DA916287EA71F3" author="Villalobos-Hiriart, JL" journalOrPublisher="Revista de la Sociedad Mexicana de Historia Natural" pageId="0" pageNumber="1" pagination="267 - 290" refId="B70" refString="Villalobos-Hiriart, JL, Diaz-Barriga, A, Lira-Fernandez, E, 1993. Los crustaceos de agua dulce de Mexico. Revista de la Sociedad Mexicana de Historia Natural 44: 267 - 290" title="Los crustaceos de agua dulce de Mexico." volume="44" year="1993">Villalobos-Hiriart et al. 1993</bibRefCitation>
: 284 (list).- 
<bibRefCitation id="4FF405FFD249EE1AB29E6787D337EC68" author="Alvarez, F" journalOrPublisher="Proceedings of the Biological Society of Washington" pageId="0" pageNumber="1" pagination="729 - 737" publicationUrl="https://www.biodiversitylibrary.org/page/35515374page/763/mode/1up" refId="B3" refString="Alvarez, F, Villalobos, JL, 1994. Two new species and one new combination of freshwater crabs from Mexico (Crustacea: Brachyura: Pseudothelphusidae). Proceedings of the Biological Society of Washington 107 (4): 729 - 737, https://www.biodiversitylibrary.org/page/35515374page/763/mode/1up" title="Two new species and one new combination of freshwater crabs from Mexico (Crustacea: Brachyura: Pseudothelphusidae)." url="https://www.biodiversitylibrary.org/page/35515374page/763/mode/1up" volume="107" year="1994">Alvarez and Villalobos 1994</bibRefCitation>
: 730.- 
<bibRefCitation id="D8A1711A88028749CFD04C8A3D427144" author="Alvarez, F" journalOrPublisher="Anales del Instituto de Biologia Universidad Nacional Autonoma de Mexico" pageId="0" pageNumber="1" pagination="1 - 27" publicationUrl="http://www.ejournal.unam.mx/zoo/070-01/ZOO70101.pdf" refId="B10" refString="Alvarez, F, Villalobos, JL, Rojas, Y, Robles, R, 1999. Listas y comentarios sobre los crustaceos decapodos de Veracruz, Mexico. Anales del Instituto de Biologia Universidad Nacional Autonoma de Mexico 70 (1): 1 - 27, http://www.ejournal.unam.mx/zoo/070-01/ZOO70101.pdf" title="Listas y comentarios sobre los crustaceos decapodos de Veracruz, Mexico." url="http://www.ejournal.unam.mx/zoo/070-01/ZOO70101.pdf" volume="70" year="1999">
<normalizedToken id="19D4BBB64D28AE483D3FC5D884B8E57E" originalValue="Álvarez">Alvarez</normalizedToken>
et al. 1999
</bibRefCitation>
: 20, fig. 3 (map), 23, box 3 (list).-Villalobos and Alvarez 2003: 2003 (in key).- 
<bibRefCitation id="F8E618D318383D2BA89574151E3450C7" author="Villalobos Hiriart, JL" editor="Alvarez, F" journalOrPublisher="UANL-PROMEP-SEP. Direccion General de Publicaciones, Universidad Autonoma de Nuevo Leon, Mexico, D. F." pageId="0" pageNumber="1" pagination="239 - 299" refId="B69" refString="Villalobos Hiriart, JL, Alvarez, F, 2008. Los cangrejos de agua dulce de la familia Pseudothelphusidae (Decapoda: Brachyura: Eubrachyura) de Mexico. Con un apendice de las especies citadas para America hasta el 2006. In: Alvarez, F, Rodriguez, G, Eds., Crustaceos de Mexico: Estado actual de su conocimiento. UANL-PROMEP-SEP. Direccion General de Publicaciones, Universidad Autonoma de Nuevo Leon, Mexico, D.F.: 239 - 299" title="Los cangrejos de agua dulce de la familia Pseudothelphusidae (Decapoda: Brachyura: Eubrachyura) de Mexico. Con un apendice de las especies citadas para America hasta el 2006." volumeTitle="Crustaceos de Mexico: Estado actual de su conocimiento." year="2008">
Villalobos Hiriart and 
<normalizedToken id="10BBE0C1FD2B19C7571CF1466649A685" originalValue="Álvarez">Alvarez</normalizedToken>
2008
</bibRefCitation>
: 280, 298 (list).- 
<bibRefCitation id="7C26CAFABDCB22FA9B8A7731F0399B28" author="Ng, PKL" journalOrPublisher="The Raffles Bulletin of Zoology" pageId="0" pageNumber="1" pagination="1 - 286" publicationUrl="https://lkcnhm.nus.edu.sg/wp-content/uploads/sites/10/app/uploads/2017/04/s17rbz.pdf" refId="B39" refString="Ng, PKL, Guinot, D, Davie, PJF, 2008. Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world. The Raffles Bulletin of Zoology 17: 1 - 286, https://lkcnhm.nus.edu.sg/wp-content/uploads/sites/10/app/uploads/2017/04/s17rbz.pdf" title="Systema Brachyurorum: Part I. An annotated checklist of extant brachyuran crabs of the world." url="https://lkcnhm.nus.edu.sg/wp-content/uploads/sites/10/app/uploads/2017/04/s17rbz.pdf" volume="17" year="2008">Ng et al. 2008</bibRefCitation>
: 177 (list).-Villalobos and Alvarez 2010: 474, 477, fig. 11 (map).- 
<bibRefCitation id="B9F022F90DDB0C38B52A9EE4305788EF" author="Alvarez, F" editor="Cruz-Angon, A" journalOrPublisher="Vol. 2. Comision Nacional para el Conocimiento y Uso de la Biodiversidad, Gobierno del Estado de Veracruz, Universidad Veracruzana, Instituto de Ecologia" pageId="0" pageNumber="1" pagination="287 - 294" refId="B12" refString="Alvarez, F, Villalobos, JL, Chazaro-Olvera, S, 2011a. Camarones y cangrejos dulceacuicolas y marinos (Crustacea: Decapoda). In: Cruz-Angon, A, Lorea-Hernandez, FG, Hernandez-Ortiz, V, Morales-Mavil, JE, Eds., La biodiversidad en Veracruz, estudio de estado, diversidad de especies: conocimiento actual. Vol. 2. Comision Nacional para el Conocimiento y Uso de la Biodiversidad, Gobierno del Estado de Veracruz, Universidad Veracruzana, Instituto de Ecologia: 287 - 294" title="Camarones y cangrejos dulceacuicolas y marinos (Crustacea: Decapoda)." volumeTitle="La biodiversidad en Veracruz, estudio de estado, diversidad de especies: conocimiento actual." year="2011 a">
<normalizedToken id="7EE86E81CC71685C0C259B221A6D90E4" originalValue="Álvarez">Alvarez</normalizedToken>
et al. 2011a
</bibRefCitation>
: appx. VIII.20, p. 13 (list).- 
<bibRefCitation id="4666A34F94E12726CBB086E692882AD2" author="Mejia-Ortiz, L" journalOrPublisher="Universidad de Quintana Roo, Unidad Cozumel, Mexico" pageId="0" pageNumber="1" refId="B33" refString="Mejia-Ortiz, L, Lopez-Mejia, M, Martinez, MI, 2011. Crustaceos decapodos del Rio Papaloapan, Veracruz. Universidad de Quintana Roo, Unidad Cozumel, Mexico" title="Crustaceos decapodos del Rio Papaloapan, Veracruz." year="2011">
<normalizedToken id="378B272131866A64878FE36B0D856357" originalValue="Mejía-Ortíz">Mejia-Ortiz</normalizedToken>
et al. 2011
</bibRefCitation>
: 97, 136 (map 2).- 
<bibRefCitation id="FC2818CBE8EE3D94803FBE5204068BE5" DOI="https://doi.org/10.22201/ib.20078706e.2012.4.1268" author="Alvarez, F" journalOrPublisher="Revista Mexicana de Biodiversidad" pageId="0" pageNumber="1" pagination="1073 - 1083" refId="B14" refString="Alvarez, F, Villalobos, JL, Armendariz, G, Hernandez, C, 2012. Relacion biogeografica entre cangrejos dulceacuicolas y acociles a lo largo de la zona mexicana de transicion: Revaluacion de la hipotesis de Rodriguez (1986). Revista Mexicana de Biodiversidad 83 (4): 1073 - 1083, DOI: https://doi.org/10.22201/ib.20078706e.2012.4.1268" title="Relacion biogeografica entre cangrejos dulceacuicolas y acociles a lo largo de la zona mexicana de transicion: Revaluacion de la hipotesis de Rodriguez (1986)." url="https://doi.org/10.22201/ib.20078706e.2012.4.1268" volume="83" year="2012">
<normalizedToken id="32FA6726090DF096F634DBA92AE3D453" originalValue="Álvarez">Alvarez</normalizedToken>
et al. 2012
</bibRefCitation>
: 1078, box 1 (list).- 
<bibRefCitation id="47200BAB3F68620FD95B0AA1B7DA3922" DOI="https://doi.org/10.3897/zookeys.457.6598" author="Cumberlidge, N" journalOrPublisher="ZooKeys" pageId="0" pageNumber="1" pagination="122 - 137" refId="B22" refString="Cumberlidge, N, Alvarez, F, Villalobos, JL, 2014. Results of the global conservation assessment of the freshwater crabs (Brachyura, Pseudothelphusidae and Trichodactylidae): The Neotropical region, with an update on diversity. ZooKeys 45: 122 - 137, DOI: https://doi.org/10.3897/zookeys.457.6598" title="Results of the global conservation assessment of the freshwater crabs (Brachyura, Pseudothelphusidae and Trichodactylidae): The Neotropical region, with an update on diversity." url="https://doi.org/10.3897/zookeys.457.6598" volume="45" year="2014">Cumberlidge et al. 2014</bibRefCitation>
: 144, tab. 3 (list).- 
<bibRefCitation id="5ADEA4FA2A20DB89E2017776230DB1B5" DOI="https://doi.org/10.1007/978-3-319-42527-6_8" author="Alvarez, F" editor="Kawai, T" journalOrPublisher="Springer, Cham" pageId="0" pageNumber="1" pagination="237 - 266" refId="B8" refString="Alvarez, F, Villalobos, JL, 2016. Chapter 8: Freshwater decapod diversity and conservation in Mexico. In: Kawai, T, Cumberlidge, N, Eds., A global overview of the conservation of freshwater decapod crustaceans. Springer, Cham: 237 - 266, DOI: https://doi.org/10.1007/978-3-319-42527-6_8" title="Chapter 8: Freshwater decapod diversity and conservation in Mexico." url="https://doi.org/10.1007/978-3-319-42527-6_8" volumeTitle="A global overview of the conservation of freshwater decapod crustaceans." year="2016">Alvarez and Villalobos 2016</bibRefCitation>
: 254, tab.8.1 (list).- 
<bibRefCitation id="EA6518CD1CA675FE17DB6BED5F109B82" DOI="https://doi.org/10.11646/zootaxa.4586.1.8" author="Villalobos-Hiriart, JL" journalOrPublisher="Zootaxa" pageId="0" pageNumber="1" pagination="151 - 161" refId="B71" refString="Villalobos-Hiriart, JL, Kamanli, SA, Alvarez, F, Garbout, A, Clark, PF, 2019. Lobithelphusa mexicana Rodriguez, 1982 (Decapoda: Pseudothelphusidae): a reassessment of key characters and systematics. Zootaxa 4586 (1): 151 - 161, DOI: https://doi.org/10.11646/zootaxa.4586.1.8" title="Lobithelphusa mexicana Rodriguez, 1982 (Decapoda: Pseudothelphusidae): a reassessment of key characters and systematics." url="https://doi.org/10.11646/zootaxa.4586.1.8" volume="4586" year="2019">Villalobos-Hiriart et al. 2019</bibRefCitation>
: 156, tab.1 (list).
</paragraph>
</subSubSection>
<subSubSection id="0AA2584EC01AFAB97A6E947EE235B7CF" pageId="0" pageNumber="1" type="materials_examined">
<paragraph id="23EB88A8E8F7773DBC358EB4DF5D7C0F" pageId="0" pageNumber="1">Material examined.</paragraph>
<paragraph id="3A307E41379525F56E0AEA2CE726E331" pageId="0" pageNumber="1">
<materialsCitation id="D1F6E95350DCEC71AA3AB77B3C18DD75" collectingDate="1953-05-02" collectorName="A. Villalobos" country="Mexico" elevation="661" latitude="18.876389" location="Paraje Nuevo" longLatPrecision="21" longitude="-96.86361" municipality="Municipality of Cordoba Ojo de Agua" specimenCount="1" specimenCount-male="1" stateProvince="Veracruz" typeStatus="holotype">
<collectingCountry id="00B9A87301FDACA673B97FE254E4E4DF" name="Mexico">Mexico</collectingCountry>
- 
<collectingRegion id="4FE68A8C7BD69CB370B186F6CD333CEB" country="Mexico" name="Veracruz de Ignacio de la Llave">Veracruz</collectingRegion>
• 
<specimenCount id="285502C14661B6C0C0ADA75327EF7BDB" type="male">1 ♂</specimenCount>
, 
<typeStatus id="AA5A6E4C98F9A8DEAB0DD02A0985EC83">holotype</typeStatus>
of 
<taxonomicName id="5ED845DC094C401DFC5240B5963C83FC" authorityName="Rodriguez &amp; Smalley" authorityYear="1972" class="Malacostraca" family="Asteraceae" genus="Pseudothelphusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudothelphusa (Tehuana) cordobensis" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="cordobensis" subGenus="Tehuana">Pseudothelphusa (Tehuana) cordobensis</taxonomicName>
<bibRefCitation id="40B69C7A52D8EA1B75042CABBE5D309A" author="Rodriguez, G" journalOrPublisher="Anales del Instituto de Biologia, Universidad Nacional Autonoma de Mexico, serie Ciencias del Mar y Limnologia" pageId="0" pageNumber="1" pagination="69 - 112" publicationUrl="https://decapoda.nhm.org/pdfs/27336/27336.pdf" refId="B59" refString="Rodriguez, G, Smalley, AE, 1972. Los cangrejos de agua dulce de Mexico de la familia Pseudothelphusidae (Crustacea, Brachyura). Anales del Instituto de Biologia, Universidad Nacional Autonoma de Mexico, serie Ciencias del Mar y Limnologia 40: 69 - 112, https://decapoda.nhm.org/pdfs/27336/27336.pdf" title="Los cangrejos de agua dulce de Mexico de la familia Pseudothelphusidae (Crustacea, Brachyura)." url="https://decapoda.nhm.org/pdfs/27336/27336.pdf" volume="40" year="1972">
<normalizedToken id="9FCAAB72B3E91BB1DCC57F5E48A22CBB" originalValue="Rodríguez">Rodriguez</normalizedToken>
and Smalley (1972)
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; LC 
<quantity id="B669478C25D684DBC98F2840B4C472D7" metricMagnitude="-2" metricUnit="m" metricValue="3.82" unit="mm" value="38.2">38.2 mm</quantity>
, AC 59.0 mm; 
<collectingMunicipality id="E6330B79D9E15CFB40C443A5CC897AFC">
Municipality of 
<normalizedToken id="D604E695B5A5CF608B82B734B56C7DEF" originalValue="Córdoba">Cordoba</normalizedToken>
Ojo de Agua
</collectingMunicipality>
, 
<location id="B32D3420B3A2AB68DB137E695BD048DC" LSID="urn:lsid:plazi:treatment:27C50A5DF13E5283A5412709123F12BE:B32D3420B3A2AB68DB137E695BD048DC" country="Mexico" latitude="18.876389" longLatPrecision="21" longitude="-96.86361" municipality="Municipality of Cordoba Ojo de Agua" name="Paraje Nuevo" stateProvince="Veracruz">Paraje Nuevo</location>
, 
<geoCoordinate id="44A6183D0E3D4887F999AFF588F7C8CD" degrees="18" direction="north" minutes="52" orientation="latitude" precision="15" seconds="35" value="18.876389">18°52'35&quot;N</geoCoordinate>
, 
<geoCoordinate id="741CCF4D602AA2D10900EC739087A6F1" degrees="96" direction="west" minutes="51" orientation="longitude" precision="15" seconds="49" value="-96.86361">96°51'49&quot;W</geoCoordinate>
; alt. 
<quantity id="5B001F1C48DC1129B5BCC0D983B13688" metricMagnitude="2" metricUnit="m" metricValue="6.609999999999999" unit="m" value="661.0">
<elevation id="ACDB69654FFEE85CBAADF34600FBD9C3" metricMagnitude="2" metricUnit="m" metricValue="6.609999999999999" unit="m" value="661.0">661 m</elevation>
</quantity>
; 
<collectingDate id="3ECC4F394CE2B23475641DC2A26C4BA6" value="1953-05-02">2 May 1953</collectingDate>
; 
<collectorName id="E116FB855392A3B9CC8624FB4ED40175">A. Villalobos</collectorName>
leg.; CNCR 311
</materialsCitation>
. 
<materialsCitation id="77581F51DF66DE901443919F97E23376" collectingDate="1996-06-13" collectorName="J. Herrera, E. Ramirez" country="Mexico" elevation="439" latitude="18.783333" location="Lourdes River Cave" longLatPrecision="21" longitude="-96.9" municipality="Municipality of Amatlan de Los Reyes" specimenCount="1" specimenCount-male="1" stateProvince="Veracruz" typeStatus="holotype">
<specimenCount id="9C46859386C66C712C6DC12E7C2C9DA8" type="male">1 ♂</specimenCount>
; LC 30.0 mm, AC 
<quantity id="D3624021EE33E1FBD6EACDA69A6D5708" metricMagnitude="-2" metricUnit="m" metricValue="4.54" unit="mm" value="45.4">45.4 mm</quantity>
; 
<collectingMunicipality id="96EF921B427F0C64F8A7F58774AB4BFC">
Municipality of 
<normalizedToken id="CD15C9D75CD308A270BF2F651396C47B" originalValue="Amatlán">Amatlan</normalizedToken>
de Los Reyes
</collectingMunicipality>
, 
<location id="A16A3AFF01A25225380E6C4AF866DD90" LSID="urn:lsid:plazi:treatment:27C50A5DF13E5283A5412709123F12BE:A16A3AFF01A25225380E6C4AF866DD90" country="Mexico" latitude="18.783333" longLatPrecision="21" longitude="-96.9" municipality="Municipality of Amatlan de Los Reyes" name="Lourdes River Cave" stateProvince="Veracruz">Lourdes River Cave</location>
; 
<geoCoordinate id="4D90A6D2A281DBFFADABD1BCA31BF34E" degrees="18" direction="north" minutes="47" orientation="latitude" precision="15" seconds="00" value="18.783333">18°47'00&quot;N</geoCoordinate>
, 
<geoCoordinate id="ABF07DFEFE9A5437A99FC4ABC933841B" degrees="96" direction="west" minutes="54" orientation="longitude" precision="15" seconds="00" value="-96.9">96°54'00&quot;W</geoCoordinate>
; alt. 
<quantity id="4DC7EBA67B5C506E5FF432349E8991C1" metricMagnitude="2" metricUnit="m" metricValue="4.39" unit="m" value="439.0">
<elevation id="6D6A37E4A0CFBC01299C18F69FC1B892" metricMagnitude="2" metricUnit="m" metricValue="4.39" unit="m" value="439.0">439 m</elevation>
</quantity>
; 
<collectingDate id="4FCC28D555935CBF2B681BD16DB71797" value="1996-06-13">13 Jun. 1996</collectingDate>
; 
<collectorName id="C2AD397E79ACEB7EBB04D06BBBD5F301">J. Herrera</collectorName>
, 
<collectorName id="598080860D36BBE22B69DE13C888F156">
E. 
<normalizedToken id="6060908BBC66EC062AB8981BFE1E60FD" originalValue="Ramírez">Ramirez</normalizedToken>
</collectorName>
leg.; CNCR 11958
</materialsCitation>
. 
<materialsCitation id="AAFC713F9FC5702CA556DFF3A5EBC03C" collectingDate="1992-06-06" collectorName="J. Herrera, E. Ramirez" country="Mexico" county="Municipality of Amatlan de Los Reyes" elevation="730" latitude="18.85639" location="Power Station" longLatPrecision="21" longitude="-96.90528" municipality="Amatlan" specimenCount="6" specimenCount-female="3" specimenCount-male="3" stateProvince="Veracruz" typeStatus="holotype">
<specimenCount id="3E07EF17FB461F5E2A9C1D4039614114" type="male">3 ♂</specimenCount>
, 
<specimenCount id="5F36359C662193707F832614E98D2611" type="female">3 ♀</specimenCount>
; LC 
<quantity id="1CF696013B129738BFD8E7595C4670FF" metricMagnitude="-2" metricUnit="m" metricValue="2.165" metricValueMax="2.79" metricValueMin="1.54" unit="mm" value="21.65" valueMax="27.9" valueMin="15.4">15.4-27.9 mm</quantity>
, AC 
<quantity id="20C60BB7BB93430CE86D6F7F3DADE513" metricMagnitude="-2" metricUnit="m" metricValue="3.28" metricValueMax="4.27" metricValueMin="2.29" unit="mm" value="32.8" valueMax="42.7" valueMin="22.9">22.9-42.7 mm</quantity>
; 
<collectingCounty id="6915412F855BA3794845793E10AEB086">
Municipality of 
<normalizedToken id="81AE4EA26233BC8F00863CD42D753116" originalValue="Amatlán">Amatlan</normalizedToken>
de Los Reyes
</collectingCounty>
, 
<normalizedToken id="73CE3D8FC41F901BD941BE05F31C3A75" originalValue="Amatlán">
<collectingMunicipality id="88A7CDB4CD4D1CBC8FA1A375D73FD4F6">Amatlan</collectingMunicipality>
</normalizedToken>
II 
<location id="4EE3A4C143D52E60225595EDCEE9E423" LSID="urn:lsid:plazi:treatment:27C50A5DF13E5283A5412709123F12BE:4EE3A4C143D52E60225595EDCEE9E423" country="Mexico" county="Municipality of Amatlan de Los Reyes" latitude="18.85639" longLatPrecision="21" longitude="-96.90528" municipality="Amatlan" name="Power Station" stateProvince="Veracruz">Power Station</location>
; 
<geoCoordinate id="4BB10C8ACE91B0C59D7C84EF9BC61A27" degrees="18" direction="north" minutes="51" orientation="latitude" precision="15" seconds="23" value="18.85639">18°51'23&quot;N</geoCoordinate>
, 
<geoCoordinate id="AA48D5DFED616A609DB32D2B5E182003" degrees="96" direction="west" minutes="54" orientation="longitude" precision="15" seconds="19" value="-96.90528">96°54'19&quot;W</geoCoordinate>
; alt. 
<quantity id="9993162B33F9A036B1D4F6EF24169547" metricMagnitude="2" metricUnit="m" metricValue="7.3" unit="m" value="730.0">
<elevation id="29270DBFB1B61E28BCF4B5302F2E89E1" metricMagnitude="2" metricUnit="m" metricValue="7.3" unit="m" value="730.0">730 m</elevation>
</quantity>
; 
<collectingDate id="EA1CF2ABE902C0F0887C4DCA6BB557B4" value="1992-06-06">6 Jun. 1992</collectingDate>
; 
<collectorName id="65CDBCEE2B126E35AAC69BE25C12AB3E">J. Herrera</collectorName>
, 
<collectorName id="34598C8AB022F08D2DA40B113063E767">
E. 
<normalizedToken id="4B1F04E452B08283DF8C1E7CEC864A51" originalValue="Ramírez">Ramirez</normalizedToken>
</collectorName>
leg.; CNCR 11957
</materialsCitation>
. 
<materialsCitation id="A537DF384BC8369D909C21F245CAC0CC" collectingDate="2022-01-01" collectingDateMax="2022-12-31" collectingDateMin="2022-01-01" collectorName="C. Pedraza, L. Garcia" country="Mexico" county="Motzorongo" elevation="271" latitude="18.641666" location="Motzorongo River" longLatPrecision="21" longitude="-96.73139" municipality="Municipality of Tezonapa" specimenCount="1" specimenCount-female="1" stateProvince="Veracruz" typeStatus="holotype">
<specimenCount id="63EE23AD7EB3E057AA203553DA0BD4AE" type="female">1 ♀</specimenCount>
, LC 
<quantity id="9E3281E86582771FCDC43B4B77D5F2E9" metricMagnitude="-2" metricUnit="m" metricValue="2.62" unit="mm" value="26.2">26.2 mm</quantity>
, AC 
<quantity id="B9E7359750E6A8A026D81978B234DF00" metricMagnitude="-2" metricUnit="m" metricValue="1.89" unit="mm" value="18.9">18.9 mm</quantity>
; 
<collectingCounty id="903FE098E65A45CEE63ACAAEE5B4C579">Motzorongo</collectingCounty>
, 
<collectingMunicipality id="0AA6AE7267A65B1030507C94080D7D7B">Municipality of Tezonapa</collectingMunicipality>
, 
<location id="5AFB1A251A54FBDF254C874591A12AA6" LSID="urn:lsid:plazi:treatment:27C50A5DF13E5283A5412709123F12BE:5AFB1A251A54FBDF254C874591A12AA6" country="Mexico" county="Motzorongo" latitude="18.641666" longLatPrecision="21" longitude="-96.73139" municipality="Municipality of Tezonapa" name="Motzorongo River" stateProvince="Veracruz">Motzorongo River</location>
; 
<geoCoordinate id="72071C7B96E8B2B5A2A7536661483E22" degrees="18" direction="north" minutes="38" orientation="latitude" precision="15" seconds="30" value="18.641666">18°38'30&quot;N</geoCoordinate>
, 
<geoCoordinate id="4B4B6D32F3CA369D115079705334EE73" degrees="96" direction="west" minutes="43" orientation="longitude" precision="15" seconds="53" value="-96.73139">96°43'53&quot;W</geoCoordinate>
; alt. 
<quantity id="C37C2167A028A4EE12A9857086A9548F" metricMagnitude="2" metricUnit="m" metricValue="2.71" unit="m" value="271.0">
<elevation id="861E66601B84DE4D8402C57B7569F573" metricMagnitude="2" metricUnit="m" metricValue="2.71" unit="m" value="271.0">271 m</elevation>
</quantity>
; 
<collectorName id="7FDD5FB6C6A68C84ECDAB863D026CF9E">C. Pedraza</collectorName>
, 
<collectorName id="0DAF2861555A6920C19906781C510263">
L. 
<normalizedToken id="3CF086537CF35CF0BF29444BBEBA23DD" originalValue="García">Garcia</normalizedToken>
</collectorName>
leg.; CNCR 34618
</materialsCitation>
.
</paragraph>
</subSubSection>
<subSubSection id="A3FBC5ED63157ED7BAF46E679D2BDE01" pageId="0" pageNumber="1" type="diagnosis">
<paragraph id="C970E1BD49985D5A984CC8A9C5F732AA" pageId="0" pageNumber="1">Diagnosis.</paragraph>
<paragraph id="EAAFE8EF053CA3B174C3E7263577326F" pageId="0" pageNumber="1">
G1, in cephalic view, with three protuberances on lateral surface, proximal one being the most developed. In mesial view, meso-distal lobe conical, with round and slender apex. In caudal view, median constriction forming a large lobe oriented proximally. Caudo-marginal projection with distal lobes separated by linear notch without leaving space in between. Distal lobe with rounded cephalic edge, shorter than proximal lobe. Proximal lobe ax-shaped, higher than distal (1.5 
<normalizedToken id="B9CD46DEB707230CD149833985B2D895" originalValue="×">x</normalizedToken>
); cephalic border circular, caudal border straight; distal crest slightly laterally oriented, lobe with sloping appearance. Internal carina well marked, circular, its length covering at least ⅓ of internal surface. Mesial process reduced, in transversal position, without lateral spine; distal edge oriented cephalad. Internal angle developed in form of lobe, two-thirds as high as mesial process, wide, touching the internal surface of distal lobe (DL) of CMP. In distal view, mesial process concave, internal angle hidden below the proximal lobe (PL) of caudo-marginal projection.
</paragraph>
</subSubSection>
<subSubSection id="E6422127D14F4EB5B22F0A7491794E1C" pageId="0" pageNumber="1" type="distribution">
<paragraph id="F4AA4FE264DBBEDA9E8379C8F28BDFDA" pageId="0" pageNumber="1">Distribution.</paragraph>
<paragraph id="5EE8C5E53CC445438DC4E2BF68519C3B" pageId="0" pageNumber="1">
<taxonomicName id="1B106CA603888737F94418CB155DFF97" baseAuthorityName="Rathbun" baseAuthorityYear="1905" class="Malacostraca" family="Pseudothelphusidae" genus="Tehuana" kingdom="Animalia" lsidName="Tehuana complanata" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="complanata">
<emphasis id="40651723E63F87C14CEE189FCEF4A2EE" italics="true" pageId="0" pageNumber="1">Tehuana complanata</emphasis>
</taxonomicName>
is distributed around the city of Cordoba, Veracruz, Mexico (Fig. 
<figureCitation id="48419F620BE69D7920C54C301EFD285A" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution of the species of the genus Tehuana in southeastern Mexico." figureDoi="10.3897/zookeys.1117.85362.figure1" httpUri="https://binary.pensoft.net/fig/725897" pageId="0" pageNumber="1">1</figureCitation>
).
</paragraph>
</subSubSection>
<subSubSection id="AEDB1B591EB9C1C90A5D1E5BF0BC9F09" pageId="0" pageNumber="1" type="remarks">
<paragraph id="09F2A5F73AD990B3AD2F0365D898E0D3" pageId="0" pageNumber="1">Remarks.</paragraph>
<paragraph id="9187CE83A93AB8EFCA2C82C05B82946D" pageId="0" pageNumber="1">
The recognition of the type locality of 
<taxonomicName id="A3C7D4FFDC858AE7FDE10549944C1A9A" baseAuthorityName="Rathbun" baseAuthorityYear="1905" class="Malacostraca" family="Pseudothelphusidae" genus="Tehuana" kingdom="Animalia" lsidName="Tehuana complanata" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="complanata">
<emphasis id="9F5121C1FDF14114AD57C302C6D81C1F" italics="true" pageId="0" pageNumber="1">Tehuana complanata</emphasis>
</taxonomicName>
has been problematic since 
<bibRefCitation id="979AC784AABC6FBF8FF1B84057261EFA" author="Rathbun, MJ" journalOrPublisher="Proceedings of the United States National Museum" pageId="0" pageNumber="1" refId="B54" refString="Rathbun, MJ, 1905. Les Crabes d'eau douce (Potamidae). Nouvelles Archives du Museum d'Histoire naturelle. Paris, series 4 7: 159-321." title="Les Crabes d'eau douce (Potamidae). Nouvelles Archives du Museum d'Histoire naturelle. Paris, series 4 7: 159 - 321." year="1905">Rathbun (1905)</bibRefCitation>
cited &quot;Coban, Alta Vera Paz, Guatemala&quot; as the type locality of 
<taxonomicName id="D9227F0C75888C810EBA164964DB3CB0" authorityName="Rathbun" authorityYear="1905" class="Malacostraca" family="Asteraceae" genus="Pseudothelphusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudothelphusa complanata." order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="complanata.">
<emphasis id="2CB43B59B8F87A8331E28F1A21C9E611" italics="true" pageId="0" pageNumber="1">Pseudothelphusa complanata.</emphasis>
</taxonomicName>
Later, 
<bibRefCitation id="BA644106FCC0AAB7BFE4F7B09FD4EAA7" author="Rodriguez, G" journalOrPublisher="Anales del Instituto de Biologia, Universidad Nacional Autonoma de Mexico, serie Ciencias del Mar y Limnologia" pageId="0" pageNumber="1" pagination="69 - 112" publicationUrl="https://decapoda.nhm.org/pdfs/27336/27336.pdf" refId="B59" refString="Rodriguez, G, Smalley, AE, 1972. Los cangrejos de agua dulce de Mexico de la familia Pseudothelphusidae (Crustacea, Brachyura). Anales del Instituto de Biologia, Universidad Nacional Autonoma de Mexico, serie Ciencias del Mar y Limnologia 40: 69 - 112, https://decapoda.nhm.org/pdfs/27336/27336.pdf" title="Los cangrejos de agua dulce de Mexico de la familia Pseudothelphusidae (Crustacea, Brachyura)." url="https://decapoda.nhm.org/pdfs/27336/27336.pdf" volume="40" year="1972">
<normalizedToken id="FD5EACBBDA6A9378C7A009857EA1F845" originalValue="Rodríguez">Rodriguez</normalizedToken>
and Smalley (1972)
</bibRefCitation>
described 
<taxonomicName id="7BD4562B2401C5A01EF77D8BA49C80D6" authorityName="Rodriguez &amp; Smalley" authorityYear="1972" class="Malacostraca" family="Asteraceae" genus="Pseudothelphusa" higherTaxonomySource="CoL" kingdom="Animalia" lsidName="Pseudothelphusa (Tehuana) cordobensis" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="cordobensis" subGenus="Tehuana">Pseudothelphusa (Tehuana) cordobensis</taxonomicName>
from &quot;Paraje Nuevo, 
<normalizedToken id="FC0ADA7C6E6947939A2A7A098F3E6A66" originalValue="Córdoba">Cordoba</normalizedToken>
, Veracruz&quot; which fits the description of 
<taxonomicName id="26603E2C369B408DEF8FA2E0DD9CEBDE" lsidName="T. complanata" pageId="0" pageNumber="1" rank="species" species="complanata">
<emphasis id="063704B79A9EC1290E7B9B65D2CC8A39" italics="true" pageId="0" pageNumber="1">T. complanata</emphasis>
</taxonomicName>
. 
<bibRefCitation id="AB18D1A7F1F2E36A0E3E88A100173A67" author="Tuerkay, M" journalOrPublisher="Lectures on Mathematics in the Life Sciences" pageId="0" pageNumber="1" refId="B65" refString="Tuerkay, M, 1978. Die identitaet von Pseudothelphusa complanata Rathbun, 1905 mit Bemerkugen zu Pseudothelphusa (Tehuana) Rodriguez &amp; Smalley, 1969 (Crustacea: Decapoda: Pseudothelphusidae). Senckenbergiana Biologica 59(1/2): 143-150." title="Die identitaet von Pseudothelphusa complanata Rathbun, 1905 mit Bemerkugen zu Pseudothelphusa (Tehuana) Rodriguez &amp; Smalley, 1969 (Crustacea: Decapoda: Pseudothelphusidae). Senckenbergiana Biologica 59 (1 / 2): 143 - 150." year="1978">
<normalizedToken id="4550FCB4F0B0523845B93C465D30D446" originalValue="Türkay">Tuerkay</normalizedToken>
(1978)
</bibRefCitation>
discussed this situation concluding that it was a labelling error by Bocourt who placed crabs from Veracruz, Mexico in a jar with specimens from Coban, Guatemala. 
<bibRefCitation id="C68BA81CDE3CBF85477C3738D3CD9CD7" author="Rodriguez, G" journalOrPublisher="Faune Tropicale, ORSTOM, Paris" pageId="0" pageNumber="1" pagination="1 - 224" publicationUrl="https://decapoda.nhm.org/pdfs/11760/11760.pdf" refId="B56" refString="Rodriguez, G, 1982. Les crabes d'eau douce d'Amerique. Famille des Pseudothelphusidae. Faune Tropicale, ORSTOM, Paris 22: 1 - 224, https://decapoda.nhm.org/pdfs/11760/11760.pdf" title="Les crabes d'eau douce d'Amerique. Famille des Pseudothelphusidae." url="https://decapoda.nhm.org/pdfs/11760/11760.pdf" volume="22" year="1982">Rodriguez (1982)</bibRefCitation>
synonymized 
<taxonomicName id="4283FAC5F38AC6B503911250D076D453" lsidName="P. (T.) cordobensis" pageId="0" pageNumber="1" rank="species" species="cordobensis" subGenus="T.">P. (T.) cordobensis</taxonomicName>
under 
<taxonomicName id="CB53CA0F1D721218A358D32A88B688F5" lsidName="T. complanata" pageId="0" pageNumber="1" rank="species" species="complanata">
<emphasis id="9B1FFA4F7A0076AC863F9B38BBE1C7F1" italics="true" pageId="0" pageNumber="1">T. complanata</emphasis>
</taxonomicName>
and designated the male from &quot;Paraje Nuevo, 
<normalizedToken id="1057F64995D4F971814033345E9F2228" originalValue="Córdoba">Cordoba</normalizedToken>
, Veracruz&quot; (CNCR 311) as the holotype.
</paragraph>
<paragraph id="B2996F313150A3963024C88BA59936CB" pageId="0" pageNumber="1">
Morphologically, the G1 of 
<taxonomicName id="B96F7BE265BBB1A45CB2DFB39F48F988" lsidName="T. complanata" pageId="0" pageNumber="1" rank="species" species="complanata">
<emphasis id="E1E64299A3F04B186E3093AF2B7ED54D" italics="true" pageId="0" pageNumber="1">T. complanata</emphasis>
</taxonomicName>
is similar to that of 
<taxonomicName id="638DA532D7B8FC8E0DB59564CEC7B0AD" lsidName="T. jacatepecensis" pageId="0" pageNumber="1" rank="species" species="jacatepecensis">
<emphasis id="5CB3FC3F81BD039876CE7AA89ADBC849" italics="true" pageId="0" pageNumber="1">T. jacatepecensis</emphasis>
</taxonomicName>
; both species have a broadly rounded to semicircular proximal lobe of the CMP, although in the former the distal and proximal lobes are subequal in length, whereas in the latter the proximal lobe is clearly shorter (Fig. 
<figureCitation id="5253D48BDA48344B9CF964D42F775032" captionStart="Figure 8" captionStartId="F8" captionText="Figure 8. Mesial view of the apical portion of the G 1 of the species of Tehuana A T. ayotzintepecensis sp. nov., CNCR 34628 B T. chontalpaensis, CNCR 17093 C T. col sp. nov., CNCR 33928 D T. complanata, CNCR 11957 E T. diabolis, CNCR 12056 F T. lamellifrons, CNCR 33939 G T. lamothei, CNCR 8812 H T. jacatepecensis, CNCR 11920 I T. poglayenorum, CNCR 33931 J T. veracruzana, CNCR 33934. Scale bars: 1 mm." figureDoi="10.3897/zookeys.1117.85362.figure8" httpUri="https://binary.pensoft.net/fig/725904" pageId="0" pageNumber="1">8D</figureCitation>
). 
<taxonomicName id="03C575A3BA81FB86DCA7DB271BF4E0A5" baseAuthorityName="Rathbun" baseAuthorityYear="1905" class="Malacostraca" family="Pseudothelphusidae" genus="Tehuana" kingdom="Animalia" lsidName="Tehuana complanata" order="Decapoda" pageId="0" pageNumber="1" phylum="Arthropoda" rank="species" species="complanata">
<emphasis id="CEC19D037FC67706DC51D931AC3623B6" italics="true" pageId="0" pageNumber="1">Tehuana complanata</emphasis>
</taxonomicName>
is also similar to 
<taxonomicName id="68CC2C63092627757BE9F75562CDB049" lsidName="T. lamellifrons" pageId="0" pageNumber="1" rank="species" species="lamellifrons">
<emphasis id="87C834BDD100C07D755D549ABF60B8D2" italics="true" pageId="0" pageNumber="1">T. lamellifrons</emphasis>
</taxonomicName>
even when they are geographically distant within the genus range (Fig. 
<figureCitation id="5AD2727E2AC2A4E37EF769FD1E19B4E1" captionStart="Figure 1" captionStartId="F1" captionText="Figure 1. Distribution of the species of the genus Tehuana in southeastern Mexico." figureDoi="10.3897/zookeys.1117.85362.figure1" httpUri="https://binary.pensoft.net/fig/725897" pageId="0" pageNumber="1">1</figureCitation>
); however, in the obtained phylogeny they appear as sister species (Fig. 
<figureCitation id="9BD052773B3A2531F2C3F4380D967682" captionStart="Figure 3" captionStartId="F3" captionText="Figure 3. Phylogenetic tree obtained for the genus Tehuana based on the concatenated analysis of three genes (H 3, 16 S, COI), using maximum likelihood and Bayesian inference. Branch supports values are posterior probability / bootstrap. Only clades with branch support above 50 % indicated. Clades a, b, and c are labelled for discussion purposes." figureDoi="10.3897/zookeys.1117.85362.figure3" httpUri="https://binary.pensoft.net/fig/725899" pageId="0" pageNumber="1">3</figureCitation>
).
</paragraph>
</subSubSection>
</treatment>
</document>