353 lines
42 KiB
XML
353 lines
42 KiB
XML
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<mods:title id="EB23857556773F51BA4529356F9C9163">Morphology and taxonomic assessment of eight genetic clades of Mercuria Boeters, 1971 (Caenogastropoda, Hydrobiidae), with the description of five new species</mods:title>
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<mods:affiliation id="E6E73EDBE6B2DC86643A4B70364FE519">Museo Nacional de Ciencias Naturales (MNCN-CSIC), José Gutiérrez Abascal 2, 28006 Madrid, Spain. Justus Liebig University Giessen, Department of Animal Ecology & Systematics, Heinrich-Buff-Ring 26 - 32 IFZD, 35392 Giessen, Germany. University of Carthage, Faculty of Sciences of Bizerte, LR 01 ES 14 Environmental Biomonitoring Laboratory, 7021 Jarzouna, Tunisia. Deceased 3 March 2023.</mods:affiliation>
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<mods:affiliation id="24698A93AC226894F1DCBB54C1694B04">Museo Nacional de Ciencias Naturales (MNCN-CSIC), José Gutiérrez Abascal 2, 28006 Madrid, Spain. Justus Liebig University Giessen, Department of Animal Ecology & Systematics, Heinrich-Buff-Ring 26 - 32 IFZD, 35392 Giessen, Germany. University of Carthage, Faculty of Sciences of Bizerte, LR 01 ES 14 Environmental Biomonitoring Laboratory, 7021 Jarzouna, Tunisia. Deceased 3 March 2023.</mods:affiliation>
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<treatment id="5C7987C4FFF9FFE5EF53F9EB3190FC9D" ID-DOI="http://doi.org/10.5281/zenodo.7891733" ID-GBIF-Taxon="209579767" ID-Zenodo-Dep="7891733" LSID="urn:lsid:plazi:treatment:5C7987C4FFF9FFE5EF53F9EB3190FC9D" httpUri="http://treatment.plazi.org/id/5C7987C4FFF9FFE5EF53F9EB3190FC9D" lastPageId="28" lastPageNumber="29" pageId="25" pageNumber="26">
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<heading id="8F2781BEFFF9FFE0EF53F9EB36DEF96A" box="[574,1013,1653,1680]" centered="true" fontSize="11" level="2" pageId="25" pageNumber="26" reason="2">
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<taxonomicName id="13D04D51FFF9FFE0EF53F9EB36DEF96A" ID-CoL="3ZTHF" authority="(Paladilhe, 1869)" baseAuthorityName="Paladilhe" baseAuthorityYear="1869" box="[574,1013,1653,1680]" class="Gastropoda" family="Hydrobiidae" genus="Mercuria" kingdom="Animalia" order="Littorinimorpha" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="maceana">
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<emphasis id="E6A4EAC0FFF9FFE0EF53F9EB360FF975" bold="true" box="[574,804,1653,1679]" italics="true" pageId="25" pageNumber="26">Mercuria maceana</emphasis>
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(
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<bibRefCitation id="B0414B23FFF9FFE0EE59F9EB36C6F96A" author="Paladilhe A." box="[820,1005,1653,1680]" pageId="25" pageNumber="26" refId="ref30481" refString="Paladilhe A. 1869. Nouvelles miscellanees malacologiques. Chez Savy, libraire-editeur, Paris. https: // doi. org / 10.5962 / bhl. title. 12995" type="book" year="1869">Paladilhe, 1869</bibRefCitation>
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)
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<subSubSection id="9CCA6559FFF9FFE0EFB5F9063677F949" box="[728,860,1688,1715]" pageId="25" pageNumber="26" type="description">
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<paragraph id="D46F36D2FFF9FFE0EFB5F9063677F949" blockId="25.[574,1013,1653,1715]" box="[728,860,1688,1715]" pageId="25" pageNumber="26">
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<figureCitation id="4CEB2A57FFF9FFE0EFB5F9063677F949" box="[728,860,1688,1715]" captionStart-0="Fig" captionStart-1="Fig" captionStartId-0="26.[189,232,1963,1989]" captionStartId-1="27.[189,232,1892,1918]" captionTargetBox-0="[189,1398,642,1923]" captionTargetBox-1="[189,1398,1117,1848]" captionTargetId-0="figure-94@26.[189,1398,642,1923]" captionTargetId-1="figure-238@27.[189,1398,1117,1848]" captionTargetPageId-0="26" captionTargetPageId-1="27" captionText-0="Fig. 15. Shells and operculum of Mercuria maceana (Paladilhe, 1869) from C’an Tunis, Barcelona, Spain. A–E. Shells. F. Operculum, inner side. G. Operculum, outer side." captionText-1="Fig. 16. Protoconch (A–C) and radula (D–F) of Mercuria maceana (Paladilhe, 1869) from C’an Tunis, Barcelona, Spain. A. Protoconch nucleus. B. Details of the protoconch. C. Protoconch microsculpture. D. Overview of radular teeth rows. E–F. Detailed view of the lateral and inner teeth." figureDoi-0="http://doi.org/10.5281/zenodo.7867393" figureDoi-1="http://doi.org/10.5281/zenodo.7867397" httpUri-0="https://zenodo.org/record/7867393/files/figure.png" httpUri-1="https://zenodo.org/record/7867397/files/figure.png" pageId="25" pageNumber="26">Figs 15–16</figureCitation>
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</paragraph>
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<paragraph id="D46F36D2FFF9FFE0EDD0F94437B4F90E" blockId="25.[189,671,1754,1780]" box="[189,671,1754,1780]" pageId="25" pageNumber="26">
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<treatmentCitationGroup id="F4C011FCFFF9FFE0EDD0F94437B4F90E" box="[189,671,1754,1780]" pageId="25" pageNumber="26">
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<treatmentCitation id="557110C3FFF9FFE0EDD0F94437B0F90E" author="Paladilhe A." box="[189,667,1754,1780]" page="103" pageId="25" pageNumber="26" year="1869">
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<taxonomicName id="13D04D51FFF9FFE0EDD0F94437B0F90E" authority="Paladilhe, 1869: 103" authorityName="Paladilhe" authorityPageNumber="103" authorityYear="1869" box="[189,667,1754,1780]" class="Gastropoda" family="Amnicolidae" genus="Amnicola" kingdom="Animalia" order="Littorinimorpha" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="maceana">
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<emphasis id="E6A4EAC0FFF9FFE0EDD0F94434B4F90E" box="[189,415,1754,1780]" italics="true" pageId="25" pageNumber="26">Amnicola maceana</emphasis>
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<bibRefCitation id="B0414B23FFF9FFE0ECCBF94437B0F90E" author="Paladilhe A." box="[422,667,1754,1780]" pageId="25" pageNumber="26" refId="ref30481" refString="Paladilhe A. 1869. Nouvelles miscellanees malacologiques. Chez Savy, libraire-editeur, Paris. https: // doi. org / 10.5962 / bhl. title. 12995" type="book" year="1869">Paladilhe, 1869: 103</bibRefCitation>
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.
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<paragraph id="D46F36D2FFF9FFE0EDD0F88237DAF8CF" blockId="25.[189,753,1819,1846]" box="[189,753,1819,1846]" pageId="25" pageNumber="26">
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<treatmentCitationGroup id="F4C011FCFFF9FFE0EDD0F88237DAF8CF" box="[189,753,1819,1846]" pageId="25" pageNumber="26">
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<taxonomicName id="13D04D51FFF9FFE0EDD0F88234B7F8CF" ID-CoL="3ZTHF" baseAuthorityName="Paladilhe" baseAuthorityYear="1869" box="[189,412,1820,1845]" class="Gastropoda" family="Hydrobiidae" genus="Mercuria" kingdom="Animalia" order="Littorinimorpha" pageId="25" pageNumber="26" phylum="Mollusca" rank="species" species="maceana">
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<emphasis id="E6A4EAC0FFF9FFE0EDD0F88234B7F8CF" box="[189,412,1820,1845]" italics="true" pageId="25" pageNumber="26">Mercuria maceana</emphasis>
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</taxonomicName>
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–
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<treatmentCitation id="557110C3FFF9FFE0ECD4F88537C6F8CF" author="Bank R. & Neubert E." box="[441,749,1819,1845]" page="16" pageId="25" pageNumber="26" year="2017">
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<bibRefCitation id="B0414B23FFF9FFE0ECD4F88537C6F8CF" author="Bank R. & Neubert E." box="[441,749,1819,1845]" pageId="25" pageNumber="26" refId="ref28115" refString="Bank R. & Neubert E. 2017. Checklist of the Land and Freshwater Gastropoda of Europe. Last update Jul. 16 th 2017. Molluscabase (Ed.)." type="book" year="2017">Bank & Neubert 2017: 16</bibRefCitation>
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</treatmentCitation>
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.
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<emphasis id="E6A4EAC0FFF9FFE0EDD0F8C234B1F88C" bold="true" box="[189,410,1884,1910]" pageId="25" pageNumber="26">Revised diagnosis</emphasis>
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</heading>
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</paragraph>
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<paragraph id="D46F36D2FFF9FFE0EDD0F81536FFF832" blockId="25.[189,1399,1931,1992]" pageId="25" pageNumber="26">Shell ovate-conic; aperture obliquely broad ovate; periostracum yellowish to pale grey; protoconch microsculpture granulated; central radular tooth formula 3-C-3/1-1.</paragraph>
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<emphasis id="E6A4EAC0FFFAFFE3EDD0FE973447FED9" bold="true" box="[189,364,265,291]" pageId="26" pageNumber="27">Type material</emphasis>
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(not examined)
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</paragraph>
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•
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(sex unknown);
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<collectingMunicipality id="340BACA8FFFAFFE3EF09FEC537F5FE8F" box="[612,734,347,373]" pageId="26" pageNumber="27">Barcelona</collectingMunicipality>
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,
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;
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|
||
<materialsCitation id="64B83C8FFFFAFFE3E9A4FEC536DFFE63" ID-GBIF-Occurrence="4098914394" collectionCode="MHNL" country="Spain" location="C'an Tunis" municipality="Barcelona" pageId="26" pageNumber="27" specimenCode="MHNL 45013950" specimenCount="1">
|
||
•
|
||
<specimenCount id="C2D6FD5BFFFAFFE3E98CFEC53012FE8F" box="[1249,1337,347,373]" pageId="26" pageNumber="27" type="generic">1 spec.</specimenCount>
|
||
(sex unknown); same collection data as for preceding;
|
||
<specimenCode id="84769EA9FFFAFFE3EE72FEE036DFFE63" box="[799,1012,382,409]" pageId="26" pageNumber="27">MHNL 45013950</specimenCode>
|
||
</materialsCitation>
|
||
<materialsCitation id="64B83C8FFFFAFFE3E96DFEE03135FE41" ID-GBIF-Occurrence="4098914435" collectionCode="UM" country="Spain" location="C'an Tunis" municipality="Barcelona" pageId="26" pageNumber="27" specimenCode="UM.PDL.006" specimenCount="1">
|
||
•
|
||
<specimenCount id="C2D6FD5BFFFAFFE3E978FEE03141FE62" box="[1045,1130,382,408]" pageId="26" pageNumber="27" type="generic">1 spec.</specimenCount>
|
||
(sex unknown); same collection data as for preceding;
|
||
<specimenCode id="84769EA9FFFAFFE3EF50FE3F37F6FE41" box="[573,733,417,443]" pageId="26" pageNumber="27">UM.PDL.006</specimenCode>
|
||
(
|
||
<bibRefCitation id="B0414B23FFFAFFE3EF80FE3F3139FE41" author="Breure A. & Audibert C." box="[749,1042,417,443]" pageId="26" pageNumber="27" pagination="28 - 39" refId="ref28746" refString="Breure A. & Audibert C. 2017. Under the spell of Bourguignat: Alcide Paladilhe's malacological work, with notes on his collection. Folia Conchyliologica 41: 28 - 39." type="journal article" year="2017">Breure & Audibert 2017</bibRefCitation>
|
||
).
|
||
</materialsCitation>
|
||
</paragraph>
|
||
<paragraph id="D46F36D2FFFAFFE3EDD0FE7D3476FE06" blockId="26.[189,349,482,509]" box="[189,349,482,509]" pageId="26" pageNumber="27">
|
||
<heading id="8F2781BEFFFAFFE3EDD0FE7D3476FE06" bold="true" box="[189,349,482,509]" fontSize="11" level="3" pageId="26" pageNumber="27" reason="3">
|
||
<emphasis id="E6A4EAC0FFFAFFE3EDD0FE7D3476FE06" bold="true" box="[189,349,482,509]" pageId="26" pageNumber="27">Type locality</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="D46F36D2FFFAFFE2EDD0FD8F34C6FE93" blockId="26.[189,1398,529,591]" lastBlockId="27.[189,1399,265,362]" lastPageId="27" lastPageNumber="28" pageId="26" pageNumber="27">
|
||
C’an
|
||
<collectingCountry id="ACC77642FFFAFFE3ED92FD8F346FFDD1" box="[255,324,529,555]" name="Tunisia" pageId="26" pageNumber="27">Tunis</collectingCountry>
|
||
, Barcelona,
|
||
<collectingCountry id="ACC77642FFFAFFE3ECB7FD8F370AFDD1" box="[474,545,529,555]" name="Spain" pageId="26" pageNumber="27">Spain</collectingCountry>
|
||
. The
|
||
<typeStatus id="0B6B8870FFFAFFE3EF09FD8C37BCFDD6" box="[612,663,530,556]" pageId="26" pageNumber="27">type</typeStatus>
|
||
locality of this species in the original description “récoltée à Antunez, près de Barcelone” refers to a series of local geographic names for a private property: C’an
|
||
<collectingCountry id="ACC77642FFFBFFE2EDD0FE973429FED9" box="[189,258,265,291]" name="Tunisia" pageId="27" pageNumber="28">Tunis</collectingCountry>
|
||
, Casa Antúnez, Camp de la Bota, Llano del Llobregat. These names have been written either conserving the original language, Catalan, sometimes using the Spanish form, or even using other local names for the same place.
|
||
</paragraph>
|
||
<caption id="80AF665AFFFAFFE3EDD0F8353139F812" ID-DOI="http://doi.org/10.5281/zenodo.7867393" ID-Zenodo-Dep="7867393" httpUri="https://zenodo.org/record/7867393/files/figure.png" pageId="26" pageNumber="27" startId="26.[189,232,1963,1989]" targetBox="[189,1398,642,1923]" targetPageId="26">
|
||
<paragraph id="D46F36D2FFFAFFE3EDD0F8353139F812" blockId="26.[189,1398,1963,2025]" pageId="26" pageNumber="27">
|
||
<emphasis id="E6A4EAC0FFFAFFE3EDD0F8353436F83F" bold="true" box="[189,285,1963,1989]" pageId="26" pageNumber="27">Fig. 15.</emphasis>
|
||
Shells and operculum of
|
||
<taxonomicName id="13D04D51FFFAFFE3EF34F8323138F83C" authority="(Paladilhe, 1869)" baseAuthorityName="Paladilhe" baseAuthorityYear="1869" box="[601,1043,1963,1990]" class="Gastropoda" family="Hydrobiidae" genus="Mercuria" kingdom="Animalia" order="Littorinimorpha" pageId="26" pageNumber="27" phylum="Mollusca" rank="species" species="maceana">
|
||
<emphasis id="E6A4EAC0FFFAFFE3EF34F8323610F83F" box="[601,827,1964,1989]" italics="true" pageId="26" pageNumber="27">Mercuria maceana</emphasis>
|
||
(
|
||
<bibRefCitation id="B0414B23FFFAFFE3EE23F8353120F83C" author="Paladilhe A." box="[846,1035,1963,1990]" pageId="26" pageNumber="27" refId="ref30481" refString="Paladilhe A. 1869. Nouvelles miscellanees malacologiques. Chez Savy, libraire-editeur, Paris. https: // doi. org / 10.5962 / bhl. title. 12995" type="book" year="1869">Paladilhe, 1869</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
from C’an Tunis, Barcelona, Spain.
|
||
<emphasis id="E6A4EAC0FFFAFFE3EC63F850346DF812" bold="true" box="[270,326,1998,2024]" pageId="26" pageNumber="27">A–E</emphasis>
|
||
. Shells.
|
||
<emphasis id="E6A4EAC0FFFAFFE3ECC6F8503496F812" bold="true" box="[427,445,1998,2024]" pageId="26" pageNumber="27">F</emphasis>
|
||
. Operculum, inner side.
|
||
<emphasis id="E6A4EAC0FFFAFFE3EF8DF85037DCF812" bold="true" box="[736,759,1998,2024]" pageId="26" pageNumber="27">G</emphasis>
|
||
. Operculum, outer side.
|
||
</paragraph>
|
||
</caption>
|
||
<paragraph id="D46F36D2FFFBFFE2EDD0FE0E3481FE50" blockId="27.[189,426,400,426]" box="[189,426,400,426]" pageId="27" pageNumber="28">
|
||
<heading id="8F2781BEFFFBFFE2EDD0FE0E3481FE50" bold="true" box="[189,426,400,426]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
|
||
<emphasis id="E6A4EAC0FFFBFFE2EDD0FE0E3481FE50" bold="true" box="[189,426,400,426]" pageId="27" pageNumber="28">Material examined</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="D46F36D2FFFBFFE2EDD0FE213783FE07" blockId="27.[189,1398,447,509]" pageId="27" pageNumber="28">
|
||
<materialsCitation id="64B83C8FFFFBFFE2EDD0FE21368FFE20" ID-GBIF-Occurrence="4098914306" box="[189,932,447,474]" collectionCode="MNCN" country="Spain" location="C'an Tunis" municipality="Barcelona" pageId="27" pageNumber="28" specimenCode="MNCN 15.05/7009" specimenCount="30">
|
||
<collectingCountry id="ACC77642FFFBFFE2EDD0FE213424FE23" box="[189,271,447,473]" name="Spain" pageId="27" pageNumber="28">SPAIN</collectingCountry>
|
||
•
|
||
<specimenCount id="C2D6FD5BFFFBFFE2EC46FE2134BEFE20" box="[299,405,447,474]" pageId="27" pageNumber="28">30 specs</specimenCount>
|
||
;
|
||
<collectingMunicipality id="340BACA8FFFBFFE2ECCFFE213737FE23" box="[418,540,447,473]" pageId="27" pageNumber="28">Barcelona</collectingMunicipality>
|
||
,
|
||
<location id="D10F6009FFFBFFE2EF47FE21379BFE23" LSID="urn:lsid:plazi:treatment:5C7987C4FFF9FFE5EF53F9EB3190FC9D:D10F6009FFFBFFE2EF47FE21379BFE23" box="[554,688,447,473]" country="Spain" municipality="Barcelona" name="C'an Tunis" pageId="27" pageNumber="28">C’an Tunis</location>
|
||
;
|
||
<specimenCode id="84769EA9FFFBFFE2EFD3FE21368FFE20" box="[702,932,447,474]" pageId="27" pageNumber="28">MNCN 15.05/7009</specimenCode>
|
||
</materialsCitation>
|
||
<materialsCitation id="64B83C8FFFFBFFE2EEC0FE5E3783FE07" ID-GBIF-Occurrence="4098914384" collectionCode="MZB" country="Spain" location="C'an Tunis" municipality="Barcelona" pageId="27" pageNumber="28" specimenCode="MZB 77-8445, MZB 77-8107" specimenCount="30">
|
||
•
|
||
<specimenCount id="C2D6FD5BFFFBFFE2EEADFE213102FE20" box="[960,1065,447,474]" pageId="27" pageNumber="28">30 specs</specimenCount>
|
||
; same collection data as for preceding;
|
||
<specimenCode id="84769EA9FFFBFFE2EC2DFE7D34C7FE07" box="[320,492,482,509]" collectionCode="MZB" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="27" pageNumber="28" type="Museum">MZB 77-8445</specimenCode>
|
||
,
|
||
<specimenCode id="84769EA9FFFBFFE2EC9AFE7D378FFE07" box="[503,676,483,509]" collectionCode="MZB" country="Indonesia" httpUri="http://grbio.org/cool/b9gz-izvg" name="Museum Zoologicum Bogoriense" pageId="27" pageNumber="28" type="Museum">MZB 77-8107</specimenCode>
|
||
.
|
||
</materialsCitation>
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9CCA6559FFFBFFE2EDD0FDBA31F6FC11" pageId="27" pageNumber="28" type="description">
|
||
<paragraph id="D46F36D2FFFBFFE2EDD0FDBA3465FDC4" blockId="27.[189,334,548,574]" box="[189,334,548,574]" pageId="27" pageNumber="28">
|
||
<heading id="8F2781BEFFFBFFE2EDD0FDBA3465FDC4" bold="true" box="[189,334,548,574]" fontSize="11" level="3" pageId="27" pageNumber="28" reason="3">
|
||
<emphasis id="E6A4EAC0FFFBFFE2EDD0FDBA3465FDC4" bold="true" box="[189,334,548,574]" pageId="27" pageNumber="28">Description</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="D46F36D2FFFBFFE2EDD0FDCD3627FCE7" blockId="27.[187,1399,594,797]" pageId="27" pageNumber="28">
|
||
<smallCapsWord id="D289A00EFFFBFFE2EDD0FDCD3429FD97" baselines="614,614" box="[189,258,595,621]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Shell" pageId="27" pageNumber="28">SHELL</smallCapsWord>
|
||
. Ovate-conic, whorls 4–5, height
|
||
<quantity id="13289B37FFFBFFE2EFC6FDCC366BFD97" box="[683,832,594,621]" metricMagnitude="-3" metricUnit="m" metricValue="4.55" metricValueMax="5.2" metricValueMin="3.9" pageId="27" pageNumber="28" unit="mm" value="4.55" valueMax="5.2" valueMin="3.9">3.9–5.2 mm</quantity>
|
||
, width
|
||
<quantity id="13289B37FFFBFFE2EEF2FDCD3137FD97" box="[927,1052,595,621]" metricMagnitude="-3" metricUnit="m" metricValue="3.4" metricValueMax="3.8" metricValueMin="3.0" pageId="27" pageNumber="28" unit="mm" value="3.4" valueMax="3.8" valueMin="3.0">3–3.8 mm</quantity>
|
||
(
|
||
<figureCitation id="4CEB2A57FFFBFFE2E95DFDCC31EFFD97" box="[1072,1220,594,621]" captionStart="Fig" captionStartId="26.[189,232,1963,1989]" captionTargetBox="[189,1398,642,1923]" captionTargetId="figure-94@26.[189,1398,642,1923]" captionTargetPageId="26" captionText="Fig. 15. Shells and operculum of Mercuria maceana (Paladilhe, 1869) from C’an Tunis, Barcelona, Spain. A–E. Shells. F. Operculum, inner side. G. Operculum, outer side." figureDoi="http://doi.org/10.5281/zenodo.7867393" httpUri="https://zenodo.org/record/7867393/files/figure.png" pageId="27" pageNumber="28">Fig. 15A–E</figureCitation>
|
||
); periostracum yellowish to pale grey; protoconch of 1.5 whorls, ca 350 µm wide, nucleus ca 200 µm wide (
|
||
<figureCitation id="4CEB2A57FFFBFFE2E996FDE8304DFD6B" box="[1275,1382,630,657]" captionStart="Fig" captionStartId="27.[189,232,1892,1918]" captionTargetBox="[189,1398,1117,1848]" captionTargetId="figure-238@27.[189,1398,1117,1848]" captionTargetPageId="27" captionText="Fig. 16. Protoconch (A–C) and radula (D–F) of Mercuria maceana (Paladilhe, 1869) from C’an Tunis, Barcelona, Spain. A. Protoconch nucleus. B. Details of the protoconch. C. Protoconch microsculpture. D. Overview of radular teeth rows. E–F. Detailed view of the lateral and inner teeth." figureDoi="http://doi.org/10.5281/zenodo.7867397" httpUri="https://zenodo.org/record/7867397/files/figure.png" pageId="27" pageNumber="28">Fig. 16A</figureCitation>
|
||
); protoconch microsculpture granulated (
|
||
<figureCitation id="4CEB2A57FFFBFFE2EFF9FD07360DFD4E" box="[660,806,665,692]" captionStart="Fig" captionStartId="27.[189,232,1892,1918]" captionTargetBox="[189,1398,1117,1848]" captionTargetId="figure-238@27.[189,1398,1117,1848]" captionTargetPageId="27" captionText="Fig. 16. Protoconch (A–C) and radula (D–F) of Mercuria maceana (Paladilhe, 1869) from C’an Tunis, Barcelona, Spain. A. Protoconch nucleus. B. Details of the protoconch. C. Protoconch microsculpture. D. Overview of radular teeth rows. E–F. Detailed view of the lateral and inner teeth." figureDoi="http://doi.org/10.5281/zenodo.7867397" httpUri="https://zenodo.org/record/7867397/files/figure.png" pageId="27" pageNumber="28">Fig. 16B–C</figureCitation>
|
||
); teleoconch whorls convex, separated by a deep suture; body whorl large, convex, occupying about two-thirds of the total shell length; aperture obliquely broad ovate, complete; inner lip thicker than outer lip; aperture margin slightly reflexed; umbilicus narrow, not covered by the inner lip (
|
||
<figureCitation id="4CEB2A57FFFBFFE2EF1DFC9C37D4FCE7" box="[624,767,770,797]" captionStart="Fig" captionStartId="26.[189,232,1963,1989]" captionTargetBox="[189,1398,642,1923]" captionTargetId="figure-94@26.[189,1398,642,1923]" captionTargetPageId="26" captionText="Fig. 15. Shells and operculum of Mercuria maceana (Paladilhe, 1869) from C’an Tunis, Barcelona, Spain. A–E. Shells. F. Operculum, inner side. G. Operculum, outer side." figureDoi="http://doi.org/10.5281/zenodo.7867393" httpUri="https://zenodo.org/record/7867393/files/figure.png" pageId="27" pageNumber="28">Fig. 15A–E</figureCitation>
|
||
).
|
||
</paragraph>
|
||
<paragraph id="D46F36D2FFFBFFE2EDD0FCDA31F6FC11" blockId="27.[188,1399,836,1004]" pageId="27" pageNumber="28">
|
||
<smallCapsWord id="D289A00EFFFBFFE2EDD0FCDA3432FCA4" baselines="855,856" box="[189,281,836,862]" lowerCaseFontSize="8" mainFontSize="11" normCase="title" normString="Radula" pageId="27" pageNumber="28">RADULA</smallCapsWord>
|
||
. Length intermediate, ca 800 µm long (35% of total shell length), containing about 63 rows of teeth. Central tooth formula 3-C-3/1-1, central cusp V shaped, cutting edge slightly concave (
|
||
<figureCitation id="4CEB2A57FFFBFFE2E990FCF93042FC78" box="[1277,1385,871,898]" captionStart="Fig" captionStartId="27.[189,232,1892,1918]" captionTargetBox="[189,1398,1117,1848]" captionTargetId="figure-238@27.[189,1398,1117,1848]" captionTargetPageId="27" captionText="Fig. 16. Protoconch (A–C) and radula (D–F) of Mercuria maceana (Paladilhe, 1869) from C’an Tunis, Barcelona, Spain. A. Protoconch nucleus. B. Details of the protoconch. C. Protoconch microsculpture. D. Overview of radular teeth rows. E–F. Detailed view of the lateral and inner teeth." figureDoi="http://doi.org/10.5281/zenodo.7867397" httpUri="https://zenodo.org/record/7867397/files/figure.png" pageId="27" pageNumber="28">Fig. 16D</figureCitation>
|
||
). Lateral tooth formula (3)4-C-4(3), central cusp V shaped and slightly longer than the central tooth one. Inner marginal teeth with 12–14 cusps; outer marginal teeth with 12–14 cusps (
|
||
<figureCitation id="4CEB2A57FFFBFFE2E915FC33302FFC32" box="[1144,1284,941,968]" captionStart="Fig" captionStartId="27.[189,232,1892,1918]" captionTargetBox="[189,1398,1117,1848]" captionTargetId="figure-238@27.[189,1398,1117,1848]" captionTargetPageId="27" captionText="Fig. 16. Protoconch (A–C) and radula (D–F) of Mercuria maceana (Paladilhe, 1869) from C’an Tunis, Barcelona, Spain. A. Protoconch nucleus. B. Details of the protoconch. C. Protoconch microsculpture. D. Overview of radular teeth rows. E–F. Detailed view of the lateral and inner teeth." figureDoi="http://doi.org/10.5281/zenodo.7867397" httpUri="https://zenodo.org/record/7867397/files/figure.png" pageId="27" pageNumber="28">Fig. 16E–F</figureCitation>
|
||
). Radular data were compiled from a dry specimen (MZB 77-8445) collected from the
|
||
<typeStatus id="0B6B8870FFFBFFE2E92BFC4F3152FC11" box="[1094,1145,977,1003]" pageId="27" pageNumber="28">type</typeStatus>
|
||
locality.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<caption id="80AF665AFFFBFFE2EDD0F8FA31B5F83F" ID-DOI="http://doi.org/10.5281/zenodo.7867397" ID-Zenodo-Dep="7867397" httpUri="https://zenodo.org/record/7867397/files/figure.png" pageId="27" pageNumber="28" startId="27.[189,232,1892,1918]" targetBox="[189,1398,1117,1848]" targetPageId="27">
|
||
<paragraph id="D46F36D2FFFBFFE2EDD0F8FA31B5F83F" blockId="27.[189,1399,1892,1990]" pageId="27" pageNumber="28">
|
||
<emphasis id="E6A4EAC0FFFBFFE2EDD0F8FA3431F885" bold="true" box="[189,282,1892,1919]" pageId="27" pageNumber="28">Fig. 16.</emphasis>
|
||
Protoconch (A–C) and radula (D–F) of
|
||
<taxonomicName id="13D04D51FFFBFFE2EF99F8F8318FF87A" authority="(Paladilhe, 1869)" baseAuthorityName="Paladilhe" baseAuthorityYear="1869" box="[756,1188,1893,1920]" class="Gastropoda" family="Hydrobiidae" genus="Mercuria" kingdom="Animalia" order="Littorinimorpha" pageId="27" pageNumber="28" phylum="Mollusca" rank="species" species="maceana">
|
||
<emphasis id="E6A4EAC0FFFBFFE2EF99F8F836F8F885" box="[756,979,1894,1919]" italics="true" pageId="27" pageNumber="28">Mercuria maceana</emphasis>
|
||
(
|
||
<bibRefCitation id="B0414B23FFFBFFE2EE8FF8FB31B7F87A" author="Paladilhe A." box="[994,1180,1893,1920]" pageId="27" pageNumber="28" refId="ref30481" refString="Paladilhe A. 1869. Nouvelles miscellanees malacologiques. Chez Savy, libraire-editeur, Paris. https: // doi. org / 10.5962 / bhl. title. 12995" type="book" year="1869">Paladilhe, 1869</bibRefCitation>
|
||
)
|
||
</taxonomicName>
|
||
from C’an Tunis, Barcelona, Spain.
|
||
<emphasis id="E6A4EAC0FFFBFFE2ECFBF8163480F858" bold="true" box="[406,427,1928,1954]" pageId="27" pageNumber="28">A</emphasis>
|
||
. Protoconch nucleus.
|
||
<emphasis id="E6A4EAC0FFFBFFE2EFDCF81637EFF858" bold="true" box="[689,708,1928,1954]" pageId="27" pageNumber="28">B</emphasis>
|
||
. Details of the protoconch.
|
||
<emphasis id="E6A4EAC0FFFBFFE2E963F8163108F858" bold="true" box="[1038,1059,1928,1954]" pageId="27" pageNumber="28">C</emphasis>
|
||
. Protoconch microsculpture.
|
||
<emphasis id="E6A4EAC0FFFBFFE2EDD3F83535F8F83F" bold="true" box="[190,211,1963,1989]" pageId="27" pageNumber="28">D</emphasis>
|
||
. Overview of radular teeth rows.
|
||
<emphasis id="E6A4EAC0FFFBFFE2EF33F83537B9F83F" bold="true" box="[606,658,1963,1989]" pageId="27" pageNumber="28">E–F</emphasis>
|
||
. Detailed view of the lateral and inner teeth.
|
||
</paragraph>
|
||
</caption>
|
||
<subSubSection id="9CCA6559FFFCFFE5EDD0FE97315BFE24" pageId="28" pageNumber="29" type="biology_ecology">
|
||
<paragraph id="D46F36D2FFFCFFE5EDD0FE9734D9FED8" blockId="28.[189,498,264,291]" box="[189,498,264,291]" pageId="28" pageNumber="29">
|
||
<heading id="8F2781BEFFFCFFE5EDD0FE9734D9FED8" bold="true" box="[189,498,264,291]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
|
||
<emphasis id="E6A4EAC0FFFCFFE5EDD0FE9734D9FED8" bold="true" box="[189,498,264,291]" pageId="28" pageNumber="29">Ecology and distribution</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="D46F36D2FFFCFFE5EDD0FEA9315BFE24" blockId="28.[188,1401,311,478]" pageId="28" pageNumber="29">
|
||
This species is only known from the
|
||
<typeStatus id="0B6B8870FFFCFFE5EF11FEA63784FEA8" box="[636,687,312,338]" pageId="28" pageNumber="29">type</typeStatus>
|
||
locality, C’an
|
||
<collectingCountry id="ACC77642FFFCFFE5EE0CFEA93688FEAB" box="[865,931,311,337]" name="Tunisia" pageId="28" pageNumber="29">Tunis</collectingCountry>
|
||
(Casa Antúnez),
|
||
<collectingRegion id="1614F830FFFCFFE5E918FEA931C3FEAB" box="[1141,1256,311,337]" country="Spain" name="Catalunya" pageId="28" pageNumber="29">Catalonia</collectingRegion>
|
||
,
|
||
<collectingCountry id="ACC77642FFFCFFE5E99BFEA93016FEAB" box="[1270,1341,311,337]" name="Spain" pageId="28" pageNumber="29">Spain</collectingCountry>
|
||
. The
|
||
<typeStatus id="0B6B8870FFFCFFE5EDD2FEC535D9FE8F" box="[191,242,347,373]" pageId="28" pageNumber="29">type</typeStatus>
|
||
locality has likely disappeared due to extensive alterations (e.g., construction, pollution) of the area as a result of port works conducted in the Barcelona harbour and the industrial zone in the 1980s and the construction work carried out for the 1992 Olympic Games (Jordi Corbella and Francesc Uribe pers. com.). Recent surveys conducted around Barcelona did not yield live specimens.
|
||
</paragraph>
|
||
</subSubSection>
|
||
<subSubSection id="9CCA6559FFFCFFE5EDD0FD9B3190FC9D" pageId="28" pageNumber="29" type="discussion">
|
||
<paragraph id="D46F36D2FFFCFFE5EDD0FD9B3404FDE5" blockId="28.[189,303,517,543]" box="[189,303,517,543]" pageId="28" pageNumber="29">
|
||
<heading id="8F2781BEFFFCFFE5EDD0FD9B3404FDE5" bold="true" box="[189,303,517,543]" fontSize="11" level="3" pageId="28" pageNumber="29" reason="3">
|
||
<emphasis id="E6A4EAC0FFFCFFE5EDD0FD9B3404FDE5" bold="true" box="[189,303,517,543]" pageId="28" pageNumber="29">Remarks</emphasis>
|
||
</heading>
|
||
</paragraph>
|
||
<paragraph id="D46F36D2FFFCFFE5EDD0FDAB3190FC9D" blockId="28.[188,1399,564,872]" pageId="28" pageNumber="29">
|
||
<taxonomicName id="13D04D51FFFCFFE5EDD0FDAB3489FDB4" baseAuthorityName="Paladilhe" baseAuthorityYear="1869" box="[189,418,565,590]" class="Gastropoda" family="Hydrobiidae" genus="Mercuria" kingdom="Animalia" order="Littorinimorpha" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="maceana">
|
||
<emphasis id="E6A4EAC0FFFCFFE5EDD0FDAB3489FDB4" box="[189,418,565,590]" italics="true" pageId="28" pageNumber="29">Mercuria maceana</emphasis>
|
||
</taxonomicName>
|
||
is only known from its
|
||
<typeStatus id="0B6B8870FFFCFFE5EFB0FDAA363BFDB4" box="[733,784,564,590]" pageId="28" pageNumber="29">type</typeStatus>
|
||
locality, which has been destroyed over the past few decades. Specimens of this species have not been found in the nearby areas, suggesting that the species is extinct (IUCN category EX). The shells and dry specimens of
|
||
<taxonomicName id="13D04D51FFFCFFE5E94DFDE53198FD6E" baseAuthorityName="Paladilhe" baseAuthorityYear="1869" box="[1056,1203,635,660]" class="Gastropoda" family="Hydrobiidae" genus="Mercuria" kingdom="Animalia" order="Littorinimorpha" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="maceana">
|
||
<emphasis id="E6A4EAC0FFFCFFE5E94DFDE53198FD6E" box="[1056,1203,635,660]" italics="true" pageId="28" pageNumber="29">M. maceana</emphasis>
|
||
</taxonomicName>
|
||
deposited in the Zoological Museum of Barcelona were examined to delineate the morphological variation presented by the species and to support its recognition. Our PCA based on shell shape differentiated this species (
|
||
<figureCitation id="4CEB2A57FFFCFFE5EDAAFD7A3427FD04" box="[199,268,740,766]" captionStart="Fig" captionStartId="7.[189,232,1299,1325]" captionTargetBox="[225,1377,281,1239]" captionTargetId="figure-86@7.[189,1399,262,1263]" captionTargetPageId="7" captionText="Fig. 3. Principal components analysis plot for species of Mercuria Boeters, 1971 based on 21 coordinates (9 landmarks and 12 semilandmarks). Each dot on the plot represents the consensus of the shape of each of the sampled populations. Warp grid deformations (TPS) show expansion (in red colour) and contraction (in blue) of the shell shape." figureDoi="http://doi.org/10.5281/zenodo.7867365" httpUri="https://zenodo.org/record/7867365/files/figure.png" pageId="28" pageNumber="29">Fig. 3</figureCitation>
|
||
). Accordingly,
|
||
<taxonomicName id="13D04D51FFFCFFE5ECD3FD7B3764FD04" baseAuthorityName="Paladilhe" baseAuthorityYear="1869" box="[446,591,741,766]" class="Gastropoda" family="Hydrobiidae" genus="Mercuria" kingdom="Animalia" order="Littorinimorpha" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="maceana">
|
||
<emphasis id="E6A4EAC0FFFCFFE5ECD3FD7B3764FD04" box="[446,591,741,766]" italics="true" pageId="28" pageNumber="29">M. maceana</emphasis>
|
||
</taxonomicName>
|
||
differs from
|
||
<taxonomicName id="13D04D51FFFCFFE5EF87FD7B36ABFD04" authority="Miller & Delicado & García-Guerrero & Khalloufi & Ramos, 2023" authorityName="Miller & Delicado & García-Guerrero & Khalloufi & Ramos" authorityYear="2023" box="[746,896,741,766]" class="Gastropoda" family="Hydrobiidae" genus="Mercuria" kingdom="Animalia" order="Littorinimorpha" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="egarensis" status="sp. nov.">
|
||
<emphasis id="E6A4EAC0FFFCFFE5EF87FD7B36ABFD04" box="[746,896,741,766]" italics="true" pageId="28" pageNumber="29">M. egarensis</emphasis>
|
||
</taxonomicName>
|
||
<taxonomicNameLabel id="FD9757BBFFFCFFE5EEEAFD7A36C9FD04" box="[903,994,740,766]" pageId="28" pageNumber="29" rank="species">sp. nov.</taxonomicNameLabel>
|
||
by its shorter and more globose or subglobose shell and is more similar to the Moroccan species
|
||
<taxonomicName id="13D04D51FFFCFFE5EEE1FC96317FFCDB" authorityName="Gloer, Boeters & Walther" authorityYear="2015" box="[908,1108,776,801]" class="Gastropoda" family="Hydrobiidae" genus="Mercuria" kingdom="Animalia" order="Littorinimorpha" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="targouasensis">
|
||
<emphasis id="E6A4EAC0FFFCFFE5EEE1FC96317FFCDB" box="[908,1108,776,801]" italics="true" pageId="28" pageNumber="29">M. targouasensis</emphasis>
|
||
</taxonomicName>
|
||
. Based on the characters observed on the radulae,
|
||
<taxonomicName id="13D04D51FFFCFFE5EC8CFCB5375AFCBE" baseAuthorityName="Paladilhe" baseAuthorityYear="1869" box="[481,625,811,836]" class="Gastropoda" family="Hydrobiidae" genus="Mercuria" kingdom="Animalia" order="Littorinimorpha" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="maceana">
|
||
<emphasis id="E6A4EAC0FFFCFFE5EC8CFCB5375AFCBE" box="[481,625,811,836]" italics="true" pageId="28" pageNumber="29">M. maceana</emphasis>
|
||
</taxonomicName>
|
||
cannot be distinguished from
|
||
<taxonomicName id="13D04D51FFFCFFE5EEBEFCB5316DFCBE" baseAuthorityName="Draparnaud" baseAuthorityYear="1805" box="[979,1094,810,836]" class="Gastropoda" family="Hydrobiidae" genus="Mercuria" kingdom="Animalia" order="Littorinimorpha" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="similis">
|
||
<emphasis id="E6A4EAC0FFFCFFE5EEBEFCB5316DFCBE" box="[979,1094,810,836]" italics="true" pageId="28" pageNumber="29">M. similis</emphasis>
|
||
</taxonomicName>
|
||
or
|
||
<taxonomicName id="13D04D51FFFCFFE5E901FCB53029FCBE" authorityName="Miller & Delicado & García-Guerrero & Khalloufi & Ramos" authorityYear="2023" box="[1132,1282,811,836]" class="Gastropoda" family="Hydrobiidae" genus="Mercuria" kingdom="Animalia" order="Littorinimorpha" pageId="28" pageNumber="29" phylum="Mollusca" rank="species" species="egarensis">
|
||
<emphasis id="E6A4EAC0FFFCFFE5E901FCB53029FCBE" box="[1132,1282,811,836]" italics="true" pageId="28" pageNumber="29">M. egarensis</emphasis>
|
||
</taxonomicName>
|
||
. We were unable to study other anatomical features due to the lack of ethanol-preserved material.
|
||
</paragraph>
|
||
</subSubSection>
|
||
</treatment>
|
||
</document> |